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<document id="1A7AA3BD6D00D8751AD15D5A98785A1F" ID-DOI="10.5281/zenodo.5178595" ID-ISSN="1942-1354" ID-Zenodo-Dep="5178595" ID-ZooBank="FAE960F6-2EE4-4D9B-95C7-395EC8E99790" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628314504382" checkinUser="felipe" docAuthor="Shelley, Rowland M. &amp; Richart, Casey H." docDate="2014" docId="03E0B33F496E1876FF12F977FA5AFD11" docLanguage="en" docName="InsectaMundi.0340.1-19.pdf" docOrigin="Insecta Mundi 2014 (340)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Diactis hedini Shelley &amp; Richart 2014, new species" docType="treatment" docVersion="3" lastPageNumber="7" masterDocId="FFD9CB474968187EFFD2FFBCFFFBFF88" masterDocTitle="Tynommatidae, n. stat., a family of western North American millipeds: Hypotheses on origins and affinities; tribal elevations; rediagnoses of Diactis Loomis, 1937, and Florea and Caliactis, both Shelley, 1996; and description of D. hedini, n. sp. (Callipodida: Schizopetalidea)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="5" updateTime="1699042317474" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="202501161D603FE01856ECFD99EDFAC6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F124BE32FBF77F53CB412201BF2ADCFD">
<mods:title id="F7A113907313D1965C2EA6416D42038F">Tynommatidae, n. stat., a family of western North American millipeds: Hypotheses on origins and affinities; tribal elevations; rediagnoses of Diactis Loomis, 1937, and Florea and Caliactis, both Shelley, 1996; and description of D. hedini, n. sp. (Callipodida: Schizopetalidea)</mods:title>
</mods:titleInfo>
<mods:name id="FBAD325C0CA999F7B20BA829207CCCF4" type="personal">
<mods:role id="827DE3A4CF96FD228A86AA8A8B11606B">
<mods:roleTerm id="33C0FE2800515F5D312933CDD1C51B78">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="430FD8FBDB410D2F69D76508842143F3">Shelley, Rowland M.</mods:namePart>
</mods:name>
<mods:name id="D2CFA785C7F723D64E906B7C2B3DD71C" type="personal">
<mods:role id="E504F99F55F5670B33A8B16FEF572A6F">
<mods:roleTerm id="B5E370CE0CDA11F820D8AD78E1B93717">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4B2E19DBBB08D95F10D431ABED9DD547">Richart, Casey H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="9AA1C05777234557F45526395007E0D8">text</mods:typeOfResource>
<mods:relatedItem id="E901FFBAFA04412368B9AD73BBE43CF2" type="host">
<mods:titleInfo id="EA88CC284FF13FFC2DA8C94DFCCCF09A">
<mods:title id="BCCBAB2D21409DDEA867BCBC0035226E">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="095ADACEFBB8F3A92EC18DA3AA088DF7">
<mods:date id="7F9C33DA1747EE2766B83E18BED62938">2014</mods:date>
<mods:detail id="5BC4AA5C09F4D7D5C3578E3B7B211589" type="pubDate">
<mods:number id="DFDDED84C0146D670A6759F32481A2CA">2014-01-31</mods:number>
</mods:detail>
<mods:detail id="7B31CE86D72EC578D9D5F034FE872087" type="volume">
<mods:number id="DBA8C4212B210BE1571213B792B7749C">2014</mods:number>
</mods:detail>
<mods:detail id="4BD516ECFE5A91928DB7AF4127F0F368" type="issue">
<mods:number id="8FC8008EA7513A038633A9D49B20C1AA">340</mods:number>
</mods:detail>
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<mods:start id="1070E6E9D9A45FF2D10E45A9D6FE407D">1</mods:start>
<mods:end id="A326E4FF79CEE3FEFD51C3C74F434BE4">19</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="20ABDF9133C99A629095BC54C7BB66BB">journal article</mods:classification>
<mods:identifier id="715C53D012D31217C70C5F26E6F3ACAA" type="DOI">10.5281/zenodo.5178595</mods:identifier>
<mods:identifier id="DD7B1BC8677795AB583761642D84D336" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="AB5D4EA46D47EAA96918590830AE4A0A" type="Zenodo-Dep">5178595</mods:identifier>
<mods:identifier id="B1ABE6F5B3A9A029B0B02C7380FD9F81" type="ZooBank">FAE960F6-2EE4-4D9B-95C7-395EC8E99790</mods:identifier>
</mods:mods>
<treatment id="03E0B33F496E1876FF12F977FA5AFD11" LSID="urn:lsid:plazi:treatment:03E0B33F496E1876FF12F977FA5AFD11" httpUri="http://treatment.plazi.org/id/03E0B33F496E1876FF12F977FA5AFD11" lastPageId="8" lastPageNumber="7" pageId="6" pageNumber="5">
<subSubSection id="C35351A2496E1878FF12F977FEABF880" pageId="6" pageNumber="5" type="nomenclature">
<paragraph id="8BF60229496E1878FF12F977FEABF880" blockId="6.[192,755,1665,1800]" pageId="6" pageNumber="5">
<emphasis id="B93DDE3B496E1878FF12F977FDCFF96B" bold="true" box="[192,564,1739,1763]" pageId="6" pageNumber="5">
<taxonomicName id="4C4979AA496E1878FF12F977FE79F96B" authority="Shelley &amp; Richart, 2014" authorityName="Shelley &amp; Richart" authorityYear="2014" box="[192,386,1739,1763]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="hedini" status="sp. nov.">
<emphasis id="B93DDE3B496E1878FF12F977FE79F96B" bold="true" box="[192,386,1739,1763]" italics="true" pageId="6" pageNumber="5">Diactis hedini</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20E6340496E1878FE40F977FDCFF96B" box="[402,564,1739,1763]" pageId="6" pageNumber="5" rank="species">new species</taxonomicNameLabel>
</emphasis>
<figureCitation id="13721EAC496E1878FF12F94CFEABF880" box="[192,336,1776,1800]" captionStart="Figures 1-6" captionStartId="7.[192,288,1270,1292]" captionTargetBox="[201,1432,206,1250]" captionTargetId="figure-174@7.[192,1440,192,1251]" captionTargetPageId="7" captionText="Figures 1-6. Diactis spp. 1) D. hedini holotype, right gonopod, caudal view. 2) D. hedini holotype, right gonopod, telopodite of the same, medial view. 3) D. hedini holotype, right gonopod, the same, lateral view. 4) D. hedini holotype, right gonopod, distal extremity of the same, anterior view. 5) Telopodite of right gonopod of D. frondifera male from Los Angeles Co., California, lateral view. 6) Telopodite of right gonopod of D. triangula male from San Bernardino Co., California, anterior view. A, process ‘A’; B, process ‘B’; pfp, prefemoral process; s, solenomere; sb, solenomere branch; tt, tibiotarsus." figureDoi="http://doi.org/10.5281/zenodo.5178597" httpUri="https://zenodo.org/record/5178597/files/figure.png" pageId="6" pageNumber="5">Figures 1–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35351A2496E1879FF12F889FAC7FA45" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5" type="materials_examined">
<paragraph id="8BF60229496E1878FF12F889FA5BF8E5" blockId="6.[192,1441,1845,1901]" pageId="6" pageNumber="5">
<emphasis id="B93DDE3B496E1878FF12F889FE60F8C5" bold="true" box="[192,411,1845,1869]" pageId="6" pageNumber="5">Type specimens</emphasis>
. M
<typeStatus id="54F2BC8B496E1878FE1AF889FDCAF8C5" box="[456,561,1845,1869]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
and F
<typeStatus id="54F2BC8B496E1878FD51F889FD0AF8C5" box="[643,753,1845,1869]" pageId="6" pageNumber="5" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED589AEC496E1878FD2DF889FCAFF8C5" box="[767,852,1845,1869]" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="6" pageNumber="5" type="Museum">NCSM</collectionCode>
) collected by M. Hedin,
<date id="FFF724E9496E1878FBA4F889FAA2F8C5" box="[1142,1369,1845,1869]" pageId="6" pageNumber="5" value="2005-12-17">
17 December
<quantity id="4CB1AFCC496E1878FAC9F889FAA2F8C5" box="[1307,1369,1845,1869]" metricMagnitude="6" metricUnit="m" metricValue="2.0054" pageId="6" pageNumber="5" unit="km" value="2005.4">2005</quantity>
</date>
, 4 km (2.5 mi) N of Interstate Highway 8 along SR 79, 1.9 km (1.2 mi) NE Descanso Jct., NW side of Guatay
</paragraph>
<caption id="DF3652A1496F1879FF12FB4AFDC5FA14" ID-DOI="http://doi.org/10.5281/zenodo.5178597" ID-Zenodo-Dep="5178597" httpUri="https://zenodo.org/record/5178597/files/figure.png" pageId="7" pageNumber="6" startId="7.[192,288,1270,1292]" targetBox="[201,1432,206,1250]" targetPageId="7">
<paragraph id="8BF60229496F1879FF12FB4AFDC5FA14" blockId="7.[192,1441,1270,1436]" pageId="7" pageNumber="6">
<emphasis id="B93DDE3B496F1879FF12FB4AFEAAFA83" bold="true" box="[192,337,1270,1292]" pageId="7" pageNumber="6">Figures 1-6.</emphasis>
<taxonomicName id="4C4979AA496F1879FE85FB4BFE16FA84" box="[343,493,1270,1292]" class="Diplopoda" family="Tynommatidae" genus="Diactis" kingdom="Animalia" order="Callipodida" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B93DDE3B496F1879FE85FB4BFE5EFA84" box="[343,421,1271,1292]" italics="true" pageId="7" pageNumber="6">Diactis</emphasis>
spp.
<emphasis id="B93DDE3B496F1879FE0DFB4AFE16FA84" bold="true" box="[479,493,1270,1292]" pageId="7" pageNumber="6">1</emphasis>
</taxonomicName>
<emphasis id="B93DDE3B496F1879FE3FFB4AFE0DFA84" bold="true" box="[493,502,1270,1292]" pageId="7" pageNumber="6">)</emphasis>
<taxonomicName id="4C4979AA496F1879FE2FFB4BFD9FFA83" authorityName="Shelley &amp; Richart" authorityYear="2014" box="[509,612,1270,1292]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="hedini">
<emphasis id="B93DDE3B496F1879FE2FFB4BFD9FFA83" box="[509,612,1270,1292]" italics="true" pageId="7" pageNumber="6">D. hedini</emphasis>
</taxonomicName>
holotype, right gonopod, caudal view.
<emphasis id="B93DDE3B496F1879FBD9FB4AFBD9FA84" bold="true" box="[1035,1058,1270,1292]" pageId="7" pageNumber="6">2)</emphasis>
<taxonomicName id="4C4979AA496F1879FBFBFB4BFB74FA83" authorityName="Shelley &amp; Richart" authorityYear="2014" box="[1065,1167,1270,1292]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="hedini">
<emphasis id="B93DDE3B496F1879FBFBFB4BFB74FA83" box="[1065,1167,1270,1292]" italics="true" pageId="7" pageNumber="6">D. hedini</emphasis>
</taxonomicName>
holotype, right gonopod, telopodite of the same, medial view.
<emphasis id="B93DDE3B496F1879FDB3FAAFFD83FAA1" bold="true" box="[609,632,1299,1321]" pageId="7" pageNumber="6">3)</emphasis>
<taxonomicName id="4C4979AA496F1879FD53FAA8FD11FAA0" authorityName="Shelley &amp; Richart" authorityYear="2014" box="[641,746,1299,1321]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="hedini">
<emphasis id="B93DDE3B496F1879FD53FAA8FD11FAA0" box="[641,746,1299,1321]" italics="true" pageId="7" pageNumber="6">D. hedini</emphasis>
</taxonomicName>
holotype, right gonopod, the same, lateral view.
<emphasis id="B93DDE3B496F1879FAC5FAAFFAD5FAA1" bold="true" box="[1303,1326,1299,1321]" pageId="7" pageNumber="6">4)</emphasis>
<taxonomicName id="4C4979AA496F1879FAE5FAA8FA5BFAA0" authorityName="Shelley &amp; Richart" authorityYear="2014" box="[1335,1440,1299,1321]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="hedini">
<emphasis id="B93DDE3B496F1879FAE5FAA8FA5BFAA0" box="[1335,1440,1299,1321]" italics="true" pageId="7" pageNumber="6">D. hedini</emphasis>
</taxonomicName>
holotype, right gonopod, distal extremity of the same, anterior view.
<emphasis id="B93DDE3B496F1879FC7EFA8CFC39FACE" bold="true" box="[940,962,1328,1350]" pageId="7" pageNumber="6">5)</emphasis>
Telopodite of right gonopod of
<taxonomicName id="4C4979AA496F1879FAC0FA8DFA5BFACD" authorityName="Loomis" authorityYear="1937" box="[1298,1440,1328,1350]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="frondifera">
<emphasis id="B93DDE3B496F1879FAC0FA8DFA5BFACD" box="[1298,1440,1328,1350]" italics="true" pageId="7" pageNumber="6">D. frondifera</emphasis>
</taxonomicName>
male from Los Angeles Co., California, lateral view.
<emphasis id="B93DDE3B496F1879FCD0FAF1FCE2FAEB" bold="true" box="[770,793,1357,1379]" pageId="7" pageNumber="6">6)</emphasis>
Telopodite of right gonopod of
<taxonomicName id="4C4979AA496F1879FBBCFAF2FB0CFAEA" authorityName="Loomis" authorityYear="1937" box="[1134,1271,1357,1379]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="triangula">
<emphasis id="B93DDE3B496F1879FBBCFAF2FB0CFAEA" box="[1134,1271,1357,1379]" italics="true" pageId="7" pageNumber="6">D. triangula</emphasis>
</taxonomicName>
male from San Bernardino Co., California, anterior view. A, process ‘A’; B, process ‘B’; pfp, prefemoral process; s, solenomere; sb, solenomere branch; tt, tibiotarsus.
</paragraph>
</caption>
<paragraph id="8BF60229496F1879FF12FA09FAC7FA45" blockId="7.[192,1340,1461,1485]" box="[192,1340,1461,1485]" pageId="7" pageNumber="6">
Mountain (
<geoCoordinate id="EE7D64EE496F1879FE9BFA09FE2EFA45" box="[329,469,1461,1485]" degrees="32" direction="north" minutes="51" orientation="latitude" pageId="7" pageNumber="6" precision="15" seconds="16" value="32.854443">32°51’16”N</geoCoordinate>
, 116°35’68”W), elevation
<quantity id="4CB1AFCC496F1879FCDFFA09FC8BFA45" box="[781,880,1461,1485]" metricMagnitude="0" metricUnit="m" metricValue="1.08" pageId="7" pageNumber="6" unit="m" value="1.08">1,080 m</quantity>
(
<quantity id="4CB1AFCC496F1879FCADFA09FC1DFA45" box="[895,998,1461,1485]" metricMagnitude="0" metricUnit="m" metricValue="1.09728" pageId="7" pageNumber="6" unit="ft" value="3.6">3,600 ft.</quantity>
), San Diego Co.,
<collectingRegion id="498DCCCB496F1879FB6AFA09FAC3FA45" box="[1208,1336,1461,1485]" country="United States of America" name="California" pageId="7" pageNumber="6">California</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C35351A2496F1879FF12FA49FB99F905" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="8BF60229496F1879FF12FA49FB99F905" blockId="7.[192,1440,1525,1677]" pageId="7" pageNumber="6">
<emphasis id="B93DDE3B496F1879FF12FA49FEB3F985" bold="true" box="[192,328,1525,1549]" pageId="7" pageNumber="6">Diagnosis</emphasis>
50 rings including epiproct. Gonosternum with dactyliform, medial projection. Telopodal stem of subequal width throughout most of length, tapering apically to acuminate tip; prefemoral process short and broad, directed distolaterad. Distal processes arising subapically; tibiotarsus a broad continuation of telopodal stem curving opposite to solenomere branch; latter bending caudolaterad, process ‘B’ absent, apparently represented by prefemoral process (
<figureCitation id="13721EAC496F1879FC3CF9C9FBAFF905" box="[1006,1108,1653,1677]" captionStart="Figures 1-6" captionStartId="7.[192,288,1270,1292]" captionTargetBox="[201,1432,206,1250]" captionTargetId="figure-174@7.[192,1440,192,1251]" captionTargetPageId="7" captionText="Figures 1-6. Diactis spp. 1) D. hedini holotype, right gonopod, caudal view. 2) D. hedini holotype, right gonopod, telopodite of the same, medial view. 3) D. hedini holotype, right gonopod, the same, lateral view. 4) D. hedini holotype, right gonopod, distal extremity of the same, anterior view. 5) Telopodite of right gonopod of D. frondifera male from Los Angeles Co., California, lateral view. 6) Telopodite of right gonopod of D. triangula male from San Bernardino Co., California, anterior view. A, process ‘A’; B, process ‘B’; pfp, prefemoral process; s, solenomere; sb, solenomere branch; tt, tibiotarsus." figureDoi="http://doi.org/10.5281/zenodo.5178597" httpUri="https://zenodo.org/record/5178597/files/figure.png" pageId="7" pageNumber="6">Fig. 1–4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35351A2496F1879FF12F909FD3EF8E5" pageId="7" pageNumber="6" type="biology_ecology">
<paragraph id="8BF60229496F1879FF12F909FD3EF8E5" blockId="7.[192,1441,1717,1901]" pageId="7" pageNumber="6">
<emphasis id="B93DDE3B496F1879FF12F909FED5F945" bold="true" box="[192,302,1717,1741]" pageId="7" pageNumber="6">Ecology</emphasis>
. The
<typeStatus id="54F2BC8B496F1879FEA4F90AFE43F946" box="[374,440,1718,1742]" pageId="7" pageNumber="6">types</typeStatus>
were retrieved from a berlesate of Coast Live Oak (
<taxonomicName id="4C4979AA496F1879FB92F909FAABF945" authority="Nee" authorityName="Nee" box="[1088,1360,1717,1741]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="6" phylum="Tracheophyta" rank="species" species="agrifolia">
<emphasis id="B93DDE3B496F1879FB92F909FAECF945" box="[1088,1303,1717,1741]" italics="true" pageId="7" pageNumber="6">Quercus agrifolia</emphasis>
Nee
</taxonomicName>
) litter adjacent to Samagatuma Creek in a Western Sycamore (
<taxonomicName id="4C4979AA496F1879FC55F969FB46F965" authority="Nutt." authorityName="Nutt." box="[903,1213,1749,1773]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="7" pageNumber="6" phylum="Tracheophyta" rank="species" species="racemosa">
<emphasis id="B93DDE3B496F1879FC55F969FB89F965" box="[903,1138,1749,1773]" italics="true" pageId="7" pageNumber="6">Platanus racemosa</emphasis>
Nutt.
</taxonomicName>
) riparian corridor forest that runs through coastal sage and chaparral habitat. This is the southernmost
<collectingCountry id="F35E42B9496F1879FB3CF949FAEEF885" box="[1262,1301,1781,1805]" name="United States of America" pageId="7" pageNumber="6">US</collectingCountry>
locality for the salamander,
<taxonomicName id="4C4979AA496F1879FE42F8AAFC50F8A5" authority="Dunn, 1929" authorityName="Dunn" authorityYear="1929" box="[400,939,1813,1837]" class="Amphibia" family="Plethodontidae" genus="Ensatina" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="6" phylum="Chordata" rank="subSpecies" species="eschscholtzii" subSpecies="klauberi">
<emphasis id="B93DDE3B496F1879FE42F8AAFCEAF8A5" box="[400,785,1813,1837]" italics="true" pageId="7" pageNumber="6">Ensatina eschscholtzii klauberi</emphasis>
Dunn, 1929
</taxonomicName>
, and investigating other localities where it is known (e.g., Sierra San Pedro Martir,
<collectingRegion id="498DCCCB496F1879FD15F889FC79F8C5" box="[711,898,1845,1869]" country="Mexico" name="Baja California" pageId="7" pageNumber="6">Baja California</collectingRegion>
Norte,
<collectingCountry id="F35E42B9496F1879FC0CF889FBCDF8C5" box="[990,1078,1845,1869]" name="Mexico" pageId="7" pageNumber="6">Mexico</collectingCountry>
) may provide more diactidine callipodidans and perhaps even
<taxonomicName id="4C4979AA496F1879FD9EF8EAFD46F8E5" authorityName="Shelley &amp; Richart" authorityYear="2014" box="[588,701,1877,1901]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="hedini">
<emphasis id="B93DDE3B496F1879FD9EF8EAFD46F8E5" box="[588,701,1877,1901]" italics="true" pageId="7" pageNumber="6">D. hedini</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35351A249601876FF12FF7EFB58FE92" pageId="8" pageNumber="7" type="etymology">
<paragraph id="8BF6022949601876FF12FF7EFB58FE92" blockId="8.[192,1440,194,282]" pageId="8" pageNumber="7">
<emphasis id="B93DDE3B49601876FF12FF7EFEAFFF52" bold="true" box="[192,340,194,218]" pageId="8" pageNumber="7">Etymology</emphasis>
. We are pleased to name this species for Dr. Hedin, who graciously allowed RMS to search through his collection, take samples of interest, and add them to the invertebrate research holdings at the NCSM. The
<typeStatus id="54F2BC8B49601876FE5AFEBEFE31FE92" box="[392,458,258,282]" pageId="8" pageNumber="7">types</typeStatus>
of this species are deposited in the primary
<typeStatus id="54F2BC8B49601876FC39FEBEFBDBFE92" box="[1003,1056,258,282]" pageId="8" pageNumber="7">type</typeStatus>
collection.
</paragraph>
</subSubSection>
<subSubSection id="C35351A249601876FF12FEFDFD4BFE12" pageId="8" pageNumber="7" type="distribution">
<paragraph id="8BF6022949601876FF12FEFDFD4BFE12" blockId="8.[192,1440,321,410]" pageId="8" pageNumber="7">
<emphasis id="B93DDE3B49601876FF12FEFDFE97FED1" bold="true" box="[192,364,321,345]" pageId="8" pageNumber="7">Distribution</emphasis>
. Known only from the
<typeStatus id="54F2BC8B49601876FD55FEFEFD47FED2" box="[647,700,322,346]" pageId="8" pageNumber="7">type</typeStatus>
locality, in the Cleveland Nat. For., southcentral San Diego Co., near the center of the generic range. The site is also the community of Guatay, approximately
<quantity id="4CB1AFCC49601876FA50FEDEFF13FE12" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="7" unit="km" value="40.0">40 km</quantity>
(
<quantity id="4CB1AFCC49601876FF25FE3DFEC5FE12" box="[247,318,385,410]" metricMagnitude="4" metricUnit="m" metricValue="4.02336" pageId="8" pageNumber="7" unit="mi" value="25.0">25 mi</quantity>
) east of downtown San Diego.
</paragraph>
</subSubSection>
<subSubSection id="C35351A249601876FF12FE7EFA5AFD11" pageId="8" pageNumber="7" type="discussion">
<paragraph id="8BF6022949601876FF12FE7EFA5AFD11" blockId="8.[192,1441,450,666]" pageId="8" pageNumber="7">
<emphasis id="B93DDE3B49601876FF12FE7EFEC7FE52" bold="true" box="[192,316,450,474]" pageId="8" pageNumber="7">Remarks</emphasis>
.
<taxonomicName id="4C4979AA49601876FE80FE7EFE04FE52" authorityName="Shelley &amp; Richart" authorityYear="2014" box="[338,511,450,474]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="hedini">
<emphasis id="B93DDE3B49601876FE80FE7EFE04FE52" box="[338,511,450,474]" italics="true" pageId="8" pageNumber="7">Diactis hedini</emphasis>
</taxonomicName>
belongs to the “
<taxonomicName id="4C4979AA49601876FD16FE7EFCA6FE52" authorityName="Loomis" authorityYear="1937" box="[708,861,450,474]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="triangula">
<emphasis id="B93DDE3B49601876FD16FE7EFCA6FE52" box="[708,861,450,474]" italics="true" pageId="8" pageNumber="7">D. triangula</emphasis>
</taxonomicName>
species group” with
<taxonomicName id="4C4979AA49601876FB89FE7EFB0FFE52" authorityName="Loomis" authorityYear="1937" box="[1115,1268,450,474]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="triangula">
<emphasis id="B93DDE3B49601876FB89FE7EFB0FFE52" box="[1115,1268,450,474]" italics="true" pageId="8" pageNumber="7">D. triangula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4979AA49601876FAE3FE7EFF02FE72" authorityName="Loomis" authorityYear="1937" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="frondifera">
<emphasis id="B93DDE3B49601876FAE3FE7EFF02FE72" italics="true" pageId="8" pageNumber="7">D. frondifera</emphasis>
</taxonomicName>
. While differences exist in the distal telopodal configurations and branches, the major distinction between the components is the position of the third projection, subbasal and a true “prefemoral process” in
<taxonomicName id="4C4979AA49601876FF0DFD9EFEABFDB2" authorityName="Shelley &amp; Richart" authorityYear="2014" box="[223,336,546,570]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="hedini">
<emphasis id="B93DDE3B49601876FF0DFD9EFEABFDB2" box="[223,336,546,570]" italics="true" pageId="8" pageNumber="7">D. hedini</emphasis>
</taxonomicName>
and shorter and progressively more distal in
<taxonomicName id="4C4979AA49601876FCA9FD9EFBE2FDB2" authorityName="Loomis" authorityYear="1937" box="[891,1049,546,570]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="frondifera">
<emphasis id="B93DDE3B49601876FCA9FD9EFBE2FDB2" box="[891,1049,546,570]" italics="true" pageId="8" pageNumber="7">D. frondifera</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4979AA49601876FB87FD9EFB17FDB2" authorityName="Loomis" authorityYear="1937" box="[1109,1260,546,570]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="triangula">
<emphasis id="B93DDE3B49601876FB87FD9EFB91FDB1" box="[1109,1130,546,569]" italics="true" pageId="8" pageNumber="7">D</emphasis>
.
<emphasis id="B93DDE3B49601876FBA5FD9EFB17FDB2" box="[1143,1260,546,570]" italics="true" pageId="8" pageNumber="7">triangula</emphasis>
</taxonomicName>
, hence labeled process ‘B’ (
<figureCitation id="13721EAC49601876FE9CFDFEFE4FFDD1" box="[334,436,577,602]" captionStart="Figures 1-6" captionStartId="7.[192,288,1270,1292]" captionTargetBox="[201,1432,206,1250]" captionTargetId="figure-174@7.[192,1440,192,1251]" captionTargetPageId="7" captionText="Figures 1-6. Diactis spp. 1) D. hedini holotype, right gonopod, caudal view. 2) D. hedini holotype, right gonopod, telopodite of the same, medial view. 3) D. hedini holotype, right gonopod, the same, lateral view. 4) D. hedini holotype, right gonopod, distal extremity of the same, anterior view. 5) Telopodite of right gonopod of D. frondifera male from Los Angeles Co., California, lateral view. 6) Telopodite of right gonopod of D. triangula male from San Bernardino Co., California, anterior view. A, process ‘A’; B, process ‘B’; pfp, prefemoral process; s, solenomere; sb, solenomere branch; tt, tibiotarsus." figureDoi="http://doi.org/10.5281/zenodo.5178597" httpUri="https://zenodo.org/record/5178597/files/figure.png" pageId="8" pageNumber="7">Fig. 5–6</figureCitation>
). Though longer in
<taxonomicName id="4C4979AA49601876FD4DFDFEFCEBFDD2" authorityName="Shelley &amp; Richart" authorityYear="2014" box="[671,784,578,602]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="hedini">
<emphasis id="B93DDE3B49601876FD4DFDFEFCEBFDD2" box="[671,784,578,602]" italics="true" pageId="8" pageNumber="7">D. hedini</emphasis>
</taxonomicName>
, the projections seem to represent the same structure in all species; its telopodal position is what changes. Consequently,
<taxonomicName id="4C4979AA49601876FC3FFDDEFBA7FDF2" authorityName="Shelley &amp; Richart" authorityYear="2014" box="[1005,1116,610,634]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="hedini">
<emphasis id="B93DDE3B49601876FC3FFDDEFBA7FDF2" box="[1005,1116,610,634]" italics="true" pageId="8" pageNumber="7">D. hedini</emphasis>
</taxonomicName>
clarifies process ‘B’, whose apparent identity was not evident when
<taxonomicName id="4C4979AA49601876FD77FD3EFCC1FD12" authorityName="Loomis" authorityYear="1937" box="[677,826,642,666]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="triangula">
<emphasis id="B93DDE3B49601876FD77FD3EFCC1FD12" box="[677,826,642,666]" italics="true" pageId="8" pageNumber="7">D. triangula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4979AA49601876FCA1FD3EFBF4FD12" authorityName="Loomis" authorityYear="1937" box="[883,1039,642,666]" class="Diplopoda" family="Tynommatidae" genus="Diactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Callipodida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="frondifera">
<emphasis id="B93DDE3B49601876FCA1FD3EFC73FD11" box="[883,904,642,665]" italics="true" pageId="8" pageNumber="7">D</emphasis>
.
<emphasis id="B93DDE3B49601876FC46FD3EFBF4FD12" box="[916,1039,642,666]" italics="true" pageId="8" pageNumber="7">frondifera</emphasis>
</taxonomicName>
were the only group components.
</paragraph>
</subSubSection>
</treatment>
</document>
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