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<document id="7FAFFD727625FF1F3E3716BD8D111938" ID-DOI="10.1649/0010-065X-69.mo4.115" ID-ISSN="1938-4394" ID-Zenodo-Dep="4908240" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623094854904" checkinUser="carolina" docAuthor="Iwan, Dariusz &amp; Matthews, Eric G." docDate="2015" docId="903023785D46FFED52B42FBAC175D0B5" docLanguage="en" docName="TheColeopBull.69.mo14.115-121.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 14) 69" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Scleropatroides Lobl and Merkl 2003" docType="treatment" docVersion="3" lastPageNumber="116" masterDocId="6C095B005D46FFEC50402A17C22DD22D" masterDocTitle="Scleropatroides Löbl and Merkl (Coleoptera: Tenebrionidae: Opatrini) Discovered in Australia, with Description of a New Species and Discussion of Related Genera" masterLastPageNumber="121" masterPageNumber="115" pageNumber="115" updateTime="1698956321997" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0C4EFCD64AEB803BB6C97DB390164CDA">Scleropatroides Löbl and Merkl (Coleoptera: Tenebrionidae: Opatrini) Discovered in Australia, with Description of a New Species and Discussion of Related Genera</mods:title>
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<mods:name id="D99DC69BC7E9304F6B42F09FA1CED151" type="personal">
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<mods:roleTerm id="157755401F34F726329A6FF98C917936">Author</mods:roleTerm>
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<mods:namePart id="E093AF8BC7F2B73FB50C874ED3796222">Iwan, Dariusz</mods:namePart>
<mods:affiliation id="2E6FE6ACF57D3BAEC19FF776C22170B6">Museum and Institute of Zoology, Polish Academy of Sciences Wilcza 64, 00 - 679, Warsaw, POLAND</mods:affiliation>
<mods:nameIdentifier id="75C5C8F1C755EB0273599591CDA43D16" type="email">darek@miiz.waw.pl</mods:nameIdentifier>
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<mods:namePart id="9FA1C429A4D55E473E870D1F2F5B1C32">Matthews, Eric G.</mods:namePart>
<mods:affiliation id="FD57829CA308FA0769B4AFFE63708C94">The South Australian Museum, North Terrace Adelaide, SA, AUSTRALIA</mods:affiliation>
<mods:nameIdentifier id="515BB294EAB3F642E719561102608D91" type="email">eric.matthews@samuseum.sa.gov.au</mods:nameIdentifier>
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<mods:title id="ED3191C5C40FAA6909441AC39FACA4F7">The Coleopterists Bulletin</mods:title>
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<mods:part id="523504293182A2E2950D2D6AAE1E850B">
<mods:date id="9E03D4CA26B733E8705316FD5F6AFD4D">2015</mods:date>
<mods:detail id="A8703D3AE3A11BD884328F4D8ABF54CA" type="series">
<mods:title id="D9295E24D15CD0CB1D6261C8164475F8">mo 14</mods:title>
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<mods:number id="7E60FB4CD1D06CCAB7CA457F744955D8">2015-12-18</mods:number>
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<mods:detail id="F23A3E5DAA93AB6102A5A9C151E2022F" type="volume">
<mods:number id="3C88CE5E49FE8142B428A6B72613449A">69</mods:number>
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<mods:classification id="5673A89F55B3C80C120DFFEC5018FDF1">journal article</mods:classification>
<mods:identifier id="6A9E57E9882C216B41594BD3DEB5104E" type="DOI">10.1649/0010-065X-69.mo4.115</mods:identifier>
<mods:identifier id="0FF49604B06DD16F1B3E660ACA9C9F52" type="ISSN">1938-4394</mods:identifier>
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<treatment id="903023785D46FFED52B42FBAC175D0B5" ID-DOI="http://doi.org/10.5281/zenodo.4908298" ID-Zenodo-Dep="4908298" LSID="urn:lsid:plazi:treatment:903023785D46FFED52B42FBAC175D0B5" httpUri="http://treatment.plazi.org/id/903023785D46FFED52B42FBAC175D0B5" lastPageId="1" lastPageNumber="116" pageId="0" pageNumber="115">
<subSubSection id="5083C1E55D46FFEC52B42FBAC655D7EF" box="[756,1144,1453,1474]" pageId="0" pageNumber="115" type="nomenclature">
<paragraph id="1826926E5D46FFEC52B42FBAC655D7EF" blockId="0.[756,1144,1453,1474]" box="[756,1144,1453,1474]" pageId="0" pageNumber="115">
<heading id="436E25025D46FFEC52B42FBAC655D7EF" box="[756,1144,1453,1474]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="115" reason="2">
<taxonomicName id="DF99E9ED5D46FFEC52B42FBAC655D7EF" authority="Lobl and Merkl, 2003" authorityName="Lobl and Merkl" authorityYear="2003" box="[756,1144,1453,1474]" class="Insecta" family="Tenebrionidae" genus="Scleropatroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis id="2AED4E7C5D46FFEC52B42FBAC655D7EF" bold="true" box="[756,1144,1453,1474]" pageId="0" pageNumber="115">
<emphasis id="2AED4E7C5D46FFEC52B42FBAC1A6D7EF" bold="true" box="[756,907,1453,1474]" italics="true" pageId="0" pageNumber="115">Scleropatroides</emphasis>
Löbl and Merkl, 2003
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="5083C1E55D46FFEC529B2FCFC1CDD42A" pageId="0" pageNumber="115" type="type_taxon">
<paragraph id="1826926E5D46FFEC529B2FCFC1CDD42A" blockId="0.[707,1193,1495,1729]" pageId="0" pageNumber="115">
<emphasis id="2AED4E7C5D46FFEC529B2FCFC173D7C0" bold="true" box="[731,862,1496,1517]" pageId="0" pageNumber="115">
<typeStatus id="C7222CCC5D46FFEC529B2FCFC127D7C0" box="[731,778,1496,1517]" pageId="0" pageNumber="115">Type</typeStatus>
Species.
</emphasis>
<taxonomicName id="DF99E9ED5D46FFEC532B2FC0C0D9D42A" authority="Baudi di Selve, 1876" authorityName="Baudi di Selve" authorityYear="1876" class="Insecta" family="Tenebrionidae" genus="Opatrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="hirtulum">
<emphasis id="2AED4E7C5D46FFEC532B2FC0C639D7C1" box="[875,1044,1495,1516]" italics="true" pageId="0" pageNumber="115">Opatrum hirtulum</emphasis>
Baudi di Selve, 1876
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="5083C1E55D46FFED529B2C1AC175D0B5" lastPageId="1" lastPageNumber="116" pageId="0" pageNumber="115" type="diagnosis">
<paragraph id="1826926E5D46FFEC529B2C1AC1C3D4EC" blockId="0.[707,1193,1495,1729]" pageId="0" pageNumber="115">
<emphasis id="2AED4E7C5D46FFEC529B2C1AC169D40F" bold="true" box="[731,836,1549,1570]" pageId="0" pageNumber="115">Diagnosis.</emphasis>
Clypeus deeply emarginate; eyes not completely divided; third antennomere short (less than two times as long as second); pronotal disc evenly convex; elytral striae with large foveate punctures; elytral intervals carinate, with one row of tubercles; wings fully developed; abdominal ventrites flat; aedeagus simple.
</paragraph>
<caption id="4CE6C2E65D47FFED50EF2AD6C1F9D2F9" box="[175,980,193,212]" pageId="1" pageNumber="116" startId="1.[175,223,193,212]">
<paragraph id="1826926E5D47FFED50EF2AD6C1F9D2F9" blockId="1.[175,980,193,212]" box="[175,980,193,212]" pageId="1" pageNumber="116">
<emphasis id="2AED4E7C5D47FFED50EF2AD6C2DAD2F9" bold="true" box="[175,247,193,212]" pageId="1" pageNumber="116">Table 1.</emphasis>
Distinguishing features of
<taxonomicName id="DF99E9ED5D47FFED51B22AD6C059D2F9" authorityName="Lobl and Merkl" authorityYear="2003" box="[498,628,193,212]" class="Insecta" family="Tenebrionidae" genus="Scleropatroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="2AED4E7C5D47FFED51B22AD6C059D2F9" box="[498,628,193,212]" italics="true" pageId="1" pageNumber="116">Scleropatroides</emphasis>
</taxonomicName>
,
<taxonomicName id="DF99E9ED5D47FFED52C12AD6C101D2F9" box="[641,812,193,212]" class="Insecta" family="Tenebrionidae" genus="Polycoelogastridion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="2AED4E7C5D47FFED52C12AD6C101D2F9" box="[641,812,193,212]" italics="true" pageId="1" pageNumber="116">Polycoelogastridion</emphasis>
</taxonomicName>
, and
<taxonomicName id="DF99E9ED5D47FFED531E2AD6C1E2D2F9" authorityName="Reitter" authorityYear="1877" box="[862,975,193,212]" class="Insecta" family="Tenebrionidae" genus="Scleropatrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="2AED4E7C5D47FFED531E2AD6C1E2D2F9" box="[862,975,193,212]" italics="true" pageId="1" pageNumber="116">Scleropatrum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="1826926E5D47FFED50A72AE3C65DD389" pageId="1" pageNumber="116">
<table id="6A9960CE5D47001350D72AE3C6BED389" box="[151,1171,244,420]" gridcols="4" gridrows="7" pageId="1" pageNumber="116">
<tr id="A6A9902C5D47001350D72AE3C6BED32A" box="[151,1171,244,263]" gridrow="0" pageId="1" pageNumber="116">
<th id="E578F9505D47001350D72AE3C3F9D32A" box="[151,468,244,263]" gridcol="0" gridrow="0" pageId="1" pageNumber="116">
<emphasis id="2AED4E7C5D47FFED50A72AE3C3A8D32A" bold="true" box="[231,389,244,263]" pageId="1" pageNumber="116">Character/Genus</emphasis>
</th>
<th id="E578F9505D470013525A2AE3C0CBD32A" box="[538,742,244,263]" gridcol="1" gridrow="0" pageId="1" pageNumber="116">
<taxonomicName id="DF99E9ED5D47FFED527D2AE3C0EED32A" authorityName="Lobl and Merkl" authorityYear="2003" box="[573,707,244,263]" class="Insecta" family="Tenebrionidae" genus="Scleropatroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="2AED4E7C5D47FFED527D2AE3C0EED32A" bold="true" box="[573,707,244,263]" italics="true" pageId="1" pageNumber="116">Scleropatroides</emphasis>
</taxonomicName>
</th>
<th id="E578F9505D470013536C2AE3C1F5D32A" box="[812,984,244,263]" gridcol="2" gridrow="0" pageId="1" pageNumber="116">
<taxonomicName id="DF99E9ED5D47FFED536C2AE3C1F5D32A" box="[812,984,244,263]" class="Insecta" family="Tenebrionidae" genus="Polycoelogastridion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="2AED4E7C5D47FFED536C2AE3C1F5D32A" bold="true" box="[812,984,244,263]" italics="true" pageId="1" pageNumber="116">Polycoelogastridion</emphasis>
</taxonomicName>
</th>
<th id="E578F9505D470013545D2AE3C6BED32A" box="[1053,1171,244,263]" gridcol="3" gridrow="0" pageId="1" pageNumber="116">
<taxonomicName id="DF99E9ED5D47FFED545D2AE3C6BED32A" authorityName="Reitter" authorityYear="1877" box="[1053,1171,244,263]" class="Insecta" family="Tenebrionidae" genus="Scleropatrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="2AED4E7C5D47FFED545D2AE3C6BED32A" bold="true" box="[1053,1171,244,263]" italics="true" pageId="1" pageNumber="116">Scleropatrum</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="A6A9902C5D47001350D72B02C6BED301" box="[151,1171,277,300]" gridrow="1" pageId="1" pageNumber="116">
<th id="E578F9505D47001350D72B02C3F9D301" box="[151,468,277,300]" gridcol="0" gridrow="1" pageId="1" pageNumber="116">Length ratio of 3rd:2nd antennomeres</th>
<td id="E578F9505D470013525A2B02C0CBD301" box="[538,742,277,300]" gridcol="1" gridrow="1" pageId="1" pageNumber="116">&lt;2.0</td>
<td id="E578F9505D470013536C2B02C1F5D301" box="[812,984,277,300]" gridcol="2" gridrow="1" pageId="1" pageNumber="116">&lt;2.0</td>
<td id="E578F9505D470013545D2B02C6BED301" box="[1053,1171,277,300]" gridcol="3" gridrow="1" pageId="1" pageNumber="116">2.5–4.0</td>
</tr>
<tr id="A6A9902C5D47001350D72B26C6BED369" box="[151,1171,305,324]" gridrow="2" pageId="1" pageNumber="116">
<th id="E578F9505D47001350D72B26C3F9D369" box="[151,468,305,324]" gridcol="0" gridrow="2" pageId="1" pageNumber="116">Median longitudinal depression on</th>
<td id="E578F9505D470013525A2B26C0CBD369" box="[538,742,305,324]" gridcol="1" gridrow="2" pageId="1" pageNumber="116">Absent</td>
<td id="E578F9505D470013536C2B26C1F5D369" box="[812,984,305,324]" gridcol="2" gridrow="2" pageId="1" pageNumber="116">Present</td>
<td id="E578F9505D470013545D2B26C6BED369" box="[1053,1171,305,324]" gridcol="3" gridrow="2" pageId="1" pageNumber="116">Absent</td>
</tr>
<tr id="A6A9902C5D47001350D72B5EC6BED371" box="[151,1171,329,348]" gridrow="3" pageId="1" pageNumber="116" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="E578F9505D47001350D72B5EC3F9D371" box="[151,468,329,348]" gridcol="0" gridrow="3" pageId="1" pageNumber="116">pronotal disc</th>
</tr>
<tr id="A6A9902C5D47001350D72B76C6BED359" box="[151,1171,353,372]" gridrow="4" pageId="1" pageNumber="116">
<th id="E578F9505D47001350D72B76C3F9D359" box="[151,468,353,372]" gridcol="0" gridrow="4" pageId="1" pageNumber="116">Punctures in elytral striae</th>
<td id="E578F9505D470013525A2B76C0CBD359" box="[538,742,353,372]" gridcol="1" gridrow="4" pageId="1" pageNumber="116">Small to large (foveate)</td>
<td id="E578F9505D470013536C2B76C1F5D359" box="[812,984,353,372]" gridcol="2" gridrow="4" pageId="1" pageNumber="116">Large (foveate)</td>
<td id="E578F9505D470013545D2B76C6BED359" box="[1053,1171,353,372]" gridcol="3" gridrow="4" pageId="1" pageNumber="116">Absent</td>
</tr>
<tr id="A6A9902C5D47001350D72B6EC6BED3A1" box="[151,1171,377,396]" gridrow="5" pageId="1" pageNumber="116">
<th id="E578F9505D47001350D72B6EC3F9D3A1" box="[151,468,377,396]" gridcol="0" gridrow="5" pageId="1" pageNumber="116">Hind wings</th>
<td id="E578F9505D470013525A2B6EC0CBD3A1" box="[538,742,377,396]" gridcol="1" gridrow="5" pageId="1" pageNumber="116">Present</td>
<td id="E578F9505D470013536C2B6EC1F5D3A1" box="[812,984,377,396]" gridcol="2" gridrow="5" pageId="1" pageNumber="116">Present</td>
<td id="E578F9505D470013545D2B6EC6BED3A1" box="[1053,1171,377,396]" gridcol="3" gridrow="5" pageId="1" pageNumber="116">Reduced</td>
</tr>
<tr id="A6A9902C5D47001350D72B86C6BED389" box="[151,1171,401,420]" gridrow="6" pageId="1" pageNumber="116">
<th id="E578F9505D47001350D72B86C3F9D389" box="[151,468,401,420]" gridcol="0" gridrow="6" pageId="1" pageNumber="116">Depression on ventrites I-III</th>
<td id="E578F9505D470013525A2B86C0CBD389" box="[538,742,401,420]" gridcol="1" gridrow="6" pageId="1" pageNumber="116">Absent</td>
<td id="E578F9505D470013536C2B86C1F5D389" box="[812,984,401,420]" gridcol="2" gridrow="6" pageId="1" pageNumber="116">Present</td>
<td id="E578F9505D470013545D2B86C6BED389" box="[1053,1171,401,420]" gridcol="3" gridrow="6" pageId="1" pageNumber="116">Absent</td>
</tr>
</table>
</paragraph>
<paragraph id="1826926E5D47FFED50EF2BF3C377D18F" blockId="1.[151,638,484,930]" pageId="1" pageNumber="116">
The genus belongs to the tribe
<taxonomicName id="DF99E9ED5D47FFED518E2BF3C033D3D4" box="[462,542,484,505]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="tribe" tribe="Opatrini">Opatrini</taxonomicName>
, as shown by the excised clypeus, antennae without complex sensoria, labrum subquadrate with tormal arms mesally bent into long extensions (
<figureCitation id="80A28EEB5D47FFED51A72823C00FD064" box="[487,546,564,585]" captionStart="Figs" captionStartId="3.[175,215,1365,1384]" captionTargetBox="[192,1159,188,1342]" captionTargetId="figure-145@3.[151,1173,183,1352]" captionTargetPageId="3" captionText="Figs. 9–14. Scleropatroides triplehorni. 9) Labrum-epipharynx; 10) Maxilla; 11) Labium; 12) Aedeagus, dorsal view; 13) Aedeagus, ventral view; 14) Ovipositor and female tract." figureDoi="http://doi.org/10.5281/zenodo.4908248" httpUri="https://zenodo.org/record/4908248/files/figure.png" pageId="1" pageNumber="116">Fig. 9</figureCitation>
), laciniae uncinate, protrochanters of opatrinoid
<typeStatus id="C7222CCC5D47FFED52562859C012D04E" box="[534,575,590,611]" pageId="1" pageNumber="116">type</typeStatus>
<taxonomicName id="DF99E9ED5D47FFED5208285AC337D053" authority="Iwan (2004)" authorityName="Iwan" authorityYear="2004" pageId="1" pageNumber="116" rank="species" species="sensu">
<emphasis id="2AED4E7C5D47FFED5208285AC050D04F" box="[584,637,589,610]" italics="true" pageId="1" pageNumber="116">sensu</emphasis>
<bibRefCitation id="7C08EF9F5D47FFED50D7287EC337D053" author="Iwan" box="[151,282,617,638]" pageId="1" pageNumber="116" pagination="735 - 765" refId="ref2656" refString="Iwan, D. 2004. A comparative study of male genitalia in Opatrinae sensu Medvedev (1968) (Coleoptera: Tenebrionidae), with notes on the reinterpreted tribal classification. Part II. Annales Zoologici 54: 735 - 765." type="journal article" year="2004">Iwan (2004)</bibRefCitation>
</taxonomicName>
, protibiae fossorial (although not strongly so) (
<figureCitation id="80A28EEB5D47FFED51652894C34BD0B5" box="[293,358,643,664]" captionStart="Figs" captionStartId="2.[197,237,1639,1658]" captionTargetBox="[176,655,1176,1617]" captionTargetId="figure-553@2.[172,659,1161,1622]" captionTargetPageId="2" captionText="Figs. 5–8. Scleropatroides triplehorni. 5) Head and prothorax, lateral view; 6) Head and prothorax, ventral view; 7) Head and pronotum, anterior view; 8) Mouthparts in situ. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4908246" httpUri="https://zenodo.org/record/4908246/files/figure.png" pageId="1" pageNumber="116">Fig. 6</figureCitation>
), and ovipositor shortened (
<figureCitation id="80A28EEB5D47FFED50DE2889C2C8D09E" box="[158,229,670,691]" captionStart="Figs" captionStartId="3.[175,215,1365,1384]" captionTargetBox="[192,1159,188,1342]" captionTargetId="figure-145@3.[151,1173,183,1352]" captionTargetPageId="3" captionText="Figs. 9–14. Scleropatroides triplehorni. 9) Labrum-epipharynx; 10) Maxilla; 11) Labium; 12) Aedeagus, dorsal view; 13) Aedeagus, ventral view; 14) Ovipositor and female tract." figureDoi="http://doi.org/10.5281/zenodo.4908248" httpUri="https://zenodo.org/record/4908248/files/figure.png" pageId="1" pageNumber="116">Fig. 14</figureCitation>
). It belongs to the subtribe
<taxonomicName id="DF99E9ED5D47FFED51A82889C014D09E" authorityName="Brulle" authorityYear="1832" box="[488,569,670,691]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="subTribe" subTribe="Opatrina">Opatrina</taxonomicName>
as evidenced by a female tract without a bursa copulatrix, with a single tubular spermatheca attached to the anterior end of the vagina (
<figureCitation id="80A28EEB5D47FFED51EF28FAC3D7D12F" box="[431,506,749,770]" captionStart="Figs" captionStartId="3.[175,215,1365,1384]" captionTargetBox="[192,1159,188,1342]" captionTargetId="figure-145@3.[151,1173,183,1352]" captionTargetPageId="3" captionText="Figs. 9–14. Scleropatroides triplehorni. 9) Labrum-epipharynx; 10) Maxilla; 11) Labium; 12) Aedeagus, dorsal view; 13) Aedeagus, ventral view; 14) Ovipositor and female tract." figureDoi="http://doi.org/10.5281/zenodo.4908248" httpUri="https://zenodo.org/record/4908248/files/figure.png" pageId="1" pageNumber="116">Fig. 14</figureCitation>
), and simple (not tripartite) aedeagus. The aedeagus (
<figureCitation id="80A28EEB5D47FFED5266291FC29DD115" captionStart="Figs" captionStartId="3.[175,215,1365,1384]" captionTargetBox="[192,1159,188,1342]" captionTargetId="figure-145@3.[151,1173,183,1352]" captionTargetPageId="3" captionText="Figs. 9–14. Scleropatroides triplehorni. 9) Labrum-epipharynx; 10) Maxilla; 11) Labium; 12) Aedeagus, dorsal view; 13) Aedeagus, ventral view; 14) Ovipositor and female tract." figureDoi="http://doi.org/10.5281/zenodo.4908248" httpUri="https://zenodo.org/record/4908248/files/figure.png" pageId="1" pageNumber="116">Figs. 12, 13</figureCitation>
) appears to have a normal apical piece of joined parameres as in most
<taxonomicName id="DF99E9ED5D47FFED5180292AC07FD17F" authorityName="Latreille" authorityYear="1802" box="[448,594,829,850]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
, but these are actually pseudoparameres
<taxonomicName id="DF99E9ED5D47FFED524B2940C2F5D1AA" authority="Iwan (2004)" authorityName="Iwan" authorityYear="2004" pageId="1" pageNumber="116" rank="species" species="sensu">
<emphasis id="2AED4E7C5D47FFED524B2940C06ED141" box="[523,579,855,876]" italics="true" pageId="1" pageNumber="116">sensu</emphasis>
<bibRefCitation id="7C08EF9F5D47FFED520C294FC2F5D1AA" author="Iwan" pageId="1" pageNumber="116" pagination="735 - 765" refId="ref2656" refString="Iwan, D. 2004. A comparative study of male genitalia in Opatrinae sensu Medvedev (1968) (Coleoptera: Tenebrionidae), with notes on the reinterpreted tribal classification. Part II. Annales Zoologici 54: 735 - 765." type="journal article" year="2004">Iwan (2004)</bibRefCitation>
</taxonomicName>
, consisting of fused original parameres plus the laciniae (clavae).
</paragraph>
<paragraph id="1826926E5D47FFED52852BF4C175D0B5" blockId="1.[685,1172,483,664]" pageId="1" pageNumber="116">
<taxonomicName id="DF99E9ED5D47FFED52852BF4C175D3D5" authorityName="Lobl and Merkl" authorityYear="2003" box="[709,856,483,504]" class="Insecta" family="Tenebrionidae" genus="Scleropatroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="2AED4E7C5D47FFED52852BF4C175D3D5" box="[709,856,483,504]" italics="true" pageId="1" pageNumber="116">Scleropatroides</emphasis>
</taxonomicName>
can be separated from the other Australian
<taxonomicName id="DF99E9ED5D47FFED53532BE9C14DD03E" box="[787,864,510,531]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="tribe" tribe="Opatrini">Opatrini</taxonomicName>
by using the key to the genera of
<taxonomicName id="DF99E9ED5D47FFED52ED280EC0D3D003" authorityName="Brulle" authorityYear="1832" box="[685,766,537,558]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="subTribe" subTribe="Opatrina">Opatrina</taxonomicName>
in Matthews and Bouchard (2008), where it appears under the erroneous name
<taxonomicName id="DF99E9ED5D47FFED544F2824C157D04E" authority="(Blackburn)" baseAuthorityName="Blackburn" class="Insecta" family="Tenebrionidae" genus="Mesomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="species" species="darwini">
<emphasis id="2AED4E7C5D47FFED544F2824C0D5D04F" italics="true" pageId="1" pageNumber="116">Mesomorphus darwini</emphasis>
(Blackburn)
</taxonomicName>
. The latter is in fact a valid species of
<taxonomicName id="DF99E9ED5D47FFED5357287FC600D053" authority="Miedel, 1880" authorityName="Miedel" authorityYear="1880" box="[791,1069,616,638]" class="Insecta" family="Tenebrionidae" genus="Mesomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="2AED4E7C5D47FFED5357287FC18DD050" box="[791,928,616,637]" italics="true" pageId="1" pageNumber="116">Mesomorphus</emphasis>
Miedel, 1880
</taxonomicName>
, as shown by
<bibRefCitation id="7C08EF9F5D47FFED528E2894C179D0B5" author="Ferrer" box="[718,852,643,664]" pageId="1" pageNumber="116" pagination="287 - 300" refId="ref2527" refString="Ferrer, J. 2008. Les especes australiennes du genre Mesomorphus Miedel, 1880, et description d' une nouvelle espece d' Afrique du Sud (Coleoptera, Tenebrionidae, Opatrini). Bulletin de la Societe entomologique de France 113 (3): 287 - 300." type="journal article" year="2008">Ferrer (2008)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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