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<document id="47EA6035E3E8E0EBE9B1C186B9F89649" ID-DOI="10.1111/j.1096-3642.2011.00737.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5442183" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630452696336" checkinUser="valdenar" docAuthor="Mazza, Paul P. A. &amp; Rustioni, Marco" docDate="2011" docId="03E0692FBC25FFE2FC59ED23D39FF92B" docLanguage="en" docName="j.1096-3642.2011.00737.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (4)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00737.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Hoplitomeryx magnus Mazza &amp; Rustioni 2011, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="1326" masterDocId="FFD91157BC31FFF4FFFDE85BD237FFB2" masterDocTitle="Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983" masterLastPageNumber="1333" masterPageNumber="1304" pageNumber="1324" updateTime="1699275809674" updateUser="plazi">
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<mods:title id="2A26BCC412070AF1FB15B6DC42ACA5FF">Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983</mods:title>
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<mods:namePart id="8E562AA65CB116841DE1CC16C300368B">Mazza, Paul P. A.</mods:namePart>
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<mods:namePart id="B96F7E1ACA8079BC486F3ED58F454DF3">Rustioni, Marco</mods:namePart>
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<mods:title id="2CBE0630830D5AC796CB01D9CCEB961D">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="72C007340B5D6FF5BA850848EA6D85A2">2011</mods:date>
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<treatment id="03E0692FBC25FFE2FC59ED23D39FF92B" LSID="urn:lsid:plazi:treatment:03E0692FBC25FFE2FC59ED23D39FF92B" httpUri="http://treatment.plazi.org/id/03E0692FBC25FFE2FC59ED23D39FF92B" lastPageId="22" lastPageNumber="1326" pageId="20" pageNumber="1324">
<subSubSection id="C3538BB2BC25FFE0FC59ED23D6D4FA06" pageId="20" pageNumber="1324" type="nomenclature">
<paragraph id="8BF6D839BC25FFE0FC59ED23D70FFA3F" blockId="20.[932,1336,1400,1421]" box="[932,1336,1400,1421]" pageId="20" pageNumber="1324">
<heading id="D0BE6F55BC25FFE0FC59ED23D70FFA3F" box="[932,1336,1400,1421]" centered="true" fontSize="9" level="2" pageId="20" pageNumber="1324" reason="4">
<taxonomicName id="4C49A3BABC25FFE0FC59ED23D6FFFA3E" authority="Mazza &amp; Rustioni, 2011" authorityName="Mazza &amp; Rustioni" authorityYear="2011" box="[932,1224,1400,1421]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="20" pageNumber="1316" phylum="Chordata" rank="species" species="magnus" status="sp. nov.">HOPLITOMERYX MAGNUS</taxonomicName>
<emphasis id="B93D042BBC25FFE0FB2EED20D70FFA3F" bold="true" box="[1235,1336,1400,1421]" pageId="20" pageNumber="1324">
<taxonomicNameLabel id="A20EB950BC25FFE0FB2EED20D70FFA3F" box="[1235,1336,1400,1421]" pageId="20" pageNumber="1324" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF6D839BC25FFE0FC04EDC5D6D4FA06" blockId="20.[1017,1251,1438,1460]" box="[1017,1251,1438,1460]" pageId="20" pageNumber="1324">
<heading id="D0BE6F55BC25FFE0FC04EDC5D6D4FA06" box="[1017,1251,1438,1460]" centered="true" fontSize="8" level="2" pageId="20" pageNumber="1324" reason="8">
(
<tableCitation id="C6CBED82BC25FFE0FBFDEDC5D657FA01" box="[1024,1120,1438,1459]" captionStart="Table 1" captionStartId="3.[144,208,467,488]" captionText="Table 1. Differential diagnoses of dental characters" pageId="20" pageNumber="1324">TABLE 1</tableCitation>
;
<figureCitation id="1372C4BCBC25FFE0FB8CEDC4D68AFA01" box="[1137,1213,1438,1460]" captionStart="Figure 4" captionStartId="19.[144,223,1446,1465]" captionTargetBox="[304,1262,195,1416]" captionTargetId="figure-58@19.[303,1263,194,1416]" captionTargetPageId="19" captionText="Figure 4. A, Hoplitomeryx apruthiensis sp. nov., right maxillary (SCT 59): labial view. B, Hoplitomeryx apruthiensis sp. nov., right maxillary (SCT 59): lingual view. C, Hoplitomeryx apruthiensis sp. nov., right maxillary (SCT 59): occlusal view. D, Hoplitomeryx apruthiensis sp. nov., right mandible (SCT 29): labial view. E, Hoplitomeryx apruthiensis sp. nov., right mandible (SCT 29): lingual view. F, Hoplitomeryx apruthiensis sp. nov., right mandible (SCT 29): occlusal view. G, Hoplitomeryx magnus sp. nov., left maxillary (SCT 86): occlusal view. H, Hoplitomeryx magnus sp. nov., left maxillary (SCT 86): lingual view. I, Hoplitomeryx magnus sp. nov., left maxillary (SCT 86): labial view. J, Hoplitomeryx magnus sp. nov., left mandible (SCT 20): labial view. K, Hoplitomeryx magnus sp. nov., left mandible (SCT 20): lingual view. L, Hoplitomeryx magnus sp. nov., left mandible (SCT 20): occlusal view. M, Hoplitomeryx magnus sp. nov., left mandible (RGM 260.951): lingual view. N, Hoplitomeryx magnus sp. nov., left mandible (RGM 260.951): labial view. O, Hoplitomeryx magnus sp. nov., left mandible (RGM 260.951): occlusal view." figureDoi="http://doi.org/10.5281/zenodo.5442191" httpUri="https://zenodo.org/record/5442191/files/figure.png" pageId="20" pageNumber="1324">FIGS 4</figureCitation>
,
<figureCitation id="1372C4BCBC25FFE0FB31EDC5D6EDFA06" box="[1228,1242,1438,1460]" captionStart="Figure 5" captionStartId="20.[164,243,890,909]" captionTargetBox="[419,1187,196,860]" captionTargetId="figure-258@20.[419,1187,196,860]" captionTargetPageId="20" captionText="Figure 5. Hoplitomeryx minutus sp. nov., left p3: occlusal view. B, Hoplitomeryx minutus sp. nov., left p4: occlusal view. C, Hoplitomeryx minutus sp. nov., left m3: occlusal view. D, Hoplitomeryx falcidens sp. nov., left p3: occlusal view. E, Hoplitomeryx falcidens sp. nov., left p4: occlusal view. F, Hoplitomeryx falcidens sp. nov., left m3: occlusal view. G, Hoplitomeryx apulicus sp. nov., right p3: occlusal view. H, Hoplitomeryx apulicus sp. nov., right p4: occlusal view. I, Hoplitomeryx apulicus sp. nov., right m3: occlusal view. J, Hoplitomeryx matthei sp. nov., left p3: occlusal view. K, Hoplitomeryx matthei sp. nov., left p4: occlusal view. L, Hoplitomeryx matthei sp. nov., left m3: occlusal view. M, Hoplitomeryx apruthiensis sp. nov., right p3: occlusal view. N, Hoplitomeryx apruthiensis sp. nov., right p4: occlusal view. O, Hoplitomeryx apruthiensis sp. nov., right m3: occlusal view. P, Hoplitomeryx magnus sp. nov., left p3: occlusal view. Q, Hoplitomeryx magnus sp. nov., left p4: occlusal view. R, Hoplitomeryx magnus sp. nov., left m3: occlusal view." figureDoi="http://doi.org/10.5281/zenodo.5442193" httpUri="https://zenodo.org/record/5442193/files/figure.png" pageId="20" pageNumber="1324">5</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3538BB2BC25FFE2FCC7ED9ED39FF92B" lastPageId="22" lastPageNumber="1326" pageId="20" pageNumber="1324" type="materials_examined">
<paragraph id="8BF6D839BC25FFE0FCC7ED9ED6C5FA68" blockId="20.[826,1266,1477,1499]" box="[826,1266,1477,1499]" pageId="20" pageNumber="1324">
<emphasis id="B93D042BBC25FFE0FCC7ED9ED193FA68" box="[826,932,1477,1498]" italics="true" pageId="20" pageNumber="1324">
<typeStatus id="54F2669BBC25FFE0FCC7ED9ED1A9FA68" box="[826,926,1477,1498]" pageId="20" pageNumber="1324" type="holotype">Holotype</typeStatus>
:
</emphasis>
Left hemimandible SCT 20.
</paragraph>
<paragraph id="8BF6D839BC25FFE0FCC7EE5DD1D6F9EA" blockId="20.[826,1441,1541,1624]" pageId="20" pageNumber="1324">
<emphasis id="B93D042BBC25FFE0FCC7EE5DD185F9A9" box="[826,946,1542,1563]" italics="true" pageId="20" pageNumber="1324">
<typeStatus id="54F2669BBC25FFE0FCC7EE5DD19CF9A9" box="[826,939,1542,1563]" pageId="20" pageNumber="1324" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<materialsCitation id="3B21D264BC25FFE0FC34EE5ED1D6F9EA" collectionCode="RGM" pageId="20" pageNumber="1324" specimenCode="RGM 260.951, RGM 261.135, RGM 261.144" specimenCount="1" typeStatus="holotype">
Maxillaries SCT 18, SCT 86,
<specimenCode id="DBEF7042BC25FFE0FA98EE5ED1A1F98B" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="1324" type="Museum">RGM 260.951</specimenCode>
; mandibles
<specimenCode id="DBEF7042BC25FFE0FBCEEE7FD6EAF98B" box="[1075,1245,1571,1593]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="1324" type="Museum">RGM 260.951</specimenCode>
,
<specimenCode id="DBEF7042BC25FFE0FB0EEE7FD7AAF98B" box="[1267,1437,1571,1593]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="1324" type="Museum">RGM 261.135</specimenCode>
,
<specimenCode id="DBEF7042BC25FFE0FCC7EE19D1EAF9EA" box="[826,989,1602,1624]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="1324" type="Museum">RGM 261.144</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF6D839BC25FFE1FCC7EED8D32AFE00" blockId="20.[826,1441,1667,1903]" lastBlockId="21.[144,759,198,434]" lastPageId="21" lastPageNumber="1325" pageId="20" pageNumber="1324">
<emphasis id="B93D042BBC25FFE0FCC7EED8D643F92A" box="[826,1140,1667,1688]" italics="true" pageId="20" pageNumber="1324">Type locality and horizon:</emphasis>
<materialsCitation id="3B21D264BC25FFE0FB7FEED8D60DF880" country="Italy" county="L'Aquila" latitude="41.75432" location="Scontrone" longLatPrecision="1" longitude="14.0369835" municipality="National Park of Abruzzi" pageId="20" pageNumber="1324" specimenCount="2" typeStatus="holotype">
<typeStatus id="54F2669BBC25FFE0FB7FEED8D6DFF92B" box="[1154,1256,1667,1689]" pageId="20" pageNumber="1324" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54F2669BBC25FFE0FACDEED8D796F92A" box="[1328,1441,1667,1688]" pageId="20" pageNumber="1324" type="paratype">paratypes</typeStatus>
tagged SCT from the Tortonian Scontrone Member of the Lithothamnium Limestone (
<bibRefCitation id="EFD8A5C8BC25FFE0FB48EE9BD7ABF967" author="Patacca E &amp; Scandone P &amp; Mazza P" box="[1205,1436,1728,1749]" pageId="20" pageNumber="1324" pagination="337 - 355" refId="ref20273" refString="Patacca E, Scandone P, Mazza P. 2008. Oligocene migration path for Apulia macromammals: the Central-Adriatic bridge. Bollettino Della Societa Geolologica Italiana 127: 337 - 355." type="journal article" year="2008">
Patacca
<emphasis id="B93D042BBC25FFE0FAE7EE9BD77DF967" box="[1306,1354,1728,1749]" italics="true" pageId="20" pageNumber="1324">et al</emphasis>
., 2008
</bibRefCitation>
;
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,
<geoCoordinate id="EE7DBEFEBC25FFE0FC10EE84D6B5F946" box="[1005,1154,1757,1780]" degrees="14" direction="east" minutes="2" orientation="longitude" pageId="20" pageNumber="1324" precision="1" seconds="13. 14" value="14.0369835">14°213.14″E</geoCoordinate>
), outskirts of
<location id="8E968EE2BC25FFE0FAD5EE84D7AAF946" LSID="urn:lsid:plazi:treatment:03E0692FBC25FFE2FC59ED23D39FF92B:8E968EE2BC25FFE0FAD5EE84D7AAF946" box="[1320,1437,1759,1780]" country="Italy" county="L'Aquila" latitude="41.75432" longLatPrecision="1" longitude="14.0369835" municipality="National Park of Abruzzi" name="Scontrone" pageId="20" pageNumber="1324">Scontrone</location>
, southern border of the
<collectingMunicipality id="6B924243BC25FFE0FB9EEEA6D7AAF8A1" box="[1123,1437,1789,1811]" pageId="20" pageNumber="1324">National Park of Abruzzi</collectingMunicipality>
,
<collectingCounty id="6297A0B5BC25FFE0FCC7EF47D1A9F880" box="[826,926,1820,1842]" pageId="20" pageNumber="1324">LAquila</collectingCounty>
, central
<collectingCountry id="F35E98A9BC25FFE0FBFEEF47D60DF880" box="[1027,1082,1820,1842]" name="Italy" pageId="20" pageNumber="1324">Italy</collectingCountry>
</materialsCitation>
.
<materialsCitation id="3B21D264BC25FFE0FBBBEF47D76EF8DC" collectionCode="RGM" collectorName="Messinian &amp; S. Giovannino" country="Italy" pageId="20" pageNumber="1324" specimenCode="RGM 260.951" specimenCount="1" typeStatus="paratype">
<typeStatus id="54F2669BBC25FFE0FBBBEF47D68EF883" box="[1094,1209,1820,1841]" pageId="20" pageNumber="1324" type="paratype">Paratypes</typeStatus>
<specimenCode id="DBEF7042BC25FFE0FB3FEF47D768F883" box="[1218,1375,1819,1841]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="1324" type="Museum">RGM 260.951</specimenCode>
come from the
<collectorName id="26BCBDEFBC25FFE0FC4DEF61D611F8E2" box="[944,1062,1850,1872]" pageId="20" pageNumber="1324">Messinian</collectorName>
karstic fissure filling called
<collectorName id="26BCBDEFBC25FFE0FA76EF61D18AF8DD" pageId="20" pageNumber="1324">S. Giovannino</collectorName>
in a homonymous limestone quarry
</materialsCitation>
;
<materialsCitation id="3B21D264BC25FFE1FA98EF02D2DAFEAB" collectionCode="RGM" country="Italy" lastPageId="21" lastPageNumber="1325" pageId="20" pageNumber="1324" specimenCode="RGM 261.135" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBEF7042BC25FFE1FA98EF02D2DEFF6E" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" lastPageId="21" lastPageNumber="1325" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="1324" type="Museum">RGM 261.135</specimenCode>
comes from the Messinian karstic fissure filling called Nazario
<quantity id="4CB175DCBC24FFE1FE5AE8BED3E9FF48" box="[423,478,228,250]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="21" pageNumber="1325" unit="in" value="4.0">4 in</quantity>
the limestone quarry Nazario
</materialsCitation>
;
<materialsCitation id="3B21D264BC24FFE1FEFDE958D32EFE00" collectionCode="RGM" country="Italy" latitude="41.803333" location="Messinian" longLatPrecision="19" longitude="15.384444" municipality="Chiro" pageId="21" pageNumber="1325" specimenCode="RGM 261.144" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBEF7042BC24FFE1FEFDE958D392FEAB" box="[256,421,259,281]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="1325" type="Museum">RGM 261.144</specimenCode>
comes from the
<location id="8E968EE2BC24FFE1FD7CE958D0C0FEAB" LSID="urn:lsid:plazi:treatment:03E0692FBC25FFE2FC59ED23D39FF92B:8E968EE2BC24FFE1FD7CE958D0C0FEAB" box="[641,759,259,281]" country="Italy" latitude="41.803333" longLatPrecision="19" longitude="15.384444" municipality="Chiro" name="Messinian" pageId="21" pageNumber="1325">Messinian</location>
karstic fissure filling called
<collectingMunicipality id="6B924243BC24FFE1FE2DE979D027FE8A" box="[464,528,290,312]" pageId="21" pageNumber="1325">Chiro</collectingMunicipality>
<quantity id="4CB175DCBC24FFE1FDE5E979D064FE8A" box="[536,595,290,312]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" pageId="21" pageNumber="1325" unit="in" value="27.0">27 in</quantity>
the limestone quarry Chiro. The three quarries are located
<locationDeviation id="9A44C11BBC24FFE1FD65E91BD0C1FEC6" firstLocation="Apricena" pageId="21" pageNumber="1325" secondLocation="Poggio Imperiale, Foggia, Gargano">between Apricena and Poggio Imperiale, Foggia, Gargano</locationDeviation>
promontory, south-eastern
<collectingCountry id="F35E98A9BC24FFE1FDF5E926D009FE21" box="[520,574,381,403]" name="Italy" pageId="21" pageNumber="1325">Italy</collectingCountry>
(
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,
<geoCoordinate id="EE7DBEFEBC24FFE1FF6DE9C7D327FE00" box="[144,272,411,434]" degrees="15" direction="east" minutes="23" orientation="longitude" pageId="21" pageNumber="1325" precision="15" seconds="04" value="15.384444">15°2304″E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF6D839BC24FFE1FF6DE981D0C2FDF9" blockId="21.[144,759,473,587]" pageId="21" pageNumber="1325">
<emphasis id="B93D042BBC24FFE1FF6DE981D029FE5C" box="[144,542,473,495]" italics="true" pageId="21" pageNumber="1325">Preservation and deposition of</emphasis>
<materialsCitation id="3B21D264BC24FFE1FDCAE981D0C6FDF9" country="Italy" county="Leiden" location="Scontrone" municipality="Museum Naturalis" pageId="21" pageNumber="1325" specimenCount="1" typeStatus="holotype">
<emphasis id="B93D042BBC24FFE1FDCAE981D0C1FE5D" box="[567,758,474,495]" italics="true" pageId="21" pageNumber="1325">
<typeStatus id="54F2669BBC24FFE1FDCAE981D052FE5D" box="[567,613,474,495]" pageId="21" pageNumber="1325">type</typeStatus>
specimens:
</emphasis>
<location id="8E968EE2BC24FFE1FF6DE9A3D335FDBF" LSID="urn:lsid:plazi:treatment:03E0692FBC25FFE2FC59ED23D39FF92B:8E968EE2BC24FFE1FF6DE9A3D335FDBF" box="[144,258,504,525]" country="Italy" county="Leiden" municipality="Museum Naturalis" name="Scontrone" pageId="21" pageNumber="1325">Scontrone</location>
specimens:
<location id="8E968EE2BC24FFE1FE5CE9A3D0C0FDBC" LSID="urn:lsid:plazi:treatment:03E0692FBC25FFE2FC59ED23D39FF92B:8E968EE2BC24FFE1FE5CE9A3D0C0FDBC" box="[417,759,504,526]" country="Italy" county="Leiden" municipality="Museum Naturalis" name="Soprintendenza Archeologica" pageId="21" pageNumber="1325">Soprintendenza Archeologica</location>
dellAbruzzo (
<location id="8E968EE2BC24FFE1FECEEA4DD34BFD9E" LSID="urn:lsid:plazi:treatment:03E0692FBC25FFE2FC59ED23D39FF92B:8E968EE2BC24FFE1FECEEA4DD34BFD9E" box="[307,380,534,556]" country="Italy" county="Leiden" municipality="Museum Naturalis" name="Chieti" pageId="21" pageNumber="1325">Chieti</location>
, central
<collectingCountry id="F35E98A9BC24FFE1FE0DEA4DD01EFD9E" box="[496,553,534,556]" name="Italy" pageId="21" pageNumber="1325">Italy</collectingCountry>
);
<location id="8E968EE2BC24FFE1FDB9EA4CD091FD9E" LSID="urn:lsid:plazi:treatment:03E0692FBC25FFE2FC59ED23D39FF92B:8E968EE2BC24FFE1FDB9EA4CD091FD9E" box="[580,678,535,556]" country="Italy" county="Leiden" municipality="Museum Naturalis" name="Gargano" pageId="21" pageNumber="1325">Gargano</location>
specimens:
<collectingMunicipality id="6B924243BC24FFE1FF20EA6ED38DFDF9" box="[221,442,565,587]" pageId="21" pageNumber="1325">Museum Naturalis</collectingMunicipality>
,
<collectingCounty id="6297A0B5BC24FFE1FE35EA6ED021FDF9" box="[456,534,565,587]" pageId="21" pageNumber="1325">Leiden</collectingCounty>
(the Netherlands)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF6D839BC24FFE1FF6DEA29D078FD15" blockId="21.[144,758,626,679]" pageId="21" pageNumber="1325">
<emphasis id="B93D042BBC24FFE1FF6DEA29D33AFD35" box="[144,269,626,647]" italics="true" pageId="21" pageNumber="1325">Etymology:</emphasis>
Refers to the particularly large dimensions of the individuals of the species.
</paragraph>
<paragraph id="8BF6D839BC24FFE1FF6DEA94D33BF9E9" blockId="21.[144,759,718,1627]" pageId="21" pageNumber="1325">
<emphasis id="B93D042BBC24FFE1FF6DEA94D331FD56" box="[144,262,719,740]" italics="true" pageId="21" pageNumber="1325">Diagnosis:</emphasis>
Apomorphies of species: mesodont cheek teeth (
<collectionCode id="ED5840FCBC24FFE1FF26EAB6D2CDFCB0" box="[219,250,749,770]" pageId="21" pageNumber="1325">HI</collectionCode>
= 0.83); presence of upper canines; M1 with very reduced entostyle; paracone supported by robust and prominent median rib; external rib of metacone absent; well-developed and lingually protruding parastyle and mesostyle; metastyle very weak; cementum absent and overall smooth enamel in upper cheek teeth. Mandible with horizontal ramus massive lateromedially, ventral profile moderately convex under cheek teeth corpus; lower cheek dental formula: unknown at the moment in the Scontrone representatives,
<quantity id="4CB175DCBC24FFE1FF0EEC44D374FB87" box="[243,323,1055,1077]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="21" pageNumber="1325" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">23 in</quantity>
Gargano specimens; in p3 and p4 paraconid barely developed or totally absent and protoconid and metaconid of same height; lower molars with widely spaced, triangular, backwardverging labial cuspids and with flattened mesial enamel walls; protoconid and hypoconid isolated in unworn molars; in worn molars protoconid connected mesially with metaconid and distally with entoconid, and hypoconid connected with postprotocristid and postentocristid; lower molars with lingual cuspids relatively less worn than labial cuspids; metastylid and postentocristid prominent, but blunt; ectostylids blunt and robust; third lower molars with tear-shaped hypoconulid and entoconulid not fused distally; labial enamel wall corrugated, lingual wall smooth in lower cheek teeth; protoconids with faint traces of cingulum at their base. Premolar to molar ratio in the lower cheek toothrows: not assessable in the available specimens.
</paragraph>
<paragraph id="8BF6D839BC24FFE1FF6DEED8D014F92A" blockId="21.[144,547,1667,1689]" box="[144,547,1667,1689]" pageId="21" pageNumber="1325">
<emphasis id="B93D042BBC24FFE1FF6DEED8D3B9F92A" box="[144,398,1667,1688]" italics="true" pageId="21" pageNumber="1325">Differential diagnosis:</emphasis>
See
<tableCitation id="C6CBED82BC24FFE1FE37EED8D029F92A" box="[458,542,1667,1689]" captionStart="Table 1" captionStartId="3.[144,208,467,488]" captionText="Table 1. Differential diagnoses of dental characters" pageId="21" pageNumber="1325">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BF6D839BC24FFE1FF6DEE9BD6D5FE5D" blockId="21.[144,759,1728,1903]" lastBlockId="21.[806,1422,198,495]" pageId="21" pageNumber="1325">
<emphasis id="B93D042BBC24FFE1FF6DEE9BD390F967" box="[144,423,1728,1749]" italics="true" pageId="21" pageNumber="1325">
Description of
<typeStatus id="54F2669BBC24FFE1FEBFEE9BD396F967" box="[322,417,1728,1749]" pageId="21" pageNumber="1325" type="holotype">holotype</typeStatus>
:
</emphasis>
Jawbone with pachyostotic horizontal ramus. m1m2. Lingual conids higher than labial conids. Mesial enamel walls flattened. Premetacristid straight and weak. Metastylid and postentocristid prominent lingually, but blunt. Postprotocristid fused to pre-entocristid. Hypoconid isolated. Labial conids fairly spaced, with blunt, robust ectostylid in between growing progressively smaller from m1 to m3. Labial enamel wall moderately corrugated, lingual wall smooth. Faint traces of cingulum at the base of protoconid. Occlusal surfaces somewhat inclined outwards; inner cuspids fairly sharp and pointed, outer cuspids affected by relatively higher degrees of wear. m3 practically a slightly enlarged replica of m2. Hypoconulid and entoconulid maintain separated from one another.
</paragraph>
<paragraph id="8BF6D839BC24FFE1FCDBEA4DD69BFC92" blockId="21.[806,1422,534,1903]" pageId="21" pageNumber="1325">
<emphasis id="B93D042BBC24FFE1FCDBEA4DD715FD9E" box="[806,1314,534,556]" italics="true" pageId="21" pageNumber="1325">
Additional characters shown by
<typeStatus id="54F2669BBC24FFE1FB50EA4CD72BFD9E" box="[1197,1308,535,556]" pageId="21" pageNumber="1325" type="paratype">paratypes</typeStatus>
:
</emphasis>
This, at the moment, is the largest of the species represented in the sample of cranial remains and mandibles from both Scontrone and Gargano. The root of a right upper canine is still preserved on the Gargano skull fragment
<collectionCode id="ED5840FCBC24FFE1FC61EAEBD1EFFD77" box="[924,984,688,709]" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="1325" type="Museum">RGM</collectionCode>
260.951. The Scontrone representatives include a very fragmental maxillary of a young specimen, SCT 18, with P3 and P4 (both erupting), DP
<quantity id="4CB175DCBC24FFE1FCB7EB50D14FFC93" box="[842,888,779,801]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="21" pageNumber="1325" unit="in" value="4.0">4 in</quantity>
substitution, M1 and M2.
</paragraph>
<paragraph id="8BF6D839BC24FFE1FCDBEB71D6D3FC6B" blockId="21.[806,1422,534,1903]" pageId="21" pageNumber="1325">DP4. Molariform, with very robust paracone labial rib; no metacone rib; well-developed parastyle and metastyle; very strong mesostyle. Protocone larger than metacone. Postprotocrista bent forward. As wear progresses, postparacrista and postprotocrista fuse to premetaconule crista. Small entostyle.</paragraph>
<paragraph id="8BF6D839BC24FFE1FCDBEBBAD71DFB1D" blockId="21.[806,1422,534,1903]" pageId="21" pageNumber="1325">P3. Strong styles and external rib of paracone. Protocone fairly small, centrally situated, lingually directed and fused to robust metaconule. Metacone relatively small. Two strong spurs issue from lateral enamel walls of lingual cusp, one from preprotocrista, the other from postmetaconule crista. Lingual enamel wall rougher than labial one. No entostyles.</paragraph>
<paragraph id="8BF6D839BC24FFE1FCDBECE3D1A7FB5E" blockId="21.[806,1422,534,1903]" pageId="21" pageNumber="1325">P4. Tooth preserved only on SCT 18, but still erupting.</paragraph>
<paragraph id="8BF6D839BC24FFE1FCDBECAED675FA16" blockId="21.[806,1422,534,1903]" pageId="21" pageNumber="1325">M1M2. Robust parastyle and even stronger mesostyle. Strong external rib of paracone, metacone with no labial rib. Postprotocrista and postmetaconule crista fairly robust, and lingual crescents closely spaced. Lingual enamel wall more corrugated than labial one. No entostyle.</paragraph>
<paragraph id="8BF6D839BC24FFE1FCDBEDF7D631F946" blockId="21.[806,1422,534,1903]" pageId="21" pageNumber="1325">p3. Tooth elongated mesiodistally, triangular in occlusal view. Parastylid robust, paraconid rudimentary and present only in unworn teeth. Protoconid and metaconid of same height, the latter markedly inclined backwards. Entoconid well developed and inclined like metaconid. Entostylid very weak. Hypoconid prominent labially but lower than entoconid. Labial enamel wall corrugated, lingual wall smooth. p4. Very similar to p3, but much larger and expanded labiolingually. Wear tends to concentrate particularly on talonid of tooth.</paragraph>
<paragraph id="8BF6D839BC24FFE2FCDBEEA6D3D0FE21" blockId="21.[806,1422,534,1903]" lastBlockId="22.[164,779,198,403]" lastPageId="22" lastPageNumber="1326" pageId="21" pageNumber="1325">m1m2. Lingual conids tendantially higher than labial ones, also in worn teeth. Premetacristid tendentially straight and weak, fused, in worn teeth, to lingually bent preprotocristid. In unworn or slightly worn first lower molars preprotocristid in touch with metaconid, and postprotocristid with pre-entocristid. As wear progresses these structures normally fuse, protoconid mesially to metaconid and distally to entoconid. Hypoconid isolated at low levels of wear; as wear increases hypoconid fused to postprotocristid and then to postentocristid.</paragraph>
<paragraph id="8BF6D839BC27FFE2FF59E9E0D3D3FE62" blockId="22.[164,779,443,1689]" box="[164,484,443,465]" pageId="22" pageNumber="1326">
<emphasis id="B93D042BBC27FFE2FF59E9E0D379FE62" box="[164,334,443,464]" italics="true" pageId="22" pageNumber="1326">Measurements:</emphasis>
See
<tableCitation id="C6CBED82BC27FFE2FE77E9E0D3E8FE62" box="[394,479,443,465]" captionStart="Table 2" captionStartId="9.[144,208,445,466]" captionText="Table 2. Odontometrics of the hoplitomerycids" pageId="22" pageNumber="1326">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BF6D839BC27FFE2FF41E982D32AFD35" blockId="22.[164,779,443,1689]" pageId="22" pageNumber="1326">
Specimen incertae sedis
<collectionCode id="ED5840FCBC27FFE2FE21E982D02FFE5C" box="[476,536,473,494]" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="1326" type="Museum">RGM</collectionCode>
261.138: a left maxillary fragment of a very small-sized ruminant from Gargano,
<collectionCode id="ED5840FCBC27FFE2FEE7EA4CD361FD9E" box="[282,342,535,556]" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="1326" type="Museum">RGM</collectionCode>
261.138, still preserves P3 and P4. The dental characteristics of this remain differ from those of any other specimen from both Gargano and Scontrone.
</paragraph>
<paragraph id="8BF6D839BC27FFE2FF41EACAD39BFC6B" blockId="22.[164,779,443,1689]" pageId="22" pageNumber="1326">The most particular feature is the presence of three lingual cusps, aligned mesiodistally but well distinguishable from one another although fused together in a sort of crest-like structure. Moving distalwards, the single cusps can be distinguished by undulations in the enamel wall. Two of the cusps must be the protocone and metaconule. We excluded that the third could be interpreted as a paraconule because this is a primitive cuspule characteristic of very archaic artiodactyls. Therefore, the third cusp has been considered a residual metaconule.</paragraph>
<paragraph id="8BF6D839BC27FFE2FF59EBB9D324FAFA" blockId="22.[164,779,443,1689]" pageId="22" pageNumber="1326">P3. Mesodont tooth. Three robust lingual cusps, aligned mesiodistally, are interpreted, moving distalwards, as the protocone and a double metaconule, fused together in a continuous crest-like structure. The protocone and parastyle are fused together mesially, as opposed to the distal metaconule, which is separated distally from the metastyle by a narrow fissure. The external rib of the paracone is strong, the styles, in contrast, are weak. The lingual and labial enamel walls are both smooth. Traces of cementum are present on the labial wall of the tooth. There is no entostyle.</paragraph>
<paragraph id="8BF6D839BC27FFE2FF59ED0AD39FF92B" blockId="22.[164,779,443,1689]" pageId="22" pageNumber="1326">P4. Mesodont tooth. As in P3, three robust lingual cusps are aligned mesiodistally and fused together in a continuous crest, but in this case the protocone is weaker than the metaconule. The protocone and parastyle are fused together mesially, whereas the metaconule is separated from the metastyle by a narrow fissure distally. The external rib of the paracone and the styles are strong. The lingual and labial enamel walls are both smooth. Traces of cementum are present on the labial wall of the tooth. There is no cingulum or entostyle.</paragraph>
</subSubSection>
</treatment>
</document>