158 lines
20 KiB
XML
158 lines
20 KiB
XML
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<mods:title id="DEFBA6EF8AA69C606EEAA7CA8B2FB3EA">A catalogue of the ants of Paraguay (Hymenoptera: Formicidae).</mods:title>
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<mods:namePart id="EAEC94A12631B393DE8C92B5F373104F">Wild, A. L.</mods:namePart>
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<mods:date id="39734D87A5EACBD9571FE7F64B6C4A66">2007</mods:date>
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<treatment id="326A12AB5D4927902F78BB69DF317F3B" ID-DOI="http://doi.org/10.5281/zenodo.6239898" ID-GBIF-Taxon="100131673" ID-Zenodo-Dep="6239898" LSID="urn:lsid:plazi:treatment:326A12AB5D4927902F78BB69DF317F3B" httpUri="http://treatment.plazi.org/id/326A12AB5D4927902F78BB69DF317F3B" lastPageNumber="46" pageNumber="45">
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<subSubSection id="894765B4CA310E26AC8EE5EFB49AF381" type="nomenclature">
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<paragraph id="0A23AB988F1F43C06FC9A8938F09EF5F" pageNumber="45">
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<taxonomicName id="467ECD19B19B354B16AFCDFAAC11CECD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:134673" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus pellitus Mayr" order="Hymenoptera" pageNumber="45" phylum="Arthropoda" rank="species" species="pellitus">Camponotus pellitus Mayr</taxonomicName>
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1862.
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<taxonomicNameLabel id="AAD13D0857F438E0E94855D22FAD3D4B">REVISED STATUS</taxonomicNameLabel>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="069543103B8A23EB01310B68E3ADC300" type="reference_group">
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<paragraph id="709AC65E7793FB78916C34A7E0210C75" pageNumber="45">
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<taxonomicName id="20975930DD6F8FA107F83A7C65C5DDF8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:134673" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus pellitus Mayr" order="Hymenoptera" pageNumber="45" phylum="Arthropoda" rank="species" species="pellitus">Camponotus pellitus Mayr</taxonomicName>
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1862:668.
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<materialsCitation id="1703864B3772132502296B947378E3A1" ID-GBIF-Occurrence="923905288" collectionCode="NHMW" country="Brazil" typeStatus="holotype">
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[
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<typeStatus id="AC34A14E233E001D4CCA3A8CDFB4CEFE">holotype</typeStatus>
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w examined,
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<collectionCode id="5FBF9AF16C9EDADC68FD987AFF057360" collectionName="Austria, Wien, Naturhistorisches Museum Wien">NHMW</collectionCode>
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;
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<collectingCountry id="81189A2111E0A3E94EE14CDF208ED1CD">Brazil</collectingCountry>
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]
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</materialsCitation>
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</paragraph>
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<paragraph id="8A825B055AB60A2A67016BFE698B4596" pageNumber="45">
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<taxonomicName id="473694E4AAA2AE27F37E4C90D8D44293" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:134676" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus pellitus r. scintillans Forel" order="Hymenoptera" pageNumber="45" phylum="Arthropoda" rank="variety" species="pellitus" variety="scintillans">Camponotus pellitus var. scintillans Forel</taxonomicName>
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1901e: 72.
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<taxonomicNameLabel id="1DD4E7F1F7D6077762029DE97B7255FC">NEW SYNONYMY</taxonomicNameLabel>
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.
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<materialsCitation id="181C13F90FAABD543F7A7F701B235AF1" ID-GBIF-Occurrence="923905289" collectionCode="MHNG" collectorName="Jerrmann" country="Paraguay" typeStatus="syntypes">
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[w
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<typeStatus id="80E178F5118A8349C68CBA8DEFF8139C">syntypes</typeStatus>
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examined,
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<collectionCode id="31492286C96BCB0F41D97FE91DE597FF" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
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;
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<collectingCountry id="E64CDF4AD2C4DC84BAFF492307614B5C">Paraguay</collectingCountry>
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(
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<collectorName id="377A251EAD1024D2AF5CBFDEBE06494E">Jerrmann</collectorName>
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)]
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</materialsCitation>
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</paragraph>
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<paragraph id="0801C3422ACACDACC39FCB321A2F7B79" pageNumber="45">
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<taxonomicName id="8AFEA35EE3F6EDBBAF11248E42A5C8E9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26268" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus blandus (Smith)" order="Hymenoptera" pageNumber="45" phylum="Arthropoda" rank="species" species="blandus">Camponotus blandus Emery (nec F. Smith)</taxonomicName>
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1903: 67 (part). Misidentification.
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</paragraph>
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<paragraph id="34091A6E9529E329D7AB14ECAD4A100C" pageNumber="45">
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<taxonomicName id="2D0A8A929418A94E57E93B78E1C6DC57" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239756" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus (Myrmamblis) pellitus" order="Hymenoptera" pageNumber="45" phylum="Arthropoda" rank="species" species="pellitus" subGenus="Myrmamblis">Camponotus (Myrmamblis) pellitus Mayr</taxonomicName>
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. Bruch 1914: 229.
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</paragraph>
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<paragraph id="2A5756BDC2A73A8867687E93873C088B" pageNumber="45">
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<taxonomicName id="B6076954A9834F315F43883A8602BE9D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:148687" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus blandus pellitus Mayr" order="Hymenoptera" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="blandus" subSpecies="pellitus">Camponotus blandus pellitus Mayr</taxonomicName>
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. Santschi 1922b: 111.
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</paragraph>
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<paragraph id="4B9ECE9FA3360ABCD3386ED9465D864A" pageNumber="46">
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<taxonomicName id="DCE7243040CA67F478896B74553E5342" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:148786" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus blandus scintillans Forel" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="blandus" subSpecies="scintillans">Camponotus blandus st. scintillans Forel</taxonomicName>
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. Santschi 1922b: 111.
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</paragraph>
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<paragraph id="9923DE2450E67562D3ABBE44F807C403" pageNumber="46">
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<taxonomicName id="0F75A3473276B2D2E5B522AA7856024E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:236985" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus (Myrmaphaenus) blandus subsp. scintillans Forel" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="blandus" subGenus="Myrmaphaenus" subSpecies="scintillans">Camponotus (Myrmaphaenus) blandus subsp. scintillans Forel</taxonomicName>
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. Emery 1925: 154.
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</paragraph>
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<paragraph id="058E41245BD421015290738F30C9F64E" pageNumber="46">
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<taxonomicName id="9BA9F8EE3F77D46EB8EE6126A3D5BD4E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:148687" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus blandus pellitus Mayr" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="variety" species="blandus" subGenus="Myrmaphaenus" variety="pellitus">Camponotus (Myrmaphaenus) blandus var. pellitus Mayr</taxonomicName>
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. Kempf 1972: 44.
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</paragraph>
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<paragraph id="96BB265BF7201C4D3507590D2596952B" pageNumber="46">
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<taxonomicName id="E9F881535E978E9E471401DBE1A5AD55" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:236985" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus (Myrmaphaenus) blandus subsp. scintillans Forel" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="blandus" subGenus="Myrmaphaenus" subSpecies="scintillans">Camponotus (Myrmaphaenus) blandus scintillans Forel</taxonomicName>
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. Kempf 1972: 45.
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</paragraph>
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</subSubSection>
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<subSubSection id="D88E7B9086DE8A6A9694EB0945F98E82" type="discussion">
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<paragraph id="48A97B1DAAEF01244B5352461BC009FF" pageNumber="46">
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The names associated with
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<taxonomicName id="20C2F2B40F04982BF5321F2AD747E1A4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26268" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus blandus (Smith)" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="blandus">Camponotus blandus</taxonomicName>
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present one of the most difficult taxonomic problems in the Paraguayan fauna, and one that is particularly frustrating given the abundance of these ants in open habitats throughout the region. Multiple closely-related species certainly occur in Paraguay, but aside from Santschi’s
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<taxonomicName id="C823990D735473753BB7764FE7E29971" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239656" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus crispulus" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="crispulus">C. crispulus</taxonomicName>
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(see separate discussion under that species), the taxonomic status of these forms is ambiguous and the arrangement in the present study is tenuous.
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</paragraph>
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<paragraph id="23BF400303633A58CE6084D210D323CB" pageNumber="46">
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In eastern Paraguay, most collections are of dark-colored ants closely matching Mayr’s type of
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<taxonomicName id="A8420CA5EACB277B4EBA652E143E595E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:134673" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus pellitus Mayr" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">C. pellitus</taxonomicName>
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from Brazil and Forel’s types of
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<taxonomicName id="5A6B112AF89334EC2D3A3E85391A6DA2" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:134676" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus pellitus r. scintillans Forel" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="variety" species="pellitus" variety="scintillans">C. scintillans</taxonomicName>
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from Paraguay. Workers from western Paraguay are almost universally bicolored with a red head and mesosoma and dark legs and gaster. The major workers of these chaco forms have finer suberect hairs on the sides of the head, spaced more densely, than do workers of
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<taxonomicName id="D6A889C5F9FDB554B1DBFF49752BD107" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:134673" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus pellitus Mayr" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">C. pellitus</taxonomicName>
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, and the postero-lateral corners of the head of the largest workers are often more produced. Minor workers of the bicolored forms have smaller eyes than do minors of
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<taxonomicName id="B8F73D64B5E7EB59A985544843FA31C1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:134673" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus pellitus Mayr" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">C. pellitus</taxonomicName>
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, and they have a slightly lower, less convex propodeum. Both forms co-occur in eastern Paraguay near the Paraguay River with little evidence of interbreeding. Forel’s types of
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<taxonomicName id="FE0DC34DB1A57018712DBD1B73DA8C3C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239646" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus rosariensis" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="rosariensis">C. rosariensis</taxonomicName>
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fall within the range of variation of these bicolored chacoan forms, so I adopt here a two-species scheme with
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<taxonomicName id="36BAAEA1286279CC57C5DF0071E57C32" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239646" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus rosariensis" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="rosariensis">C. rosariensis</taxonomicName>
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representing the bulk of the chacoan collections and
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<taxonomicName id="A697E0C3DE0753978FE9770FDE093F63" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:134673" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus pellitus Mayr" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">C. pellitus</taxonomicName>
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representing most of the eastern collections.
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</paragraph>
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<paragraph id="A6FA83A4B6A4976B2A4E8CDA700F1AF3" pageNumber="46">
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The chacoan forms, while difficult to subdivide further on the basis of external worker morphology, show two distinct nest phenotypes in reproductively mature colonies. I have observed these nest forms within a few meters of each other. One form constructs large, conspicuous crater mounds while the other has cryptic nest entrances in the soil consisting of a single small hole with a slight scattering of excavated earth placed up to half a meter away, a pattern similar to the nests of
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<taxonomicName id="E7256CF23B5DFB1DD1FFC66A76AF1471" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:134673" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus pellitus Mayr" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">C. pellitus</taxonomicName>
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. This sympatric difference in nesting behavior suggests the existence of cryptic species.
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</paragraph>
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<paragraph id="62C3EB2975DEC1984353FEA979A02F77" pageNumber="46">
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F. Smith’s holotype of
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<taxonomicName id="27A43D8CCB4C2136B6AD7F28C7B3A0F6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26268" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus blandus (Smith)" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="blandus">C. blandus</taxonomicName>
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, a minor worker from
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<normalizedToken id="43BB3801BFC53038B7FE7E4BF9ECA927" originalValue="Pará">Para</normalizedToken>
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(BMNH, examined), is not a close match to any Paraguayan material. This ant has the smaller eyes and the lower propodeum more similar to
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<taxonomicName id="7463D3CDCB56286B240BC2DDB49F0519" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239646" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus rosariensis" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="rosariensis">C. rosariensis</taxonomicName>
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than to
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<taxonomicName id="6FECAE3BA98412EB447B8E052F9C28E9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:134673" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus pellitus Mayr" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">C. pellitus</taxonomicName>
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, but with denser pubescence on gastric tergites 3 and 4 (= abd. tergites 5 and 6) than in most specimens of
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<taxonomicName id="E262B61ECC4A2D0A43A02C9D4E4DB39A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239646" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus rosariensis" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="rosariensis">C. rosariensis</taxonomicName>
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. The
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<taxonomicName id="769F8941DBAB975A7110D0EF50E9CD99" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26268" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus blandus (Smith)" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="blandus">C. blandus</taxonomicName>
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type is also colored differently, with the posterior portion of the head quite dark, as in
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<taxonomicName id="84A442B36AB81FE10FF495D7679D62D8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:134673" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus pellitus Mayr" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">C. pellitus</taxonomicName>
|
||
, the mesosoma lighter, as in
|
||
<taxonomicName id="3E29BAFB4B27386DA82BCC4AB0C66996" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239646" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus rosariensis" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="rosariensis">C. rosariensis</taxonomicName>
|
||
, and the first gastric tergite concolorous with the mesosoma and infuscated posteriorly. However, I have not examined any material occurring between the type locality in
|
||
<normalizedToken id="72837B3128636A081322F43C6B25A6CD" originalValue="Pará">Para</normalizedToken>
|
||
and Paraguay, so I cannot dismiss the eventuality that the name
|
||
<taxonomicName id="42BCE3F4532CD698BD7701ED651AA13B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26268" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus blandus (Smith)" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="blandus">C. blandus</taxonomicName>
|
||
may apply one of these subtropical populations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |