298 lines
21 KiB
XML
298 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.70.54149" ID-GBIF-Dataset="7f3d4c9e-33e0-4ca0-8e78-11b78478538a" ID-PMC="PMC7406542" ID-Pensoft-Pub="1314-4049-70-89" ID-Pensoft-UUID="65FFE6B3A0F25CEF8D1B84BCEB9F6D24" ID-PubMed="32821216" ModsDocID="1314-4049-70-89" checkinTime="1596086616378" checkinUser="pensoft" docAuthor="Huang, Hong-Yan, Zhao, Jie, Zhang, Ping, Ge, Zai-Wei, Li, Xian & Tang, Li-Ping" docDate="2020" docId="2C3EA552C1E1514EB95C9AFC1A002716" docLanguage="en" docName="MycoKeys 70: 89-121" docOrigin="MycoKeys 70" docSource="http://dx.doi.org/10.3897/mycokeys.70.54149" docTitle="Clavariadelphus gansuensis J. Zhao & L. P. Tang 2020, sp. nov." docType="treatment" docVersion="4" id="65FFE6B3A0F25CEF8D1B84BCEB9F6D24" lastPageNumber="89" masterDocId="65FFE6B3A0F25CEF8D1B84BCEB9F6D24" masterDocTitle="The genus Clavariadelphus (Clavariadelphaceae, Gomphales) in China" masterLastPageNumber="121" masterPageNumber="89" pageNumber="89" updateTime="1668136509504" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Clavariadelphus (Clavariadelphaceae, Gomphales) in China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huang, Hong-Yan</mods:namePart>
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<mods:affiliation>School of Pharmaceutical Sciences and Yunnan Key Laboratory of Pharmacology for Natural Products, Kunming Medical University, Kunming, 650500, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhao, Jie</mods:namePart>
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<mods:affiliation>School of Pharmaceutical Sciences and Yunnan Key Laboratory of Pharmacology for Natural Products, Kunming Medical University, Kunming, 650500, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Ping</mods:namePart>
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<mods:affiliation>College of Life Science, Hunan Normal University, Changsha, 410081, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ge, Zai-Wei</mods:namePart>
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<mods:affiliation>Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, 650201, China</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-3184-4604</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Xian</mods:namePart>
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<mods:affiliation>School of Pharmaceutical Sciences and Yunnan Key Laboratory of Pharmacology for Natural Products, Kunming Medical University, Kunming, 650500, China</mods:affiliation>
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<mods:nameIdentifier type="email">xianlikm@163.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tang, Li-Ping</mods:namePart>
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<mods:affiliation>School of Pharmaceutical Sciences and Yunnan Key Laboratory of Pharmacology for Natural Products, Kunming Medical University, Kunming, 650500, China</mods:affiliation>
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<mods:nameIdentifier type="email">lipingtang11@qq.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>MycoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>70</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>89</mods:start>
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<mods:end>121</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.70.54149</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.70.54149</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-4049-70-89</mods:identifier>
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<mods:identifier type="Pensoft-UUID">65FFE6B3A0F25CEF8D1B84BCEB9F6D24</mods:identifier>
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<treatment ID-GBIF-Taxon="166017068" LSID="urn:lsid:plazi:treatment:2C3EA552C1E1514EB95C9AFC1A002716" httpUri="http://treatment.plazi.org/id/2C3EA552C1E1514EB95C9AFC1A002716" lastPageNumber="89" pageId="0" pageNumber="89">
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<subSubSection pageId="0" pageNumber="89" type="nomenclature">
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<paragraph pageId="0" pageNumber="89">
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4.
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<taxonomicName LSID="2C3EA552-C1E1-514E-B95C-9AFC1A002716" authority="J. Zhao & L. P. Tang" authorityName="J. Zhao & L. P. Tang" authorityYear="2020" class="Agaricomycetes" family="Clavariadelphaceae" genus="Clavariadelphus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Clavariadelphus gansuensis" order="Gomphales" pageId="0" pageNumber="89" phylum="Basidiomycota" rank="species" species="gansuensis" status="sp. nov.">Clavariadelphus gansuensis J. Zhao & L.P. Tang</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="89">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Clavariadelphus species in China. a C. alpinus (HKAS 57396, holotype) b, c C. amplus (HKAS 54876, holotype) d, e C. elongatus (d from HKAS 50742 e from HKAS 76589) f C. gansuensis (HKAS 76487, holotype) g C. himalayensis (HKAS 58811) h, i C. khinganensis (h from MHHNU 7789, holotype i from MHKMU H. Y. Huang 368) j C. sachalinensis (MHHNU 7816) k, l C. yunnanensis (k from HKAS 49398 l from HKAS 58789)." figureDoi="10.3897/mycokeys.70.54149.figure2" httpUri="https://binary.pensoft.net/fig/437013" pageId="0" pageNumber="89">Figs 2f</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Basidiospores of Clavariadelphus under light microscope. a C. alpinus (HKAS 57396, holotype) b C. amplus (HKAS 54876, holotype) c C. elongatus (HKAS 76589) d C. gansuensis (HKAS 76487, holotype) e C. himalayensis (HKAS 58811) f C. khinganensis (MHHNU 7789, holotype) g C. ligula (HKAS 35954) h C. sachalinensis (MHHNU 7816) i C. yunnanensis (HKAS 57659)." figureDoi="10.3897/mycokeys.70.54149.figure3" httpUri="https://binary.pensoft.net/fig/437014" pageId="0" pageNumber="89">, 3d</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Basidiospores of Clavariadelphus under SEM. a C. alpinus (HKAS 57396, holotype) b C. amplus (HKAS 54876, holotype) c C. elongatus (HKAS 76589) d C. gansuensis (HKAS 76487, holotype); e, f C. himalayensis (HKAS 58811) g C. khinganensis (MHHNU 7789, holotype) h C. yunnanensis (HKAS 57659)." figureDoi="10.3897/mycokeys.70.54149.figure4" httpUri="https://binary.pensoft.net/fig/437015" pageId="0" pageNumber="89">, 4d</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hyphae of basal mycelium from Clavariadelphus under SEM. a C. alpinus (HKAS 57396, holotype) b C. amplus (HKAS 54876, holotype) c C. elongatus (HKAS 76589) d C. gansuensis (HKAS 76487, holotype) e C. himalayensis (HKAS 58811) f, g C. sachalinensis (f from HKAS 33844; g from MHHNU 7816) h C. yunnanensis (HKAS 57659)." figureDoi="10.3897/mycokeys.70.54149.figure5" httpUri="https://binary.pensoft.net/fig/437016" pageId="0" pageNumber="89">, 5d</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Microscopic features of Clavariadelphus gansuensis (HKAS 76487, holotype). a Leptocystidia and immature basidia b Basidia." figureDoi="10.3897/mycokeys.70.54149.figure9" httpUri="https://binary.pensoft.net/fig/437020" pageId="0" pageNumber="89">, 9a, b</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="diagnosis">
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<paragraph pageId="0" pageNumber="89">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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This species is characterised by its orange, clavate basidiomes with slightly enlarged, truncate, sterile apex, broadly ellipsoid to ellipsoid basidiospores, hyphae of the basal mycelium with nipple-shaped protuberances and prism-like crystals and basidiomes that turn pink or light cherry-red in KOH. It differs from
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<taxonomicName lsidName="C. truncatus" pageId="0" pageNumber="89" rank="species" species="truncatus">
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<emphasis italics="true" pageId="0" pageNumber="89">C. truncatus</emphasis>
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</taxonomicName>
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by the
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<normalizedToken originalValue="latter’s">latter's</normalizedToken>
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robust, darker basidiomes with enlarged apices, and larger basidiospores.
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</paragraph>
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<caption doi="10.3897/mycokeys.70.54149.figure9" httpUri="https://binary.pensoft.net/fig/437020" pageId="0" pageNumber="89" start="Figure 9" startId="F9">
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<paragraph pageId="0" pageNumber="89">
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<emphasis bold="true" pageId="0" pageNumber="89">Figure 9.</emphasis>
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Microscopic features of
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<taxonomicName authorityName="J. Zhao & L. P. Tang" authorityYear="2020" class="Agaricomycetes" family="Clavariadelphaceae" genus="Clavariadelphus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Clavariadelphus gansuensis" order="Gomphales" pageId="0" pageNumber="89" phylum="Basidiomycota" rank="species" species="gansuensis">
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<emphasis italics="true" pageId="0" pageNumber="89">Clavariadelphus gansuensis</emphasis>
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</taxonomicName>
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(HKAS 76487, holotype).
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<emphasis bold="true" pageId="0" pageNumber="89">a</emphasis>
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<taxonomicName genus="Leptocystidia" lsidName="Leptocystidia" pageId="0" pageNumber="89" rank="genus">Leptocystidia</taxonomicName>
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and immature basidia
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<emphasis bold="true" pageId="0" pageNumber="89">b</emphasis>
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Basidia.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="etymology">
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<paragraph pageId="0" pageNumber="89">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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Latin "
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<taxonomicName genus="Leptocystidia" lsidName="gansuensis" pageId="0" pageNumber="89" rank="species" species="gansuensis">
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<emphasis italics="true" pageId="0" pageNumber="89">gansuensis</emphasis>
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</taxonomicName>
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" refers to the holotype location in Gansu Province.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="description">
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<paragraph pageId="0" pageNumber="89">Description.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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<taxonomicName genus="Basidiomes" lsidName="Basidiomes" pageId="0" pageNumber="89" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="89">Basidiomes</emphasis>
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</taxonomicName>
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up to 9 cm high, enlarged upwards to 1.6 cm diam., simple, clavate;
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<emphasis italics="true" pageId="0" pageNumber="89">hymenium</emphasis>
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longitudinally rugose, pruinose, light yellow to greyish-orange at maturity;
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<emphasis italics="true" pageId="0" pageNumber="89">apex</emphasis>
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initially obtuse or broadly rounded, flattening laterally, then truncate, slightly rugose, light orange or melon-orange (5A5-7) to orange (6A6-7) in age;
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<emphasis italics="true" pageId="0" pageNumber="89">base</emphasis>
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terete, smooth, pruinose, dirty white or pallid where covered, otherwise pruinose, pale orange or light orange (5A3-4);
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<emphasis italics="true" pageId="0" pageNumber="89">mycelial hyphae</emphasis>
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white;
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<emphasis italics="true" pageId="0" pageNumber="89">flesh</emphasis>
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initially solid, then soft and spongy upwards as the apex enlarges, white to pallid.
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<emphasis italics="true" pageId="0" pageNumber="89">Odour</emphasis>
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and
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<emphasis italics="true" pageId="0" pageNumber="89">taste</emphasis>
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not recorded.
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</paragraph>
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<paragraph pageId="0" pageNumber="89">
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<emphasis italics="true" pageId="0" pageNumber="89">Hymenium</emphasis>
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limited to the side of basidiomata, composed of basidia and leptocystidia; the apex of basidiomata composed of sterile elements 15-25
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<normalizedToken originalValue="×">x</normalizedToken>
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5-7
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, clavate, thin-walled, smooth, clamped.
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<emphasis italics="true" pageId="0" pageNumber="89">Basidia</emphasis>
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75-90
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<normalizedToken originalValue="×">x</normalizedToken>
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8-10
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, clavate, hyaline, thin-walled to thick-walled, 4-spored, sterigmata 7-10
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<normalizedToken originalValue="μm">μm</normalizedToken>
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in length.
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<emphasis italics="true" pageId="0" pageNumber="89">Basidiospores</emphasis>
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[20/1/1] 8.3-10.1 (-10.3)
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<normalizedToken originalValue="×">x</normalizedToken>
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5.3-6.3 (-6.4)
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<normalizedToken originalValue="μm">μm</normalizedToken>
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,
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<emphasis italics="true" pageId="0" pageNumber="89">Q</emphasis>
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= (1.34-) 1.47 -1.78 (-1.83),
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<emphasis italics="true" pageId="0" pageNumber="89">Q</emphasis>
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m = 1.60
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.09, ellipsoid to broadly ellipsoid, ovate or amygdaliform, with a small apiculus, inamyloid, thin-walled, hyaline in KOH.
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<taxonomicName genus="Leptocystidia" lsidName="Leptocystidia" pageId="0" pageNumber="89" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="89">Leptocystidia</emphasis>
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</taxonomicName>
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50-65
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<normalizedToken originalValue="×">x</normalizedToken>
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3-5
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, scattered amongst and scarcely projecting beyond the basidia, cylindrical to narrowly clavate, thin-walled, smooth, hyaline, non-pigmented, clamped, inflated apically at maturity, at times with apical or sub-apical branches.
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<emphasis italics="true" pageId="0" pageNumber="89">Mycelial hyphae</emphasis>
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2-3
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<normalizedToken originalValue="μm">μm</normalizedToken>
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diam., interwoven or aggregated into rhizomorphic strands, branched, clamped; the hyphal walls echinulate with light microscopy, covered with massive nipple-shaped protuberances, as well as encrusted with prism-like crystals up 5
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<normalizedToken originalValue="μm">μm</normalizedToken>
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long that are insoluble in KOH.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="chemical reactions">
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<paragraph pageId="0" pageNumber="89">Chemical reactions.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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(dried basidiomes): KOH = positive, pink, light coral or light cherry-red; FeCl3 = positive, green-yellow; NH4OH = positive, golden-rod or vivid yellow; phenol = positive, yellow; ethanol, FeSO4 and
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<normalizedToken originalValue="Melzer’s">Melzer's</normalizedToken>
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reagent = negative.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="distribution">
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<paragraph pageId="0" pageNumber="89">Known distribution and ecology.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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NW China, Gansu Province. Solitary on the ground in coniferous woods (
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<taxonomicName class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Abies" order="Pinales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="89">Abies</emphasis>
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</taxonomicName>
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spp.) or mixed with broad-leaved trees (
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<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Betula" order="Fagales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="89">Betula</emphasis>
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</taxonomicName>
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spp. and
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<taxonomicName family="Rosaceae" lsidName="" pageId="0" pageNumber="89" rank="family">Rosaceae</taxonomicName>
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) at elevations of approximately 3000 m.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="materials_examined">
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<paragraph pageId="0" pageNumber="89">Materials examined.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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China. Gansu Province: Lintan Prefecture, Yeliguan National Forest Park, coniferous woods (
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<taxonomicName class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Abies" order="Pinales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="89">Abies</emphasis>
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</taxonomicName>
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spp.) or mixed with
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<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Betula" order="Fagales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="89">Betula</emphasis>
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</taxonomicName>
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spp. and
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<taxonomicName family="Rosaceae" lsidName="" pageId="0" pageNumber="89" rank="family">Rosaceae</taxonomicName>
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plants, alt. 3000 m, 10 August 2012,
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<emphasis italics="true" pageId="0" pageNumber="89">X.T. Zhu 638</emphasis>
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(HKAS 76487,
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<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Holotype</emphasis>
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); Wudu Prefecture, September 1992,
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<emphasis italics="true" pageId="0" pageNumber="89">M.L. Tian M6465</emphasis>
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(HMAS 63052).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="comments">
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<paragraph pageId="0" pageNumber="89">Comments.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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<taxonomicName authorityName="J. Zhao & L. P. Tang" authorityYear="2020" class="Agaricomycetes" family="Clavariadelphaceae" genus="Clavariadelphus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Clavariadelphus gansuensis" order="Gomphales" pageId="0" pageNumber="89" phylum="Basidiomycota" rank="species" species="gansuensis">
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<emphasis italics="true" pageId="0" pageNumber="89">Clavariadelphus gansuensis</emphasis>
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</taxonomicName>
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, currently known only from NW China, is distinct by its slender, clavate, orange basidiomes with truncate apex, ellipsoid basidiospores (8.3-10.1
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<normalizedToken originalValue="×">x</normalizedToken>
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5.3-6.3
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<normalizedToken originalValue="μm">μm</normalizedToken>
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), pink staining reaction to KOH, hyphae of the basal mycelium with nipple-shaped protuberances and prism-like crystals and solitary growth habit in coniferous or mixed forests.
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</paragraph>
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<paragraph pageId="0" pageNumber="89">
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This species is most likely to be confused with several taxa, including
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<taxonomicName lsidName="C. amplus" pageId="0" pageNumber="89" rank="species" species="amplus">
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<emphasis italics="true" pageId="0" pageNumber="89">C. amplus</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="C. pallido-incarnatus" pageId="0" pageNumber="89" rank="species" species="pallido-incarnatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="89">C. pallido-incarnatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="C. pakistanicus" pageId="0" pageNumber="89" rank="species" species="pakistanicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="89">C. pakistanicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="C. truncatus" pageId="0" pageNumber="89" rank="species" species="truncatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="89">C. truncatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="C. unicolor" pageId="0" pageNumber="89" rank="species" species="unicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="89">C. unicolor</emphasis>
|
||
</taxonomicName>
|
||
. The comparison between
|
||
<taxonomicName lsidName="C. gansuensis" pageId="0" pageNumber="89" rank="species" species="gansuensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="89">C. gansuensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="C. amplus" pageId="0" pageNumber="89" rank="species" species="amplus">
|
||
<emphasis italics="true" pageId="0" pageNumber="89">C. amplus</emphasis>
|
||
</taxonomicName>
|
||
can be found in our treatment of
|
||
<taxonomicName lsidName="C. amplus" pageId="0" pageNumber="89" rank="species" species="amplus">
|
||
<emphasis italics="true" pageId="0" pageNumber="89">C. amplus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="89">
|
||
According to our phylogenetic analyses,
|
||
<taxonomicName lsidName="C. gansuensis" pageId="0" pageNumber="89" rank="species" species="gansuensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="89">C. gansuensis</emphasis>
|
||
</taxonomicName>
|
||
is allied with the sequence of "
|
||
<taxonomicName lsidName="C. truncatus" pageId="0" pageNumber="89" rank="species" species="truncatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="89">C. truncatus</emphasis>
|
||
</taxonomicName>
|
||
" from GenBank with strong support (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic tree of Clavariadelphus based on ITS sequence data. RAxML BP values (≥ 50 %) are shown above branches, Bayesian posterior probabilities (≥ 0.90) are shown above branches; new taxa are marked in red." figureDoi="10.3897/mycokeys.70.54149.figure1" httpUri="https://binary.pensoft.net/fig/437012" pageId="0" pageNumber="89">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |