187 lines
30 KiB
XML
187 lines
30 KiB
XML
<document id="EB56E3DA6D9224D4D11AB3CB8519B02D" ID-CLB-Dataset="8118" ID-DOI="10.5252/z2010n1a2" ID-GBIF-Dataset="4cecd899-5900-4411-8446-0d9552b1b098" ID-ISSN="1638-9387" ID-Zenodo-Dep="4520753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612808309234" checkinUser="felipe" docAuthor="Randriamanantsoa, Richard, Aberlenc, Henri-Pierre, Ralisoa, Olga Bakoly, Ratnadass, Alain & Vercambre, Bernard" docDate="2010" docId="901F8794FF98746CFD63FBA2FC5CFC03" docLanguage="fr" docName="Zoosystema.32.1.19-72.pdf" docOrigin="Zoosystema 32 (1)" docSource="http://www.bioone.org/doi/abs/10.5252/z2010n1a2" docStyle="DocumentStyle:E638A3C99FA728B529662B46F84D75FD.3:Zoosystema.2009-2010.journal_article" docStyleId="E638A3C99FA728B529662B46F84D75FD" docStyleName="Zoosystema.2009-2010.journal_article" docStyleVersion="3" docTitle="Heteroconus paradoxus" docType="treatment" docVersion="10" lastPageNumber="32" masterDocId="6C26FFECFF947461FFB5FFEDFF94FFE8" masterDocTitle="Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar" masterLastPageNumber="72" masterPageNumber="19" pageNumber="31" updateTime="1698928337295" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="091350890C05EA9EAE5E76E8503720A3">Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar</mods:title>
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<mods:roleTerm id="F1844428F94544009267299DE35C463A">Author</mods:roleTerm>
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<mods:namePart id="656953EF2EDBA4C5AD455912BE64B09A">Randriamanantsoa, Richard</mods:namePart>
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<mods:affiliation id="24F0E6284390A3EC1C7B7FE718910900">Fofifa - URP Scrid, 110 Antsirabe (Madagascar) r _ randriamanantsoa @ yahoo. fr</mods:affiliation>
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<mods:namePart id="4AE5D880B4D6DA4DE4F2025F264D51F7">Aberlenc, Henri-Pierre</mods:namePart>
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<mods:affiliation id="393B43C2FC16E5696E30FDF076CFBCB2">Cirad Bios, UMR CBGP (INRA / IRD / Cirad / Montpellier SupAgro), Campus international de Baillarguet, Ta A- 55 / L, F- 34398 Montpellier cedex 5 (France) henri-pierre. aberlenc @ cirad. fr</mods:affiliation>
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<mods:nameIdentifier id="F931B612CFC856CB01FDFDC073D4D73A" type="email">aberlenc@cirad.fr</mods:nameIdentifier>
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<mods:namePart id="27C2481517E94A2E515A99F8E818D5B4">Ralisoa, Olga Bakoly</mods:namePart>
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<mods:affiliation id="59CEE051FB43A7EA9733C900E21EB6F1">Département d’Entomologie, Faculté des Sciences, Université d’Antananarivo, Antananarivo 101 (Madagascar)</mods:affiliation>
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<mods:namePart id="3CB17882499E35D278F318D35C4589B2">Ratnadass, Alain</mods:namePart>
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<mods:affiliation id="675F78400A8477433FABA430F46F6641">Cirad, UPR HortSys - ICRISAT BP 12404, Niamey (Niger) a. ratnadass @ cgiar. org</mods:affiliation>
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<mods:nameIdentifier id="2E78C2AA26CB197F789385BB05FD714F" type="email">ratnadass@cgiar.org</mods:nameIdentifier>
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<mods:namePart id="CDBAA0DC89672F8108ABA94FB5CEEF76">Vercambre, Bernard</mods:namePart>
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<mods:affiliation id="A8CB2D575170784C2A858829374EF1B9">Cirad, UPR Systèmes de Cultures annuels, avenue Agropolis, F- 34398 Montpellier cedex 5 (France) bernard. vercambre @ cirad. fr</mods:affiliation>
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<mods:nameIdentifier id="D3F83BDC220B9BD238507EEB8B67754E" type="email">vercambre@cirad.fr</mods:nameIdentifier>
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<mods:title id="3475B95D6F851DCCC14EE9B268880BCA">Zoosystema</mods:title>
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<mods:part id="2798B9C430354F870B41DBB68986DD62">
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<mods:date id="3FF93DBFF1D0450463F9591BCB89FDE1">2010</mods:date>
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<mods:number id="920A95F56C64B182B17F6252A50C7DB9">2010-03-31</mods:number>
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<mods:number id="5A75B33EFCCE2691280762575627417E">32</mods:number>
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<mods:start id="CC6B994027FBFDD97D5101CAE3A5C355">19</mods:start>
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<mods:url id="DCEFA4D780A15657C99115CF2CB6E622">http://www.bioone.org/doi/abs/10.5252/z2010n1a2</mods:url>
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<mods:classification id="EFE174DCD5426F9E76BD50E53AADAB55">journal article</mods:classification>
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<mods:identifier id="6C679FB98D41DC5B8F10900AB5708535" type="DOI">10.5252/z2010n1a2</mods:identifier>
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<treatment id="901F8794FF98746CFD63FBA2FC5CFC03" ID-DOI="http://doi.org/10.5281/zenodo.4520719" ID-GBIF-Taxon="177425652" ID-Zenodo-Dep="4520719" LSID="urn:lsid:plazi:treatment:901F8794FF98746CFD63FBA2FC5CFC03" httpUri="http://treatment.plazi.org/id/901F8794FF98746CFD63FBA2FC5CFC03" lastPageId="13" lastPageNumber="32" pageId="12" pageNumber="31">
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<subSubSection id="50AC6509FF98746DFD63FBA2FCE1FB0F" pageId="12" pageNumber="31" type="nomenclature">
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<paragraph id="18093682FF98746DFD63FBA2FB1CFB82" blockId="12.[726,1160,1102,1193]" box="[726,1160,1102,1130]" pageId="12" pageNumber="31">
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<taxonomicName id="DFB64D01FF98746DFD63FBA2FB1CFB82" authority="(Endrodi, 1968)" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[726,1160,1102,1130]" class="Insecta" family="Dynastidae" genus="Heteroconus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="31" phylum="Arthropoda" rank="species" species="paradoxus">
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<emphasis id="2AC2EA90FF98746DFD63FBA2FC44FB81" bold="true" box="[726,976,1102,1129]" italics="true" pageId="12" pageNumber="31">Heteroconus paradoxus</emphasis>
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(Endrödi, 1968)
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</taxonomicName>
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</paragraph>
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<paragraph id="18093682FF98746DFD51FB82FBD8FB41" blockId="12.[726,1160,1102,1193]" pageId="12" pageNumber="31">
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(
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<figureCitation id="808D2A07FF98746DFD5EFB82FCA2FB62" box="[747,822,1135,1162]" captionStart="FIG" captionStartId="29.[113,124,1567,1584]" captionTargetBox="[152,1172,222,1545]" captionTargetId="figure-18@29.[148,709,692,1169]" captionTargetPageId="29" captionText="FIG. 2. — Larves de Scarabaeoidea de Madagascar, habitus: A, Bricoptis variolosa (Gory & Percheron, 1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paramorphochelus cornutus (Nonfried,1892); K, Encya sikorai (Brenske,1891); L, Apicencya waterloti (Dewailly, 1950); M, Empecta scutata (Fairmaire,1901); N, Hoplochelus betanimena (Künckel,1887); O, Triodontus nitidulus (Guérin, 1844). Échelles: A-I, K, L, N, 5 mm; J, O, 2 mm; M, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4520757" httpUri="https://zenodo.org/record/4520757/files/figure.png" pageId="12" pageNumber="31">Figs 2I</figureCitation>
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;
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<figureCitation id="808D2A07FF98746DFCFDFB82FCF8FB62" box="[840,876,1135,1162]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="12" pageNumber="31">4D</figureCitation>
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;
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<figureCitation id="808D2A07FF98746DFCC8FB82FC09FB62" box="[893,925,1135,1162]" captionStart="FIG" captionStartId="35.[113,124,1635,1652]" captionTargetBox="[141,1181,217,1589]" captionTargetId="figure-14@35.[141,1181,217,1589]" captionTargetPageId="35" captionText="FIG. 8. — Larves de Scarabaeoidea de Madagascar, épipharynx: A, Heteroconus paradoxus (Endrödi,1968); B, Paranodon coquerelii (Fairmaire, 1871); C, Paramorphochelus cornutus (Nonfried,1892); D, sp. M1; E, Encya sikorai (Brenske,1891); F, Enaria melanictera (Klug, 1833); G, sp. M2. Échelles: A, F, 0,5 mm; B, C, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520775" httpUri="https://zenodo.org/record/4520775/files/figure.png" pageId="12" pageNumber="31">8A</figureCitation>
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||
;
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<figureCitation id="808D2A07FF98746DFC1BFB82FC40FB62" box="[942,980,1135,1162]" captionStart="FIG" captionStartId="37.[113,124,1589,1606]" captionTargetBox="[146,1173,219,1547]" captionTargetId="figure-14@37.[146,1173,218,1547]" captionTargetPageId="37" captionText="FIG. 10. — Larves de Scarabaeoidea de Madagascar,hypopharynx: A, Bricoptis variolosa (Gory & Percheron,1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K, Paramorphochelus cornutus (Nonfried,1892); L, Encya sikorai (Brenske, 1891). Échelles: A, J, 0,2 mm; C, D, F, G, I, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520783" httpUri="https://zenodo.org/record/4520783/files/figure.png" pageId="12" pageNumber="31">10I</figureCitation>
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;
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<figureCitation id="808D2A07FF98746DFC50FB82FB9FFB62" box="[997,1035,1135,1162]" captionStart="FIG" captionStartId="39.[113,124,1567,1584]" captionTargetBox="[118,1207,216,1486]" captionTargetId="figure-14@39.[118,1207,216,1486]" captionTargetPageId="39" captionText="FIG. 12. — Larves de Scarabaeoidea de Madagascar: A-E, hypopharynx; A, Enaria melanictera (Klug, 1833); B, Apicencya waterloti (Dewailly, 1950); C, Empecta scutata (Fairmaire, 1901); D, Hoplochelus betanimena (Künckel, 1887); E, Triodontus nitidulus (Guérin, 1844); F-O, antenne; F, Anochilia bifida (Olivier, 1789); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K-M, Encya sikorai (Brenske,1891); L, M, plages sensorielles de l’article apical; N, Apicencya waterloti (Dewailly, 1950); O, Empecta scutata (Fairmaire, 1901). Échelles: A, F, G, I, J, O, 0,5 mm; E, L, M, 0,2 mm; H, K, N, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520787" httpUri="https://zenodo.org/record/4520787/files/figure.png" pageId="12" pageNumber="31">12I</figureCitation>
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;
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<figureCitation id="808D2A07FF98746DFBA9FB82FBA1FB62" box="[1052,1077,1135,1162]" captionStart="FIG" captionStartId="41.[113,124,1589,1606]" captionTargetBox="[161,1157,219,1557]" captionTargetId="figure-14@41.[157,1165,214,1560]" captionTargetPageId="41" captionText="FIG. 14. — Larves de Scarabaeoidea de Madagascar,mandibule: A, D, H, L, P, T, X, gauche,face dorsale; B, E, I, M, Q, U, Y, gauche, face ventrale; C, F, J, N, R, V, Z1, droite, face dorsale; G, K, O, S, W, Z2, droite, face ventrale; A-C, Bricoptis variolosa (Gory & Percheron, 1833); D-G, Anochilia bifida (Olivier, 1789); H-K, Euryomia argentea (Olivier, 1789); L-O, Heteronychus arator rugifrons (Fairmaire,1871); P-S, Heteronychus bituberculatus (Kolbe,1900); T-W, Hexodon unicolor (Olivier,1789); X-Z2, Heteroconus paradoxus (Endrödi, 1968). Échelles: A-C, H-O, 0,5 mm; D-G, P-Z2, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520795" httpUri="https://zenodo.org/record/4520795/files/figure.png" pageId="12" pageNumber="31">14</figureCitation>
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X-Z
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<subScript id="843234C7FF98746DFBD0FB96FBFBFB64" attach="left" box="[1125,1135,1147,1164]" fontSize="7" pageId="12" pageNumber="31">2</subScript>
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; 16O; 19E; 21U-W; 25N-P)
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</paragraph>
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<paragraph id="18093682FF98746DFD10FB23FCE1FB0F" blockId="12.[674,1210,1230,1478]" box="[677,885,1230,1257]" pageId="12" pageNumber="31">MATÉRIEL EXAMINÉ</paragraph>
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</subSubSection>
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<subSubSection id="50AC6509FF98746CFD10FB00FC5CFC03" lastPageId="13" lastPageNumber="32" pageId="12" pageNumber="31" type="description">
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<paragraph id="18093682FF98746DFD10FB00FC02FA2E" blockId="12.[674,1210,1230,1478]" pageId="12" pageNumber="31">
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Larves provenant d’Ankevobe (
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<quantity id="DF4E9B67FF98746DFC59FB00FBA5FAE0" box="[1004,1073,1261,1288]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="31" unit="km" value="13.0">13 km</quantity>
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d’Antsirabe) et d’Ampandrotrarana (
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<quantity id="DF4E9B67FF98746DFC20FAE0FC4DFAC0" box="[917,985,1293,1320]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="31" unit="km" value="20.0">20 km</quantity>
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d’Antsirabe) dans des cultures de Riz pluvial (
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<taxonomicName id="DFB64D01FF98746DFC28FAC0FB8AFAAE" box="[925,1054,1325,1351]" class="Liliopsida" family="Poaceae" genus="Oryza" kingdom="Plantae" order="Poales" pageId="12" pageNumber="31" phylum="Tracheophyta" rank="species" species="sativa">
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<emphasis id="2AC2EA90FF98746DFC28FAC0FB8AFAAE" box="[925,1054,1325,1351]" italics="true" pageId="12" pageNumber="31">Oryza sativa</emphasis>
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</taxonomicName>
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) et d’Arachide (
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<taxonomicName id="DFB64D01FF98746DFD19FAA1FCD9FA8E" box="[684,845,1356,1382]" class="Magnoliopsida" family="Fabaceae" genus="Arachis" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="31" phylum="Tracheophyta" rank="species" species="hypogea">
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<emphasis id="2AC2EA90FF98746DFD19FAA1FCD9FA8E" box="[684,845,1356,1382]" italics="true" pageId="12" pageNumber="31">Arachis hypogea</emphasis>
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</taxonomicName>
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L.) en haut de pente.
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<typeStatus id="C70D8820FF98746DFB80FAA1FBFEFA8F" box="[1077,1130,1356,1383]" pageId="12" pageNumber="31">Type</typeStatus>
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de sol: ferrallitique. Spécimens conservés dans les collections du Fofifa (20 larves), du Cirad-CBGP (3 larves) et du MNHN (2 larves).
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</paragraph>
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<paragraph id="18093682FF98746DFD10FA06FCA1F9EC" blockId="12.[677,1001,1515,1573]" box="[677,821,1515,1542]" pageId="12" pageNumber="31">DESCRIPTION</paragraph>
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<paragraph id="18093682FF98746DFD10F9E7FC7DF9CD" blockId="12.[677,1001,1515,1573]" box="[677,1001,1546,1573]" pageId="12" pageNumber="31">
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Couleur blanc ivoire (
|
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<figureCitation id="808D2A07FF98746DFC25F9E7FC48F9CD" box="[912,988,1546,1573]" captionStart="FIG" captionStartId="29.[113,124,1567,1584]" captionTargetBox="[152,1172,222,1545]" captionTargetId="figure-18@29.[148,709,692,1169]" captionTargetPageId="29" captionText="FIG. 2. — Larves de Scarabaeoidea de Madagascar, habitus: A, Bricoptis variolosa (Gory & Percheron, 1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paramorphochelus cornutus (Nonfried,1892); K, Encya sikorai (Brenske,1891); L, Apicencya waterloti (Dewailly, 1950); M, Empecta scutata (Fairmaire,1901); N, Hoplochelus betanimena (Künckel,1887); O, Triodontus nitidulus (Guérin, 1844). Échelles: A-I, K, L, N, 5 mm; J, O, 2 mm; M, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4520757" httpUri="https://zenodo.org/record/4520757/files/figure.png" pageId="12" pageNumber="31">Fig. 2I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746DFD10F9A7FD5BF98C" blockId="12.[677,1210,1610,1700]" box="[677,719,1610,1636]" pageId="12" pageNumber="31">
|
||
<emphasis id="2AC2EA90FF98746DFD10F9A7FD5BF98C" box="[677,719,1610,1636]" italics="true" pageId="12" pageNumber="31">Tête</emphasis>
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746CFD10F984FF6DFE9A" blockId="12.[677,1210,1610,1700]" lastBlockId="13.[113,648,216,370]" lastPageId="13" lastPageNumber="32" pageId="12" pageNumber="31">Couleur brun jaunâtre. Épicrâne avec 6 soies dorsoépicraniales et 6 soies épicraniales latéro-externes disposées symétriquement. 5 soies de chaque côté de l’angle frontal. 2 soies frontales antérieures disposées symétriquement. Clypéus transverse avec 5 soies clypéales externes et 1 soie interne de chaque côté.</paragraph>
|
||
<paragraph id="18093682FF99746CFFC4FE7BFED4FD87" blockId="13.[111,649,406,623]" pageId="13" pageNumber="32">
|
||
<emphasis id="2AC2EA90FF99746CFFC4FE7BFF4DFE58" bold="true" box="[113,217,406,432]" pageId="13" pageNumber="32">Antennes</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF99746CFF52FE7BFEABFE59" box="[231,319,406,433]" captionStart="FIG" captionStartId="39.[113,124,1567,1584]" captionTargetBox="[118,1207,216,1486]" captionTargetId="figure-14@39.[118,1207,216,1486]" captionTargetPageId="39" captionText="FIG. 12. — Larves de Scarabaeoidea de Madagascar: A-E, hypopharynx; A, Enaria melanictera (Klug, 1833); B, Apicencya waterloti (Dewailly, 1950); C, Empecta scutata (Fairmaire, 1901); D, Hoplochelus betanimena (Künckel, 1887); E, Triodontus nitidulus (Guérin, 1844); F-O, antenne; F, Anochilia bifida (Olivier, 1789); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K-M, Encya sikorai (Brenske,1891); L, M, plages sensorielles de l’article apical; N, Apicencya waterloti (Dewailly, 1950); O, Empecta scutata (Fairmaire, 1901). Échelles: A, F, G, I, J, O, 0,5 mm; E, L, M, 0,2 mm; H, K, N, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520787" httpUri="https://zenodo.org/record/4520787/files/figure.png" pageId="13" pageNumber="32">Fig. 12I</figureCitation>
|
||
). Les longueurs respectives des articles sont: I = 1,75; II = 1,91; III = 1,58; IV = 1. Une aire sensorielle apico-latérale sur l’avant-dernier article. Dernier article antennaire avec 4 plages sensorielles: 1 petite plage apicale ronde, 2 ventrales et 1 dorsale.7 longues soies sur l’article I et 3 longues soies sur l’article II.
|
||
</paragraph>
|
||
<paragraph id="18093682FF99746CFFC4FD7EFDFEFC23" blockId="13.[111,649,659,971]" pageId="13" pageNumber="32">
|
||
<emphasis id="2AC2EA90FF99746CFFC4FD7EFF20FD45" bold="true" box="[113,180,659,685]" pageId="13" pageNumber="32">Labre.</emphasis>
|
||
Transverse, asymétrique, avec de longues soies. Zone médiane du bord antérieur saillante, formant presque un lobe médian, légèrement différencié du côté droit, avec 4 soies antérieures externes et 2 soies antérieures internes rapprochées. De part et d’autre, 3 à 4 soies latéro-antérieures externes, 1 à 2 soies latéro-antérieures internes, puis 1 soie latérale. 2 longues soies labro-internes en avant du milieu. Une rangée irrégulière de 15 soies postérieures en 2 groupes de 8 et 7, la zone centrale étant glabre.
|
||
</paragraph>
|
||
<paragraph id="18093682FF99746CFFC4FC1DFECEFAE0" blockId="13.[111,649,1008,1288]" pageId="13" pageNumber="32">
|
||
<emphasis id="2AC2EA90FF99746CFFC4FC1DFF60FBE3" bold="true" box="[113,244,1008,1035]" pageId="13" pageNumber="32">Épipharynx</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF99746CFEB3FC1DFECAFBE3" box="[262,350,1008,1035]" captionStart="FIG" captionStartId="35.[113,124,1635,1652]" captionTargetBox="[141,1181,217,1589]" captionTargetId="figure-14@35.[141,1181,217,1589]" captionTargetPageId="35" captionText="FIG. 8. — Larves de Scarabaeoidea de Madagascar, épipharynx: A, Heteroconus paradoxus (Endrödi,1968); B, Paranodon coquerelii (Fairmaire, 1871); C, Paramorphochelus cornutus (Nonfried,1892); D, sp. M1; E, Encya sikorai (Brenske,1891); F, Enaria melanictera (Klug, 1833); G, sp. M2. Échelles: A, F, 0,5 mm; B, C, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520775" httpUri="https://zenodo.org/record/4520775/files/figure.png" pageId="13" pageNumber="32">Fig. 8A</figureCitation>
|
||
). Haptomère formant une saillie redressée sclérifiée arrondie. Acroparia avec de fortes soies très denses. Aire pédiale glabre, avec un sillon anguleux, en arc de cercle vers la base. Chaetoparia dissymétriques, à longues et fortes soies convergeant vers l’aire pédiale, les plus fortes soies étant à proximité de l’haptomère. Acanthoparia avec 21 à 22 soies lamelliformes, de taille croissante, incurvées vers l’avant.
|
||
</paragraph>
|
||
<paragraph id="18093682FF99746CFFC4FAC1FE99F94C" blockId="13.[112,649,1324,1700]" pageId="13" pageNumber="32">
|
||
<emphasis id="2AC2EA90FF99746CFFC4FAC1FF62FAAE" bold="true" box="[113,246,1324,1350]" pageId="13" pageNumber="32">Mandibules</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF99746CFEB2FAC0FEC0FAA0" box="[263,340,1325,1352]" captionStart="FIG" captionStartId="41.[113,124,1589,1606]" captionTargetBox="[161,1157,219,1557]" captionTargetId="figure-14@41.[157,1165,214,1560]" captionTargetPageId="41" captionText="FIG. 14. — Larves de Scarabaeoidea de Madagascar,mandibule: A, D, H, L, P, T, X, gauche,face dorsale; B, E, I, M, Q, U, Y, gauche, face ventrale; C, F, J, N, R, V, Z1, droite, face dorsale; G, K, O, S, W, Z2, droite, face ventrale; A-C, Bricoptis variolosa (Gory & Percheron, 1833); D-G, Anochilia bifida (Olivier, 1789); H-K, Euryomia argentea (Olivier, 1789); L-O, Heteronychus arator rugifrons (Fairmaire,1871); P-S, Heteronychus bituberculatus (Kolbe,1900); T-W, Hexodon unicolor (Olivier,1789); X-Z2, Heteroconus paradoxus (Endrödi, 1968). Échelles: A-C, H-O, 0,5 mm; D-G, P-Z2, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520795" httpUri="https://zenodo.org/record/4520795/files/figure.png" pageId="13" pageNumber="32">Fig. 14</figureCitation>
|
||
X-Z
|
||
<subScript id="843234C7FF99746CFE31FAD7FE1AFAA3" attach="left" box="[388,398,1338,1355]" fontSize="7" pageId="13" pageNumber="32">2</subScript>
|
||
). Aire scissoriale de la mandibule gauche avec 3 dents développées, la dent apicale très forte et tronquée, la seconde dent arrondie et la troisième aiguë. Aire scissoriale de la mandibule droite avec une forte dent apicale émoussée, la seconde dent arrondie. 19 à 20 soies soies baso-latérales et 3 soies dorso-externes. Face ventrale des deux mandibules avec une aire stridulatoire proche du lobe molaire et d’une touffe de courtes soies, très réduite, difficile à observer, avec seulement 4 à 5 bourrelets transverses, subconvergents, espacés.
|
||
</paragraph>
|
||
<paragraph id="18093682FF99746CFD10FF35FC0EFE9A" blockId="13.[677,1211,216,370]" pageId="13" pageNumber="32">
|
||
<emphasis id="2AC2EA90FF99746CFD10FF35FC93FF1A" bold="true" box="[677,775,216,242]" pageId="13" pageNumber="32">Maxilles</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF99746CFCADFF35FC12FF1B" box="[792,902,216,243]" captionStart="FIG" captionStartId="43.[113,124,1567,1584]" captionTargetBox="[152,1180,217,1546]" captionTargetId="figure-14@43.[151,1175,219,1547]" captionTargetPageId="43" captionText="FIG. 16. — Larves de Scarabaeoidea de Madagascar: A-D, mandibule, Triodontus nitidulus (Guérin, 1844); A, gauche, face dorsale; B, gauche, face ventrale; C, droite, face dorsale; D, droite, face ventrale; E-O, maxille, face dorsale; E, Bricoptis variolosa (Gory & Percheron, 1833); F, Celidota parvula (Janson, 1881); G, Anochilia bifida (Olivier, 1789); H, Euryomia argentea (Olivier, 1789); I, J, Heteronychus arator rugifrons (Fairmaire, 1871); J, dents stridulatoires; K, L, Heteronychus bituberculatus (Kolbe,1900); L, dents stridulatoires; M, Heteronychus plebeius (Klug,1833); N, Hexodon unicolor (Olivier,1789); O, Heteroconus paradoxus (Endrödi,1968). Échelles: A-D, N, O, 0,5 mm; E, F, H, I, K, 0,4 mm; G, 1 mm; J, 0,01 mm; L, 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.4520805" httpUri="https://zenodo.org/record/4520805/files/figure.png" pageId="13" pageNumber="32">Fig. 16O</figureCitation>
|
||
). Galea et lacinia soudées. Lacinia à uncus avec deux dents sclérifiées soudées à la base. Galea avec uncus sclérifié. Sur la face dorsale, aire stridulatoire composée de 12 à 15 dents à sommet aplati.
|
||
</paragraph>
|
||
<paragraph id="18093682FF99746CFD10FE7BFC70FDF8" blockId="13.[677,1212,406,528]" pageId="13" pageNumber="32">
|
||
<emphasis id="2AC2EA90FF99746CFD10FE7BFCD4FE58" bold="true" box="[677,832,406,432]" pageId="13" pageNumber="32">Hypopharynx</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF99746CFCE4FE7BFC39FE59" box="[849,941,406,433]" captionStart="FIG" captionStartId="37.[113,124,1589,1606]" captionTargetBox="[146,1173,219,1547]" captionTargetId="figure-14@37.[146,1173,218,1547]" captionTargetPageId="37" captionText="FIG. 10. — Larves de Scarabaeoidea de Madagascar,hypopharynx: A, Bricoptis variolosa (Gory & Percheron,1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K, Paramorphochelus cornutus (Nonfried,1892); L, Encya sikorai (Brenske, 1891). Échelles: A, J, 0,2 mm; C, D, F, G, I, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520783" httpUri="https://zenodo.org/record/4520783/files/figure.png" pageId="13" pageNumber="32">Fig. 10I</figureCitation>
|
||
). Glosses recouvertes au milieu et à la base par des soies courtes, fortes et pointues disposées en triangle et par de longues soies sur le reste de la surface.
|
||
</paragraph>
|
||
<paragraph id="18093682FF99746CFD10FDD9FD79FDA6" blockId="13.[675,1210,564,718]" box="[677,749,564,590]" pageId="13" pageNumber="32">
|
||
<emphasis id="2AC2EA90FF99746CFD10FDD9FD79FDA6" box="[677,749,564,590]" italics="true" pageId="13" pageNumber="32">Thorax</emphasis>
|
||
</paragraph>
|
||
<paragraph id="18093682FF99746CFD10FDB9FB20FD26" blockId="13.[675,1210,564,718]" pageId="13" pageNumber="32">
|
||
<emphasis id="2AC2EA90FF99746CFD10FDB9FD73FD86" bold="true" box="[677,743,596,622]" pageId="13" pageNumber="32">Pattes</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF99746CFD41FDB9FCC0FD87" box="[756,852,596,623]" captionStart="FIG" captionStartId="46.[113,124,1635,1652]" captionTargetBox="[121,1206,215,1527]" captionTargetId="figure-21@46.[117,1206,214,1542]" captionTargetPageId="46" captionText="FIG. 19. — Larves de Scarabaeoidea de Madagascar, pattes: A, Heteronychus arator rugifrons (Fairmaire, 1871); B, Heteronychus bituberculatus (Kolbe, 1900); C, Heteronychus plebeius (Klug, 1833); D, Hexodon unicolor (Olivier, 1789); E, Heteroconus paradoxus (Endrödi, 1968); F, Paranodon coquerelii (Fairmaire, 1871). Échelles: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520815" httpUri="https://zenodo.org/record/4520815/files/figure.png" pageId="13" pageNumber="32">Fig. 19E</figureCitation>
|
||
). Longues, de longueur croissante de P1 à P3, toutes de structure analogue: coxa, trochanter, fémur, tibiotarse, griffe longue, aiguë, incurvée et sclérifiée à l’apex. Coxa 3 très longue.
|
||
</paragraph>
|
||
<paragraph id="18093682FF99746CFD10FD1FFC93FCE4" blockId="13.[675,1210,754,1003]" box="[677,775,754,780]" pageId="13" pageNumber="32">
|
||
<emphasis id="2AC2EA90FF99746CFD10FD1FFC93FCE4" box="[677,775,754,780]" italics="true" pageId="13" pageNumber="32">Abdomen</emphasis>
|
||
</paragraph>
|
||
<paragraph id="18093682FF99746CFD10FCFFFC5CFC03" blockId="13.[675,1210,754,1003]" pageId="13" pageNumber="32">
|
||
<emphasis id="2AC2EA90FF99746CFD10FCFFFD7DFCC4" bold="true" box="[677,745,786,812]" pageId="13" pageNumber="32">Raster</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF99746CFD40FCFFFCDDFCC5" box="[757,841,786,813]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="13" pageNumber="32">Fig. 4D</figureCitation>
|
||
). Fente anale transverse. Une étroite bande glabre le long du côté dorsal de la fente anale. Du côté dorsal de la fente anale, nombreuses soies courtes et épaisses avec quelques très longues soies fines. Six longues soies du côté ventral de la fente anale.Teges avec plus d’une trentaine de très fortes soies recourbées et aplaties.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |