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<document id="051DE47F9ED3F90D9566763D3A322CCE" ID-DOI="10.1111/j.1096-3642.2008.00440.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5445984" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630459985039" checkinUser="felipe" docAuthor="Inger, Robert F., Stuart, Bryan L. &amp; Iskandar, Djoko T." docDate="2009" docId="F8158E66D4751437399AFD1C3C2E4958" docLanguage="en" docName="j.1096-3642.2008.00440.x.pdf" docOrigin="Zoological Journal of the Linnean Society 155 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00440.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Rana eschatia Inger &amp; Stuart &amp; Iskandar 2009, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="135" masterDocId="042CF61ED47E143B3A4DFFC9386C4A7A" masterDocTitle="Systematics of a widespread Southeast Asian frog, Rana chalconota (Amphibia: Anura: Ranidae)" masterLastPageNumber="147" masterPageNumber="123" pageNumber="134" updateTime="1699278897583" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="10B7337BDA6D9687F9F907CC821204CC">Systematics of a widespread Southeast Asian frog, Rana chalconota (Amphibia: Anura: Ranidae)</mods:title>
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<mods:namePart id="C21B8B91A711D694FC8439F6CC7FB545">Inger, Robert F.</mods:namePart>
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<mods:namePart id="B20FE9796450D67EFAB1EDCB2CCD8623">Stuart, Bryan L.</mods:namePart>
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<mods:namePart id="A6526CE118C2604BCF2E8ECBC4FAE380">Iskandar, Djoko T.</mods:namePart>
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<mods:date id="DC71E2795D8146B9BAC0BAC07D49360B">2009</mods:date>
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<heading id="2B4B881CD4751430399AFD1C3D684890" box="[983,1284,725,746]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="134" reason="2">
<taxonomicName id="B7BC44F3D4751430399AFD1C3CFA4893" ID-CoL="4RDPH" authority="Inger &amp; Stuart &amp; Iskandar, 2009" authorityName="Inger &amp; Stuart &amp; Iskandar" authorityYear="2009" box="[983,1174,725,746]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="eschatia" status="sp. nov.">RANA ESCHATIA</taxonomicName>
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<taxonomicNameLabel id="59FB5E19D47514303ED2FD113D684890" box="[1183,1284,725,746]" pageId="11" pageNumber="134" rank="species">SP. NOV.</taxonomicNameLabel>
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<paragraph id="70033F70D4751430391EFD323DEB496B" blockId="11.[851,1415,763,785]" box="[851,1415,763,785]" pageId="11" pageNumber="134">
(Previously referred to as
<collectingCountry id="08AB7FE0D47514303ECBFD323C81496B" box="[1158,1261,763,785]" name="Thailand" pageId="11" pageNumber="134">Thailand</collectingCountry>
morphotype)
</paragraph>
</subSubSection>
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<paragraph id="70033F70D47514303977FCE33CC34945" blockId="11.[826,1199,809,831]" box="[826,1199,809,831]" pageId="11" pageNumber="134">
<treatmentCitationGroup id="50AC185ED47514303977FCE33CC34945" box="[826,1199,809,831]" pageId="11" pageNumber="134">
<treatmentCitation id="F11D1961D47514303977FCE33CC74945" author="Smith MA" box="[826,1195,809,831]" page="168" pageId="11" pageNumber="134" year="1916">
<taxonomicName id="B7BC44F3D47514303977FCE33CC74945" ID-CoL="7935R" authority="Smith, 1916: 168" authorityName="Smith" authorityPageNumber="168" authorityYear="1916" box="[826,1195,809,831]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="labialis">
<emphasis id="42C8E362D47514303977FCE33BB44944" box="[826,984,809,831]" italics="true" pageId="11" pageNumber="134">Rana labialis</emphasis>
<bibRefCitation id="142D4281D475143039ACFCE03CC74945" author="Smith MA" box="[993,1195,809,831]" pageId="11" pageNumber="134" pagination="148 - 171" refId="ref17175" refString="Smith MA. 1916. On a collection of reptiles and batrachians from peninsular Thailand. Journal of Natural History Society of Siam 2: 148 - 171." type="journal article" year="1916">Smith, 1916: 168</bibRefCitation>
</taxonomicName>
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.
</treatmentCitationGroup>
</paragraph>
<paragraph id="70033F70D47514303977FC9E3CBF4916" blockId="11.[826,1235,855,877]" box="[826,1235,855,877]" pageId="11" pageNumber="134">
<treatmentCitationGroup id="50AC185ED47514303977FC9E3CBF4916" box="[826,1235,855,877]" pageId="11" pageNumber="134">
<treatmentCitation id="F11D1961D47514303977FC9E3CA34916" author="Smith MA" box="[826,1231,855,877]" page="109" pageId="11" pageNumber="134" year="1930">
<taxonomicName id="B7BC44F3D47514303977FC9E3CA34916" ID-CoL="4RDLG" authority="Smith, 1930: 109" authorityName="Smith" authorityPageNumber="109" authorityYear="1930" box="[826,1231,855,877]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="42C8E362D47514303977FC9E3B904916" box="[826,1020,855,876]" italics="true" pageId="11" pageNumber="134">Rana chalconota</emphasis>
<bibRefCitation id="142D4281D47514303E48FC9E3CA34916" author="Smith MA" box="[1029,1231,855,877]" pageId="11" pageNumber="134" pagination="1 - 149" refId="ref17204" refString="Smith MA. 1930. The Reptilia and Amphibia of the Malay Peninsula. Bulletin of the Raffles Museum 3: 1 - 149." type="journal article" year="1930">Smith, 1930: 109</bibRefCitation>
</taxonomicName>
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.
</treatmentCitationGroup>
</paragraph>
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<emphasis id="42C8E362D47514303977FC583BF149DC" box="[826,925,913,934]" italics="true" pageId="11" pageNumber="134">
<typeStatus id="AF0781D2D47514303977FC583BF149DC" box="[826,925,913,934]" pageId="11" pageNumber="134" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="70033F70D47514303977FC793BC24E25" blockId="11.[826,1441,913,1119]" pageId="11" pageNumber="134">
<materialsCitation id="C0D4352DD47514303977FC793BC24E25" collectingDate="2004-11-26" collectionCode="THNHM" country="Thailand" elevation="1" latitude="9.933333" location="Ngao Falls National Park" longLatPrecision="1298" longitude="98.71667" pageId="11" pageNumber="134" specimenCode="THNHM 05677" specimenCount="1" stateProvince="Ranong" typeStatus="holotype">
<specimenCode id="201A970BD47514303977FC793B8649BC" box="[826,1002,944,966]" collectionCode="THNHM" pageId="11" pageNumber="134">THNHM 05677</specimenCode>
(field number 66721), an adult female from
<location id="756369ABD475143039CFFC063CA7499E" LSID="urn:lsid:plazi:treatment:F8158E66D4751437399AFD1C3C2E4958:756369ABD475143039CFFC063CA7499E" box="[898,1227,974,996]" country="Thailand" latitude="9.933333" longLatPrecision="1298" longitude="98.71667" name="Ngao Falls National Park" pageId="11" pageNumber="134" stateProvince="Ranong">Ngao Falls National Park</location>
(
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/
<geoCoordinate id="158859B7D47514303F71FC073DF8499E" box="[1340,1428,973,996]" degrees="98" direction="east" minutes="43" orientation="longitude" pageId="11" pageNumber="134" precision="925" value="98.71667">98°43E</geoCoordinate>
),
<collectingRegion id="B278F192D47514303977FC243C6B4E79" box="[826,1031,1005,1027]" country="Thailand" name="Ranong" pageId="11" pageNumber="134">Ranong Province</collectingRegion>
,
<collectingCountry id="08AB7FE0D47514303E56FC243CEA4E79" box="[1051,1158,1005,1027]" name="Thailand" pageId="11" pageNumber="134">Thailand</collectingCountry>
. Collected on a gravel bank
<quantity id="B7449295D47514303933FBC53BD24E5B" box="[894,958,1036,1057]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="134" unit="m" value="0.1">
<elevation id="FB91D843D47514303933FBC53BD24E5B" box="[894,958,1036,1057]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="134" unit="m" value="0.1">0.1 m</elevation>
</quantity>
from the edge of a stream in secondary forest,
<date id="040219B0D475143039C2FBE33C6B4E3A" box="[911,1031,1066,1088]" pageId="11" pageNumber="134" value="2004-11-26">
<collectingDate id="1446E058D475143039C2FBE33C6B4E3A" box="[911,1031,1066,1088]" pageId="11" pageNumber="134" value="2004-11-26">26.xi.2004</collectingDate>
</date>
, by Jennifer Sheridan and Tanya Chan-ard.
</materialsCitation>
</paragraph>
<paragraph id="70033F70D47514303977FB4D3BC74EE3" blockId="11.[826,1441,1156,1299]" box="[826,939,1156,1177]" pageId="11" pageNumber="134">
<emphasis id="42C8E362D47514303977FB4D3BC74EE3" box="[826,939,1156,1177]" italics="true" pageId="11" pageNumber="134">
<typeStatus id="AF0781D2D47514303977FB4D3BC74EE3" box="[826,939,1156,1177]" pageId="11" pageNumber="134" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="70033F70D47514303977FB6B3D4B4F69" blockId="11.[826,1441,1156,1299]" pageId="11" pageNumber="134">
From the
<typeStatus id="AF0781D2D475143039F6FB6B3B804ECD" box="[955,1004,1186,1207]" pageId="11" pageNumber="134">type</typeStatus>
locality FMNH 268523, 268526-28, 268530 (adult males with nuptial pads) FMNH 268524, 268529 (adult females with convoluted oviducts), FMNH 268521, 268525 (juveniles).
</paragraph>
<paragraph id="70033F70D47514303977FAF13BDD4F37" blockId="11.[826,1441,1336,1450]" box="[826,945,1336,1357]" pageId="11" pageNumber="134">
<emphasis id="42C8E362D47514303977FAF13BDD4F37" box="[826,945,1336,1357]" italics="true" pageId="11" pageNumber="134">Etymology</emphasis>
</paragraph>
<paragraph id="70033F70D47514303977FA9F3BDE4FD3" blockId="11.[826,1441,1336,1450]" pageId="11" pageNumber="134">
Specific name from
<taxonomicName id="B7BC44F3D47514303E57FA9F3CC64F11" authority=", Gr." authorityName="Gr." box="[1050,1194,1366,1388]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="eschatia">
<emphasis id="42C8E362D47514303E57FA9F3C1B4F11" box="[1050,1143,1366,1387]" italics="true" pageId="11" pageNumber="134">eschatia</emphasis>
, Gr.
</taxonomicName>
, outskirt, referring to distribution at the edge of the geographical range of the group.
</paragraph>
<paragraph id="70033F70D47514303977FA073C694F99" blockId="11.[826,1441,1486,1783]" box="[826,1029,1486,1507]" pageId="11" pageNumber="134">
<emphasis id="42C8E362D47514303977FA073C694F99" box="[826,1029,1486,1507]" italics="true" pageId="11" pageNumber="134">Referred material</emphasis>
</paragraph>
<paragraph id="70033F70D47514303977FA253B164C8D" blockId="11.[826,1441,1486,1783]" pageId="11" pageNumber="134">
<materialsCitation id="C0D4352DD47514303977FA253BA64C45" collectionCode="FMNH" country="Thailand" latitude="8.5" location="Khao Luang National Park" longLatPrecision="1301" longitude="99.75" pageId="11" pageNumber="134" specimenCode="FMNH 268852-54, 268856-57" specimenCount="1" stateProvince="Nakhon Si Thammarat">
<emphasis id="42C8E362D47514303977FA253BC24C78" bold="true" box="[826,942,1516,1538]" pageId="11" pageNumber="134">
<collectingCountry id="08AB7FE0D47514303977FA253BC24C78" box="[826,942,1516,1538]" name="Thailand" pageId="11" pageNumber="134">Thailand</collectingCountry>
</emphasis>
:
<specimenCode id="201A970BD475143039F1FA243CE74C78" box="[956,1163,1516,1538]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="134" type="Museum">FMNH 268852-54</specimenCode>
,
<specimenCode id="201A970BD47514303EDAFA253D614C78" box="[1175,1293,1516,1538]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="134" type="Museum">268856-57</specimenCode>
<location id="756369ABD47514303F59FA253B894C5B" LSID="urn:lsid:plazi:treatment:F8158E66D4751437399AFD1C3C2E4958:756369ABD47514303F59FA253B894C5B" country="Thailand" latitude="8.5" longLatPrecision="1301" longitude="99.75" name="Khao Luang National Park" pageId="11" pageNumber="134" stateProvince="Nakhon Si Thammarat">Khao Luang National Park</location>
(
<geoCoordinate id="158859B7D475143039B1F9C23C254C5B" box="[1020,1097,1546,1569]" degrees="8" direction="north" minutes="30" orientation="latitude" pageId="11" pageNumber="134" precision="925" value="8.5">8°30N</geoCoordinate>
/
<geoCoordinate id="158859B7D47514303E1FF9C23CC64C5B" box="[1106,1194,1546,1569]" degrees="99" direction="east" minutes="45" orientation="longitude" pageId="11" pageNumber="134" precision="925" value="99.75">99°45E</geoCoordinate>
),
<collectingRegion id="B278F192D47514303E8BF9C23BA64C45" country="Thailand" name="Nakhon Si Thammarat" pageId="11" pageNumber="134">Nakhon Si Thammarat Prov.</collectingRegion>
</materialsCitation>
;
<materialsCitation id="C0D4352DD47514303996F9E33B1A4C06" collectionCode="FMNH" country="Thailand" latitude="8.233334" location="Khao Phanom Bencha National Park" longLatPrecision="0" longitude="99.0" pageId="11" pageNumber="134" specimenCode="FMNH 268858, 268860, 268869" specimenCount="1" stateProvince="Krabi">
<specimenCode id="201A970BD47514303996F9E33CE14C3A" box="[987,1165,1578,1600]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="134" type="Museum">FMNH 268858</specimenCode>
,
<specimenCode id="201A970BD47514303ED2F9E33C984C3A" box="[1183,1268,1578,1600]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="134" type="Museum">268860</specimenCode>
,
<specimenCode id="201A970BD47514303F48F9E33D3B4C3A" box="[1285,1367,1578,1600]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="134" type="Museum">268869</specimenCode>
<location id="756369ABD47514303F28F9E33CC54C24" LSID="urn:lsid:plazi:treatment:F8158E66D4751437399AFD1C3C2E4958:756369ABD47514303F28F9E33CC54C24" country="Thailand" latitude="8.233334" longLatPrecision="0" longitude="99.0" name="Khao Phanom Bencha National Park" pageId="11" pageNumber="134" stateProvince="Krabi">Khao Phanom Bencha National Park</location>
(
<geoCoordinate id="158859B7D47514303EF6F9813D654C24" box="[1211,1289,1607,1630]" degrees="8" direction="north" minutes="14" orientation="latitude" pageId="11" pageNumber="134" precision="925" value="8.233334">8°14N</geoCoordinate>
/
<geoCoordinate id="158859B7D47514303F5FF9813D254C24" box="[1298,1353,1608,1630]" degrees="99" direction="east" orientation="longitude" pageId="11" pageNumber="134" precision="55555" value="99.0">99°E</geoCoordinate>
),
<collectingRegion id="B278F192D47514303F2DF9813B1A4C06" country="Thailand" name="Krabi" pageId="11" pageNumber="134">Krabi Prov.</collectingRegion>
</materialsCitation>
;
<materialsCitation id="C0D4352DD475143039C9F9AE3D504CE1" collectionCode="FMNH" country="Thailand" latitude="8.933333" location="Khao Sok National Park" longLatPrecision="1300" longitude="98.566666" pageId="11" pageNumber="134" specimenCode="FMNH 268872, 268874-84" specimenCount="1" stateProvince="Surat Thani">
<specimenCode id="201A970BD475143039C9F9AE3C584C07" box="[900,1076,1639,1661]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="134" type="Museum">FMNH 268872</specimenCode>
,
<specimenCode id="201A970BD47514303E0FF9AE3CDB4C07" box="[1090,1207,1639,1661]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="134" type="Museum">268874-84</specimenCode>
<location id="756369ABD47514303E8FF9AE3B1E4CE1" LSID="urn:lsid:plazi:treatment:F8158E66D4751437399AFD1C3C2E4958:756369ABD47514303E8FF9AE3B1E4CE1" country="Thailand" latitude="8.933333" longLatPrecision="1300" longitude="98.566666" name="Khao Sok National Park" pageId="11" pageNumber="134" stateProvince="Surat Thani">Khao Sok National Park</location>
(
<geoCoordinate id="158859B7D475143039C6F94C3BB44CE1" box="[907,984,1668,1691]" degrees="8" direction="north" minutes="56" orientation="latitude" pageId="11" pageNumber="134" precision="925" value="8.933333">8°56N</geoCoordinate>
/
<geoCoordinate id="158859B7D475143039ACF94F3C564CE1" box="[993,1082,1668,1691]" degrees="98" direction="east" minutes="34" orientation="longitude" pageId="11" pageNumber="134" precision="925" value="98.566666">98°34E</geoCoordinate>
),
<collectingRegion id="B278F192D47514303E15F94F3D504CE1" box="[1112,1340,1669,1691]" country="Thailand" name="Surat Thani" pageId="11" pageNumber="134">Surat Thani Prov.</collectingRegion>
</materialsCitation>
;
<materialsCitation id="C0D4352DD47514303F1CF94F3C4B4CC0" collectionCode="FMNH" country="Thailand" pageId="11" pageNumber="134" specimenCode="FMNH 268531-4, 268536-9" specimenCount="1">
<specimenCode id="201A970BD47514303F1CF94F3BC94CC0" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="134" type="Museum">FMNH 268531-4</specimenCode>
,
<specimenCode id="201A970BD475143039F5F96D3C4F4CC0" box="[952,1059,1700,1722]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="134" type="Museum">268536-9</specimenCode>
,
</materialsCitation>
<materialsCitation id="C0D4352DD47514303E7BF96C3DCD4CA3" collectionCode="THNHM" country="Thailand" latitude="9.566667" location="Kaeng Krung National Park" longLatPrecision="1299" longitude="98.816666" pageId="11" pageNumber="134" specimenCode="THNHM 05690, 05695" specimenCount="1" stateProvince="Surat Thani">
<specimenCode id="201A970BD47514303E7BF96C3C9D4CC0" box="[1078,1265,1700,1722]" collectionCode="THNHM" pageId="11" pageNumber="134">THNHM 05690</specimenCode>
,
<specimenCode id="201A970BD47514303F49F96D3D244CC0" box="[1284,1352,1700,1722]" collectionCode="THNHM" pageId="11" pageNumber="134">05695</specimenCode>
<location id="756369ABD47514303F15F96C3C5A4CA3" LSID="urn:lsid:plazi:treatment:F8158E66D4751437399AFD1C3C2E4958:756369ABD47514303F15F96C3C5A4CA3" country="Thailand" latitude="9.566667" longLatPrecision="1299" longitude="98.816666" name="Kaeng Krung National Park" pageId="11" pageNumber="134" stateProvince="Surat Thani">Kaeng Krung National Park</location>
(
<geoCoordinate id="158859B7D47514303E07F90A3CFB4CA2" box="[1098,1175,1729,1752]" degrees="9" direction="north" minutes="34" orientation="latitude" pageId="11" pageNumber="134" precision="925" value="9.566667">9°34N</geoCoordinate>
/
<geoCoordinate id="158859B7D47514303EEDF90A3C944CA2" box="[1184,1272,1729,1752]" degrees="98" direction="east" minutes="49" orientation="longitude" pageId="11" pageNumber="134" precision="925" value="98.816666">98°49E</geoCoordinate>
),
<collectingRegion id="B278F192D47514303F5CF90A3DCD4CA3" box="[1297,1441,1731,1753]" country="Thailand" name="Surat Thani" pageId="11" pageNumber="134">Surat Thani</collectingRegion>
</materialsCitation>
Prov..
</paragraph>
<paragraph id="70033F70D47514303977F8D53BC64D4B" blockId="11.[826,1441,1820,1903]" box="[826,938,1820,1841]" pageId="11" pageNumber="134">
<emphasis id="42C8E362D47514303977F8D53BC64D4B" box="[826,938,1820,1841]" italics="true" pageId="11" pageNumber="134">Diagnosis</emphasis>
</paragraph>
<paragraph id="70033F70D47514373977F8F33A8D4B63" blockId="11.[826,1441,1820,1903]" lastBlockId="12.[144,759,198,281]" lastPageId="12" lastPageNumber="135" pageId="11" pageNumber="134">
A moderate-sized species of the
<taxonomicName id="B7BC44F3D47514303EE9F8F33D724D35" authorityName="Smith" authorityYear="1930" box="[1188,1310,1850,1871]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="42C8E362D47514303EE9F8F33D724D35" box="[1188,1310,1850,1871]" italics="true" pageId="11" pageNumber="134">chalconota</emphasis>
</taxonomicName>
group with males up to
<quantity id="B7449295D47514303998F8903C484D14" box="[981,1060,1881,1902]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="134" unit="mm" value="40.0">40 mm</quantity>
<collectionCode id="16ADA7B5D47514303E7DF8903C084D14" box="[1072,1124,1881,1902]" pageId="11" pageNumber="134">SVL</collectionCode>
, females up to
<quantity id="B7449295D47514303F69F8903D194D14" box="[1316,1397,1881,1903]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="11" pageNumber="134" unit="mm" value="57.0">57 mm</quantity>
, no dorsal spotting, relatively wide head (
<collectionCode id="16ADA7B5D4721437387AFF0F3A084AA1" box="[567,612,198,219]" pageId="12" pageNumber="135">HW</collectionCode>
/
<collectionCode id="16ADA7B5D47214373821FF0F3AF14AA1" box="[620,669,198,219]" pageId="12" pageNumber="135">SVL</collectionCode>
usually&gt; 0.305), relatively long leg (
<collectionCode id="16ADA7B5D47214373BABFF2C399A4A80" box="[486,502,229,250]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="135">T</collectionCode>
/
<collectionCode id="16ADA7B5D47214373BB2FF2C3A5D4A80" box="[511,561,229,250]" pageId="12" pageNumber="135">SVL</collectionCode>
usually&gt; 0.575) and males with constricted or divided nuptial pads.
</paragraph>
<paragraph id="70033F70D47214373ADDFE88397D4B2C" blockId="12.[144,759,321,1658]" box="[144,273,321,342]" pageId="12" pageNumber="135">
<emphasis id="42C8E362D47214373ADDFE88397D4B2C" box="[144,273,321,342]" italics="true" pageId="12" pageNumber="135">Description</emphasis>
</paragraph>
<paragraph id="70033F70D47214373ADDFE963884489E" blockId="12.[144,759,321,1658]" pageId="12" pageNumber="135">Habitus slender, head slightly wider than trunk, legs long. Head triangular; snout obtusely pointed, rounded in profile, projecting beyond lower jaw, longer than diameter of eye; nostril lateral, very close to tip of snout; canthus angular, not constricted; lores concave, vertical; interobital wider than upper eyelid and internarial; tympanum distinct, about two-thirds eye diameter in females, slightly larger in males, inside its rim the tympanum is slightly depressed relative to the surface of the temporal region; vomerine teeth in oblique groups, gap between groups less than length of one group and equal to distance from choana.</paragraph>
<paragraph id="70033F70D47214373AE5FD243A9B4F70" blockId="12.[144,759,321,1658]" pageId="12" pageNumber="135">Fingers long, third finger longer than snout; fingers without webbing; second and third fingers with narrow, movable fold of skin along medial margins; tips of three outer fingers with wide discs, that of third finger almost equal diameter of tympanum in female; disc of first finger much narrow than that of second; all discs with circummarginal grooves; subarticular tubercles conspicuous; third finger with two small supernumerary tubercles, bases of second and fourth fingers with a single supernumerary tubercle. Tips of toes expanded into discs smaller than those of outer fingers, but with circummarginal grooves; webbing extensive, reaching discs of first three toes on lateral margins and disc of fifth toe medially; fourth toe webbed to distal subarticular tubercle medially and slightly beyond that laterally; no dermal ridge along outer margins of first and fifth toes; a low oval inner metatarsal tubercle and a round outer one.</paragraph>
<paragraph id="70033F70D47214373AE5FADA3A6C4FFF" blockId="12.[144,759,321,1658]" pageId="12" pageNumber="135">Skin of back granular, in males granules weakly spinose; dorsolateral fold distinct, low; ventral surfaces smooth, except weakly rugose at rear of abdomen; rictal glands present.</paragraph>
<paragraph id="70033F70D47214373AE5FA4739144C46" blockId="12.[144,759,321,1658]" pageId="12" pageNumber="135">Colour in preservative dark brown dorsally and laterally; no black spots on dorsal surfaces; ventral surfaces cream-coloured or white; in some individuals throat with round dark spots; limbs without dark crossbars; rear of thigh dark brown with indistinct lighter round areas.</paragraph>
<paragraph id="70033F70D47214373AE5F98F3A3C4C03" blockId="12.[144,759,321,1658]" pageId="12" pageNumber="135">
Measurements (mm) of
<typeStatus id="AF0781D2D47214373BF1F98C3A4C4C21" box="[444,544,1605,1627]" pageId="12" pageNumber="135" type="holotype">holotype</typeStatus>
:
<collectionCode id="16ADA7B5D47214373860F98C3A334C20" box="[557,607,1605,1626]" pageId="12" pageNumber="135">SVL</collectionCode>
55.6,
<collectionCode id="16ADA7B5D472143738EBF98F3ADA4C21" box="[678,694,1606,1627]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="135">T</collectionCode>
31.0, HW 16.0, HL 19.8, TYM 4.6, DF3 3.2.
</paragraph>
<paragraph id="70033F70D47214373ADDF96B38964CCD" blockId="12.[144,759,1698,1903]" box="[144,250,1698,1719]" pageId="12" pageNumber="135">
<emphasis id="42C8E362D47214373ADDF96B38964CCD" box="[144,250,1698,1719]" italics="true" pageId="12" pageNumber="135">Variation</emphasis>
</paragraph>
<paragraph id="70033F70D47214373ADDF9093BCA4B42" blockId="12.[144,759,1698,1903]" lastBlockId="12.[806,1422,198,312]" pageId="12" pageNumber="135">
Adult females
<quantity id="B7449295D47214373B73F909398C4CAF" box="[318,480,1728,1750]" metricMagnitude="-2" metricUnit="m" metricValue="4.97" metricValueMax="5.66" metricValueMin="4.279999999999999" pageId="12" pageNumber="135" unit="mm" value="49.7" valueMax="56.6" valueMin="42.8">42.856.6 mm</quantity>
, mean 47.57 ±
<quantity id="B7449295D472143738D9F9093A9B4CAF" box="[660,759,1728,1750]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="12" pageNumber="135" unit="mm" value="0.96">0.96 mm</quantity>
(
<emphasis id="42C8E362D47214373AD5F91638C04C8E" box="[152,172,1759,1780]" italics="true" pageId="12" pageNumber="135">N</emphasis>
= 17), males
<quantity id="B7449295D47214373B09F917398A4C8E" box="[324,486,1758,1780]" metricMagnitude="-2" metricUnit="m" metricValue="3.51" metricValueMax="3.96" metricValueMin="3.06" pageId="12" pageNumber="135" unit="mm" value="35.1" valueMax="39.6" valueMin="30.6">30.639.6 mm</quantity>
, mean 34.14 ±
<quantity id="B7449295D472143738D9F9173A9B4C8E" box="[660,759,1758,1780]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="12" pageNumber="135" unit="mm" value="0.39">0.39 mm</quantity>
(
<emphasis id="42C8E362D47214373AD5F93738C04D69" box="[152,172,1790,1811]" italics="true" pageId="12" pageNumber="135">N</emphasis>
= 22). Variation in body proportions given in
<tableCitation id="3D3E0ACBD47214373ADDF8D538894D4B" box="[144,229,1820,1842]" captionStart="Table 8" captionStartId="10.[144,208,200,219]" captionText="Table 8. Comparison of southern Thailand frogs with Rana labialis from Selangor, Peninsular Malaysia" pageId="12" pageNumber="135">Table 8</tableCitation>
. Relative tympanum diameter in females
<geoCoordinate id="158859B7D47214373ADDF8F338A34D2A" box="[144,207,1850,1872]" degrees="0.077" direction="north" orientation="latitude" pageId="12" pageNumber="135" precision="55" value="0.077">0.077</geoCoordinate>
<geoCoordinate id="158859B7D47214373A83F8F339774D2A" box="[206,283,1850,1872]" degrees="0.095" direction="west" orientation="longitude" pageId="12" pageNumber="135" precision="55" value="-0.095">0.095</geoCoordinate>
, in males
<geoCoordinate id="158859B7D47214373BDAF8F339BA4D2A" box="[407,470,1850,1872]" degrees="0.096" direction="north" orientation="latitude" pageId="12" pageNumber="135" precision="55" value="0.096">0.096</geoCoordinate>
<geoCoordinate id="158859B7D47214373B9BF8F33A484D2A" box="[470,548,1850,1872]" degrees="0.127" direction="west" orientation="longitude" pageId="12" pageNumber="135" precision="55" value="-0.127">0.127</geoCoordinate>
. The sexes do not differ in relative head width; HW/
<collectionCode id="16ADA7B5D47214373877F8903A074D14" box="[570,619,1881,1902]" pageId="12" pageNumber="135">SVL</collectionCode>
in females
<geoCoordinate id="158859B7D4721437396BFF0F3B0F4AA6" box="[806,867,198,220]" degrees="0.288" direction="north" orientation="latitude" pageId="12" pageNumber="135" precision="55" value="0.288">0.288</geoCoordinate>
<geoCoordinate id="158859B7D47214373929FF0F3BC34AA6" box="[868,943,198,220]" degrees="0.325" direction="west" orientation="longitude" pageId="12" pageNumber="135" precision="55" value="-0.325">0.325</geoCoordinate>
(median 0.308), in males
<geoCoordinate id="158859B7D47214373F49FF0F3D2D4AA6" box="[1284,1345,198,220]" degrees="0.275" direction="north" orientation="latitude" pageId="12" pageNumber="135" precision="55" value="0.275">0.275</geoCoordinate>
<geoCoordinate id="158859B7D47214373F0CFF0F3DE14AA6" box="[1345,1421,198,220]" degrees="0.326" direction="west" orientation="longitude" pageId="12" pageNumber="135" precision="55" value="-0.326">0.326</geoCoordinate>
(median 0.310). All males have constricted or divided nuptial pads and vocal sac openings at the corners of the mouth.
