154 lines
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154 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.443.7880" ID-GBIF-Dataset="33ef48a3-7145-4103-9159-e11be2ba9adc" ID-PMC="PMC4205504" ID-Pensoft-Pub="1313-2970-443-61" ID-PubMed="25349497" ID-ZBK="E33841396A6E426C840D85BC32A12E78" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-443-61" ModsDocOrigin="ZooKeys 443" ModsDocTitle="The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation" checkinTime="1451245273153" checkinUser="pensoft" docAuthor="Sokolov, Igor M. & Kavanaugh, David H." docDate="2014" docId="F81585EF8485CBC080CA35D5B4EBA803" docLanguage="en" docName="ZooKeys 443: 61-118" docOrigin="ZooKeys 443" docSource="http://dx.doi.org/10.3897/zookeys.443.7880" docTitle="Geocharidius longinoi Sokolov & Kavanaugh, 2014, sp. n." docType="treatment" docUuid="2037D9B1-5260-4836-8ED4-0D0AE950111F" docUuidSource="ZooBank" docVersion="4" lastPageNumber="79" masterDocId="3546FFA8CF78F064801706227F76A615" masterDocTitle="The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation" masterLastPageNumber="118" masterPageNumber="61" pageNumber="79" updateTime="1668159453511" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sokolov, Igor M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kavanaugh, David H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>443</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>61</mods:start>
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<mods:end>118</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.443.7880</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.443.7880</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-443-61</mods:identifier>
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<mods:identifier type="ZBK">E33841396A6E426C840D85BC32A12E78</mods:identifier>
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<mods:identifier type="ZooBank">E33841396A6E426C840D85BC32A12E78</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152055685" LSID="urn:lsid:zoobank.org:act:2037D9B1-5260-4836-8ED4-0D0AE950111F" httpUri="http://treatment.plazi.org/id/F81585EF8485CBC080CA35D5B4EBA803" lastPageNumber="79" pageId="18" pageNumber="79">
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<subSubSection pageId="18" pageNumber="79" type="multiple">
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<paragraph pageId="18" pageNumber="79">Taxon classification Animalia Coleoptera Carabidae</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="79" type="nomenclature">
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<paragraph pageId="18" pageNumber="79">
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<taxonomicName LSID="http://zoobank.org/2037D9B1-5260-4836-8ED4-0D0AE950111F" class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius longinoi" order="Coleoptera" pageId="18" pageNumber="79" phylum="Arthropoda" rank="species" species="longinoi">Geocharidius longinoi</taxonomicName>
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<taxonomicNameLabel pageId="18" pageNumber="79">sp. n.</taxonomicNameLabel>
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Figs 5E, 7C, G, 12C, 13
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<normalizedToken originalValue="R–T">R-T</normalizedToken>
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, 16F, 17F, 22
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="79" type="type material">
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<paragraph pageId="18" pageNumber="79">Type material.</paragraph>
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<paragraph pageId="18" pageNumber="79">
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HOLOTYPE, a male, in KUNHM, point-mounted, labeled: \ GUATEMALA: El Progreso: Cerro
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<normalizedToken originalValue="Pinalón">Pinalon</normalizedToken>
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, 15.08392-89.93013
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<normalizedToken originalValue="±">+/-</normalizedToken>
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55m, 2750m, 1-V-2009 ex. sifted leaf litter, cloud forest, LLAMA09 Wm-B-01-1-04 \ KUNHM \ HOLOTYPE
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius longinoi" order="Coleoptera" pageId="18" pageNumber="79" phylum="Arthropoda" rank="species" species="longinoi">Geocharidius longinoi</taxonomicName>
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Sokolov and Kavanaugh 2014 [red label] \. PARATYPES: A total of 13 specimens (2 males and 1 female were dissected), deposited in CAS, CMNC and KUNHM; 5 specimens labeled same as holotype; 7 specimens labeled: \ GUATEMALA: El Progreso: Cerro
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<normalizedToken originalValue="Pinalón">Pinalon</normalizedToken>
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, 15.08411-89.93239
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<normalizedToken originalValue="±">+/-</normalizedToken>
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57m, 2715m, 1-V-2009 ex. sifted leaf litter, cloud forest, LLAMA09 Wm-B-01-1-05 \ KUNHM \; 1 specimen labeled: \ GUAT.: EL PROGRESO: 19.6km.N.Estancia de la Virgen, 2000m, Finca la Illuciones, 24.VI.1993, R.Anderson, cloud for. litter, 93-13C \ CMNC \.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="79" type="type locality">
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<paragraph pageId="18" pageNumber="79">Type locality.</paragraph>
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<paragraph pageId="18" pageNumber="79">
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Guatemala, El Progreso Department, Cerro
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<normalizedToken originalValue="Pinalón">Pinalon</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="79" type="etymology">
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<paragraph pageId="18" pageNumber="79">Etymology.</paragraph>
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<paragraph pageId="18" pageNumber="79">
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The specific epithet is a Latinized eponym in the genitive case, and is based on the surname of John T. (Jack) Longino, Professor of the Biology Department of the University of Utah, and one of
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<normalizedToken originalValue="Co-PI’s">Co-PI's</normalizedToken>
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of the LLAMA project, which provided the material on which the description of this species is based.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="79" type="recognition">
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<paragraph pageId="18" pageNumber="79">Recognition.</paragraph>
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<paragraph pageId="18" pageNumber="79">
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Adults of this new species are distinguished from those of other members of the integripennis species group by the large size, distinctive shape of the pronotum with very wide basal margin, and the proepisternum with evident microsculpture. Males and females of
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius longinoi" order="Coleoptera" pageId="18" pageNumber="79" phylum="Arthropoda" rank="species" species="longinoi">Geocharidius longinoi</taxonomicName>
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are distinguished from those of other members of the integripennis species group by the structure of the median lobe and the shape of spermatheca, respectively.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="79" type="description">
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<paragraph pageId="18" pageNumber="79">Description.</paragraph>
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<paragraph pageId="18" pageNumber="79">
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Size. Medium to large for genus (SBL range 1.34-1.51 mm, mean 1.41
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.071mm, n=12).
