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<document ID-DOI="10.11646/zootaxa.5263.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7800978" ID-ZooBank="B07051F2-A15B-49EC-9A2F-91B9420FEA4A" approvalRequired="33" approvalRequired_for_taxonomicNames="7" approvalRequired_for_textStreams="15" approvalRequired_for_treatments="11" checkinTime="1680677567416" checkinUser="plazi" docAuthor="Gupta, Devanshu, Keith, Denis, Bhunia, Debika, Das, Priyanka, Ghosh, Joyjit &amp; Chandra, Kailash" docDate="2023" docId="BC1F87FF136BFFEDFF74FA67FA6EBF85" docLanguage="en" docName="zootaxa.5263.2.2.pdf" docOrigin="Zootaxa 5263 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Melolontha incana" docType="key" docVersion="2" lastPageNumber="212" masterDocId="4026FF87137EFFF8FFE3FFD9FFF2B92E" masterDocTitle="Review of Melolontha Fabricius, 1775 (Coleoptera: Scarabaeidae: Melolonthinae) from India with the description of two new species" masterLastPageNumber="216" masterPageNumber="191" pageNumber="212" updateTime="1680678180874" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Review of Melolontha Fabricius, 1775 (Coleoptera: Scarabaeidae: Melolonthinae) from India with the description of two new species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gupta, Devanshu</mods:namePart>
<mods:affiliation>Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India.</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Keith, Denis</mods:namePart>
<mods:affiliation>120 rue Gabriel Péri, F- 28000 Chartres, France.</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bhunia, Debika</mods:namePart>
<mods:affiliation>Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India.</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Das, Priyanka</mods:namePart>
<mods:affiliation>Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India.</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ghosh, Joyjit</mods:namePart>
<mods:affiliation>Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India.</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Chandra, Kailash</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-9076-5442</mods:nameIdentifier>
<mods:affiliation>Former Director, Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India. kailash 611 @ rediffmail. com; https: // orcid. org / 0000 - 0001 - 9076 - 5442</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-04-05</mods:number>
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<mods:number>5263</mods:number>
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<mods:number>2</mods:number>
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Key to the Indian species of
<taxonomicName baseAuthorityName="Motschulsky" baseAuthorityYear="1854" box="[503,586,1470,1496]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="212" phylum="Arthropoda" rank="species" species="incana">
<emphasis bold="true" box="[503,586,1470,1496]" italics="true" pageId="21" pageNumber="212">incana</emphasis>
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group
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<emphasis bold="true" box="[151,170,1539,1560]" pageId="21" pageNumber="212">1.</emphasis>
Setae on elytra sparse; costae not demarcated (
<figureCitation box="[675,745,1539,1561]" captionStart="FIGURES 912" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[203,1419,198,1899]" captionTargetId="figure-17@5.[167,1420,181,1919]" captionTargetPageId="5" captionText="FIGURES 912. Habitus of Melolontha species. 9, M. carinata; 10, M. indica; 11, M. flabellata; 12, M. furcicauda." figureDoi="http://doi.org/10.5281/zenodo.7800984" httpUri="https://zenodo.org/record/7800984/files/figure.png" pageId="21" pageNumber="212">Fig. 11</figureCitation>
); pygidium bifurcate at apex, apical edges straight at outer margin (
<figureCitation captionStart="FIGURES 1924" captionStartId="10.[151,264,1757,1781]" captionTargetBox="[152,1435,182,1724]" captionTargetId="figure-25@10.[151,1436,181,1733]" captionTargetPageId="10" captionText="FIGURES 1924. Pygidium of Melolontha species. 19, M. chinensis; 20, M. nepalensis; 21, M. carinata; 22, M. indica; 23, M. flabellata; 24, M. furcicauda." figureDoi="http://doi.org/10.5281/zenodo.7800988" httpUri="https://zenodo.org/record/7800988/files/figure.png" pageId="21" pageNumber="212">Fig. 23</figureCitation>
); parameres broader medially, outer edge semicircular, apex straight (in dorsal view) (
<figureCitation box="[1115,1188,1568,1590]" captionStart="FIGURES 4752" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[234,1342,182,1912]" captionTargetId="figure-25@14.[233,1354,181,1920]" captionTargetPageId="14" captionText="FIGURES 4752. Parameres of Melolontha species in dorsal view. 47, M. chinensis; 48, M. nepalensis; 49, M. carinata; 50, M. indica; 51, M. flabellata; 52, M. furcicauda." figureDoi="http://doi.org/10.5281/zenodo.7801004" httpUri="https://zenodo.org/record/7801004/files/figure.png" pageId="21" pageNumber="212">Fig. 51</figureCitation>
), apex pointed (in lateral view) (
<figureCitation box="[294,364,1597,1619]" captionStart="FIGURES 5364" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[209,1358,186,1908]" captionTargetId="figure-17@15.[209,1379,181,1920]" captionTargetPageId="15" captionText="FIGURES 5364. Parameres of Melolontha species in lateral view. 53, M. guttigera; 54, M. virescens; 55, M. arunachalensis new species (holotype); 56, M. lachungensis new species (holotype); 57, M. aeneicollis; 58, M. cuprescens; 59, M. chinensis; 60, M. nepalensis; 61, M. carinata; 62, M. indica; 63, M. flabellata; 64, M. furcicauda." figureDoi="http://doi.org/10.5281/zenodo.7801008" httpUri="https://zenodo.org/record/7801008/files/figure.png" pageId="21" pageNumber="212">Fig. 63</figureCitation>
)...................................................................
<taxonomicName authority="Sharp, 1876" authorityName="Sharp" authorityYear="1876" box="[1188,1436,1597,1619]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="212" phylum="Arthropoda" rank="species" species="flabellata">
<emphasis bold="true" box="[1188,1314,1597,1619]" pageId="21" pageNumber="212">
<emphasis bold="true" box="[1188,1209,1598,1619]" italics="true" pageId="21" pageNumber="212">M</emphasis>
.
<emphasis bold="true" box="[1221,1314,1597,1619]" italics="true" pageId="21" pageNumber="212">flabellata</emphasis>
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<bibRefCitation author="Sharp, D." box="[1320,1436,1597,1619]" pageId="21" pageNumber="212" pagination="65 - 90" refId="ref15828" refString="Sharp, D. (1876) Descriptions of some new species of Scarabaeidae from tropical Asia and Malaysia. Coleopterologische Hefte, 15, 65 - 90." type="journal article" year="1876">Sharp, 1876</bibRefCitation>
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<paragraph blockId="21.[151,1437,1539,1707]" pageId="21" pageNumber="212">
<keyLead pageId="21" pageNumber="212">
Setae on elytra dense; costae demarcated (
<figureCitation box="[632,704,1627,1649]" captionStart="FIGURES 912" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[203,1419,198,1899]" captionTargetId="figure-17@5.[167,1420,181,1919]" captionTargetPageId="5" captionText="FIGURES 912. Habitus of Melolontha species. 9, M. carinata; 10, M. indica; 11, M. flabellata; 12, M. furcicauda." figureDoi="http://doi.org/10.5281/zenodo.7800984" httpUri="https://zenodo.org/record/7800984/files/figure.png" pageId="21" pageNumber="212">Fig. 12</figureCitation>
); pygidium bifurcate at apex, apical edges curved outwardly at outer margin (
<figureCitation box="[233,306,1656,1678]" captionStart="FIGURES 1924" captionStartId="10.[151,264,1757,1781]" captionTargetBox="[152,1435,182,1724]" captionTargetId="figure-25@10.[151,1436,181,1733]" captionTargetPageId="10" captionText="FIGURES 1924. Pygidium of Melolontha species. 19, M. chinensis; 20, M. nepalensis; 21, M. carinata; 22, M. indica; 23, M. flabellata; 24, M. furcicauda." figureDoi="http://doi.org/10.5281/zenodo.7800988" httpUri="https://zenodo.org/record/7800988/files/figure.png" pageId="21" pageNumber="212">Fig. 24</figureCitation>
); parameres relatively narrow medially, outer edge and apex nearly curved (in dorsal view) (
<figureCitation box="[1205,1278,1656,1678]" captionStart="FIGURES 4752" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[234,1342,182,1912]" captionTargetId="figure-25@14.[233,1354,181,1920]" captionTargetPageId="14" captionText="FIGURES 4752. Parameres of Melolontha species in dorsal view. 47, M. chinensis; 48, M. nepalensis; 49, M. carinata; 50, M. indica; 51, M. flabellata; 52, M. furcicauda." figureDoi="http://doi.org/10.5281/zenodo.7801004" httpUri="https://zenodo.org/record/7801004/files/figure.png" pageId="21" pageNumber="212">Fig. 52</figureCitation>
), apex broad (in lateral view) (
<figureCitation box="[360,429,1685,1707]" captionStart="FIGURES 5364" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[209,1358,186,1908]" captionTargetId="figure-17@15.[209,1379,181,1920]" captionTargetPageId="15" captionText="FIGURES 5364. Parameres of Melolontha species in lateral view. 53, M. guttigera; 54, M. virescens; 55, M. arunachalensis new species (holotype); 56, M. lachungensis new species (holotype); 57, M. aeneicollis; 58, M. cuprescens; 59, M. chinensis; 60, M. nepalensis; 61, M. carinata; 62, M. indica; 63, M. flabellata; 64, M. furcicauda." figureDoi="http://doi.org/10.5281/zenodo.7801008" httpUri="https://zenodo.org/record/7801008/files/figure.png" pageId="21" pageNumber="212">Fig. 64</figureCitation>
)............................................................
<taxonomicName authority="Ancey, 1881" authorityName="Ancey" authorityYear="1881" box="[1168,1436,1685,1707]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="212" phylum="Arthropoda" rank="species" species="furcicauda">
<emphasis bold="true" box="[1168,1309,1685,1707]" pageId="21" pageNumber="212">
<emphasis bold="true" box="[1168,1189,1686,1707]" italics="true" pageId="21" pageNumber="212">M</emphasis>
.
<emphasis bold="true" box="[1201,1309,1685,1707]" italics="true" pageId="21" pageNumber="212">furcicauda</emphasis>
</emphasis>
<bibRefCitation author="Ancey, F." box="[1315,1436,1686,1707]" pageId="21" pageNumber="212" pagination="412 - 413" refId="ref12774" refString="Ancey, F. (1881) Descriptions des Coleopteres nouveaux. Le Naturaliste, 1, 412 - 413." type="journal article" year="1881">Ancey, 1881</bibRefCitation>
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