203 lines
20 KiB
XML
203 lines
20 KiB
XML
<document id="73D29A073F9DF67B293E56C8A30D4FCF" ID-CLB-Dataset="26691" ID-DOI="http://dx.doi.org/10.3897/evolsyst.2.30778" ID-GBIF-Dataset="00106674-20d9-4d70-bb0f-8f000fb311f2" ID-Pensoft-Pub="2535-0730-2-169" ID-ZooBank="C0E89FEC8BE54DE9803D784FF6727BA0" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="2535-0730-2-169" ModsDocOrigin="Evolutionary Systematics 2 (2)" ModsDocTitle="The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae)" checkinTime="1558621757476" checkinUser="pensoft" docAuthor="Framenau, Volker W. & Baehr, Barbara C." docDate="2018" docId="1275A20343036E702F3591EBF79AB45E" docLanguage="en" docName="EvolutSyst 2(2): 169-241" docOrigin="Evolutionary Systematics 2 (2)" docSource="http://dx.doi.org/10.3897/evolsyst.2.30778" docTitle="Artoria belfordensis Framenau & Baehr, 2018, sp. n." docType="treatment" docUuid="8B0092B0-56B6-4AA8-BD11-BC63FB3CD772" docUuidSource="ZooBank" docVersion="6" lastPageNumber="169" masterDocId="FFE8FFB72743554D943BFFB83D324B46" masterDocTitle="The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae)" masterLastPageNumber="241" masterPageNumber="169" pageNumber="169" updateTime="1701381204780" updateUser="ExternalLinkService">
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<mods:titleInfo id="8FC177ED4B017CD2BCCE6C660B53A8E9">
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<mods:title id="C2F8E3BC452F5FEF389017584620FFE7">The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae)</mods:title>
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<mods:namePart id="7C04B538424C9A41FD94D6A076392E00">Framenau, Volker W.</mods:namePart>
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<mods:name id="7BA5E6ECE6837E719D742E20D10625FC" type="personal">
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<mods:roleTerm id="1728DE95B3D8B9CC8CF14B78554F7AC9">Author</mods:roleTerm>
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<mods:namePart id="46BBF461E73214289EBF7C5A6E92ECC4">Baehr, Barbara C.</mods:namePart>
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<mods:typeOfResource id="0B701B973B2F14A3B0C236B0758ADD82">text</mods:typeOfResource>
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<mods:title id="73423604CD13A788995267F29AB08768">Evolutionary Systematics</mods:title>
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<mods:part id="7DABF05FAB2BE0F6EA621AF09E747C87">
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<mods:date id="F48E675ECE21FF38E884AA54F5F3FA22">2018</mods:date>
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<mods:number id="D657035866CCCDC9E9018F014DFD9863">2</mods:number>
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<mods:detail id="8252AC9673F61CE5B9A1D5C8869E4492" type="issue">
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<mods:number id="43E29FD2A544E2F252FAFA4A4B6A916A">2</mods:number>
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<mods:start id="B0430300E33B48E23D80EC5A5B783B8D">169</mods:start>
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<mods:url id="300263CBA36EFFB978DD8E8E48A47D6D">http://dx.doi.org/10.3897/evolsyst.2.30778</mods:url>
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<mods:classification id="EDB6D60F4773795334C4CC286E695F1B">journal article</mods:classification>
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<mods:identifier id="6B1A0DD39F42E3C589591E5B2B50EF2A" type="DOI">http://dx.doi.org/10.3897/evolsyst.2.30778</mods:identifier>
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<treatment id="1275A20343036E702F3591EBF79AB45E" ID-GBIF-Taxon="157251914" LSID="urn:lsid:zoobank.org:act:8B0092B0-56B6-4AA8-BD11-BC63FB3CD772" httpUri="http://treatment.plazi.org/id/1275A20343036E702F3591EBF79AB45E" lastPageNumber="169" pageId="0" pageNumber="169">
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<subSubSection id="A9B6A5C3845435596D7EED28E5C22A8F" pageId="0" pageNumber="169" type="nomenclature">
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<paragraph id="18AD0CC0A4F5E5C106D2A2AF97AD320E" pageId="0" pageNumber="169">
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<taxonomicName id="2F04940D242B95F2BEE57121C0EAA3BF" ID-CoL="H2BD" LSID="http://zoobank.org/8B0092B0-56B6-4AA8-BD11-BC63FB3CD772" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria belfordensis" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="belfordensis">Artoria belfordensis</taxonomicName>
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<taxonomicNameLabel id="8DF7C326D0663F8426D316FFD9726D25" pageId="0" pageNumber="169">sp. n.</taxonomicNameLabel>
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Figs 2D, 8, 9
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<normalizedToken id="1C2573CC5DD871374954BE29064C56AE" originalValue="A–H">A-H</normalizedToken>
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, 47C Belford Forest Runner
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</paragraph>
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</subSubSection>
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<subSubSection id="07DBAE1D6C8486CA3FEC72D8994940AF" pageId="0" pageNumber="169" type="materials_examined">
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<paragraph id="24D234A5A60E9F160E8F064B39FD89CE" pageId="0" pageNumber="169">Material examined.</paragraph>
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<paragraph id="D6E40AE9137C6A3E2DB6DE6499E7F00D" pageId="0" pageNumber="169">
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Holotype male, Belford National Park, 60 m from Kirkton Road and 2.5 km east of Belford (
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<geoCoordinate id="C13851B8286513CF9647E9E466C958C4" direction="south" orientation="latitude" precision="925" value="-32.65">32°39'S</geoCoordinate>
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,
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<geoCoordinate id="656172A367FC9B94FD38E4E757B045B5" direction="east" orientation="longitude" precision="925" value="151.3">151°18'E</geoCoordinate>
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, New South Wales, AUSTRALIA), 17-27 June 2012, J.R. Gollan, M.A. Ashcroft, pitfall trap, canopy of trees but moderate condition, (AM KS127759). Paratypes: 5 males, same data as holotype (AM KS122425); 2 males, 1 female, Maroota State Forest (
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<geoCoordinate id="CC204E6AD566C2CCFF225B8C68FF327A" direction="south" orientation="latitude" precision="925" value="-33.516666">33°31'S</geoCoordinate>
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,
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<geoCoordinate id="D11784D1767952B35CBE5313EA61ED4E" direction="east" orientation="longitude" precision="925" value="150.98334">150°59'E</geoCoordinate>
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, New South Wales, AUSTRALIA), 26 October 1979, G.A. Webb, NSW States Forest Survey H/8 (AM KS73425).
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</paragraph>
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</subSubSection>
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<subSubSection id="E93BE8DE8D1CE3A405DC6047B052FA76" pageId="0" pageNumber="169" type="materials_examined">
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<paragraph id="F73CC0D8FFEAFC3EDFEAF1FE30CE560C" pageId="0" pageNumber="169">Other material examined.</paragraph>
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<paragraph id="C72E8350E46D8231DE61872F10A5DB2E" pageId="0" pageNumber="169">
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18 males 13 records (all NSW). AUSTRALIA: New South Wales: 1 male, Currawong,
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<geoCoordinate id="DC30B6DBA401F0DC7F2FE05F9562A1F1" direction="south" orientation="latitude" precision="925" value="-33.6">33°36'S</geoCoordinate>
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,
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<geoCoordinate id="E9165F12AC90FF58F67E90D3DCD588FD" direction="east" orientation="longitude" precision="925" value="151.3">151°18'E</geoCoordinate>
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(AM KS86426); 3 males, Ferntree Gully Reserve, approx. 170 m from Bylong Valley Way and 4 km SW of Ginghi,
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<geoCoordinate id="A1E1922509C0166EBDBAF1F529364F48" direction="south" orientation="latitude" precision="15" value="-32.62222">
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32°37
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<normalizedToken id="996EC98F7C75D4575C8FC2955F2AB3F8" originalValue="’20”">'20"</normalizedToken>
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S
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</geoCoordinate>
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,
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<geoCoordinate id="C1FC64DBACECA4EF19DA94EA26F56262" direction="east" orientation="longitude" precision="15" value="150.06029">
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150°03
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<normalizedToken id="E39610A49E1F9916C9089A74F013A40A" originalValue="’37”">'37"</normalizedToken>
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E
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</geoCoordinate>
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(AM KS122454); 1 male, Goulburn River National Park, approx. 120 m from Bylong Valley Way and 6 km E of Bylong,
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<geoCoordinate id="3639F45C28588417284A2ACD8BC18F1E" direction="south" orientation="latitude" precision="15" value="-32.405003">
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32°24
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<normalizedToken id="4A9EC833225AE7DB57BF6526555E5523" originalValue="’18”">'18"</normalizedToken>
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S
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</geoCoordinate>
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,
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<geoCoordinate id="AF1D4DC9B9D58C07723296CFF7BC9F3A" direction="east" orientation="longitude" precision="15" value="150.18611">
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150°11
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<normalizedToken id="47256AB1FDA83B84C7319C61AB89E8D8" originalValue="’10”">'10"</normalizedToken>
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E
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</geoCoordinate>
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(AM KS122446); 13 males, Maroota State Forest,
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<geoCoordinate id="9A5CB22B7BD275CF196D1330D7AFE30D" direction="south" orientation="latitude" precision="925" value="-33.516666">33°31'S</geoCoordinate>
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,
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<geoCoordinate id="B88DF3BA41E398107434C60D6D23D458" direction="east" orientation="longitude" precision="925" value="150.98334">150°59'E</geoCoordinate>
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(AM KS73313, KS73327, KS73338, KS73384, KS73388, KS73397, KS73407, KS73441, KS84096); 1 female, same locality (AM KS 73452); 1 female, Wallaroo State Forest, Flaggy Creek,
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<geoCoordinate id="C412E96DE5BB4B2D2C806302617F3F20" direction="south" orientation="latitude" precision="15" value="-32.60083">
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32°36
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<normalizedToken id="6C03AC6B92F2568C5B93618DB1BF7AC7" originalValue="’03”">'03"</normalizedToken>
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S
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</geoCoordinate>
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,
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<geoCoordinate id="DE2C35EB09A00216381EB69A890B9DA1" direction="east" orientation="longitude" precision="15" value="151.80194">
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151°48
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<normalizedToken id="E01438D651931DFEAF0F3B6B8D3A8768" originalValue="’07”">'07"</normalizedToken>
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E
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</geoCoordinate>
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(AM KS39707); 1 female, Yengo National Park, approx. 90 m from Howes Trail Road,
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<geoCoordinate id="B751DEC7954764A03E78389FBAEF9453" direction="south" orientation="latitude" precision="15" value="-32.892223">
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32°53
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<normalizedToken id="5591CF2522147868B8DB89BD3464FDFB" originalValue="’32”">'32"</normalizedToken>
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S
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</geoCoordinate>
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,
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<geoCoordinate id="A7AB71107235F87841100232E326BB05" direction="east" orientation="longitude" precision="15" value="150.87973">
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150°52
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<normalizedToken id="A738AB17D8A4B9E1336F2545FAF019D4" originalValue="’47”">'47"</normalizedToken>
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E
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</geoCoordinate>
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(AM KS122319); Yesabah Caves, near Kempsey,
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<geoCoordinate id="2CCBB0404C8125A59BB0C4BAB3612009" direction="south" orientation="latitude" precision="925" value="-31.083334">31°05'S</geoCoordinate>
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,
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<geoCoordinate id="81990C6E5D17A3A2B1519EAE97E23277" direction="east" orientation="longitude" precision="925" value="152.71666">152°43'E</geoCoordinate>
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(AM KS84058).
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</paragraph>
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</subSubSection>
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<subSubSection id="10CC17FEA18551287180811EA9594D23" pageId="0" pageNumber="169" type="etymology">
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<paragraph id="379DF5D00FB75DD80C52647A2A57E37A" pageId="0" pageNumber="169">Etymology.</paragraph>
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<paragraph id="70DB16BDEDE05B1846744BECB67D6912" pageId="0" pageNumber="169">The specific name is an adjective in apposition derived from the type locality, Belford National Park.</paragraph>
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</subSubSection>
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<subSubSection id="37EBC1A04DD921447D37FEEE65ACD122" pageId="0" pageNumber="169" type="diagnosis">
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<paragraph id="38F47023B94C1D3B1ECA821C2BD538DB" pageId="0" pageNumber="169">Diagnosis.</paragraph>
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<paragraph id="2900F59783F25BD3E2D69414D9505F8A" pageId="0" pageNumber="169">
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Males of
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<taxonomicName id="345A4920634099E7ABD8642B1B23A1C5" lsidName="A. belfordensis" pageId="0" pageNumber="169" rank="species" species="belfordensis">A. belfordensis</taxonomicName>
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sp. n. differ from all other
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<taxonomicName id="E10A1FAD4BD5E648A348614B84096860" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="genus">Artoria</taxonomicName>
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species, except
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<taxonomicName id="49A8718351CAE1522C6AE238A1F72078" lsidName="A. taeniifera" pageId="0" pageNumber="169" rank="species" species="taeniifera">A. taeniifera</taxonomicName>
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, by the base of the basoembolic apophysis, which is drawn out into an apical tip (Fig. 47C). It differs from
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<taxonomicName id="6639B21EEDE02D993CFB8646F75E8A10" lsidName="A. taeniifera" pageId="0" pageNumber="169" rank="species" species="taeniifera">A. taeniifera</taxonomicName>
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by the shape of the tegular apophysis, distinctly two-pronged in
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<taxonomicName id="61B36588760FA60C55A0BA87A63A158A" lsidName="A. taeniifera" pageId="0" pageNumber="169" rank="species" species="taeniifera">A. taeniifera</taxonomicName>
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, but three-lobed in
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<taxonomicName id="B2169C55E19DEE9221DE1882F198EB3E" lsidName="A. belfordensis" pageId="0" pageNumber="169" rank="species" species="belfordensis">A. belfordensis</taxonomicName>
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sp. n. There is some variation in this character, but also in the shape of the tegular apophysis within this species. Too little material is available to judge if this represents intra- or interspecific variation. We therefore consider this variation capable of occurring within one species. Females are most similar to females of
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<taxonomicName id="C65D8CD3EB9EDE17DECADD09FA33D4AC" lsidName="A. taeniifera" pageId="0" pageNumber="169" rank="species" species="taeniifera">A. taeniifera</taxonomicName>
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but differs in the shape of the atrium which is inverted U-shaped in
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<taxonomicName id="C99DC5B564043E3BC9F79E60EA34797F" lsidName="A. taeniifera" pageId="0" pageNumber="169" rank="species" species="taeniifera">A. taeniifera</taxonomicName>
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(Fig. 38G) and obscure in
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<taxonomicName id="1D343FE91502A97395F914126BC49591" lsidName="A. belfordensis" pageId="0" pageNumber="169" rank="species" species="belfordensis">A. belfordensis</taxonomicName>
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(Fig. 9G).
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</paragraph>
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</subSubSection>
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<subSubSection id="1114AEAE33BDD69E4B732F83416956E4" pageId="0" pageNumber="169" type="description">
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<paragraph id="F900FDA25B5637AEA9D397053B770171" pageId="0" pageNumber="169">Description.</paragraph>
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<paragraph id="B1FA00CCB1056EE5C2F112529DCB1D52" pageId="0" pageNumber="169">Male (based on holotype AM KS127759; palea AM KS122425).</paragraph>
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<paragraph id="A117ACD29357E1133DC3042F173E5281" pageId="0" pageNumber="169">Total length 4.3.</paragraph>
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<paragraph id="45B356C590D7B4CBA2E5935A06254D6E" pageId="0" pageNumber="169">Prosoma. Length 2.5, width 1.8; carapace light yellow-brown with indistinct dark radial and black V-shaped pattern between cephalic and thoracic region; covered with dark setae centrally and marginally with a band of lighter setae (Fig. 9A); sternum medium brown, dusted dark grey (Fig. 9B).</paragraph>
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<paragraph id="881E757918C6A04F5BB991F02726C99F" pageId="0" pageNumber="169">Eyes (Fig. 2D). Diameter of AME: 0.09; ALE: 0.08; PME: 0.26; PLE: 0.21.</paragraph>
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<paragraph id="3E70A62C114D5C1EFA3D3EA0E402BF5C" pageId="0" pageNumber="169">Anterior eye row. Strongly procurved, evenly spaced.</paragraph>
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<paragraph id="DC9871899820070A5E92FCB02750A17D" pageId="0" pageNumber="169">Chelicerae. Dark brown darker apically.</paragraph>
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<paragraph id="84A38DF8729208EF8F375FE0DA31D83A" pageId="0" pageNumber="169">Labium. Dark brown, with lighter anterior rim (Fig. 9B)</paragraph>
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<paragraph id="EC2E9C48F22540F058FC291770F3E623" pageId="0" pageNumber="169">Pedipalp coxae. Dark brown, with lighter anterior rim (Fig. 9B).</paragraph>
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<paragraph id="CC109BA452888C500BA610E82BADE41B" pageId="0" pageNumber="169">Legs. Yellow brown with darker annulations tibiae, metatarsi and tarsi darker, less annulated (Fig. 9A).</paragraph>
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<paragraph id="CF3C91E9D607F2E4E8457CEBEFC7416E" pageId="0" pageNumber="169">Opisthosoma. Length 1.8, width 1.3; yellow-brown with light yellow-brown anterior cardiac mark covered with white setae, and dark grey irregular pattern mainly in posterior half (Fig. 9A). Venter yellow-brown with darker pattern (Fig. 9A); spinnerets dark grey.</paragraph>
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<paragraph id="35D025442576060916437CECDF84F4FF" pageId="0" pageNumber="169">Pedipalps. Tibia globular, as long as broad; Cymbium tip without macrosetae (Fig. 9E, F); dorsal scopula patch absent; tegular apophysis long, nearly rectangular distally slightly scooped, basally narrowed to 2/3, retrolateral tip pointed, not reaching margin of cymbium (Fig. 9E); basoembolic apophysis nearly twice as long as broad, broadly rectangular indented medially; embolus broad, widely semicircular; terminal apophysis broad, tip pointed (Fig. 47C).</paragraph>
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<paragraph id="A15D2429B88C3F413421ACE8146E8C84" pageId="0" pageNumber="169">Female (based on paratype, AM KS73425).</paragraph>
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<paragraph id="BF5F38A14BB06B55D8F1D8E521C6C6D3" pageId="0" pageNumber="169">Total length 3.4.</paragraph>
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<paragraph id="C6F043E6A83E6F129B777F9A43DFA651" pageId="0" pageNumber="169">Prosoma. Length 1.9, width 1.3; carapace and sternum colouration as male, although somewhat lighter (Fig. 9C, D).</paragraph>
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<paragraph id="FEA51F55443FF29A740A90EB616D2864" pageId="0" pageNumber="169">Eyes. Diameter of AME 0.05, ALE 0.04, PME 0.19, PLE 0.16.</paragraph>
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<paragraph id="BFA8BFB97E94C0498E26053943CF0F03" pageId="0" pageNumber="169">Anterior eye row. Strongly procurved, evenly spaced.</paragraph>
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<paragraph id="AB00F7E27C92E428DEAB2A8AAB45695B" pageId="0" pageNumber="169">Chelicerae, labium, pedipalp coxae, legs and opisthosoma. Opisthosoma length 1.9, width 1.2; otherwise as male, but legs I not darker and opisthosoma pattern more obscure (opisthosoma deflated due to preservation) (Fig. 9C, D).</paragraph>
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<paragraph id="06115794EE1AD8FD13D8C480241EC880" pageId="0" pageNumber="169">Epigyne slightly wider than long, obscure, with heavily sclerotized posterior tips (Fig. 9G); spermathecal heads ovoid about 1/2 of diameter apart, spermathecal stalks attached postero-laterally and slightly S-shaped (Fig. 9H).</paragraph>
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<caption id="734DE3CC3D1E4FEBB3F75C24C1DA4D3A" pageId="0" pageNumber="169">
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<paragraph id="7C1CA556ED9C5FA27FC15C8A253DFBE2" pageId="0" pageNumber="169">
|
||
Figure 9.
|
||
<taxonomicName id="A0F44AB6330F282C6D69D944725C0353" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria belfordensis" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="belfordensis">Artoria belfordensis</taxonomicName>
|
||
sp. n., male holotype (AM KS127759), female paratype (AM KS73425): A, male habitus, dorsal view; B, male habitus, ventral view; C, female habitus, dorsal view; D, female habitus, ventral view; E, male pedipalp, ventral view; F, male pedipalp, retrolateral view; G, epigyne, ventral view; H, epigyne, dorsal view. Scale bars: habitus 1.0 mm; pedipalp, epigyne 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="EA14525CDB05E6BD429C0B45D0067C9E" pageId="0" pageNumber="169" type="life history and habitat preferences">
|
||
<paragraph id="C3DEAC3DF05CB66823083337B3B7B16A" pageId="0" pageNumber="169">Life history and habitat preferences.</paragraph>
|
||
<paragraph id="31B7FA7CE64095F781E2F0ADDD2B7779" pageId="0" pageNumber="169">
|
||
<taxonomicName id="41FC569B769433426C687E25667420EB" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria belfordensis" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="belfordensis">Artoria belfordensis</taxonomicName>
|
||
sp. n. was collected in undisturbed or moderately disturbed forest. Mature males were collected in June, October and November and the single female in October, indicating winter to spring maturity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D463A9C6CB2493E289A115E920BE8408" pageId="0" pageNumber="169" type="distribution">
|
||
<paragraph id="29541A27E40CC2F3EC038BF9B6C3FD8D" pageId="0" pageNumber="169">Distribution.</paragraph>
|
||
<paragraph id="C70EAEAAA210C2A47F19F9A94445E353" pageId="0" pageNumber="169">This species is known from central eastern NSW, principally the Sydney Basin (SYB) and NSW North Coast (NNC) IBRA regions (Fig. 8).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |