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<document id="A1DBAEC10E565528CB5BE04EE5793FE4" ID-CLB-Dataset="298071" ID-DOI="10.5281/zenodo.10979877" ID-GBIF-Dataset="13070b4b-afe1-4eae-8c82-2afda9e0bfe5" ID-ISSN="1407-8953" ID-Zenodo-Dep="10979877" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1713275363296" checkinUser="felipe" docAuthor="Alekseev, Vitalii I." docDate="2018" docId="03F88A05FFA8C51DFDCBFC51FE57FE44" docLanguage="en" docName="ActaBiolUnivDaugavp.18.2.111-152.pdf" docOrigin="Acta Biologica Universitatis Daugavpiliensis 18 (2)" docStyle="DocumentStyle:DD7D35B4ACFEF3C20AEB77B8656EEADF.5:ActaBiolUnivDaugavp.2009-.journal_article.type1" docStyleId="DD7D35B4ACFEF3C20AEB77B8656EEADF" docStyleName="ActaBiolUnivDaugavp.2009-.journal_article.type1" docStyleVersion="5" docTitle="Onthophagus medius" docType="treatment" docVersion="5" lastPageNumber="135" masterDocId="FFC1F27DFFBFC505FFB4FFC2FFF4FF88" masterDocTitle="Scarabaeoidea (Insecta: Coleoptera) Of The Kaliningrad Region (Russia): The Commented Actual Checklist, Assessment Of Rarity And Notes To Regional Protection" masterLastPageNumber="152" masterPageNumber="111" pageNumber="134" updateTime="1717508418951" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="65FD7DA247674AC0B2EB38469D86C4C1">Scarabaeoidea (Insecta: Coleoptera) Of The Kaliningrad Region (Russia): The Commented Actual Checklist, Assessment Of Rarity And Notes To Regional Protection</mods:title>
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<mods:namePart id="FA68092A930A46020325F14727728AA4">Alekseev, Vitalii I.</mods:namePart>
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<mods:date id="2C9711C906CB0E0A94C18D7AB9BC0D14">2018</mods:date>
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<treatment id="03F88A05FFA8C51DFDCBFC51FE57FE44" ID-DOI="http://doi.org/10.5281/zenodo.11447556" ID-GBIF-Taxon="232131685" ID-Zenodo-Dep="11447556" LSID="urn:lsid:plazi:treatment:03F88A05FFA8C51DFDCBFC51FE57FE44" httpUri="http://treatment.plazi.org/id/03F88A05FFA8C51DFDCBFC51FE57FE44" lastPageId="24" lastPageNumber="135" pageId="23" pageNumber="134">
<subSubSection id="C34B6898FFA8C512FDCBFC51FC05FC23" box="[639,1009,915,939]" pageId="23" pageNumber="134" type="nomenclature">
<paragraph id="8BEE3B13FFA8C512FDCBFC51FC05FC23" blockId="23.[637,1150,915,1675]" box="[639,1009,915,939]" pageId="23" pageNumber="134">
(59).
<taxonomicName id="4C514090FFA8C512FD0CFC51FC05FC23" ID-CoL="49MP3" authority="(Kugelann, 1792)" baseAuthorityName="Kugelann" baseAuthorityYear="1792" box="[696,1009,915,939]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="medius">
<emphasis id="B925E701FFA8C512FD0CFC51FC05FC23" bold="true" box="[696,1009,915,939]" pageId="23" pageNumber="134">
<emphasis id="B925E701FFA8C512FD0CFC51FCDCFC23" bold="true" box="[696,808,915,939]" italics="true" pageId="23" pageNumber="134">O. medius</emphasis>
(Kugelann, 1792)
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C34B6898FFA8C512FDCBFC76FC02FC63" pageId="23" pageNumber="134" type="reference_group">
<paragraph id="8BEE3B13FFA8C512FDCBFC76FC02FC63" blockId="23.[637,1150,915,1675]" pageId="23" pageNumber="134">
<bibRefCitation id="EFC046E2FFA8C512FDCBFC76FD0FFC43" author="Lentz F. L." box="[639,763,947,971]" pageId="23" pageNumber="134" refId="ref26437" refString="Lentz F. L. 1853. The beetles of Preussen. Written for the collecting young people. Grafe und Unzer, Konigsberg (in German)." type="book" year="1853">Lentz 1853</bibRefCitation>
;
<bibRefCitation id="EFC046E2FFA8C512FCBCFC76FC76FC43" author="Lentz F. L." box="[776,898,947,971]" pageId="23" pageNumber="134" refId="ref26466" refString="Lentz F. L. 1879. The catalogue of the Preussishe beetles. Konigsberg (in German)." type="book" year="1879">Lentz 1879</bibRefCitation>
(
<taxonomicName id="4C514090FFA8C512FC27FC76FC25FC43" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[915,977,948,971]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="vacca">
<emphasis id="B925E701FFA8C512FC27FC76FC25FC43" box="[915,977,948,971]" italics="true" pageId="23" pageNumber="134">vacca</emphasis>
</taxonomicName>
L.
<taxonomicName id="4C514090FFA8C512FC4DFC71FB84FC43" authority="Pz." authorityName="Pz." box="[1017,1136,947,971]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="medius">
<emphasis id="B925E701FFA8C512FC4DFC71FBB1FC43" box="[1017,1093,947,971]" italics="true" pageId="23" pageNumber="134">medius</emphasis>
Pz.
</taxonomicName>
); Bercio &amp; Folwaczny 1979 (
<taxonomicName id="4C514090FFA8C512FC1CFC16FC12FC63" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[936,998,980,1003]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="vacca">
<emphasis id="B925E701FFA8C512FC1CFC16FC12FC63" box="[936,998,980,1003]" italics="true" pageId="23" pageNumber="134">vacca</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B6898FFA8C512FDCBFC31FBF2FBC3" pageId="23" pageNumber="134" type="distribution">
<paragraph id="8BEE3B13FFA8C512FDCBFC31FBF2FBC3" blockId="23.[637,1150,915,1675]" pageId="23" pageNumber="134">
<emphasis id="B925E701FFA8C512FDCBFC31FC8EFB83" bold="true" box="[639,890,1011,1035]" pageId="23" pageNumber="134">
<collectingRegion id="4995F5F1FFA8C512FDCBFC31FC8EFB83" box="[639,890,1011,1035]" country="Russia" name="Kaliningrad" pageId="23" pageNumber="134">Kaliningrad Region</collectingRegion>
</emphasis>
: Gur. (Königsberg, Metgethen, Spittelhof, Wilkie), Nest. (Rominten), Zel. (Cranz, Rauschen).
</paragraph>
</subSubSection>
<subSubSection id="C34B6898FFA8C51DFDCBFB91FE57FE44" lastPageId="24" lastPageNumber="135" pageId="23" pageNumber="134" type="discussion">
<paragraph id="8BEE3B13FFA8C51DFDCBFB91FE22FF04" blockId="23.[637,1150,915,1675]" lastBlockId="24.[136,470,116,140]" lastPageId="24" lastPageNumber="135" pageId="23" pageNumber="134">
<emphasis id="B925E701FFA8C512FDCBFB91FCF7FBE3" bold="true" box="[639,771,1107,1131]" pageId="23" pageNumber="134">Comments:</emphasis>
The species “
<taxonomicName id="4C514090FFA8C512FC10FB96FBF2FBE3" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[932,1030,1108,1131]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="vacca">
<emphasis id="B925E701FFA8C512FC10FB96FBF2FBE3" box="[932,1030,1108,1131]" italics="true" pageId="23" pageNumber="134">O. vacca</emphasis>
</taxonomicName>
L.
<taxonomicName id="4C514090FFA8C512FB85FB91FD55FB03" authority="Pz." authorityName="Pz." class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="medius">
<emphasis id="B925E701FFA8C512FB85FB91FB89FBE3" box="[1073,1149,1107,1131]" italics="true" pageId="23" pageNumber="134">medius</emphasis>
Pz.
</taxonomicName>
” or “
<taxonomicName id="4C514090FFA8C512FD5CFBB6FCB9FB03" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[744,845,1140,1163]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="vacca">
<emphasis id="B925E701FFA8C512FD5CFBB6FCB9FB03" box="[744,845,1140,1163]" italics="true" pageId="23" pageNumber="134">O. vacca</emphasis>
</taxonomicName>
L.” is reported from the Region according to
<bibRefCitation id="EFC046E2FFA8C512FCDFFB56FC0EFB23" author="Lentz F. L." box="[875,1018,1171,1195]" pageId="23" pageNumber="134" refId="ref26466" refString="Lentz F. L. 1879. The catalogue of the Preussishe beetles. Konigsberg (in German)." type="book" year="1879">Lentz (1879)</bibRefCitation>
and Bercio &amp; Folwaczny (1979). In the recent research of
<bibRefCitation id="EFC046E2FFA8C512FD2BFB16FC72FB63" author="Rossner E. &amp; Schonfeld J. &amp; Ahrens D." box="[671,902,1235,1259]" pageId="23" pageNumber="134" pagination="1 - 28" refId="ref27546" refString="Rossner E., Schonfeld J., Ahrens D. 2010. Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) - a good Western Palaearctic species in the Onthophagus vacca complex (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini). Zootaxa, 2629: 1 - 28." type="journal article" year="2010">Rössner et al. (2010)</bibRefCitation>
the former species
<taxonomicName id="4C514090FFA8C512FBD7FB16FD49FA83" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="vacca">
<emphasis id="B925E701FFA8C512FBD7FB16FD49FA83" italics="true" pageId="23" pageNumber="134">O. vacca</emphasis>
</taxonomicName>
(L.) is considered to consist of two sibling species:
<taxonomicName id="4C514090FFA8C512FD69FAD6FBE2FAA3" authority="(Kugelann, 1792)" baseAuthorityName="Kugelann" baseAuthorityYear="1792" box="[733,1046,1299,1323]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="medius">
<emphasis id="B925E701FFA8C512FD69FAD6FCB9FAA3" box="[733,845,1299,1323]" italics="true" pageId="23" pageNumber="134">O. medius</emphasis>
(Kugelann, 1792)
</taxonomicName>
occuring further north and reaching at least 54° of northern latitude and
<taxonomicName id="4C514090FFA8C512FCEAFA96FB8DFAE3" authority="(Linnaeus, 1767)" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[862,1145,1363,1387]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="vacca">
<emphasis id="B925E701FFA8C512FCEAFA96FC4FFAE3" box="[862,955,1364,1387]" italics="true" pageId="23" pageNumber="134">O. vacca</emphasis>
(Linnaeus, 1767)
</taxonomicName>
, seemingly to be absent north of 50° latitude. The locality of the
<taxonomicName id="4C514090FFA8C512FCF9FA56FC4EFA23" baseAuthorityName="Kugelann" baseAuthorityYear="1792" box="[845,954,1427,1451]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="medius">
<emphasis id="B925E701FFA8C512FCF9FA56FC4EFA23" box="[845,954,1427,1451]" italics="true" pageId="23" pageNumber="134">O. medius</emphasis>
</taxonomicName>
<typeStatus id="54EA85B1FFA8C512FC74FA56FBE0FA24" box="[960,1044,1428,1452]" pageId="23" pageNumber="134" type="neotype">neotype</typeStatus>
is “Lyck, Ostpreussen” [now Ełk, Warmian-Masurian region, northeastern
<collectingCountry id="F3467B83FFA8C512FCC5FA11FC35FA63" box="[881,961,1491,1515]" name="Poland" pageId="23" pageNumber="134">Poland</collectingCountry>
]. In accordance with mentioned above, in the Kaliningrad Region should be expected only one species –
<taxonomicName id="4C514090FFA8C512FD22F9F6FCADF9C3" authority="(Kug.)" baseAuthorityName="Kugelann" baseAuthorityYear="1792" box="[662,857,1587,1611]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="medius">
<emphasis id="B925E701FFA8C512FD22F9F6FCF2F9C3" box="[662,774,1587,1611]" italics="true" pageId="23" pageNumber="134">O. medius</emphasis>
(Kug.)
</taxonomicName>
. Therefore, the species
<taxonomicName id="4C514090FFA8C512FBD7F9F6FD49F9E3" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="134" phylum="Arthropoda" rank="species" species="vacca">
<emphasis id="B925E701FFA8C512FBD7F9F6FD49F9E3" italics="true" pageId="23" pageNumber="134">O. vacca</emphasis>
</taxonomicName>
(L.) has been removed from the species list and the old data concerning its distribution was used for
<taxonomicName id="4C514090FFA7C51DFEA2FFB6FE25FF04" authority="(Kug.)" baseAuthorityName="Kugelann" baseAuthorityYear="1792" box="[278,465,116,140]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="135" phylum="Arthropoda" rank="species" species="medius">
<emphasis id="B925E701FFA7C51DFEA2FFB6FE76FF04" box="[278,386,116,140]" italics="true" pageId="24" pageNumber="135">O. medius</emphasis>
(Kug.)
</taxonomicName>
.
</paragraph>
<paragraph id="8BEE3B13FFA7C51DFF3CFF76FE57FE44" blockId="24.[134,647,180,460]" pageId="24" pageNumber="135">
Formerly widespread in the Region species [“often” (
<bibRefCitation id="EFC046E2FFA7C51DFF58FF16FE93FF63" author="Lentz F. L." box="[236,359,211,236]" pageId="24" pageNumber="135" refId="ref26437" refString="Lentz F. L. 1853. The beetles of Preussen. Written for the collecting young people. Grafe und Unzer, Konigsberg (in German)." type="book" year="1853">Lentz 1853</bibRefCitation>
) or “everywhere common” (
<bibRefCitation id="EFC046E2FFA7C51DFF3AFF36FEECFE84" author="Lentz F. L." box="[142,280,244,268]" pageId="24" pageNumber="135" refId="ref26466" refString="Lentz F. L. 1879. The catalogue of the Preussishe beetles. Konigsberg (in German)." type="book" year="1879">Lentz 1879</bibRefCitation>
)]. The speceis
<taxonomicName id="4C514090FFA7C51DFE67FF36FDEAFE83" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[467,542,244,268]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="135" phylum="Arthropoda" rank="species" species="vacca">
“
<emphasis id="B925E701FFA7C51DFE54FF36FDEAFE83" box="[480,542,244,267]" italics="true" pageId="24" pageNumber="135">vacca</emphasis>
</taxonomicName>
(L.)” is reported in
<collectingCountry id="F3467B83FFA7C51DFF4BFED6FEA4FEA4" box="[255,336,276,300]" name="Belarus" pageId="24" pageNumber="135">Belarus</collectingCountry>
(Alexandovitch &amp; Pisanenko 1991),
<collectingCountry id="F3467B83FFA7C51DFF66FEF6FECCFEC4" box="[210,312,308,332]" name="Lithuania" pageId="24" pageNumber="135">Lithuania</collectingCountry>
(
<bibRefCitation id="EFC046E2FFA7C51DFEF2FEF6FDE3FEC4" author="Tamutis V. &amp; Tamute B. &amp; Ferenca R." box="[326,535,308,332]" pageId="24" pageNumber="135" pagination="1 - 494" refId="ref27786" refString="Tamutis V., Tamute B., Ferenca R. 2011. A catalogue of Lithuanian beetles (Insecta: Coleoptera). ZooKeys, 121: 1 - 494." type="journal article" year="2011">Tamutis et al. 2011</bibRefCitation>
),
<collectingCountry id="F3467B83FFA7C51DFD9DFEF6FD74FEC4" box="[553,640,308,332]" name="Sweden" pageId="24" pageNumber="135">Sweden</collectingCountry>
,
<collectingCountry id="F3467B83FFA7C51DFF3CFE96FF1FFEE4" box="[136,235,340,364]" name="Denmark" pageId="24" pageNumber="135">Denmark</collectingCountry>
(
<bibRefCitation id="EFC046E2FFA7C51DFEB6FE96FE30FEE4" author="Silfverberg H." box="[258,452,340,364]" pageId="24" pageNumber="135" refId="ref27598" refString="Silfverberg H. 2010. Enumeratio renovata Coleopterorum Fennoscandiae, Daniae et" type="book" year="2010">Silfverberg 2010</bibRefCitation>
),
<collectingCountry id="F3467B83FFA7C51DFE54FE96FDD0FEE4" box="[480,548,340,364]" name="Latvia" pageId="24" pageNumber="135">Latvia</collectingCountry>
(
<bibRefCitation id="EFC046E2FFA7C51DFD8FFE96FF35FE04" author="Telnov D." pageId="24" pageNumber="135" pagination="1" refId="ref27818" refString="Telnov D. 2004. Checklist of Latvian beetles (Insecta: Coleoptera). In: Compendium of Latvian Coleoptera, edited by D. Telnov, 2 nd ed., vol. 1. Riga: Pertovskis." type="book chapter" year="2004">Telnov 2004</bibRefCitation>
), the Moscow Region (
<bibRefCitation id="EFC046E2FFA7C51DFE02FEB6FDA6FE04" author="Nikitsky N. B." box="[438,594,372,396]" pageId="24" pageNumber="135" refId="ref26838" refString="Nikitsky N. B. 2009. New and interesting finds of xylophilous and several other coleopteran insects (Coleoptera) in the Moscow Region and in Moscow. Bulleten Moskovskogo Obschestva Ispytatelei Prirody, Otdel" type="book" year="2009">Nikitsky 2009</bibRefCitation>
), but it is very possible, part of these reports really concern
<taxonomicName id="4C514090FFA7C51DFF57FE76FE6BFE44" authority="(Kug.)" baseAuthorityName="Kugelann" baseAuthorityYear="1792" box="[227,415,436,460]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="135" phylum="Arthropoda" rank="species" species="medius">
<emphasis id="B925E701FFA7C51DFF57FE76FEA4FE44" box="[227,336,436,460]" italics="true" pageId="24" pageNumber="135">O. medius</emphasis>
(Kug.)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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