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</paragraph>
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<paragraph pageId="18" pageNumber="79">
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Habitus. Body form (Fig. 12C) moderately convex, elongate ovoid, general proportions (WE/SBL 0.38
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.008) and proportions of head (WH/WPm 0.71
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.012) average for group, pronotum markedly wide compared to elytra (WPm/WE 0.80
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.013).
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</paragraph>
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<paragraph pageId="18" pageNumber="79">Color. Body brunneorufous, appendages testaceous.</paragraph>
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<paragraph pageId="18" pageNumber="79">Microsculpture. Mesh pattern of irregularly isodiametric sculpticells present over all dorsal surfaces of head and elytra. Pronotum smooth (without evident microsculpture). Proepisternum with evident microsculpture.</paragraph>
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<paragraph pageId="18" pageNumber="79">Mouthparts. Maxillae and labium (Fig. 5E).</paragraph>
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<paragraph pageId="18" pageNumber="79">
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Prothorax. Pronotum slightly transverse (WPm/LP 1.25
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.019), with lateral margins slightly constricted posteriorly (WPm/WPp 1.29
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.018). Posterior angles slightly obtuse (100-110°). Width between posterior angles greater than between anterior angles (WPa/WPp 0.94
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.020).
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</paragraph>
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<paragraph pageId="18" pageNumber="79">
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Elytra. Moderately convex, slightly depressed along suture, moderately wide (WE/LE 0.66
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.020), without traces of striae. Humeri rounded, in outline forming right angle with longitudinal axis of body. Lateral margins convex, evenly divergent at basal forth, evenly rounded to apex in apical third.
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</paragraph>
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<paragraph pageId="18" pageNumber="79">Legs. Mesotibia (Fig. 7C). Metatibia (Fig. 7G).</paragraph>
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<paragraph pageId="18" pageNumber="79">Male genitalia. Median lobe (Fig. 13R) with shaft long, apically slightly widened, apex small and narrowly rounded. Ventral margin straight. Dorsal sclerites of internal sac (Fig. 13R) in form of long waved ribbon, tapered apically and slightly dilated basally, where sclerite forms small hook-like extension. Right paramere with short apical constriction (Fig. 13T). Left paramere with rather short and narrow apical constriction (Fig. 13S). Ring sclerite with triangularly rounded handle (Fig. 16F).</paragraph>
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<paragraph pageId="18" pageNumber="79">Female internal genitalia. Spermatheca sclerotized, elongate, subparallel, almost straight, with long cornu and short nodulus (Fig. 17F). Length of spermathecal gland less than length of spermatheca. Spermathecal duct not coiled.</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="79" type="geographical distribution">
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<paragraph pageId="18" pageNumber="79">Geographical distribution.</paragraph>
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<paragraph pageId="18" pageNumber="79">
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This species is known only from Cerro
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<normalizedToken originalValue="Pinalón">Pinalon</normalizedToken>
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, part of the Sierra de las Minas range of Guatemala (Fig. 22, green squares).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="79" type="way of life">
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<paragraph pageId="18" pageNumber="79">Way of life.</paragraph>
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<paragraph pageId="18" pageNumber="79">Specimens were extracted from cloud forest litter at elevations of 2000-2750 m.</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="79" type="relationships">
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<paragraph pageId="18" pageNumber="79">Relationships.</paragraph>
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<paragraph pageId="18" pageNumber="79">
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The shape of handle of the ring sclerite (Fig. 16F) and the structure of dorsal sclerites of the internal sac (Fig. 13R) suggest a relationship with the Honduran
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius celaquensis" order="Coleoptera" pageId="18" pageNumber="79" phylum="Arthropoda" rank="species" species="celaquensis">Geocharidius celaquensis</taxonomicName>
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(Figs 20B and 19E), described below.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |