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<document id="EBDC4C0B65FD2855A3C4821E42D6D7C3" ID-CLB-Dataset="4301" ID-DOI="10.18590/euscorpius.2004.vol2004.iss14.1" ID-GBIF-Dataset="8ea0a922-0437-483c-abc9-4d008c950628" ID-ISSN="1536-9307" ID-Zenodo-Dep="4648104" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617134081705" checkinUser="carolina" docAuthor="Teruel, Rolando, Fet, Victor &amp; de Armas, Luis F." docDate="2004" docId="B92D6A760E4F2F524E06C37215C01CFF" docLanguage="en" docName="Euscorpius.14.1-11.pdf" docOrigin="Euscorpius 14 (14)" docSource="https://mds.marshall.edu/euscorpius/vol2004/iss14/1/" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Euscorpius mingrelicus Thorell 1876" docType="treatment" docVersion="9" lastPageNumber="8" masterDocId="4514120E0E472F584F0EC52B141B194D" masterDocTitle="A note on the scorpions from the Pirin Mountains, south- western Bulgaria (Scorpiones: Buthidae, Euscorpiidae)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="7" updateTime="1698935365401" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="2A98EE64A32E6A6A2DFAB81FD16FAA37">A note on the scorpions from the Pirin Mountains, south- western Bulgaria (Scorpiones: Buthidae, Euscorpiidae)</mods:title>
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<mods:namePart id="B1AB53B191DDB4800524A5EFF52DBC98">Teruel, Rolando</mods:namePart>
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<mods:namePart id="E8D69B720B1082BB6B0CDC8B0A92B773">Fet, Victor</mods:namePart>
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<mods:namePart id="12A5923952A79A10051B607F7A16238C">de Armas, Luis F.</mods:namePart>
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<mods:title id="3D0774F6759FD735A2D837E9A09D78D2">Euscorpius</mods:title>
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<treatment id="B92D6A760E4F2F524E06C37215C01CFF" ID-DOI="http://doi.org/10.5281/zenodo.5507057" ID-GBIF-Taxon="187973103" ID-Zenodo-Dep="5507057" LSID="urn:lsid:plazi:treatment:B92D6A760E4F2F524E06C37215C01CFF" httpUri="http://treatment.plazi.org/id/B92D6A760E4F2F524E06C37215C01CFF" lastPageId="10" lastPageNumber="8" pageId="8" pageNumber="7">
<subSubSection id="799E88EB0E4F2F504E06C37216C11F3C" box="[264,730,1625,1649]" pageId="8" pageNumber="7" type="nomenclature">
<paragraph id="313BDB600E4F2F504E06C37216C11F3C" blockId="8.[264,730,1625,1649]" box="[264,730,1625,1649]" pageId="8" pageNumber="7">
<heading id="6A736C0C0E4F2F504E06C37216C11F3C" box="[264,730,1625,1649]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="7" reason="3">
<emphasis id="03F007720E4F2F504E06C37216C11F3C" bold="true" box="[264,730,1625,1649]" pageId="8" pageNumber="7">
<taxonomicName id="F684A0E30E4F2F504E06C37216C91F3C" ID-CoL="4H4X" authorityName="Thorell" authorityYear="1876" baseAuthorityName="Kessler" baseAuthorityYear="1874" box="[264,722,1625,1649]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="03F007720E4F2F504E06C372159E1F3C" bold="true" box="[264,389,1625,1649]" italics="true" pageId="8" pageNumber="7">Euscorpius</emphasis>
sp. 2 (“
<emphasis id="03F007720E4F2F504ED2C37216451F3C" bold="true" box="[476,606,1625,1649]" italics="true" pageId="8" pageNumber="7">mingrelicus</emphasis>
complex”
</taxonomicName>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="799E88EB0E4F2F504E8EC352167A1FDD" box="[384,609,1657,1680]" pageId="8" pageNumber="7" type="description">
<paragraph id="313BDB600E4F2F504E8EC352167A1FDD" blockId="8.[384,609,1657,1680]" box="[384,609,1657,1680]" pageId="8" pageNumber="7">
<emphasis id="03F007720E4F2F504E8EC352167A1FDD" bold="true" box="[384,609,1657,1680]" pageId="8" pageNumber="7">
<figureCitation id="A9BFC7E50E4F2F504E8EC352161A1FDD" box="[384,513,1657,1680]" captionStart-0="Figure 18" captionStart-1="Figure 19" captionStartId-0="10.[1095,1171,199,222]" captionStartId-1="10.[1149,1225,1177,1200]" captionTargetBox-0="[171,1071,213,381]" captionTargetBox-1="[195,1126,468,1317]" captionTargetId-0="figure-264@10.[163,1071,211,381]" captionTargetId-1="figure-265@10.[192,1126,468,1322]" captionTargetPageId-0="10" captionTargetPageId-1="10" captionText-0="Figure 18: Subadult male of Euscorpius sp. 2 (“mingrelicus complex”) from Pirin Mountains. Telson and metasomal segments IV-V, ventral view, showing partially developed ventral median and ventral lateral carinae on segment V." captionText-1="Figure 19: Trichobothrial patterns of the external aspect of left and right pedipalp patellae of the same specimen as in Fig. 18." figureDoi-0="http://doi.org/10.5281/zenodo.4648118" figureDoi-1="http://doi.org/10.5281/zenodo.4648120" httpUri-0="https://zenodo.org/record/4648118/files/figure.png" httpUri-1="https://zenodo.org/record/4648120/files/figure.png" pageId="8" pageNumber="7">Figs. 1819</figureCitation>
,
<tableCitation id="7C06EEDB0E4F2F504D03C352167A1FDD" box="[525,609,1657,1680]" captionStart="Table 1" captionStartId="9.[192,257,955,978]" captionTargetPageId="9" captionText="Table 1: Measurements (in mm) of two undetermined species of Euscorpius from Pirin Mts. L = length, W = width, D = depth." httpUri="http://table.plazi.org/id/65FB8BE80E4E2F514FCEC690118F1A9F" pageId="8" pageNumber="7" tableUuid="65FB8BE80E4E2F514FCEC690118F1A9F">Table 1</tableCitation>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="799E88EB0E4F2F504FFEC39D109D1CD7" pageId="8" pageNumber="7" type="materials_examined">
<paragraph id="313BDB600E4F2F504FFEC39D163D1E47" blockId="8.[192,802,1718,1894]" pageId="8" pageNumber="7">
<emphasis id="03F007720E4F2F504FFEC39D15D41F80" bold="true" box="[240,463,1718,1741]" pageId="8" pageNumber="7">Material examined.</emphasis>
<materialsCitation id="81ECD13D0E4F2F504ED8C39D16391E47" ID-GBIF-Occurrence="3385271304" collectingDate="1972-07" collectionCode="IES" collectorName="Gallia" country="Bulgaria" elevation="1250" location="Pirin Mountains" pageId="8" pageNumber="7" specimenCount="2" specimenCount-male="1" specimenCount-subadult="1">
<collectingCountry id="49939BF00E4F2F504ED8C39D167F1F80" box="[470,612,1718,1741]" name="Bulgaria" pageId="8" pageNumber="7">BULGARIA</collectingCountry>
:
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, under stones in open areas,
<elevation id="BAA93C530E4F2F504EE9C3FF16B51FA6" box="[487,686,1748,1771]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="8" pageNumber="7" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">
<quantity id="F67C76850E4F2F504EE9C3FF16931FA6" box="[487,648,1748,1771]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="8" pageNumber="7" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1,2001,300 m</quantity>
asl
</elevation>
,
<date id="453AFDA00E4F2F504DB7C3FF17391FA6" box="[697,802,1748,1771]" pageId="8" pageNumber="7" value="1972-07">
<collectingDate id="557E04480E4F2F504DB7C3FF17391FA6" box="[697,802,1748,1771]" pageId="8" pageNumber="7" value="1972-07">July 1972</collectingDate>
</date>
(coll.
<collectorName id="9C71BEB60E4F2F504FF0C3D815581E47" box="[254,323,1779,1802]" pageId="8" pageNumber="7">Gallia</collectorName>
),
<specimenCount id="278210E90E4F2F504E5BC3D815DF1E47" box="[341,452,1779,1802]" pageId="8" pageNumber="7" type="subadult">1 subadult</specimenCount>
<specimenCount id="278210E90E4F2F504EC5C3D815C41E47" box="[459,479,1779,1802]" pageId="8" pageNumber="7" type="male"></specimenCount>
(
<collectionCode id="579543A50E4F2F504EE3C3D816011E47" box="[493,538,1779,1802]" pageId="8" pageNumber="7">IES</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="313BDB600E4F2F504FFEC239109D1CD7" blockId="8.[192,802,1718,1894]" lastBlockId="8.[830,1441,1196,1894]" pageId="8" pageNumber="7">
<emphasis id="03F007720E4F2F504FFEC239159F1E64" bold="true" box="[240,388,1810,1833]" pageId="8" pageNumber="7">Morphology.</emphasis>
Small scorpion (subadult male about
<quantity id="F67C76850E4F2F504FCEC21B15171E0A" box="[192,268,1840,1863]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="7" unit="mm" value="23.0">23 mm</quantity>
long, adults probably under
<quantity id="F67C76850E4F2F504D47C21B16801E0A" box="[585,667,1840,1863]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="7" unit="mm" value="30.0">30 mm</quantity>
). Body light orange brown, with prosoma, tergites, metasoma and legs strongly marbled with dark brown; pedipalps reddish with carinae conspicuously darker; chelicerae, legs and telson yellowish. Metasoma short, slender and cylindrical in shape, with all carinae obsolete to absent; telson vesicle moderately enlarged. Trichobothrial patellar pattern:
<emphasis id="03F007720E4F2F504CFFC06D10111C10" box="[1009,1034,1350,1373]" italics="true" pageId="8" pageNumber="7">eb</emphasis>
= 4,
<emphasis id="03F007720E4F2F504B46C06D10751C10" box="[1096,1134,1350,1373]" italics="true" pageId="8" pageNumber="7">eba</emphasis>
= 4,
<emphasis id="03F007720E4F2F504BA2C06D10D41C10" box="[1196,1231,1350,1373]" italics="true" pageId="8" pageNumber="7">esb</emphasis>
= 2,
<emphasis id="03F007720E4F2F504A00C06D11361C10" box="[1294,1325,1350,1373]" italics="true" pageId="8" pageNumber="7">em</emphasis>
= 3,
<emphasis id="03F007720E4F2F504A64C06D11931C10" box="[1386,1416,1350,1373]" italics="true" pageId="8" pageNumber="7">est</emphasis>
= 34,
<emphasis id="03F007720E4F2F504C76C04F17901C36" box="[888,907,1380,1403]" italics="true" pageId="8" pageNumber="7">et</emphasis>
= 4,
<emphasis id="03F007720E4F2F504CC0C04F17C11C36" box="[974,986,1380,1403]" italics="true" pageId="8" pageNumber="7">v</emphasis>
= 6; fixed finger
<emphasis id="03F007720E4F2F504BA7C04F10F91C36" box="[1193,1250,1380,1403]" italics="true" pageId="8" pageNumber="7">et -est</emphasis>
/
<emphasis id="03F007720E4F2F504BF1C04F11501C36" box="[1279,1355,1380,1403]" italics="true" pageId="8" pageNumber="7">est -dsb</emphasis>
ratio = 2.17. Pectinal tooth count 77.
</paragraph>
</subSubSection>
<subSubSection id="799E88EB0E4F2F524C60C08915C01CFF" lastPageId="10" lastPageNumber="9" pageId="8" pageNumber="7" type="discussion">
<paragraph id="313BDB600E4F2F504C60C08910851F80" blockId="8.[830,1441,1196,1894]" pageId="8" pageNumber="7">
<emphasis id="03F007720E4F2F504C60C08917EB1CF4" bold="true" box="[878,1008,1442,1465]" pageId="8" pageNumber="7">Comments.</emphasis>
Even though the studied specimen is not an adult, it clearly shows all characters defining the “
<taxonomicName id="F684A0E30E4F2F504C44C0F417D31CBB" baseAuthorityName="Kessler" baseAuthorityYear="1874" box="[842,968,1503,1526]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="mingrelicus">
<emphasis id="03F007720E4F2F504C44C0F417D31CBB" box="[842,968,1503,1526]" italics="true" pageId="8" pageNumber="7">mingrelicus</emphasis>
</taxonomicName>
complex” of the subgenus
<taxonomicName id="F684A0E30E4F2F504A17C0F411BB1CBB" box="[1305,1440,1503,1526]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="03F007720E4F2F504A17C0F411BB1CBB" box="[1305,1440,1503,1526]" italics="true" pageId="8" pageNumber="7">Alpiscorpius</emphasis>
</taxonomicName>
(obsolescence of metasomal carination,
<emphasis id="03F007720E4F2F504A0BC0D5115B1F58" box="[1285,1344,1534,1557]" italics="true" pageId="8" pageNumber="7">et -est</emphasis>
/
<emphasis id="03F007720E4F2F504A5AC0D511BB1F58" box="[1364,1440,1534,1557]" italics="true" pageId="8" pageNumber="7">est -dsb</emphasis>
ratio ≥ 1.5), which otherwise are not age correlated. This represents the first record of this species complex and subgenus from
<collectingCountry id="49939BF00E4F2F504CE4C37110521F3C" box="[1002,1097,1626,1649]" name="Bulgaria" pageId="8" pageNumber="7">Bulgaria</collectingCountry>
. Its presence in this country is not surprising, however, as members of this complex are widely distributed over the western Balkan Peninsula (Fet, 2000; Fet &amp; Sissom, 2000).
</paragraph>
<paragraph id="313BDB600E4F2F514C60C3FF16621FFC" blockId="8.[830,1441,1196,1894]" lastBlockId="9.[192,802,1015,1897]" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
Fet (1993) reviewed all known distribution and taxonomic composition of the species (
<emphasis id="03F007720E4F2F504BF8C3D8117E1E47" box="[1270,1381,1779,1802]" italics="true" pageId="8" pageNumber="7">sensu lato</emphasis>
)
<taxonomicName id="F684A0E30E4F2F504A77C3D810C31E64" authority="(Kessler, 1874)" baseAuthorityName="Kessler" baseAuthorityYear="1874" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="mingrelicus">
<emphasis id="03F007720E4F2F504A77C3D810381E64" italics="true" pageId="8" pageNumber="7">Euscorpius mingrelicus</emphasis>
(Kessler, 1874)
</taxonomicName>
which was originally described from
<collectingCountry id="49939BF00E4F2F504B26C21B10651E0A" box="[1064,1150,1840,1863]" name="Georgia" pageId="8" pageNumber="7">Georgia</collectingCountry>
(Caucasus) but later redefined (Bonacina, 1980) as a part of the former species
<taxonomicName id="F684A0E30E4E2F514FCEC6DC16BA1D43" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[192,673,1015,1038]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="germanus">
<emphasis id="03F007720E4E2F514FCEC6DC15B41D43" box="[192,431,1015,1038]" italics="true" pageId="9" pageNumber="8">Euscorpius germanus</emphasis>
(C. L. Koch, 1837)
</taxonomicName>
. The latter species is in fact limited to the Alpine zone of Europe (Gantenbein et al., 2000a). Traditionally treated as one species widespread from the Alps to Caucasus (Bonacina, 1980; see Fet &amp; Sissom, 2000 for the taxonomic history), “
<taxonomicName id="F684A0E30E4E2F514E3FC1BB15CA1DEA" baseAuthorityName="Kessler" baseAuthorityYear="1874" box="[305,465,1168,1191]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="mingrelicus">
<emphasis id="03F007720E4E2F514E3FC1BB15CA1DEA" box="[305,465,1168,1191]" italics="true" pageId="9" pageNumber="8">E. mingrelicus</emphasis>
</taxonomicName>
complex” is currently under revision using both morphological and molecular techniques (Fet, 2000; Gantenbein et al., 2000a; Sherabon et al., 2000; Fet et al., 2003; etc.). Currently, three species are recognized in this complex, with
<taxonomicName id="F684A0E30E4E2F514D70C02015771C0C" authority="(Kessler, 1874)" baseAuthorityName="Kessler" baseAuthorityYear="1874" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" sensu="stricto" species="mingrelicus">
<emphasis id="03F007720E4E2F514D70C02017391C6F" box="[638,802,1291,1314]" italics="true" pageId="9" pageNumber="8">E. mingrelicus</emphasis>
(Kessler, 1874)
</taxonomicName>
<emphasis id="03F007720E4E2F514E77C001161D1C0C" box="[377,518,1322,1345]" italics="true" pageId="9" pageNumber="8">
<taxonomicNameLabel id="18C3BA090E4E2F514E77C001161D1C0C" box="[377,518,1322,1345]" pageId="9" pageNumber="8" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
ranging from
<collectingCountry id="49939BF00E4E2F514DBBC00116E41C0C" box="[693,767,1322,1345]" name="Bosnia and Herzegovina" pageId="9" pageNumber="8">Bosnia</collectingCountry>
to Caucasus (Fet, 1993; Fet &amp; Sissom, 2000); however, this division is clearly not satisfactory. Fet (2000) described
<taxonomicName id="F684A0E30E4E2F514FCEC0AD15321CD0" authorityName="Fet" authorityYear="2000" box="[192,297,1414,1437]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="beroni">
<emphasis id="03F007720E4E2F514FCEC0AD15321CD0" box="[192,297,1414,1437]" italics="true" pageId="9" pageNumber="8">E. beroni</emphasis>
</taxonomicName>
from the high mountains of
<collectingCountry id="49939BF00E4E2F514D73C0AD16C91CD0" box="[637,722,1414,1437]" name="Albania" pageId="9" pageNumber="8">Albania</collectingCountry>
(Prokletije), and Scherabon et al. (2000) demonstrated a separate status of
<taxonomicName id="F684A0E30E4E2F514E58C0E8165C1C97" authority="Caporiacco" authorityName="Caporiacco" box="[342,583,1475,1498]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="gamma">
<emphasis id="03F007720E4E2F514E58C0E815DF1C97" box="[342,452,1475,1498]" italics="true" pageId="9" pageNumber="8">E. gamma</emphasis>
Caporiacco
</taxonomicName>
from the northeastern part of the geographic range of this complex (northeastern
<collectingCountry id="49939BF00E4E2F514E18C32B15511F5A" box="[278,330,1536,1559]" name="Italy" pageId="9" pageNumber="8">Italy</collectingCountry>
,
<collectingCountry id="49939BF00E4E2F514E59C32B15AC1F5A" box="[343,439,1536,1559]" name="Slovenia" pageId="9" pageNumber="8">Slovenia</collectingCountry>
,
<collectingCountry id="49939BF00E4E2F514ECBC32B160D1F5A" box="[453,534,1536,1559]" name="Croatia" pageId="9" pageNumber="8">Croatia</collectingCountry>
, and
<collectingCountry id="49939BF00E4E2F514D5BC32B16B31F5A" box="[597,680,1536,1559]" name="Austria" pageId="9" pageNumber="8">Austria</collectingCountry>
). For ecology and distribution of
<taxonomicName id="F684A0E30E4E2F514ECDC33416291F7B" box="[451,562,1567,1590]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="gamma">
<emphasis id="03F007720E4E2F514ECDC33416291F7B" box="[451,562,1567,1590]" italics="true" pageId="9" pageNumber="8">E. gamma</emphasis>
</taxonomicName>
in
<collectingCountry id="49939BF00E4E2F514D56C33416AE1F7B" box="[600,693,1567,1590]" name="Slovenia" pageId="9" pageNumber="8">Slovenia</collectingCountry>
and
<collectingCountry id="49939BF00E4E2F514DE0C33414FD1F18" name="Austria" pageId="9" pageNumber="8">Austria</collectingCountry>
, see also Scherabon (1987), Fet et al. (2001), and Komposch et al. (2001). Together with “
<taxonomicName id="F684A0E30E4E2F514D74C37716F81F3E" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[634,739,1628,1651]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="germanus">
<emphasis id="03F007720E4E2F514D74C37716F81F3E" box="[634,739,1628,1651]" italics="true" pageId="9" pageNumber="8">germanus</emphasis>
</taxonomicName>
complex”, the “
<taxonomicName id="F684A0E30E4E2F514E4FC35015A41FDF" baseAuthorityName="Kessler" baseAuthorityYear="1874" box="[321,447,1659,1682]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="mingrelicus">
<emphasis id="03F007720E4E2F514E4FC35015A41FDF" box="[321,447,1659,1682]" italics="true" pageId="9" pageNumber="8">mingrelicus</emphasis>
</taxonomicName>
complex” comprises the subgenus
<taxonomicName id="F684A0E30E4E2F514FE2C3B1166E1FFC" authority="Gantenbein et al., 1999" authorityName="Gantenbein" authorityYear="1999" box="[236,629,1690,1713]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="03F007720E4E2F514FE2C3B115681FFC" box="[236,371,1690,1713]" italics="true" pageId="9" pageNumber="8">Alpiscorpius</emphasis>
Gantenbein et al., 1999
</taxonomicName>
.
</paragraph>
<paragraph id="313BDB600E4E2F514DBAC5C610EC1ADE" pageId="9" pageNumber="8">
<table id="438429C00E4ED0A74FC4C5C6117A1ADE" box="[202,1377,237,915]" gridcols="4" gridrows="4" pageId="9" pageNumber="8">
<tr id="8FB4D9220E4ED0A74FC4C5C6117A186E" box="[202,1377,237,291]" gridrow="0" pageId="9" pageNumber="8" rowspan-0="1">
<th id="CC65B05E0E4ED0A74D46C5C617F8186E" box="[584,995,237,291]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="9" pageNumber="8">
<emphasis id="03F007720E4E2F514DBAC5C61788186E" bold="true" italics="true" pageId="9" pageNumber="8">
<taxonomicName id="F684A0E30E4E2F514DBAC5C6172A1848" authorityName="Thorell" authorityYear="1876" box="[692,817,237,261]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">Euscorpius</taxonomicName>
sp. 1, “
<taxonomicName id="F684A0E30E4E2F514DAEC420173A186E" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[672,801,267,291]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="carpathicus">
<emphasis id="03F007720E4E2F514DAEC420173A186E" bold="true" box="[672,801,267,291]" italics="true" pageId="9" pageNumber="8">carpathicus</emphasis>
</taxonomicName>
complex”
</emphasis>
</th>
<th id="CC65B05E0E4ED0A74B50C5C6117A186E" box="[1118,1377,237,291]" gridcol="3" gridrow="0" pageId="9" pageNumber="8">
<emphasis id="03F007720E4E2F514B8FC5C6117A186E" bold="true" italics="true" pageId="9" pageNumber="8">
<taxonomicName id="F684A0E30E4E2F514B8FC5C610E51848" authorityName="Thorell" authorityYear="1876" box="[1153,1278,237,261]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">Euscorpius</taxonomicName>
sp. 2, “
<taxonomicName id="F684A0E30E4E2F514B62C42010F5186E" baseAuthorityName="Kessler" baseAuthorityYear="1874" box="[1132,1262,267,291]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="mingrelicus">
<emphasis id="03F007720E4E2F514B62C42010F5186E" bold="true" box="[1132,1262,267,291]" italics="true" pageId="9" pageNumber="8">mingrelicus</emphasis>
</taxonomicName>
complex”
</emphasis>
</th>
</tr>
<tr id="8FB4D9220E4ED0A74FC4C404117A180B" box="[202,1377,303,326]" gridrow="1" pageId="9" pageNumber="8" rowspan-0="1">
<td id="CC65B05E0E4ED0A74D46C40416CE180B" box="[584,725,303,326]" gridcol="1" gridrow="1" pageId="9" pageNumber="8">
<emphasis id="03F007720E4E2F514D5DC40416D7180B" bold="true" box="[595,716,303,326]" pageId="9" pageNumber="8">adult male</emphasis>
</td>
<td id="CC65B05E0E4ED0A74C5BC40417F8180B" box="[853,995,303,326]" gridcol="2" gridrow="1" pageId="9" pageNumber="8">
<emphasis id="03F007720E4E2F514C5BC40417F9180B" bold="true" box="[853,994,303,326]" pageId="9" pageNumber="8">adult female</emphasis>
</td>
<td id="CC65B05E0E4ED0A74B50C404117A180B" box="[1118,1377,303,326]" gridcol="3" gridrow="1" pageId="9" pageNumber="8">
<emphasis id="03F007720E4E2F514B81C404112B180B" bold="true" box="[1167,1328,303,326]" pageId="9" pageNumber="8">subadult male</emphasis>
</td>
</tr>
<tr id="8FB4D9220E4ED0A74FC4C479117A1A3F" box="[202,1377,338,882]" gridrow="2" pageId="9" pageNumber="8">
<th id="CC65B05E0E4ED0A74FC4C47915A21A3F" box="[202,441,338,882]" gridcol="0" gridrow="2" pageId="9" pageNumber="8">
<emphasis id="03F007720E4E2F514FC4C47915341824" bold="true" box="[202,303,338,361]" pageId="9" pageNumber="8">Prosoma</emphasis>
(L/W)
<emphasis id="03F007720E4E2F514FC4C45A155918C5" bold="true" box="[202,322,369,392]" pageId="9" pageNumber="8">Mesosoma</emphasis>
Tergite VII (L/W)
<emphasis id="03F007720E4E2F514FC4C485155B1888" bold="true" box="[202,320,430,453]" pageId="9" pageNumber="8">Metasoma</emphasis>
Segment I (L/W) Segment II Segment III Segment IV Segment V Telson Vesicle (L/W/D) Aculeus
<emphasis id="03F007720E4E2F514FC4C7E915341B94" bold="true" box="[202,303,706,729]" pageId="9" pageNumber="8">Pedipalp</emphasis>
Femur (L/W) Patela Chela Palm (L/D/W) Movable finger
</th>
<td id="CC65B05E0E4ED0A74D46C47916CE1A3F" box="[584,725,338,882]" gridcol="1" gridrow="2" pageId="9" pageNumber="8">4.7 / 4.0 9.0 2.1 / 3.7 17.9 1.8 / 1.6 2.1 / 1.5 2.3 / 1.5 2.8 / 1.4 4.3 / 1.4 4.6 3.5 / 1.8 / 2.0 1.1 17.1 4.1 / 1.5 4.1 / 2.1 8.9 4.1 / 3.3 / 3.1 4.8</td>
<td id="CC65B05E0E4ED0A74C5BC47917F81A3F" box="[853,995,338,882]" gridcol="2" gridrow="2" pageId="9" pageNumber="8">5.0 / 5.0 12.0 2.4 / 4.4 17.2 1.8 / 1.9 2.1 / 1.6 2.2 / 1.5 2.7 / 1.4 4.2 / 1.4 4.2 2.8 / 1.4 / 1.4 1.4 18.0 4.4 / 1.7 4.3 / 2.2 9.3 4.4 / 3.2 / 2.7 4.9</td>
<td id="CC65B05E0E4ED0A74B50C479117A1A3F" box="[1118,1377,338,882]" gridcol="3" gridrow="2" pageId="9" pageNumber="8">3.4 / 3.5 8.7 1.8 / 3.9 10.6 1.0 / 1.3 1.3 / 1.3 1.4 / 1.2 1.4 / 1.2 2.7 / 1.1 2.8 2.0 / 1.1 / 0.9 0.8 11.2 2.6 / 1.0 3.2 / 1.2 5.4 2.7 / 2.0 / 1.4 2.7</td>
</tr>
<tr id="8FB4D9220E4ED0A74FC4C657117A1ADE" box="[202,1377,892,915]" gridrow="3" pageId="9" pageNumber="8">
<th id="CC65B05E0E4ED0A74FC4C65715A21ADE" box="[202,441,892,915]" gridcol="0" gridrow="3" pageId="9" pageNumber="8">
<emphasis id="03F007720E4E2F514FC4C657157B1ADE" bold="true" box="[202,352,892,915]" pageId="9" pageNumber="8">Total Length</emphasis>
</th>
<td id="CC65B05E0E4ED0A74D46C65716CE1ADE" box="[584,725,892,915]" gridcol="1" gridrow="3" pageId="9" pageNumber="8">31.6</td>
<td id="CC65B05E0E4ED0A74C5BC65717F81ADE" box="[853,995,892,915]" gridcol="2" gridrow="3" pageId="9" pageNumber="8">34.4</td>
<td id="CC65B05E0E4ED0A74B50C657117A1ADE" box="[1118,1377,892,915]" gridcol="3" gridrow="3" pageId="9" pageNumber="8">22.7</td>
</tr>
</table>
</paragraph>
<caption id="65FB8BE80E4E2F514FCEC690118F1A9F" ID-Table-UUID="65FB8BE80E4E2F514FCEC690118F1A9F" box="[192,1428,955,978]" httpUri="http://table.plazi.org/id/65FB8BE80E4E2F514FCEC690118F1A9F" pageId="9" pageNumber="8" startId="9.[192,257,955,978]" targetBox="[202,1377,237,915]" targetIsTable="true" targetPageId="9">
<paragraph id="313BDB600E4E2F514FCEC690118F1A9F" blockId="9.[192,1428,955,978]" box="[192,1428,955,978]" pageId="9" pageNumber="8">
<emphasis id="03F007720E4E2F514FCEC69015051A9F" bold="true" box="[192,286,955,978]" pageId="9" pageNumber="8">Table 1:</emphasis>
Measurements (in mm) of two undetermined species of
<taxonomicName id="F684A0E30E4E2F514C33C69617B21A9F" authorityName="Thorell" authorityYear="1876" box="[829,937,957,978]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="03F007720E4E2F514C33C69617B21A9F" box="[829,937,957,978]" italics="true" pageId="9" pageNumber="8">Euscorpius</emphasis>
</taxonomicName>
from Pirin Mts. L = length, W = width, D = depth.
</paragraph>
</caption>
<paragraph id="313BDB600E4E2F514FFEC39310931C6F" blockId="9.[192,802,1015,1897]" lastBlockId="9.[830,1441,1015,1314]" pageId="9" pageNumber="8">
For the large part of the “
<taxonomicName id="F684A0E30E4E2F514D0BC39316981F82" baseAuthorityName="Kessler" baseAuthorityYear="1874" box="[517,643,1720,1743]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="mingrelicus">
<emphasis id="03F007720E4E2F514D0BC39316981F82" box="[517,643,1720,1743]" italics="true" pageId="9" pageNumber="8">mingrelicus</emphasis>
</taxonomicName>
complex”, the taxonomy is not defined or is defined partially. There are formally seven valid subspecies of
<taxonomicName id="F684A0E30E4E2F514D48C3DD16FF1E40" baseAuthorityName="Kessler" baseAuthorityYear="1874" box="[582,740,1782,1805]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="mingrelicus">
<emphasis id="03F007720E4E2F514D48C3DD16FF1E40" box="[582,740,1782,1805]" italics="true" pageId="9" pageNumber="8">E. mingrelicus</emphasis>
</taxonomicName>
from Balkans to
<collectingCountry id="49939BF00E4E2F514E35C23F158C1E66" box="[315,407,1812,1835]" name="Turkey" pageId="9" pageNumber="8">Anatolia</collectingCountry>
(Fet &amp; Sissom, 2000), and status of these forms is still unclear. The high genetic diversity of populations within
<collectingCountry id="49939BF00E4E2F514E9FC27915C51E24" box="[401,478,1874,1897]" name="Turkey" pageId="9" pageNumber="8">Turkey</collectingCountry>
(Fet et al., 2003a) indicates a possibility of cryptic species as recently discovered in the related Alpine “
<taxonomicName id="F684A0E30E4E2F514B14C13D10981D60" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[1050,1155,1046,1069]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="germanus">
<emphasis id="03F007720E4E2F514B14C13D10981D60" box="[1050,1155,1046,1069]" italics="true" pageId="9" pageNumber="8">germanus</emphasis>
</taxonomicName>
complex” (Gantenbein et al., 2000a). Further investigation of the “
<taxonomicName id="F684A0E30E4E2F514A2CC11E11BB1D01" baseAuthorityName="Kessler" baseAuthorityYear="1874" box="[1314,1440,1077,1100]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="mingrelicus">
<emphasis id="03F007720E4E2F514A2CC11E11BB1D01" box="[1314,1440,1077,1100]" italics="true" pageId="9" pageNumber="8">mingrelicus</emphasis>
</taxonomicName>
complex” from the Balkans and
<collectingCountry id="49939BF00E4E2F514B90C17810E11D27" box="[1182,1274,1107,1130]" name="Turkey" pageId="9" pageNumber="8">Anatolia</collectingCountry>
will be needed to establish into the species structure of this complex. The Pirin specimen falls into the range of “
<taxonomicName id="F684A0E30E4E2F514A2CC1BB11BB1DEA" baseAuthorityName="Kessler" baseAuthorityYear="1874" box="[1314,1440,1168,1191]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="mingrelicus">
<emphasis id="03F007720E4E2F514A2CC1BB11BB1DEA" box="[1314,1440,1168,1191]" italics="true" pageId="9" pageNumber="8">mingrelicus</emphasis>
</taxonomicName>
complex” for which taxonomic identity of populations is not yet determined. It is an important biogeographic find—the first specimen of this complex from
<collectingCountry id="49939BF00E4E2F514A33C1C711871C4E" box="[1341,1436,1260,1283]" name="Bulgaria" pageId="9" pageNumber="8">Bulgaria</collectingCountry>
, and most likely a glacial relict.
</paragraph>
<paragraph id="313BDB600E4E2F514C30C065102A1C27" blockId="9.[830,1073,1358,1386]" box="[830,1073,1358,1386]" pageId="9" pageNumber="8">
<heading id="6A736C0C0E4E2F514C30C065102A1C27" box="[830,1073,1358,1386]" pageId="9" pageNumber="8">
<emphasis id="03F007720E4E2F514C30C065102A1C27" bold="true" box="[830,1073,1358,1386]" pageId="9" pageNumber="8">General Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="313BDB600E4E2F524C60C08915C01CFF" blockId="9.[830,1441,1442,1894]" lastBlockId="10.[192,802,1404,1458]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
Even though the examined material is scarce (only
<specimenCount id="278210E90E4E2F514C30C0EA17CE1C95" box="[830,981,1473,1496]" pageId="9" pageNumber="8" type="generic">12 specimens</specimenCount>
), it demonstrates a rather high species diversity (four species) for a single locality in the Pirin Mountains of
<collectingCountry id="49939BF00E4E2F514CD2C0D510201F58" box="[988,1083,1534,1557]" name="Bulgaria" pageId="9" pageNumber="8">Bulgaria</collectingCountry>
. While it is not unusual to find two or three different species of
<taxonomicName id="F684A0E30E4E2F514BB8C33611351F79" authorityName="Thorell" authorityYear="1876" box="[1206,1326,1565,1588]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="03F007720E4E2F514BB8C33611351F79" box="[1206,1326,1565,1588]" italics="true" pageId="9" pageNumber="8">Euscorpius</emphasis>
</taxonomicName>
sympatrically in Europe (Fet, 2000; Fet &amp; Braunwalder, 2000), their precise cohabitation depends on ecological heterogeneity of the area, and the Alpine-like Pirin massif in southwestern
<collectingCountry id="49939BF00E4E2F514CDAC3BC102B1FE3" box="[980,1072,1687,1710]" name="Bulgaria" pageId="9" pageNumber="8">Bulgaria</collectingCountry>
indeed provides such diversity of habitats. In addition,
<taxonomicName id="F684A0E30E4E2F514B25C39D110F1F80" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[1067,1300,1718,1741]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="03F007720E4E2F514B25C39D110F1F80" box="[1067,1300,1718,1741]" italics="true" pageId="9" pageNumber="8">Mesobuthus gibbosus</emphasis>
</taxonomicName>
record from Pirin represents the most peripheral record of this species in the northern part of its geographic range. Combination of
<taxonomicName id="F684A0E30E4E2F514CBEC239102A1E64" authorityName="Vachon" authorityYear="1950" box="[944,1073,1810,1833]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="03F007720E4E2F514CBEC239102A1E64" box="[944,1073,1810,1833]" italics="true" pageId="9" pageNumber="8">Mesobuthus</emphasis>
</taxonomicName>
and
<taxonomicName id="F684A0E30E4E2F514B7EC23910F31E64" authorityName="Thorell" authorityYear="1876" box="[1136,1256,1810,1833]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="03F007720E4E2F514B7EC23910F31E64" box="[1136,1256,1810,1833]" italics="true" pageId="9" pageNumber="8">Euscorpius</emphasis>
</taxonomicName>
scorpion faunal elements is a typical Aegean-Anatolian feature, as representatives of these two genera are very common and sympatric all over
<collectingCountry id="49939BF00E4D2F524E98C05715FA1CDE" box="[406,481,1404,1427]" name="Greece" pageId="10" pageNumber="9">Greece</collectingCountry>
and
<collectingCountry id="49939BF00E4D2F524D11C05716741CDE" box="[543,623,1404,1427]" name="Turkey" pageId="10" pageNumber="9">Turkey</collectingCountry>
, including most islands of the Aegean Sea.
</paragraph>
</subSubSection>
</treatment>
</document>