222 lines
26 KiB
XML
222 lines
26 KiB
XML
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<mods:title id="6030B42F14AB29001C05F896D38DC347">Two new species of Opheliidae (Annelida: Polychaeta): Euzonus papillatus sp. n. from a northeastern Brazilian sandy beach and Euzonus mammillatus sp. n. from the continental shelf of southeastern Brazil</mods:title>
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<mods:namePart id="36B3D07A5F6AE003313C3C3961B6D289">Santos, Cinthya S. G.</mods:namePart>
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<mods:namePart id="A3AFF9777A75DEC09E48A123148AE686">Nonato, Edmundo F.</mods:namePart>
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<mods:namePart id="027902C0A5F65E88AA684636831D9CD9">Petersen, Mary E.</mods:namePart>
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<mods:date id="732DE73247E71DA5E802BB18BF484480">2004</mods:date>
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<heading id="E92DE72DFF83FFF5FEF9FB4BFD1E5BA4" bold="true" box="[264,746,1214,1240]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
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<emphasis id="80AE8C53FF83FFF5FEF9FB4BFD1E5BA4" bold="true" box="[264,746,1214,1240]" pageId="3" pageNumber="4">
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<taxonomicName id="75DA2BC2FF83FFF5FEF9FB4BFDE35BA4" ID-CoL="3DG4N" authority="Santos & Nonato & Petersen, 2004" authorityName="Santos & Nonato & Petersen" authorityYear="2004" box="[264,535,1214,1240]" class="Polychaeta" family="Opheliidae" genus="Euzonus" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="mammillatus" status="sp. nov.">
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<emphasis id="80AE8C53FF83FFF5FEF9FB4BFDE35BA4" bold="true" box="[264,535,1214,1240]" italics="true" pageId="3" pageNumber="4">Euzonus mammillatus</emphasis>
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<taxonomicNameLabel id="9B9D3128FF83FFF5FDEEFB4BFD955BA4" box="[543,609,1214,1240]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
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(
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<figureCitation id="2AE14CC4FF83FFF5FD81FB4BFD165BA4" box="[624,738,1214,1240]" captionStart="FIGURES 4–7" captionStartId="4.[264,383,1088,1112]" captionTargetBox="[361,1248,334,1007]" captionTargetId="figure-271@4.[309,1257,302,1087]" captionTargetPageId="4" captionText="FIGURES 4–7. Euzonus mammillatus sp. n., specimen from SE Brazil, Rio de Janeiro State, from Nonato (1981): 4. right lateral view of whole body; 5. details of lateral papillae on chaetiger 10 and branchiae; 6. posterior end, dorsal view; 7. posterior end, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5030068" httpUri="https://zenodo.org/record/5030068/files/figure.png" pageId="3" pageNumber="4">Figs. 4–7</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="B2655041FF83FFF5FEF9FAF1FD8B5A47" blockId="3.[264,1323,1284,1339]" pageId="3" pageNumber="4">
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<taxonomicName id="75DA2BC2FF83FFF5FEF9FAF1FE215A67" authorityName="Nonato" box="[264,469,1284,1307]" class="Polychaeta" family="Opheliidae" genus="Euzonus" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="mamillata">
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<emphasis id="80AE8C53FF83FFF5FEF9FAF1FE215A67" box="[264,469,1284,1307]" italics="true" pageId="3" pageNumber="4">Euzonus mamillata</emphasis>
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</taxonomicName>
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[sic]
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<treatmentCitation id="337B7650FF83FFF5FDE5FAF1FCFB5A67" author="Nonato, E." box="[532,783,1284,1307]" page="182" pageId="3" pageNumber="4" year="1981">
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<bibRefCitation id="D64B2DB0FF83FFF5FDE5FAF1FCFB5A67" author="Nonato, E." box="[532,783,1284,1307]" pageId="3" pageNumber="4" refId="ref6588" refString="Nonato, E. (1981) Contribuicao ao conhecimento dos anelideos poliquetos bentonicos da plataforma continental brasileira, entre Cabo Frio e o Arroio Chui. Tese de Livre Docencia, USP- IO. 246 pp." type="book" year="1981">Nonato, 1981: 182184</bibRefCitation>
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</treatmentCitation>
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, figs. 202–203 [Manuscript name not validly published according to the ICZN].
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="FAC003CAFF83FFF5FEF9FA9EFBA5595A" pageId="3" pageNumber="4" type="materials_examined">
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<paragraph id="B2655041FF83FFF5FEF9FA9EFD975AAA" blockId="3.[264,1323,1387,1814]" pageId="3" pageNumber="4">
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<emphasis id="80AE8C53FF83FFF5FEF9FA9EFC2C5AF9" bold="true" box="[264,984,1387,1413]" pageId="3" pageNumber="4">Material originally examined by E.F. Nonato (lost): SE</emphasis>
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<materialsCitation id="02B25A1CFF83FFF5FC14FA9EFDAB5AAA" ID-GBIF-Occurrence="3337330301" collectionCode="R" country="Brazil" elevation="33" location="Lopes Mendes Bay" municipality="Ilha Grande" pageId="3" pageNumber="4" specimenCount="3" stateProvince="Rio de Janeiro">
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<collectingCountry id="CACD10D1FF83FFF5FC14FA9EFBA15AF9" box="[997,1109,1387,1413]" name="Brazil" pageId="3" pageNumber="4">BRAZIL</collectingCountry>
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</emphasis>
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:
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<collectingRegion id="701E9EA3FF83FFF5FB9BFA9EFEB25AD1" country="Brazil" name="Rio de Janeiro" pageId="3" pageNumber="4">Rio de Janeiro State</collectingRegion>
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</emphasis>
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:
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<collectingMunicipality id="5201CA3BFF83FFF5FEA6FA61FE125AD2" box="[343,486,1428,1454]" pageId="3" pageNumber="4">Ilha Grande</collectingMunicipality>
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,
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,
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<elevation id="39F7B772FF83FFF5FD16FA61FCA65AD2" box="[743,850,1428,1454]" metricMagnitude="1" metricUnit="m" metricValue="3.25" metricValueMax="4.3" metricValueMin="2.2" pageId="3" pageNumber="4" unit="m" value="32.5" valueMax="43.0" valueMin="22.0">22–43 m</elevation>
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</quantity>
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, gravel and coarse sand with mud, coll. L.
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<collectionCode id="D4CBC884FF83FFF5FED0FA49FECC5AAA" box="[289,312,1468,1494]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="4">R</collectionCode>
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. Tommasi (
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<specimenCount id="A4DC9BC8FF83FFF5FE30FA49FDA25AAA" box="[449,598,1468,1494]" pageId="3" pageNumber="4" type="generic">3 specimens</specimenCount>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="B2655041FF83FFF5FEC9FA16FBA5595A" blockId="3.[264,1323,1387,1814]" pageId="3" pageNumber="4">
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:
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<collectorName id="1F2F3597FF83FFF5FE31FA16FD2C5A81" box="[448,728,1507,1533]" pageId="3" pageNumber="4">Continental Shelf, SE</collectorName>
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<collectingCountry id="CACD10D1FF83FFF5FD14FA16FCC25A81" box="[741,822,1507,1533]" name="Brazil" pageId="3" pageNumber="4">Brazil</collectingCountry>
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:
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<collectingRegion id="701E9EA3FF83FFF5FCB9FA16FBA75A81" box="[840,1107,1507,1533]" country="Brazil" name="Rio de Janeiro" pageId="3" pageNumber="4">Rio de Janeiro State</collectingRegion>
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</emphasis>
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: 22º57’–
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<geoCoordinate id="D7EE3686FF83FFF5FB39FA11FAD15A82" box="[1224,1317,1508,1534]" degrees="23" direction="south" minutes="00" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="-23.0">23º00’S</geoCoordinate>
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; 47º07’–
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<geoCoordinate id="D7EE3686FF83FFF5FE94F9F9FE38595A" box="[357,460,1548,1574]" degrees="42" direction="west" minutes="11" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="-42.183334">42º11’W</geoCoordinate>
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,
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<quantity id="7522FDA4FF83FFF5FE29F9F9FDE1595A" box="[472,533,1548,1574]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="4" unit="m" value="45.0">
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<elevation id="39F7B772FF83FFF5FE29F9F9FDE1595A" box="[472,533,1548,1574]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="4" unit="m" value="45.0">45 m</elevation>
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</quantity>
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, coarse sand, coll.
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<collectorName id="1F2F3597FF83FFF5FD02F9F9FCA4595A" box="[755,848,1548,1574]" pageId="3" pageNumber="4">P. Paiva</collectorName>
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(
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<specimenCode id="E27CF83AFF83FFF5FCAEF9F9FBB0595A" box="[863,1092,1548,1574]" collectionCode="MCEM-BPO" pageId="3" pageNumber="4">MCEMBPO 1630</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="FAC003CAFF83FFF5FEC9F9C6FC9C586A" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="B2655041FF83FFF5FEC9F9C6FC9C586A" blockId="3.[264,1323,1387,1814]" pageId="3" pageNumber="4">
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<emphasis id="80AE8C53FF83FFF5FEC9F9C6FE435931" bold="true" box="[312,439,1587,1613]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
|
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<taxonomicName id="75DA2BC2FF83FFF5FE4EF9C1FDD65931" authorityName="Grube" authorityYear="1866" box="[447,546,1588,1613]" class="Polychaeta" family="Opheliidae" genus="Euzonus" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
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<emphasis id="80AE8C53FF83FFF5FE4EF9C1FDD65931" box="[447,546,1588,1613]" italics="true" pageId="3" pageNumber="4">Euzonus</emphasis>
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</taxonomicName>
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||
with slender body clearly divided into three regions by weak constrictions; body formula 11a+18b+6a = 35 chaetigers. Cephalic region with 2 chaetigers, chaetiger 1 uniramous; thoracic region with 8 chaetigers, last of which (chaetiger 10) with 2+5 mammilliform papillae in a long, dorsoventrally oriented band just above notopodium of each side, reaching nearly to middorsum. Last abranchiate chaetigers uniramous; anal cirri with ventral pair stout, longer than dorsal ones.
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="E6A500C9FF84FFF2FEF9FBB5FC0D5BE0" ID-DOI="http://doi.org/10.5281/zenodo.5030068" ID-Zenodo-Dep="5030068" httpUri="https://zenodo.org/record/5030068/files/figure.png" pageId="4" pageNumber="5" startId="4.[264,383,1088,1112]" targetBox="[361,1248,334,1007]" targetPageId="4">
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<paragraph id="B2655041FF84FFF2FEF9FBB5FC0D5BE0" blockId="4.[264,1323,1088,1180]" pageId="4" pageNumber="5">
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<emphasis id="80AE8C53FF84FFF2FEF9FBB5FE405B24" bold="true" box="[264,436,1088,1112]" pageId="4" pageNumber="5">FIGURES 4–7.</emphasis>
|
||
<taxonomicName id="75DA2BC2FF84FFF2FE4DFBB4FD535B24" authority="Santos & Nonato & Petersen, 2004" authorityName="Santos & Nonato & Petersen" authorityYear="2004" box="[444,679,1089,1112]" class="Polychaeta" family="Opheliidae" genus="Euzonus" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="mammillatus" status="sp. nov.">
|
||
<emphasis id="80AE8C53FF84FFF2FE4DFBB4FD535B24" box="[444,679,1089,1112]" italics="true" pageId="4" pageNumber="5">Euzonus mammillatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="9B9D3128FF84FFF2FD5FFBB5FD1D5B2B" box="[686,745,1088,1111]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
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, specimen from SE Brazil, Rio de Janeiro State, from
|
||
<bibRefCitation id="D64B2DB0FF84FFF2FEF9FB96FE535B06" author="Nonato, E." box="[264,423,1123,1146]" pageId="4" pageNumber="5" refId="ref6588" refString="Nonato, E. (1981) Contribuicao ao conhecimento dos anelideos poliquetos bentonicos da plataforma continental brasileira, entre Cabo Frio e o Arroio Chui. Tese de Livre Docencia, USP- IO. 246 pp." type="book" year="1981">Nonato (1981)</bibRefCitation>
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||
: 4. right lateral view of whole body; 5. details of lateral papillae on chaetiger 10 and branchiae; 6. posterior end, dorsal view; 7. posterior end, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="FAC003CAFF84FFF3FEC9FB0DFCA15EAA" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
|
||
<paragraph id="B2655041FF84FFF3FEC9FB0DFC485EFA" blockId="4.[264,1325,1272,1898]" lastBlockId="5.[264,1323,284,830]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
<emphasis id="80AE8C53FF84FFF2FEC9FB0DFE245A6E" bold="true" box="[312,464,1272,1298]" pageId="4" pageNumber="5">Description.</emphasis>
|
||
Preserved material colorless; color in life unknown.
|
||
<typeStatus id="6D61EEE3FF84FFF2FB99FB0DFB215A6E" box="[1128,1237,1272,1298]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
|
||
<quantity id="7522FDA4FF84FFF2FB13FB0DFADF5A6E" box="[1250,1323,1272,1298]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="5" unit="mm" value="6.0">6 mm</quantity>
|
||
long; material examined by
|
||
<bibRefCitation id="D64B2DB0FF84FFF2FDA3FAD5FD0A5A46" author="Nonato, E." box="[594,766,1312,1338]" pageId="4" pageNumber="5" refId="ref6588" refString="Nonato, E. (1981) Contribuicao ao conhecimento dos anelideos poliquetos bentonicos da plataforma continental brasileira, entre Cabo Frio e o Arroio Chui. Tese de Livre Docencia, USP- IO. 246 pp." type="book" year="1981">Nonato (1981)</bibRefCitation>
|
||
up to
|
||
<quantity id="7522FDA4FF84FFF2FCB9FAD5FC6F5A46" box="[840,923,1312,1338]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="5" unit="mm" value="30.0">30 mm</quantity>
|
||
long, all specimens with 35 chaetigers. Body slender, clearly divided into three regions by weak constrictions. Segments of thoracic region fairly well delimited, faintly annulated; segments of abdominal region multiannulate, with segmental limits less distinct. Cephalic region with oval nuchal organs and 2 chaetigers, thoracic region with 8 chaetigers, last of which (chaetiger 10) with 2+5 mammilliform papillae in a long, dorsoventrally oriented band just above notopodium of each side, reaching nearly to middorsum; chaetigers 9–16 with 2 mamilliform papillae more or less anterior to neuropodia, uppermost papilla of which sometimes interramal (
|
||
<figureCitation id="2AE14CC4FF84FFF2FEE0F995FE705906" box="[273,388,1632,1658]" captionStart="FIGURES 4–7" captionStartId="4.[264,383,1088,1112]" captionTargetBox="[361,1248,334,1007]" captionTargetId="figure-271@4.[309,1257,302,1087]" captionTargetPageId="4" captionText="FIGURES 4–7. Euzonus mammillatus sp. n., specimen from SE Brazil, Rio de Janeiro State, from Nonato (1981): 4. right lateral view of whole body; 5. details of lateral papillae on chaetiger 10 and branchiae; 6. posterior end, dorsal view; 7. posterior end, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5030068" httpUri="https://zenodo.org/record/5030068/files/figure.png" pageId="4" pageNumber="5">Figs. 4–5</figureCitation>
|
||
). Branchiae bifid, with 2 subequal, long smooth, slender branches. Chaetae all capillaries, without distal modification; chaetiger 1 uniramous, with single bundle of chaetae, chaetigers 229 biramous; notochaetae of 2–28 only slightly longer and more numerous than neurochaetae; notochaetae of 29 becoming much longer than neurochaetae (
|
||
<figureCitation id="2AE14CC4FF84FFF2FB01F92DFECC5866" captionStart="FIGURES 4–7" captionStartId="4.[264,383,1088,1112]" captionTargetBox="[361,1248,334,1007]" captionTargetId="figure-271@4.[309,1257,302,1087]" captionTargetPageId="4" captionText="FIGURES 4–7. Euzonus mammillatus sp. n., specimen from SE Brazil, Rio de Janeiro State, from Nonato (1981): 4. right lateral view of whole body; 5. details of lateral papillae on chaetiger 10 and branchiae; 6. posterior end, dorsal view; 7. posterior end, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5030068" httpUri="https://zenodo.org/record/5030068/files/figure.png" pageId="4" pageNumber="5">Figs. 6–7</figureCitation>
|
||
); chaetigers 29–
|
||
<emphasis id="80AE8C53FF84FFF2FE06F8F5FDE05866" bold="true" box="[503,532,1792,1818]" pageId="4" pageNumber="5">34</emphasis>
|
||
(= last branchiate + all posterior abranchiate) uniramous, with only very long notochaetae, about twice as long as on chaetiger 28; notochaetae several times as long as those of preceding segments except for chaetiger 34, where they are only about half as long as on chaetigers 30–34. Extent of ventral groove unknown. Pygidium broad at base, tapering to double midventral anal cirri (
|
||
<figureCitation id="2AE14CC4FF85FFF3FCC8FEB1FC775E22" box="[825,899,324,350]" captionStart="FIGURES 4–7" captionStartId="4.[264,383,1088,1112]" captionTargetBox="[361,1248,334,1007]" captionTargetId="figure-271@4.[309,1257,302,1087]" captionTargetPageId="4" captionText="FIGURES 4–7. Euzonus mammillatus sp. n., specimen from SE Brazil, Rio de Janeiro State, from Nonato (1981): 4. right lateral view of whole body; 5. details of lateral papillae on chaetiger 10 and branchiae; 6. posterior end, dorsal view; 7. posterior end, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5030068" httpUri="https://zenodo.org/record/5030068/files/figure.png" pageId="5" pageNumber="6">Fig. 7</figureCitation>
|
||
). Dorsal anal cirri disposed as a V over ventral plate,
|
||
<quantity id="7522FDA4FF85FFF3FE12FE99FDBF5EFA" box="[483,587,364,390]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="4.572" metricValueMin="3.048" pageId="5" pageNumber="6" unit="in" value="15.0" valueMax="18.0" valueMin="12.0">12–18 in</quantity>
|
||
all, with 6–9 cirri on each side.
|
||
</paragraph>
|
||
<paragraph id="B2655041FF85FFF3FEC9FE66FCA15EAA" blockId="5.[264,1323,284,830]" pageId="5" pageNumber="6">
|
||
<emphasis id="80AE8C53FF85FFF3FEC9FE66FE515ED1" bold="true" box="[312,421,403,429]" pageId="5" pageNumber="6">Records.</emphasis>
|
||
Known from Lopes Mendes Bay,
|
||
<collectingCountry id="CACD10D1FF85FFF3FCB3FE61FC9B5ED2" box="[834,879,404,430]" name="Iceland" pageId="5" pageNumber="6">Ilha</collectingCountry>
|
||
Grande and continental shelf off
|
||
<collectingRegion id="701E9EA3FF85FFF3FAF1FE61FE305EAA" country="Brazil" name="Rio de Janeiro" pageId="5" pageNumber="6">Rio de Janeiro State</collectingRegion>
|
||
, SE
|
||
<collectingCountry id="CACD10D1FF85FFF3FE08FE49FDB05EAA" box="[505,580,444,470]" name="Brazil" pageId="5" pageNumber="6">Brazil</collectingCountry>
|
||
,
|
||
<quantity id="7522FDA4FF85FFF3FDA1FE49FD4E5EAA" box="[592,698,444,470]" metricMagnitude="1" metricUnit="m" metricValue="3.35" metricValueMax="4.5" metricValueMin="2.2" pageId="5" pageNumber="6" unit="m" value="33.5" valueMax="45.0" valueMin="22.0">22–45 m</quantity>
|
||
, coarse sand.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FAC003CAFF85FFF3FEC9FE16FCFD5C42" pageId="5" pageNumber="6" type="discussion">
|
||
<paragraph id="B2655041FF85FFF3FEC9FE16FCFD5C42" blockId="5.[264,1323,284,830]" pageId="5" pageNumber="6">
|
||
<emphasis id="80AE8C53FF85FFF3FEC9FE16FE455E81" bold="true" box="[312,433,483,509]" pageId="5" pageNumber="6">Remarks.</emphasis>
|
||
Material of
|
||
<taxonomicName id="75DA2BC2FF85FFF3FDB7FE11FD275E82" box="[582,723,484,510]" class="Polychaeta" family="Opheliidae" genus="Euzonus" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="undetermined">
|
||
<emphasis id="80AE8C53FF85FFF3FDB7FE11FD5D5E81" box="[582,681,484,509]" italics="true" pageId="5" pageNumber="6">Euzonus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
from southeastern
|
||
<collectingCountry id="CACD10D1FF85FFF3FC48FE11FBF55E82" box="[953,1025,484,510]" name="Brazil" pageId="5" pageNumber="6">Brazil</collectingCountry>
|
||
was referred to as a new species,
|
||
<taxonomicName id="75DA2BC2FF85FFF3FE81FDF9FDA25D59" box="[368,598,524,549]" class="Polychaeta" family="Opheliidae" genus="Euzonus" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="mamillata">
|
||
<emphasis id="80AE8C53FF85FFF3FE81FDF9FDA25D59" box="[368,598,524,549]" italics="true" pageId="5" pageNumber="6">Euzonus mamillata</emphasis>
|
||
</taxonomicName>
|
||
[sic], by
|
||
<bibRefCitation id="D64B2DB0FF85FFF3FD22FDF9FC775D5A" author="Nonato, E." box="[723,899,524,550]" pageId="5" pageNumber="6" refId="ref6588" refString="Nonato, E. (1981) Contribuicao ao conhecimento dos anelideos poliquetos bentonicos da plataforma continental brasileira, entre Cabo Frio e o Arroio Chui. Tese de Livre Docencia, USP- IO. 246 pp." type="book" year="1981">Nonato (1981)</bibRefCitation>
|
||
in an unpublished thesis, but was never published formally. The original material has been lost and it is therefore not possible to reexamine it or to fix the name on a
|
||
<typeStatus id="6D61EEE3FF85FFF3FCC5FDA9FC925D0A" box="[820,870,604,630]" pageId="5" pageNumber="6">type</typeStatus>
|
||
from one of the original specimens. According to the new International Code of Zoological Nomenclature (ICZN 1999, effective from
|
||
<date id="C6647681FF85FFF3FE8CFD59FDC25DBA" box="[381,566,684,710]" pageId="5" pageNumber="6" value="2000-01-01">1 January 2000</date>
|
||
), a species described after 1999 can only be validly described if the name can be fixed on a
|
||
<typeStatus id="6D61EEE3FF85FFF3FDB7FD21FD8C5D92" box="[582,632,724,750]" pageId="5" pageNumber="6">type</typeStatus>
|
||
specimen (ICZN 1999 Art. 72.3). Dr. Paulo Paiva’s find of new material of the species from the same general area as the worms studied by Nonato makes it possible to fulfill this requirement.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |