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<document id="30A554AADAA7B52353B632C76A4C94CB" ID-DOI="10.26107/RBZ-2019-0041" ID-ISSN="2345-7600" ID-Zenodo-Dep="4576444" ID-ZooBank="AAA866A1-FBAF-486D-AB84-3AB989A269C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614778122966" checkinUser="carolina" docAuthor="Páll-Gergely, Barna, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Hunyadi, András &amp; Jochum, Adrienne" docDate="2019" docId="03BF447DFF8DFFBAFF77FD0BFF53FF31" docLanguage="en" docName="RafflesBZool.67.517-535.pdf" docOrigin="Raffles Bulletin of Zoology 67" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Tonkinospira triangulata Pall-Gergely &amp; Vermeulen 2019, new species" docType="treatment" docVersion="3" lastPageNumber="534" masterDocId="FF863C05FF9DFFABFFEEFA75FFE6FFC4" masterDocTitle="New Tonkinospira Jochum, Slapnik &amp; Páll-Gergely, 2014 species from Laos and Vietnam (Gastropoda: Pulmonata: Hypselostomatidae)" masterLastPageNumber="535" masterPageNumber="517" pageNumber="533" updateTime="1698878161501" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1A605109B2B13D07AFF2629275EE1D84">New Tonkinospira Jochum, Slapnik &amp; Páll-Gergely, 2014 species from Laos and Vietnam (Gastropoda: Pulmonata: Hypselostomatidae)</mods:title>
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<mods:name id="B6A7CD4AE1EF6CC22971667BE6304281" type="personal">
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<mods:namePart id="E9DAA11AD5365E32D79BF01144F9375B">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="32EBA0B9474E9824BA6840FD4539F505">Grego, Jozef</mods:namePart>
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<mods:namePart id="2AFF0DA2E37B6B6A64889197043E109D">Vermeulen, Jaap J.</mods:namePart>
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<mods:namePart id="EAA3CC3F80B4766180F70836B5C7AF23">Reischütz, Alexander</mods:namePart>
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<mods:namePart id="898FD3E7D680F04BEC2CE3A9FF813AB1">Hunyadi, András</mods:namePart>
</mods:name>
<mods:name id="AF41035B070B31C8C164BCFC05EB731E" type="personal">
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<mods:roleTerm id="840FC45DAB16F60463032C0BB05AB2F4">Author</mods:roleTerm>
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<mods:namePart id="07B56425760D155DE3BA829D29BF3DA7">Jochum, Adrienne</mods:namePart>
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<mods:typeOfResource id="A03BACEC8C23583A373B66FBC05E0A07">text</mods:typeOfResource>
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<mods:title id="834202F3584F03D65303C3FA3535BEE5">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="3B3FB2EC44430950A0D97D25D719FDE8">2019</mods:date>
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<mods:number id="3BB1C40381148A0EBCDC404755B19EF5">2019-11-08</mods:number>
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<mods:number id="D6FB0D1BD9425457A447051762471337">67</mods:number>
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<mods:identifier id="B4C46ABE5157730245DE333C47CF744F" type="ISSN">2345-7600</mods:identifier>
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<treatment id="03BF447DFF8DFFBAFF77FD0BFF53FF31" LSID="urn:lsid:plazi:treatment:03BF447DFF8DFFBAFF77FD0BFF53FF31" httpUri="http://treatment.plazi.org/id/03BF447DFF8DFFBAFF77FD0BFF53FF31" lastPageId="17" lastPageNumber="534" pageId="16" pageNumber="533">
<subSubSection id="C30CA6E0FF8DFFBBFF77FD0BFDEAF872" pageId="16" pageNumber="533" type="nomenclature">
<paragraph id="8BA9F56BFF8DFFBBFF77FD0BFDEAF872" blockId="16.[153,758,1917,2006]" pageId="16" pageNumber="533">
<heading id="D0E14207FF8DFFBBFF77FD0BFDEAF872" bold="true" centered="true" fontSize="10" level="2" pageId="16" pageNumber="533" reason="2">
<emphasis id="B9622979FF8DFFBBFF77FD0BFDEAF872" bold="true" pageId="16" pageNumber="533">
<taxonomicName id="4C168EE8FF8DFFBBFF77FD0BFD17F852" ID-CoL="7TCV3" authority="Pall-Gergely &amp; Vermeulen" authorityName="Pall-Gergely &amp; Vermeulen" authorityYear="2019" box="[153,753,1917,1942]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="533" phylum="Mollusca" rank="species" species="triangulata" status="sp. nov.">
<emphasis id="B9622979FF8DFFBBFF77FD0BFE57F852" bold="true" box="[153,433,1918,1942]" italics="true" pageId="16" pageNumber="533">Tonkinospira triangulata</emphasis>
Páll-Gergely &amp; Vermeulen
</taxonomicName>
,
<taxonomicNameLabel id="A2519402FF8DFFBBFE69FDEBFDEAF872" box="[391,524,1950,1974]" pageId="16" pageNumber="533" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF8DFFBBFE90FDCBFDF7F812" box="[382,529,1982,2006]" pageId="16" pageNumber="533" type="description">
<paragraph id="8BA9F56BFF8DFFBBFE90FDCBFDF7F812" blockId="16.[153,758,1917,2006]" box="[382,529,1982,2006]" pageId="16" pageNumber="533">
(
<figureCitation id="132DE9EEFF8DFFBBFE6BFDCBFE07F812" box="[389,481,1982,2006]" captionStart="Fig" captionStartId="15.[113,148,2091,2113]" captionTargetBox="[131,1430,189,2067]" captionTargetId="figure-10@15.[131,1430,189,2067]" captionTargetPageId="15" captionText="Fig. 12. Tonkinospira triangulata Páll-Gergely &amp; Vermeulen, new species, NHMUK 20190687 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.4576484" httpUri="https://zenodo.org/record/4576484/files/figure.png" pageId="16" pageNumber="533">Figs. 12</figureCitation>
,
<figureCitation id="132DE9EEFF8DFFBBFE00FDCBFDEFF812" box="[494,521,1982,2006]" captionStart="Fig" captionStartId="16.[135,170,689,711]" captionTargetBox="[135,1474,189,674]" captionTargetId="figure-569@16.[135,1474,189,674]" captionTargetPageId="16" captionText="Fig. 13. Variability within Tonkinospira triangulata Páll-Gergely &amp; Vermeulen, new species (JJV 6265)." figureDoi="http://doi.org/10.5281/zenodo.4576486" httpUri="https://zenodo.org/record/4576486/files/figure.png" pageId="16" pageNumber="533">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF8DFFBBFF66FD8BFC52FBD2" pageId="16" pageNumber="533" type="materials_examined">
<paragraph id="8BA9F56BFF8DFFBBFF66FD8BFCE1F7F2" blockId="16.[136,775,2046,2102]" pageId="16" pageNumber="533">
<emphasis id="B9622979FF8DFFBBFF66FD8BFED5F7D2" bold="true" box="[136,307,2046,2070]" pageId="16" pageNumber="533">Type material.</emphasis>
<materialsCitation id="3B7EFF36FF8DFFBBFED2FD8BFCE1F7F2" collectionCode="NHMUK" country="Vietnam" location="Quang Ninh Province" pageId="16" pageNumber="533" specimenCode="NHMUK 20190687" specimenCount="1" stateProvince="Quang Ninh" typeStatus="holotype">
<typeStatus id="54AD4BC9FF8DFFBBFED2FD8BFE79F7D2" box="[316,415,2046,2070]" pageId="16" pageNumber="533" type="holotype">Holotype</typeStatus>
(H:
<quantity id="4CEE588EFF8DFFBBFE3BFD8BFDDFF7D2" box="[469,569,2046,2070]" metricMagnitude="-3" metricUnit="m" metricValue="1.83" pageId="16" pageNumber="533" unit="mm" value="1.83">1.83 mm</quantity>
, D:
<quantity id="4CEE588EFF8DFFBBFD85FD8BFD29F7D2" box="[619,719,2046,2070]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="16" pageNumber="533" unit="mm" value="1.42">1.42 mm</quantity>
,
<figureCitation id="132DE9EEFF8DFFBBFD33FD8BFF45F7F2" captionStart="Fig" captionStartId="15.[113,148,2091,2113]" captionTargetBox="[131,1430,189,2067]" captionTargetId="figure-10@15.[131,1430,189,2067]" captionTargetPageId="15" captionText="Fig. 12. Tonkinospira triangulata Páll-Gergely &amp; Vermeulen, new species, NHMUK 20190687 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.4576484" httpUri="https://zenodo.org/record/4576484/files/figure.png" pageId="16" pageNumber="533">Fig. 12</figureCitation>
) (
<specimenCode id="DBB05D10FF8DFFBBFF54F26BFE7EF7F2" box="[186,408,2078,2102]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="533" type="Museum">NHMUK 20190687</specimenCode>
),
<collectingCountry id="F301B5FBFF8DFFBBFE45F26BFDECF7F2" box="[427,522,2078,2102]" name="Vietnam" pageId="16" pageNumber="533">Vietnam</collectingCountry>
,
<collectingRegion id="49D23B89FF8DFFBBFDFBF26BFCE5F7F2" box="[533,771,2078,2102]" country="Vietnam" name="Quang Ninh" pageId="16" pageNumber="533">Quang Ninh Province</collectingRegion>
,
</materialsCitation>
</paragraph>
<paragraph id="8BA9F56BFF8DFFBBFCADF88BFC52FBD2" blockId="16.[835,1474,766,1046]" pageId="16" pageNumber="533">
<materialsCitation id="3B7EFF36FF8DFFBBFCADF88BFBDBFC72" collectingDate="1998-10-02" collectionCode="JJV" collectorName="Vermeulen, J. J. &amp; Whitten, A. J." county="Song Sot Cave" latitude="20.847218" location="Halong Bay area" longLatPrecision="1" longitude="107.09445" municipality="Dao Bo Hon" pageId="16" pageNumber="533" specimenCode="JJV 6265" specimenCount="2" typeStatus="paratype">
<location id="8EC9A3B0FF8DFFBBFCADF88BFBE9FCD2" LSID="urn:lsid:plazi:treatment:03BF447DFF8DFFBAFF77FD0BFF53FF31:8EC9A3B0FF8DFFBBFCADF88BFBE9FCD2" box="[835,1039,766,791]" county="Song Sot Cave" latitude="20.847218" longLatPrecision="1" longitude="107.09445" municipality="Dao Bo Hon" name="Halong Bay area" pageId="16" pageNumber="533">Halong Bay area</location>
,
<collectingMunicipality id="6BCD6F11FF8DFFBBFBF1F88AFB5FFCD2" box="[1055,1209,767,790]" pageId="16" pageNumber="533">Dao Bo Hon</collectingMunicipality>
,
<collectingCounty id="62C88DE7FF8DFFBBFB24F88BFA99FCD2" box="[1226,1407,766,790]" pageId="16" pageNumber="533">Song Sot Cave</collectingCounty>
, drift material washed together over sinkhole in cave,
<geoCoordinate id="EE2293ACFF8DFFBBFAD8F96BFA58FCF2" box="[1334,1470,797,822]" degrees="20" direction="north" minutes="50.833" orientation="latitude" pageId="16" pageNumber="533" precision="1" value="20.847218">20°50.833N</geoCoordinate>
,
<geoCoordinate id="EE2293ACFF8DFFBBFCADF94BFC37FC92" box="[835,977,829,854]" degrees="107" direction="east" minutes="5.667" orientation="longitude" pageId="16" pageNumber="533" precision="1" value="107.09445">107°5.667E</geoCoordinate>
, coll.
<collectorName id="26E390BDFF8DFFBBFBF0F94BFB29FC92" box="[1054,1231,830,854]" pageId="16" pageNumber="533">Vermeulen, J.J.</collectorName>
&amp;
<collectorName id="26E390BDFF8DFFBBFB15F94BFA70FC93" box="[1275,1430,830,855]" pageId="16" pageNumber="533">Whitten, A.J.</collectorName>
,
<date id="FFA8D3ABFF8DFFBBFA49F94BFC07FCB2" pageId="16" pageNumber="533" value="1998-10-02">
<collectingDate id="EFEC2A43FF8DFFBBFA49F94BFC07FCB2" pageId="16" pageNumber="533" value="1998-10-02">02 October 1998</collectingDate>
</date>
; 3 cleaned
<typeStatus id="54AD4BC9FF8DFFBBFB87F92AFB35FCB3" box="[1129,1235,863,887]" pageId="16" pageNumber="533" type="paratype">paratypes</typeStatus>
(
<figureCitation id="132DE9EEFF8DFFBBFB09F92BFADDFCB2" box="[1255,1339,862,886]" captionStart="Fig" captionStartId="16.[135,170,689,711]" captionTargetBox="[135,1474,189,674]" captionTargetId="figure-569@16.[135,1474,189,674]" captionTargetPageId="16" captionText="Fig. 13. Variability within Tonkinospira triangulata Páll-Gergely &amp; Vermeulen, new species (JJV 6265)." figureDoi="http://doi.org/10.5281/zenodo.4576486" httpUri="https://zenodo.org/record/4576486/files/figure.png" pageId="16" pageNumber="533">Fig. 13</figureCitation>
) + 96 noncleaned
<typeStatus id="54AD4BC9FF8DFFBBFC70F90AFBE2FC53" box="[926,1028,895,919]" pageId="16" pageNumber="533" type="paratype">paratypes</typeStatus>
+ some fragments/juveniles (
<specimenCode id="DBB05D10FF8DFFBBFAA4F90BFA50FC52" box="[1354,1462,894,918]" collectionCode="JJV" pageId="16" pageNumber="533">JJV 6265</specimenCode>
), same data as preceding
</materialsCitation>
;
<materialsCitation id="3B7EFF36FF8DFFBBFBA9F9EBFC36FC12" collectingDate="1998-10-02" collectionCode="JJV" collectorName="Vermeulen, J. J. &amp; Whitten, A. J." county="Song Sot Cave" latitude="20.847218" location="Halong Bay area" longLatPrecision="1" longitude="107.09445" municipality="Dao Bo Hon" pageId="16" pageNumber="533" specimenCode="JJV 6244" specimenCount="84" typeStatus="paratype">
<specimenCount id="9D103EE2FF8DFFBBFBA9F9EBFB29FC73" box="[1095,1231,926,951]" pageId="16" pageNumber="533" type="generic" typeStatus="paratypes">84 paratypes</specimenCount>
(
<specimenCode id="DBB05D10FF8DFFBBFB33F9EBFAA0FC72" box="[1245,1350,926,950]" collectionCode="JJV" pageId="16" pageNumber="533">JJV 6244</specimenCode>
), same data as preceding
</materialsCitation>
;
<materialsCitation id="3B7EFF36FF8DFFBBFC30F9CBFC37FC32" collectingDate="1998-10-02" collectionCode="JJV" collectorName="Vermeulen, J. J. &amp; Whitten, A. J." county="Song Sot Cave" latitude="20.847218" location="Halong Bay area" longLatPrecision="1" longitude="107.09445" municipality="Dao Bo Hon" pageId="16" pageNumber="533" specimenCount="5" typeStatus="paratype">
<specimenCount id="9D103EE2FF8DFFBBFC30F9CBFBBDFC13" box="[990,1115,958,983]" pageId="16" pageNumber="533" type="generic" typeStatus="paratypes">5 paratypes</specimenCount>
(ZRC.MOL.015615), same data as preceding
</materialsCitation>
;
<materialsCitation id="3B7EFF36FF8DFFBBFC30F9ABFC56FBD2" collectingDate="1998-10-02" collectionCode="HNHM" collectorName="Vermeulen, J. J. &amp; Whitten, A. J." county="Song Sot Cave" latitude="20.847218" location="Halong Bay area" longLatPrecision="1" longitude="107.09445" municipality="Dao Bo Hon" pageId="16" pageNumber="533" specimenCode="HNHM 104433" specimenCount="5" typeStatus="paratype">
<specimenCount id="9D103EE2FF8DFFBBFC30F9ABFBBAFC33" box="[990,1116,990,1015]" pageId="16" pageNumber="533" type="generic" typeStatus="paratypes">5 paratypes</specimenCount>
(
<specimenCode id="DBB05D10FF8DFFBBFB83F9ABFAF9FC32" box="[1133,1311,990,1014]" collectionCode="HNHM" country="Hungary" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="16" pageNumber="533">HNHM 104433</specimenCode>
), same data as preceding
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF8DFFBBFCADFE4BFC6FFB72" pageId="16" pageNumber="533" type="diagnosis">
<paragraph id="8BA9F56BFF8DFFBBFCADFE4BFC6FFB72" blockId="16.[835,1474,1086,1206]" pageId="16" pageNumber="533">
<emphasis id="B9622979FF8DFFBBFCADFE4BFC5CFB92" bold="true" box="[835,954,1086,1110]" pageId="16" pageNumber="533">Diagnosis.</emphasis>
A medium sized
<taxonomicName id="4C168EE8FF8DFFBBFB69FE4BFAFEFB92" authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" box="[1159,1304,1086,1110]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="533" phylum="Mollusca" rank="genus">
<emphasis id="B9622979FF8DFFBBFB69FE4BFAFEFB92" box="[1159,1304,1086,1110]" italics="true" pageId="16" pageNumber="533">Tonkinospira</emphasis>
</taxonomicName>
species with a convexly conic shell, rounded body whorl, dense spiral striation, and a peristome that is not adnate to penultimate whorl.
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF8DFFBBFCADFEABFC71F8D2" pageId="16" pageNumber="533" type="description">
<paragraph id="8BA9F56BFF8DFFBBFCADFEABFC0BF972" blockId="16.[834,1475,1246,1718]" pageId="16" pageNumber="533">
<emphasis id="B9622979FF8DFFBBFCADFEABFC29FB32" bold="true" box="[835,975,1246,1270]" pageId="16" pageNumber="533">Description.</emphasis>
Shell medium sized for the genus, convexly conic with pointed apex, greyish-white; protoconch consisting of 11.25 whorls, nearly smooth, at the last quarter whorl and near the suture granular with some elevated, serrated calcareous ridges that are partly arranged in a spiral line; entire shell consisting of 5.56 whorls, suture deep; teleoconch retiform with some faint, spiral threads; spaces between spiral striae variable in width on frontal view of body whorl, but striae are more equidistant on the umbilical side; aperture almost rounded, peristome very slightly expanded, especially in direction of umbilicus, not thickened, sharp; aperture slightly detached from body whorl in ventral view, turning slightly downwards at ca. 1/5 the diameter of the shell; umbilicus narrow and deep, only very slightly covered by reflected lip.
</paragraph>
<paragraph id="8BA9F56BFF8DFFBBFCADFCABFC71F8D2" blockId="16.[835,1474,1758,1814]" pageId="16" pageNumber="533">
<emphasis id="B9622979FF8DFFBBFCADFCABFBB9F932" bold="true" box="[835,1119,1758,1782]" pageId="16" pageNumber="533">Measurements (in mm).</emphasis>
H = 1.692.01, D = 1.421.46 (n = 4).
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF8DFFBBFCADFD4BFBBDF7F2" pageId="16" pageNumber="533" type="diagnosis">
<paragraph id="8BA9F56BFF8DFFBBFCADFD4BFBBDF7F2" blockId="16.[834,1475,1854,2102]" pageId="16" pageNumber="533">
<emphasis id="B9622979FF8DFFBBFCADFD4BFBD0F892" bold="true" box="[835,1078,1854,1878]" pageId="16" pageNumber="533">Differential diagnosis.</emphasis>
<taxonomicName id="4C168EE8FF8DFFBBFBD5FD4BFAA7F892" authority="Páll-Gergely &amp; Grego &amp; Vermeulen &amp; Reischütz &amp; Hunyadi &amp; Jochum, 2019" authorityName="Páll-Gergely &amp; Grego &amp; Vermeulen &amp; Reischütz &amp; Hunyadi &amp; Jochum" authorityYear="2019" box="[1083,1345,1854,1878]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="533" phylum="Mollusca" rank="species" species="triangulata" status="sp. nov.">
<emphasis id="B9622979FF8DFFBBFBD5FD4BFAA7F892" box="[1083,1345,1854,1878]" italics="true" pageId="16" pageNumber="533">Tonkinospira triangulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2519402FF8DFFBBFAA8FD4AFA25F892" box="[1350,1475,1854,1878]" pageId="16" pageNumber="533" rank="species">new species</taxonomicNameLabel>
differs from
<taxonomicName id="4C168EE8FF8DFFBBFC21FD2AFB9DF8B2" authority="Páll-Gergely &amp; Grego &amp; Vermeulen &amp; Reischütz &amp; Hunyadi &amp; Jochum, 2019" authorityName="Páll-Gergely &amp; Grego &amp; Vermeulen &amp; Reischütz &amp; Hunyadi &amp; Jochum" authorityYear="2019" box="[975,1147,1887,1910]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="533" phylum="Mollusca" rank="species" species="crassicostata" status="sp. nov.">
<emphasis id="B9622979FF8DFFBBFC21FD2AFB9DF8B2" box="[975,1147,1887,1910]" italics="true" pageId="16" pageNumber="533">T. crassicostata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2519402FF8DFFBBFB6BFD2AFAE1F8B2" box="[1157,1287,1886,1910]" pageId="16" pageNumber="533" rank="species">new species</taxonomicNameLabel>
by the smoother shell and thinner peristome, from
<taxonomicName id="4C168EE8FF8DFFBBFB26FD0AFA8FF852" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1909" box="[1224,1385,1919,1942]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="533" phylum="Mollusca" rank="species" species="pauperrima">
<emphasis id="B9622979FF8DFFBBFB26FD0AFA8FF852" box="[1224,1385,1919,1942]" italics="true" pageId="16" pageNumber="533">T. pauperrima</emphasis>
</taxonomicName>
by less bulging whorls and less oblique aperture, and
<taxonomicName id="4C168EE8FF8DFFBBFAA9FDEAFA24F872" authorityName="Pall-Gergely &amp; Jochum" authorityYear="2017" box="[1351,1474,1951,1974]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="533" phylum="Mollusca" rank="species" species="tomasini">
<emphasis id="B9622979FF8DFFBBFAA9FDEAFA24F872" box="[1351,1474,1951,1974]" italics="true" pageId="16" pageNumber="533">T. tomasini</emphasis>
</taxonomicName>
by the smaller size and stronger sculpture and the horizontal alignment of the peristome to the body whorl versus the more oblique alignment in
<taxonomicName id="4C168EE8FF8DFFBBFB9DFD8AFB08F7D2" authorityName="Pall-Gergely &amp; Jochum" authorityYear="2017" box="[1139,1262,2047,2070]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="533" phylum="Mollusca" rank="species" species="tomasini">
<emphasis id="B9622979FF8DFFBBFB9DFD8AFB08F7D2" box="[1139,1262,2047,2070]" italics="true" pageId="16" pageNumber="533">T. tomasini</emphasis>
</taxonomicName>
. See also under
<taxonomicName id="4C168EE8FF8DFFBBFA42FD8AFC2EF7F2" authority="Páll-Gergely &amp; Grego &amp; Vermeulen &amp; Reischütz &amp; Hunyadi &amp; Jochum, 2019" authorityName="Páll-Gergely &amp; Grego &amp; Vermeulen &amp; Reischütz &amp; Hunyadi &amp; Jochum" authorityYear="2019" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="533" phylum="Mollusca" rank="species" species="danangensis" status="sp. nov.">
<emphasis id="B9622979FF8DFFBBFA42FD8AFC2EF7F2" italics="true" pageId="16" pageNumber="533">T. danangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2519402FF8DFFBBFC3FF26AFBB0F7F2" box="[977,1110,2078,2102]" pageId="16" pageNumber="533" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF8CFFBAFF9FFAC8FF53FF31" pageId="17" pageNumber="534" type="etymology">
<paragraph id="8BA9F56BFF8CFFBAFF9FFAC8FF53FF31" blockId="17.[113,751,189,245]" pageId="17" pageNumber="534">
<emphasis id="B9622979FF8CFFBAFF9FFAC8FF15FF11" bold="true" box="[113,243,189,213]" pageId="17" pageNumber="534">Etymology.</emphasis>
This species is named after its triangular shell shape.
</paragraph>
</subSubSection>
</treatment>
</document>