</paragraph>
<paragraph id="70033F70D4721437396BFEA83BD74B0C" blockId="12.[806,1421,353,803]" box="[806,955,353,374]" pageId="12" pageNumber="135">
<emphasis id="42C8E362D4721437396BFEA83BD74B0C" box="[806,955,353,374]" italics="true" pageId="12" pageNumber="135">Comparisons</emphasis>
</paragraph>
<paragraph id="70033F70D4721437396BFEB63C2E4958" blockId="12.[806,1421,353,803]" pageId="12" pageNumber="135">
Males of
<taxonomicName id="B7BC44F3D472143739D5FE493C524BEE" box="[920,1086,383,405]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="eschatia">
<emphasis id="42C8E362D472143739D5FE493C524BEE" box="[920,1086,383,405]" italics="true" pageId="12" pageNumber="135">Rana eschatia</emphasis>
</taxonomicName>
are most similar in size to those of
<taxonomicName id="B7BC44F3D472143739D5FE573CF14BC9" authority="(Selangor)" authorityName="Smith" authorityYear="1916" baseAuthorityName="Selangor" box="[920,1181,413,435]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="labialis">
<emphasis id="42C8E362D472143739D5FE573C744BC8" box="[920,1048,413,435]" italics="true" pageId="12" pageNumber="135">R. labialis</emphasis>
(
<collectingRegion id="B278F192D47214373E7DFE543CFA4BC9" box="[1072,1174,413,435]" country="Malaysia" name="Selangor" pageId="12" pageNumber="135">Selangor</collectingRegion>
)
</taxonomicName>
and
<taxonomicName id="B7BC44F3D47214373EA4FE573DE14BC8" authorityName="Inger &amp; Stuart &amp; Iskandar" authorityYear="2009" box="[1257,1421,413,435]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="parvaccola">
<emphasis id="42C8E362D47214373EA4FE573DE14BC8" box="[1257,1421,413,435]" italics="true" pageId="12" pageNumber="135">R. parvaccola</emphasis>
</taxonomicName>
(
<tableCitation id="3D3E0ACBD47214373963FE753BEF4BA8" box="[814,899,444,466]" captionStart="Table 2" captionStartId="4.[144,208,200,219]" captionText="Table 2. Summary of mensural features of morphotypes of the Rana chalconota species complex" pageId="12" pageNumber="135">Table 2</tableCitation>
), but females of
<taxonomicName id="B7BC44F3D47214373E2FFE753CD34BAB" box="[1122,1215,444,465]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="eschatia">
<emphasis id="42C8E362D47214373E2FFE753CD34BAB" box="[1122,1215,444,465]" italics="true" pageId="12" pageNumber="135">eschatia</emphasis>
</taxonomicName>
are larger than females of those two (
<tableCitation id="3D3E0ACBD47214373E6BFE123CE94B8A" box="[1062,1157,475,497]" captionStart="Table 2" captionStartId="4.[144,208,200,219]" captionText="Table 2. Summary of mensural features of morphotypes of the Rana chalconota species complex" pageId="12" pageNumber="135">Tables 2</tableCitation>
and
<tableCitation id="3D3E0ACBD47214373E8EFE123CBF4B8B" box="[1219,1235,475,497]" captionStart="Table 6" captionStartId="8.[144,208,200,219]" captionText="Table 6. P values of morphological comparisons between clades of the Rana chalconota species complex" pageId="12" pageNumber="135">6</tableCitation>
).
<taxonomicName id="B7BC44F3D47214373EA4FE123DE14B8A" box="[1257,1421,475,496]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="eschatia">
<emphasis id="42C8E362D47214373EA4FE123DE14B8A" box="[1257,1421,475,496]" italics="true" pageId="12" pageNumber="135">Rana eschatia</emphasis>
</taxonomicName>
also differs from those two species in the absence of black spots on the back and in having a wider head in both sexes (
<tableCitation id="3D3E0ACBD472143739FEFDFF3C7E4836" box="[947,1042,566,588]" captionStart="Table 2" captionStartId="4.[144,208,200,219]" captionText="Table 2. Summary of mensural features of morphotypes of the Rana chalconota species complex" pageId="12" pageNumber="135">Tables 2</tableCitation>
and
<tableCitation id="3D3E0ACBD47214373E1FFDFF3C0E4836" box="[1106,1122,566,588]" captionStart="Table 6" captionStartId="8.[144,208,200,219]" captionText="Table 6. P values of morphological comparisons between clades of the Rana chalconota species complex" pageId="12" pageNumber="135">6</tableCitation>
).
<taxonomicName id="B7BC44F3D47214373E37FDFE3D734831" box="[1146,1311,566,588]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="eschatia">
<emphasis id="42C8E362D47214373E37FDFE3D734831" box="[1146,1311,566,588]" italics="true" pageId="12" pageNumber="135">Rana eschatia</emphasis>
</taxonomicName>
is larger than
<taxonomicName id="B7BC44F3D47214373925FD9F3B814811" authorityName="Peters" authorityYear="1871" box="[872,1005,598,619]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="raniceps">
<emphasis id="42C8E362D47214373925FD9F3B814811" box="[872,1005,598,619]" italics="true" pageId="12" pageNumber="135">R. raniceps</emphasis>
</taxonomicName>
(both sexes) and has a longer leg (
<collectionCode id="16ADA7B5D47214373960FDBD3B5148F0" box="[813,829,628,650]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="135">T</collectionCode>
/
<collectionCode id="16ADA7B5D4721437390BFDBD3B1548F0" box="[838,889,628,650]" pageId="12" pageNumber="135">SVL</collectionCode>
) and a smaller tympanum in males (
<tableCitation id="3D3E0ACBD47214373F63FDBD3DE148F3" box="[1326,1421,628,650]" captionStart="Table 2" captionStartId="4.[144,208,200,219]" captionText="Table 2. Summary of mensural features of morphotypes of the Rana chalconota species complex" pageId="12" pageNumber="135">Tables 2</tableCitation>
and
<tableCitation id="3D3E0ACBD47214373914FD5B3B0548D2" box="[857,873,658,680]" captionStart="Table 6" captionStartId="8.[144,208,200,219]" captionText="Table 6. P values of morphological comparisons between clades of the Rana chalconota species complex" pageId="12" pageNumber="135">6</tableCitation>
).
<taxonomicName id="B7BC44F3D47214373932FD5A3C4D48DD" box="[895,1057,658,680]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="eschatia">
<emphasis id="42C8E362D47214373932FD5A3C4D48DD" box="[895,1057,658,680]" italics="true" pageId="12" pageNumber="135">Rana eschatia</emphasis>
</taxonomicName>
is smaller than
<taxonomicName id="B7BC44F3D47214373EACFD5A3DEA48DD" authorityName="Inger &amp; Stuart &amp; Iskandar" authorityYear="2009" box="[1249,1414,658,680]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="megalonesa">
<emphasis id="42C8E362D47214373EACFD5A3DEA48DD" box="[1249,1414,658,680]" italics="true" pageId="12" pageNumber="135">R. megalonesa</emphasis>
</taxonomicName>
,
<taxonomicName id="B7BC44F3D4721437396BFD783BF748BC" authorityName="Inger &amp; Stuart &amp; Iskandar" authorityYear="2009" box="[806,923,689,710]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="rufipes">
<emphasis id="42C8E362D4721437396BFD783BF748BC" box="[806,923,689,710]" italics="true" pageId="12" pageNumber="135">R. rufipes</emphasis>
</taxonomicName>
and
<taxonomicName id="B7BC44F3D472143739AEFD783CE948BC" authorityName="Smith" authorityYear="1930" box="[995,1157,689,710]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="42C8E362D472143739AEFD783CE948BC" box="[995,1157,689,710]" italics="true" pageId="12" pageNumber="135">R. chalconota</emphasis>
</taxonomicName>
and differs from the latter two in the length of the tibia (
<collectionCode id="16ADA7B5D47214373EAEFD193C9F489C" box="[1251,1267,720,742]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="135">T</collectionCode>
/
<collectionCode id="16ADA7B5D47214373EB6FD193D42489C" box="[1275,1326,720,742]" pageId="12" pageNumber="135">SVL</collectionCode>
) and in the form of the nuptial pad, which is constricted or divided only in
<taxonomicName id="B7BC44F3D47214373993FCC43C574958" box="[990,1083,781,802]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="eschatia">
<emphasis id="42C8E362D47214373993FCC43C574958" box="[990,1083,781,802]" italics="true" pageId="12" pageNumber="135">eschatia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>