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<mods:title id="62805071085552917DA205180E817160">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XIII. Review of Pandinops hawkeri, P. peeli, P. platycheles, and P. pugilator (Scorpionidae)</mods:title>
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<mods:namePart id="CD7B5FA5BCA805481694C512CBF9B6EB">Elmi, Hassan Sh Abdirahman</mods:namePart>
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<mods:title id="D2ED443B62240DA018D1F72E32FC4862">Euscorpius</mods:title>
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<mods:date id="CD49B757F0DE2693ACA85A5F242055E0">2017</mods:date>
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<paragraph id="8BF636AF7A251775FC53F9EEFA8A0D1C" blockId="2.[1018,1301,1618,1675]" box="[1018,1301,1618,1646]" pageId="2" pageNumber="1">
<heading id="D0BE81C37A251775FC53F9EEFA8A0D1C" box="[1018,1301,1618,1646]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="1" reason="4">
<taxonomicName id="4C494D2C7A251775FC53F9EEFA8A0D1C" ID-CoL="6D6C" authority="Birula, 1913" authorityName="Birula" authorityYear="1913" box="[1018,1301,1618,1646]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEABD7A251775FC53F9EEFBE40D1F" bold="true" box="[1018,1147,1618,1645]" italics="true" pageId="2" pageNumber="1">Pandinops</emphasis>
Birula, 1913
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C35365247A251775FBBEF9C8FC3C0C57" pageId="2" pageNumber="1" type="description">
<paragraph id="8BF636AF7A251775FBBEF9C8FB670DF9" blockId="2.[1018,1301,1618,1675]" box="[1047,1272,1652,1675]" pageId="2" pageNumber="1">
(
<figureCitation id="13722A2A7A251775FBB6F9C8FB0D0DF9" box="[1055,1170,1652,1675]" captionStart-0="Figures 12" captionStart-1="Figures 310" captionStart-10="Figures 4648" captionStart-11="Figure 49" captionStart-12="Figure 50" captionStart-13="Figures 5156" captionStart-14="Figures 5758" captionStart-15="Figures 5960" captionStart-2="Figures 1114" captionStart-3="Figures 1518" captionStart-4="Figures 1924" captionStart-5="Figures 2530" captionStart-6="Figures 3136" captionStart-7="Figures 3740" captionStart-8="Figures 4142" captionStart-9="Figures 4345" captionStartId-0="3.[192,278,1479,1502]" captionStartId-1="4.[192,278,1639,1662]" captionStartId-10="13.[192,278,1863,1886]" captionStartId-11="14.[192,268,1054,1077]" captionStartId-12="15.[192,268,1422,1445]" captionStartId-13="16.[192,278,1095,1118]" captionStartId-14="17.[192,278,1800,1823]" captionStartId-15="18.[192,278,1835,1858]" captionStartId-2="5.[192,278,1298,1321]" captionStartId-3="6.[192,278,1809,1832]" captionStartId-4="7.[192,278,1809,1832]" captionStartId-5="8.[192,278,1656,1679]" captionStartId-6="9.[192,278,1330,1353]" captionStartId-7="10.[192,278,874,897]" captionStartId-8="11.[192,278,1853,1876]" captionStartId-9="12.[192,278,1830,1853]" captionTargetBox-0="[191,1437,191,1450]" captionTargetBox-1="[214,1428,186,1605]" captionTargetBox-10="[191,1440,191,1834]" captionTargetBox-11="[191,1437,167,1024]" captionTargetBox-12="[196,1436,197,1390]" captionTargetBox-13="[191,1437,191,1066]" captionTargetBox-14="[226,1393,191,1770]" captionTargetBox-15="[242,1369,191,1805]" captionTargetBox-2="[191,1437,191,1268]" captionTargetBox-3="[209,1435,197,1772]" captionTargetBox-4="[194,1397,195,1778]" captionTargetBox-5="[191,1440,191,1626]" captionTargetBox-6="[213,1424,213,1278]" captionTargetBox-7="[200,1433,179,844]" captionTargetBox-8="[191,1440,191,1824]" captionTargetBox-9="[191,1440,171,1801]" captionTargetId-0="figure-6@3.[192,1437,192,1449]" captionTargetId-1="figure-9@4.[192,1439,168,1609]" captionTargetId-10="figure-6@13.[192,1439,192,1833]" captionTargetId-11="figure-9@14.[192,1437,168,1023]" captionTargetId-12="figure-6@15.[192,1439,192,1393]" captionTargetId-13="figure-40@16.[192,1437,192,1065]" captionTargetId-14="figure-113@17.[227,1393,192,1769]" captionTargetId-15="figure-41@18.[243,1368,192,1804]" captionTargetId-2="figure-6@5.[192,1437,192,1267]" captionTargetId-3="figure-9@6.[192,1439,192,1779]" captionTargetId-4="figure-6@7.[192,1439,192,1779]" captionTargetId-5="figure-9@8.[192,1439,192,1626]" captionTargetId-6="figure-6@9.[192,1439,192,1300]" captionTargetId-7="figure-9@10.[192,1437,168,844]" captionTargetId-8="figure-6@11.[192,1439,192,1823]" captionTargetId-9="figure-9@12.[192,1439,172,1800]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="13" captionTargetPageId-11="14" captionTargetPageId-12="15" captionTargetPageId-13="16" captionTargetPageId-14="17" captionTargetPageId-15="18" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="Figures 12: Pandinops platycheles, male holotype in dorsal (1) and ventral (2) aspects. In the plate there are also original labels. Scale bar: 10 mm." captionText-1="Figures 310: Pandinops platycheles, male holotype, pedipalp segments. Chela dorsoexternal (3), ventroexternal (4) and ventrointernal (5). Patella dorsal (6), external (7) and ventral (8). Femur and trochanter dorsal (9) and ventral (10). Trichobothrial pattern is indicated." captionText-10="Figures 4648: Pandinops pugilator, burrows at locality No. 17SR." captionText-11="Figure 49: Somaliland, Jifa Uri hill (09°43'19&quot;N 43°23'33&quot;E), the type locality of P. hawkeri." captionText-12="Figure 50: Map showing distribution of Pandinops spp. with corrected confirmed distribution of Pandinops pugilator. Scorpion in photos inside map is the holotype of P. hawkeri." captionText-13="Figures 5156: Pandinops pugilator, hemispermatophores from 3 specimens (Nos. 1317, 1318, 1333). Ventral (concave) (51, 53, 55) and dorsal (convex) (52, 54, 56) views. No. 1317 is left hemispermatophore shown in mirror image for comparison to others, Nos. 1318, 1333 are right hemispermatophores. Scale bars: 1 mm." captionText-14="Figures 5758: Pandinops pugilator, left hemispermatophore, capsule region (No. 1317). Internal (sperm duct margin) (57) and external (distal lamina margin) (58) cross-stereoscopic views. Scale bar: 500 µm. Label abbreviations: bl, basal lobe; dt, dorsal trough; h, hook; ibr, internobasal reflection of sperm duct (= median transverse trough); il, inner lobe, ml, median lobe; tf, truncal flexure (= median transverse cleavage)." captionText-15="Figures 5960: Pandinops pugilator, left hemispermatophore, capsule region (No. 1317). Dorsal (convex) (59) and ventral (concave) (60) cross-stereoscopic views. Scale bar: 500 µm. Label abbreviations: see legend for Figs. 5758." captionText-2="Figures 1114: Pandinops platycheles. Figure 11. Male paratype, metasoma and telson ventral and original labels. Figures 1214. Male holotype, metasoma and telson lateral (12), metasoma, telson ventral and sternite VII (13), and metasoma and telson dorsal (14) views. Scale bar: 10 mm." captionText-3="Figures 1518: Pandinops pugilator from locality No. 17SR. Figures 1516. Male in dorsal (15) and ventral (16) views. Figures 1718. Female in dorsal (17) and ventral (18) views. Scale bar: 10 mm." captionText-4="Figures 1924: Pandinops pugilator from locality No. 17SR. Figures 1921. Female, pedipalp chela dorsoexternal (19), ventroexternal (20), and ventrointernal (21). Figures 2224. Male, pedipalp chela and patella dorsoexternal (22), ventroexternal (23), and ventrointernal (24). Trichobothrial pattern is indicated in Figures 2224." captionText-5="Figures 2530: Pandinops pugilator from locality No. 17SR. Figures 25, 27. Male, carapace and tergites IIII (25) and coxosternal area and sternites IIIV (27). Figures 26, 2830. Female, Carapace and tergite I (26), coxosternal area and sternite III (30), right chelicera dorsal (29) and ventral (30)." captionText-6="Figures 3136: Pandinops pugilator from locality No. 17SR, metasoma, telson, and sternite VII. Figures 3133. Male, lateral (31), dorsal (32), and ventral (33) views. Figures 3436. Female, lateral (34), dorsal (35), and ventral (36) views. Scale bar: 10 mm." captionText-7="Figures 3740: Pandinops pugilator from locality No. 17SR, female, tarsomeres I and II of left legs IIV, retrolateral aspects." captionText-8="Figures 4142: Pandinops pugilator from locality No. 17SR, in vivo habitus. Figure 41. Male. Figure 42. Female." captionText-9="Figures 4345: Pandinops pugilator. Figure 43. Juvenile topotype of P. hawkeri from locality No. 17SP, in vivo habitus. Figure 44. Locality No. 17SP, Somaliland, near Jifa Uri hill, 09°43'35&quot;N 43°24'51&quot;E. Figure 45. Locality No. 17SR, Somaliland, Borama, campus Amoud University, 09°56'49&quot;N 43°13'23&quot;E." pageId="2" pageNumber="1">Figs. 160</figureCitation>
,
<tableCitation id="C6CB03147A251775FB37F9C8FB720DF9" box="[1182,1261,1652,1675]" captionStart="Table 1" captionStartId="10.[192,257,1543,1566]" captionText="Table 1: Comparative measurements of adults of Pandinops platycheles and P. turieli Kovařík, 2016. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="2" pageNumber="1">Table 1</tableCitation>
)
</paragraph>
<paragraph id="8BF636AF7A251775FCC7F90EFC3C0C57" blockId="2.[830,1441,1714,1829]" pageId="2" pageNumber="1">
<taxonomicName id="4C494D2C7A251775FCC7F90EFAF30DBB" authority="Birula, 1913: 419 - 422" authorityName="Birula" authorityPageNumber="419 - 422" authorityYear="1913" box="[878,1388,1714,1737]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Pandinops">
<emphasis id="B93DEABD7A251775FCC7F90EFC4C0DBB" box="[878,979,1714,1737]" italics="true" pageId="2" pageNumber="1">Pandinus</emphasis>
(
<emphasis id="B93DEABD7A251775FC41F90EFBC50DBB" box="[1000,1114,1714,1737]" italics="true" pageId="2" pageNumber="1">Pandinops</emphasis>
) Birula, 1913: 419422
</taxonomicName>
, fig. b;
<bibRefCitation id="EFD84B5E7A251775FCF6F96CFBA70D95" author="VACHON" box="[863,1080,1744,1767]" pageId="2" pageNumber="1" pagination="857 - 958" refId="ref6634" refString="VACHON, M. 1974. Etudes des caracteres utilises pour classer les familles et les genres des scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, 3 e serie, 140 (Zoologie, 104): 857 - 958." type="journal article" year="1974">Vachon, 1974: 921</bibRefCitation>
, 953, figs. 116118; Fet, 2000: 469; Kovařík, 2009: 5153, 115118, figs. 294298, 304327.
</paragraph>
</subSubSection>
<caption id="DF3666277A241774FF69FA7BFE2B0E85" pageId="3" pageNumber="2" startId="3.[192,278,1479,1502]" targetBox="[191,1437,191,1450]" targetPageId="3">
<paragraph id="8BF636AF7A241774FF69FA7BFE2B0E85" blockId="3.[192,1441,1479,1527]" pageId="3" pageNumber="2">
<emphasis id="B93DEABD7A241774FF69FA7BFECF0EAC" bold="true" box="[192,336,1479,1502]" pageId="3" pageNumber="2">Figures 12:</emphasis>
<taxonomicName id="4C494D2C7A241774FEF0FA75FDAA0EAC" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[345,565,1481,1502]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="platycheles">
<emphasis id="B93DEABD7A241774FEF0FA75FDAA0EAC" box="[345,565,1481,1502]" italics="true" pageId="3" pageNumber="2">Pandinops platycheles</emphasis>
</taxonomicName>
, male holotype in dorsal (1) and ventral (2) aspects. In the plate there are also original labels. Scale bar: 10 mm.
</paragraph>
</caption>
<subSubSection id="C35365247A241774FF59F990FD530D31" box="[240,716,1580,1603]" pageId="3" pageNumber="2" type="materials_examined">
<paragraph id="8BF636AF7A241774FF59F990FD530D31" blockId="3.[240,716,1580,1603]" box="[240,716,1580,1603]" pageId="3" pageNumber="2">
<materialsCitation id="3B213CF27A241774FF59F990FD5A0D31" box="[240,709,1580,1603]" collectionCode="SPECIES" pageId="3" pageNumber="2" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F2880D7A241774FF59F990FEB70D30" box="[240,296,1580,1603]" pageId="3" pageNumber="2">TYPE</typeStatus>
<collectionCode id="ED58AE6A7A241774FE87F993FE1E0D30" box="[302,385,1583,1602]" pageId="3" pageNumber="2">SPECIES</collectionCode>
.
<taxonomicName id="4C494D2C7A241774FE27F990FD5A0D31" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" box="[398,709,1580,1603]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="peeli">
<emphasis id="B93DEABD7A241774FE27F990FDB10D31" box="[398,558,1580,1603]" italics="true" pageId="3" pageNumber="2">Pandinus peeli</emphasis>
Pocock, 1900
</taxonomicName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35365247A241773FF59F9D5FB920C3A" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="8BF636AF7A241773FF59F9D5FB920C3A" blockId="3.[192,802,1641,1909]" lastBlockId="4.[830,1440,1749,1864]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
DIAGNOSIS. Total length
<quantity id="4CB19B4A7A241774FDA7F9D5FD180DF2" box="[526,647,1641,1664]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="9.5" metricValueMin="5.5" pageId="3" pageNumber="2" unit="mm" value="75.0" valueMax="95.0" valueMin="55.0">5595 mm</quantity>
. External trichobothria on patella number 1316 (56
<emphasis id="B93DEABD7A241774FD22F934FD3C0DED" box="[651,675,1672,1695]" italics="true" pageId="3" pageNumber="2">eb</emphasis>
, 24
<emphasis id="B93DEABD7A241774FD4AF934FC9A0DED" box="[739,773,1672,1695]" italics="true" pageId="3" pageNumber="2">esb</emphasis>
, 2
<emphasis id="B93DEABD7A241774FF68F91AFF400DCF" box="[193,223,1702,1725]" italics="true" pageId="3" pageNumber="2">em</emphasis>
, 12
<emphasis id="B93DEABD7A241774FE99F91AFED10DCF" box="[304,334,1702,1725]" italics="true" pageId="3" pageNumber="2">est</emphasis>
, 3
<emphasis id="B93DEABD7A241774FE2DF91AFE080DCF" box="[388,407,1702,1725]" italics="true" pageId="3" pageNumber="2">et</emphasis>
); ventral trichobothria on patella number 2235; internal trichobothria on chela number 68; ventral trichobothria on chela number 913. Pedipalp chela manus lobiform. Movable fingers of pedipalp, length of segments of pedipalps, and telson without noticeable sexual dimorphism. Pectines with fulcra. Pectinal teeth number 1121. Sternum sub- pentagonal, longer than wide. Carapace without distinct carinae. Dentate margin of pedipalp chela movable finger with distinct granules divided into 57 rows. Tergites IVI of mesosoma bear one carina. Stridulation organ located on pedipalp coxae and first pair of legs, but can be reduced. Metasomal segments IIV with paired parallel ventral median carinae or without carinae. Telson without subaculear tubercle. Legs with one pedal spur, retrolateral spur absent.
<emphasis id="B93DEABD7A241774FB52F89DFCF60C25" italics="true" pageId="3" pageNumber="2">Tarsomere I of legs</emphasis>
. Spiniform macroseta
<emphasis id="B93DEABD7A241774FBC1F8FCFB1C0C25" box="[1128,1155,1856,1879]" italics="true" pageId="3" pageNumber="2">pd</emphasis>
,
<emphasis id="B93DEABD7A241774FB3DF8FCFB380C25" box="[1172,1191,1856,1879]" italics="true" pageId="3" pageNumber="2">vt</emphasis>
,
<emphasis id="B93DEABD7A241774FB11F8FCFB550C25" box="[1208,1226,1856,1879]" italics="true" pageId="3" pageNumber="2">rt</emphasis>
,
<emphasis id="B93DEABD7A241774FB72F8FCFB660C25" box="[1243,1273,1856,1879]" italics="true" pageId="3" pageNumber="2">vst</emphasis>
are present on legs IIV, but
<emphasis id="B93DEABD7A241774FC70F8E2FC6B0C07" box="[985,1012,1886,1909]" italics="true" pageId="3" pageNumber="2">pd</emphasis>
can be replaced by seta as intraspecific variability;
<emphasis id="B93DEABD7A231773FE96F969FEC10D9E" box="[319,350,1749,1772]" italics="true" pageId="4" pageNumber="3">pst</emphasis>
is present on legs IIIIV;
<emphasis id="B93DEABD7A231773FD20F969FD010D9E" box="[649,670,1749,1772]" italics="true" pageId="4" pageNumber="3">pt</emphasis>
and
<emphasis id="B93DEABD7A231773FD71F969FD680D9E" box="[728,759,1749,1772]" italics="true" pageId="4" pageNumber="3">vm</emphasis>
are absent on all legs;
<emphasis id="B93DEABD7A231773FE2EF948FE3A0C79" box="[391,421,1780,1803]" italics="true" pageId="4" pageNumber="3">rm</emphasis>
is present on legs IIV, but is often replaced by seta or spiniform seta.
<emphasis id="B93DEABD7A231773FD96F8AEFC840C5B" box="[575,795,1810,1833]" italics="true" pageId="4" pageNumber="3">Tarsomere II of legs</emphasis>
. Spiniform formula is 3/4: 3/4: 3/4-5: 3/4-5. Tarsomere II with 2 spines on inclined anteroventral surface, but a seta on leg III can be transformed to &quot;spiniform seta&quot; which indicates a poorly developed third spine as intraspecific variability.
</paragraph>
</subSubSection>
</treatment>
</document>