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<document id="890C82DC5F146A2C1EE8EBB1D9F7E536" ID="10.11646/zootaxa.4244.3.10SLASH10573" ID-CLB-Dataset="36224" ID-DOI="10.11646/zootaxa.4244.3.10" ID-GBIF-Dataset="5ea8eae5-1706-497f-a745-78852f555e13" ID-ISSN="1175-5326" ID-Zenodo-Dep="432117" ID-ZooBank="A36107A7-7E9F-4995-94A4-E62175719D91" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490088649618" checkinUser="plazi" docAuthor="Hsueh, Pan-Wen &amp; Li, Kuo-Rong" docDate="2017" docId="0386D149FFE79D7BFF27FD5FFDE8C6A3" docLanguage="en" docName="zootaxa.4244.3.10SLASH10573.pdf" docOrigin="Zootaxa 4244 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thelepus hemeiensis Hsueh &amp; Li, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="432" masterDocId="FFBFA931FFE69D78FFB0FFDBFFC8C523" masterDocTitle="Additions of new species to Thelepus (Thelepodidae), with description of a new Terebellides (Trichobranchidae) from Taiwan" masterLastPageNumber="439" masterPageNumber="429" pageNumber="430" updateTime="1698438227721" updateUser="ExternalLinkService">
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<mods:title id="86D4A9DA34FB2504B014E9A13C4149BA">Additions of new species to Thelepus (Thelepodidae), with description of a new Terebellides (Trichobranchidae) from Taiwan</mods:title>
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<mods:namePart id="7F0E49C5AB5F219FE3E5D09EB2DC3EA5">Hsueh, Pan-Wen</mods:namePart>
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<mods:namePart id="106E4180EBD78B61F5F7FF20D797B02C">Li, Kuo-Rong</mods:namePart>
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<mods:date id="A38848ED2FFCBC259319D4BD47D2906F">2017</mods:date>
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<treatment id="0386D149FFE79D7BFF27FD5FFDE8C6A3" ID-DOI="http://doi.org/10.5281/zenodo.6048904" ID-GBIF-Taxon="127848175" ID-Zenodo-Dep="6048904" LSID="urn:lsid:plazi:treatment:0386D149FFE79D7BFF27FD5FFDE8C6A3" httpUri="http://treatment.plazi.org/id/0386D149FFE79D7BFF27FD5FFDE8C6A3" lastPageId="3" lastPageNumber="432" pageId="1" pageNumber="430">
<subSubSection id="C33533D4FFE79D79FF27FD5FFE9FC79C" pageId="1" pageNumber="430" type="nomenclature">
<paragraph id="8B90605FFFE79D79FF27FD5FFE3FC7BD" blockId="1.[151,503,644,703]" box="[151,503,644,670]" pageId="1" pageNumber="430">
<heading id="D0D8D733FFE79D79FF27FD5FFE3FC7BD" bold="true" box="[151,503,644,670]" fontSize="11" level="1" pageId="1" pageNumber="430" reason="1">
<emphasis id="B95BBC4DFFE79D79FF27FD5FFE3FC7BD" bold="true" box="[151,503,644,670]" pageId="1" pageNumber="430">
<taxonomicName id="4C2F1BDCFFE79D79FF27FD5FFE5BC7BD" ID-CoL="7C6HJ" box="[151,403,644,670]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="430" phylum="Annelida" rank="species" species="hemeiensis" status="sp. nov.">
<emphasis id="B95BBC4DFFE79D79FF27FD5FFE5BC7BD" bold="true" box="[151,403,644,670]" italics="true" pageId="1" pageNumber="430">Thelepus hemeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2680136FFE79D79FE2AFD5FFE3FC7BD" box="[410,503,644,670]" pageId="1" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B90605FFFE79D79FF27FD7DFE9FC79C" blockId="1.[151,503,644,703]" box="[151,343,678,703]" pageId="1" pageNumber="430">
<figureCitation id="13147CDAFFE79D79FF27FD7DFF1FC79C" box="[151,215,678,703]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Figs 1</figureCitation>
AJ, 2AF
</paragraph>
</subSubSection>
<subSubSection id="C33533D4FFE79D79FF27FD35FCFCC608" pageId="1" pageNumber="430" type="materials_examined">
<paragraph id="8B90605FFFE79D79FF27FD35FCFCC608" blockId="1.[151,1436,750,1999]" pageId="1" pageNumber="430">
<materialsCitation id="3B476A02FFE79D79FF27FD35FCFCC608" ID-GBIF-Occurrence="1440784453" collectingDate="2016-10-17" collectionCode="NMNS" country="Taiwan" location="Hemei" pageId="1" pageNumber="430" specimenCode="NMNS7743-1" specimenCount="1" stateProvince="New Taipei City" typeStatus="holotype">
<emphasis id="B95BBC4DFFE79D79FF27FD35FE4DC624" bold="true" box="[151,389,750,775]" pageId="1" pageNumber="430">Material examined.</emphasis>
<typeStatus id="5494DEFDFFE79D79FE21FD35FE31C624" box="[401,505,750,775]" pageId="1" pageNumber="430" type="holotype">Holotype</typeStatus>
(
<specimenCode id="DB89C824FFE79D79FDBCFD34FD65C624" box="[524,685,750,775]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="1" pageNumber="430">
<collectionCode id="ED3EF89AFFE79D79FDBCFD34FD96C624" box="[524,606,751,775]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="1" pageNumber="430">NMNS</collectionCode>
7743
<emphasis id="B95BBC4DFFE79D79FD27FD35FD68C624" bold="true" box="[663,672,750,775]" pageId="1" pageNumber="430">-</emphasis>
1
</specimenCode>
),
<location id="8EF03684FFE79D79FD7AFD35FCDDC624" LSID="urn:lsid:plazi:treatment:0386D149FFE79D7BFF27FD5FFDE8C6A3:8EF03684FFE79D79FD7AFD35FCDDC624" box="[714,789,750,775]" country="Taiwan" name="Hemei" pageId="1" pageNumber="430" stateProvince="New Taipei City">Hemei</location>
(25°04´54˝N, 121°54´58˝E),
<collectingRegion id="49EBAEBDFFE79D79FBC0FD34FAFBC624" box="[1136,1331,750,775]" country="Taiwan" name="New Taipei" pageId="1" pageNumber="430">
New
<collectingRegion id="49EBAEBDFFE79D79FB00FD35FAFBC624" box="[1200,1331,750,775]" inferredBySuffix="true" pageId="1" pageNumber="430">Taipei City</collectingRegion>
</collectingRegion>
,
<collectingCountry id="F33820CFFFE79D79FAF3FD35FA50C624" box="[1347,1432,750,775]" name="Taiwan" pageId="1" pageNumber="430">Taiwan</collectingCountry>
, intertidal under rock on fine sand bottom,
<collectingDate id="EFD5BF77FFE79D79FDC0FCC8FCF8C608" box="[624,816,786,811]" pageId="1" pageNumber="430" value="2016-10-17">
<date id="FF91469FFFE79D79FDC0FCC8FCF8C608" box="[624,816,786,811]" pageId="1" pageNumber="430" value="2016-10-17">17 October 2016</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33533D4FFE79D79FF77FCEDFAEEC240" pageId="1" pageNumber="430" type="description">
<paragraph id="8B90605FFFE79D79FF77FCEDFE47C698" blockId="1.[151,1436,750,1999]" pageId="1" pageNumber="430">
<emphasis id="B95BBC4DFFE79D79FF77FCEDFE91C66C" bold="true" box="[199,345,822,847]" pageId="1" pageNumber="430">Description.</emphasis>
<typeStatus id="5494DEFDFFE79D79FED3FCEDFE06C66C" box="[355,462,822,847]" pageId="1" pageNumber="430" type="holotype">Holotype</typeStatus>
, complete, ovigerous female; live specimen body uniformly orange to reddish with conspicuous blood vessels visible through body wall in anterior body, preserved body uniformly beige in alcohol, without distinct patterns of pigmentation (
<figureCitation id="13147CDAFFE79D79FDC8FCA5FD70C6B4" box="[632,696,894,919]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
AB); body length
<quantity id="4CD7CDBAFFE79D79FC2BFCA4FBC9C6B5" box="[923,1025,895,919]" metricMagnitude="-2" metricUnit="m" metricValue="4.42" pageId="1" pageNumber="430" unit="mm" value="44.2">44.2 mm</quantity>
with 84 segments, maximum width
<quantity id="4CD7CDBAFFE79D79FF27FC78FF25C699" box="[151,237,931,955]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="430" unit="mm" value="3.8">3.8 mm</quantity>
on segment 6.
</paragraph>
<paragraph id="8B90605FFFE79D79FF77FC1DFCD5C194" blockId="1.[151,1436,750,1999]" pageId="1" pageNumber="430">
Prostomium at base of upper lip; about twenty-two grooved buccal tentacles, thin, moderately elongate, arising along the distal part of prostomium, with long ones reaching to segments 4144 (
<figureCitation id="13147CDAFFE79D79FB9AFC31FBA0C120" box="[1066,1128,1002,1027]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
CD, H); eyespots present, forming a row at anterior base of prostomium, more concentrated latero-ventrally, becoming scattered at dorsal area of prostomial base (
<figureCitation id="13147CDAFFE79D79FE1EFBE9FE26C168" box="[430,494,1074,1099]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
CE). Peristomium forming lips and continuing dorsally; upper lip large and thick, horseshoe-shaped; lower lip smooth, moderately large and thick (
<figureCitation id="13147CDAFFE79D79FCC6FB8DFC7CC14C" box="[886,948,1110,1135]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
CD); pharyngeal organ exposed. Segment 1 with ventral lobe posterior to lower lip (
<figureCitation id="13147CDAFFE79D79FDC0FBA1FD67C1B0" box="[624,687,1146,1171]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
CD). Ventral surface distinctly glandular, swollen, up to segment 10, more markedly glandular on segment 5 to 7 (
<figureCitation id="13147CDAFFE79D79FD09FB45FD30C194" box="[697,760,1182,1207]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
E).
</paragraph>
<paragraph id="8B90605FFFE79D79FF77FB19FB5BC048" blockId="1.[151,1436,750,1999]" pageId="1" pageNumber="430">
Lateral lobe absent. Three pairs of branchiae, on segment 2 to 4, branchial filaments moderately thick and elongate, mostly not exceeding four body segments in length (
<figureCitation id="13147CDAFFE79D79FCC1FB3DFC7BC1DC" box="[881,947,1254,1279]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
EF); pairs arranged in transversal bands, arising from a glandular area, first pair lateral to the first pair of notopodia, following pairs placed progressively more dorsal, with wide medial gap between each pair, widest gap in the third pair (
<figureCitation id="13147CDAFFE79D79FBF6FAF5FB4EC064" box="[1094,1158,1326,1351]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
EH); pairs of segments 24 with 1112, 5 and 4 simple branchial filaments on each side, respectively (
<figureCitation id="13147CDAFFE79D79FBBCFA89FB83C048" box="[1036,1099,1362,1387]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
EH).
</paragraph>
<paragraph id="8B90605FFFE79D79FF77FAADFF35C360" blockId="1.[151,1436,750,1999]" pageId="1" pageNumber="430">
Notopodia present from segment 3 to 39; first eight pairs similar in length, becoming longer from segment 11 to 21, thereafter shorter gradually to the last thoracic segment (
<figureCitation id="13147CDAFFE79D79FCDAFA41FC63C090" box="[874,939,1434,1459]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
B, E), all notopodia aligned vertically (
<figureCitation id="13147CDAFFE79D79FADEFA41FF69C0F4" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
HI). Trapezoidal notopodia, with glandular forming pre-and postchaetal lobe from segment 3 to 11, notochaetae emerging from between lobes (
<figureCitation id="13147CDAFFE79D79FDB1FA39FD8CC0D8" box="[513,580,1506,1531]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
E, HI), thereafter notopodia becoming less glandular. Notochaetae of all thoracic segments as two rows of winged chaetae, anterior row slightly shorter than posterior row, tapering to tips (
<figureCitation id="13147CDAFFE79D79FF2FF9F1FF28C360" box="[159,224,1578,1603]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
J).
</paragraph>
<paragraph id="8B90605FFFE79D79FF77F995FBABC21C" blockId="1.[151,1436,750,1999]" pageId="1" pageNumber="430">
Neuropodia present from segment 5 to 84, glandular and rectangular; neuropodia gradually wider in width up to segment 11 (
<figureCitation id="13147CDAFFE79D79FEFCF9A9FE47C3A8" box="[332,399,1650,1675]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
E, H), thereafter gradually narrower to posterior thoracic segment; abdominal neuropodia cylindrical progressively smaller to last segment prior to pygidium, as fleshy pinnules slightly raised from the surface of the body (
<figureCitation id="13147CDAFFE79D79FE32F961FE0BC3F0" box="[386,451,1722,1747]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
B); anterior neuropodia with small gap between pairs, wider from mid-body. Uncini in straight row, but slightly curved at both distal ends on anterior to mid-thoracic segments (
<figureCitation id="13147CDAFFE79D79FBC9F905FB71C3D4" box="[1145,1209,1758,1783]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">
Fig.
<date id="FF91469FFFE79D79FB1DF904FB71C3D4" box="[1197,1209,1759,1783]" pageId="1" pageNumber="430">1</date>
</figureCitation>
I, 2A), becoming Cshaped on posterior thoracic segments to the last abdominal segment (
<figureCitation id="13147CDAFFE79D79FC17F8D9FC23C238" box="[935,1003,1794,1819]" captionStart="FIGURE 2" captionStartId="3.[151,250,1927,1949]" captionTargetBox="[151,1435,942,1905]" captionTargetId="figure@3.[151,1436,937,1905]" captionTargetPageId="3" captionText="FIGURE 2. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, neuropodia of thoracic segments, mid-body; B, neuropodia of abdominal segments; C, lateral view of thoracic uncini, segment 8; D, lateral view line drawing of thoracic uncini, segment 8. Scale: A B, 0.5 mm; C D, 0.01 mm." httpUri="https://zenodo.org/record/432128/files/figure.png" pageId="1" pageNumber="430">Fig. 2</figureCitation>
B); dental formula MF:2:1 throughout, uncini with pointed prow and slightly upturned subterminal dorsal button (
<figureCitation id="13147CDAFFE79D79FC6BF8FDFBD5C21C" box="[987,1053,1830,1855]" captionStart="FIGURE 2" captionStartId="3.[151,250,1927,1949]" captionTargetBox="[151,1435,942,1905]" captionTargetId="figure@3.[151,1436,937,1905]" captionTargetPageId="3" captionText="FIGURE 2. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, neuropodia of thoracic segments, mid-body; B, neuropodia of abdominal segments; C, lateral view of thoracic uncini, segment 8; D, lateral view line drawing of thoracic uncini, segment 8. Scale: A B, 0.5 mm; C D, 0.01 mm." httpUri="https://zenodo.org/record/432128/files/figure.png" pageId="1" pageNumber="430">Fig. 2</figureCitation>
CD).
</paragraph>
<paragraph id="8B90605FFFE79D79FF77F891FAEEC240" blockId="1.[151,1436,750,1999]" box="[199,1318,1866,1891]" pageId="1" pageNumber="430">
Genital and nephridial papillae not observed. Pygidium rounded, with crenulated opening (
<figureCitation id="13147CDAFFE79D79FB72F891FAC9C240" box="[1218,1281,1866,1891]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[167,1408,204,1852]" captionTargetId="figure@2.[167,1408,195,1852]" captionTargetPageId="2" captionText="FIGURE 1. Thelepus hemeiensis sp. nov. Holotype (NMNS 7743 - 1): A, life animal; B, complete specimen in alcohol; C D, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; C E, 1.0 mm; F I, 0.5 mm; J, 0.1 mm." httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="1" pageNumber="430">Fig. 1</figureCitation>
B).
</paragraph>
</subSubSection>
<subSubSection id="C33533D4FFE79D79FF77F8B5FAC2C2A4" box="[199,1290,1902,1927]" pageId="1" pageNumber="430" type="etymology">
<paragraph id="8B90605FFFE79D79FF77F8B5FAC2C2A4" blockId="1.[151,1436,750,1999]" box="[199,1290,1902,1927]" pageId="1" pageNumber="430">
<emphasis id="B95BBC4DFFE79D79FF77F8B5FE86C2A4" bold="true" box="[199,334,1902,1927]" pageId="1" pageNumber="430">Etymology.</emphasis>
The name is derived from the name of nearby village where the worm was collected.
</paragraph>
</subSubSection>
<subSubSection id="C33533D4FFE79D79FF77F849FD2BC288" box="[199,739,1938,1963]" pageId="1" pageNumber="430" type="materials_examined">
<paragraph id="8B90605FFFE79D79FF77F849FD2BC288" blockId="1.[151,1436,750,1999]" box="[199,739,1938,1963]" pageId="1" pageNumber="430">
<materialsCitation id="3B476A02FFE79D79FF77F849FD2BC288" ID-GBIF-Occurrence="1440784457" box="[199,739,1938,1963]" country="Taiwan" location="Hemei" pageId="1" pageNumber="430" specimenCount="1" stateProvince="New Taipei City" typeStatus="holotype">
<emphasis id="B95BBC4DFFE79D79FF77F849FEADC288" bold="true" box="[199,357,1938,1963]" pageId="1" pageNumber="430">
<typeStatus id="5494DEFDFFE79D79FF77F849FECAC288" box="[199,258,1938,1963]" pageId="1" pageNumber="430">Type</typeStatus>
locality.
</emphasis>
<location id="8EF03684FFE79D79FEDCF849FE72C288" LSID="urn:lsid:plazi:treatment:0386D149FFE79D7BFF27FD5FFDE8C6A3:8EF03684FFE79D79FEDCF849FE72C288" box="[364,442,1938,1963]" country="Taiwan" name="Hemei" pageId="1" pageNumber="430" stateProvince="New Taipei City">Hemei</location>
,
<collectingRegion id="49EBAEBDFFE79D79FE75F848FDB6C288" box="[453,638,1938,1963]" country="Taiwan" name="New Taipei" pageId="1" pageNumber="430">
New
<collectingRegion id="49EBAEBDFFE79D79FDB0F849FDB6C288" box="[512,638,1938,1963]" inferredBySuffix="true" pageId="1" pageNumber="430">Taipei City</collectingRegion>
</collectingRegion>
,
<collectingCountry id="F33820CFFFE79D79FD39F849FD17C288" box="[649,735,1938,1963]" name="Taiwan" pageId="1" pageNumber="430">Taiwan</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33533D4FFE79D79FF77F86DFD39C2EC" box="[199,753,1974,1999]" pageId="1" pageNumber="430" type="distribution">
<paragraph id="8B90605FFFE79D79FF77F86DFD39C2EC" blockId="1.[151,1436,750,1999]" box="[199,753,1974,1999]" pageId="1" pageNumber="430">
<emphasis id="B95BBC4DFFE79D79FF77F86DFEA8C2EC" bold="true" box="[199,352,1974,1999]" pageId="1" pageNumber="430">Distribution.</emphasis>
Only known from the
<typeStatus id="5494DEFDFFE79D79FDD0F86CFD58C2EC" box="[608,656,1975,1999]" pageId="1" pageNumber="430">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="DF5030D7FFE49D7AFF27F88CFBDAC2CB" httpUri="https://zenodo.org/record/432121/files/figure.png" pageId="2" pageNumber="431" targetBox="[167,1408,204,1852]" targetPageId="2">
<paragraph id="8B90605FFFE49D7AFF27F88CFBDAC2CB" blockId="2.[151,1436,1879,2024]" pageId="2" pageNumber="431">
<emphasis id="B95BBC4DFFE49D7AFF27F88CFEDCC24F" bold="true" box="[151,276,1879,1901]" pageId="2" pageNumber="431">FIGURE 1.</emphasis>
<taxonomicName id="4C2F1BDCFFE49D7AFEACF88CFE26C24E" box="[284,494,1879,1901]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="431" phylum="Annelida" rank="species" species="hemeiensis" status="sp. nov.">
<emphasis id="B95BBC4DFFE49D7AFEACF88CFE26C24E" box="[284,494,1879,1901]" italics="true" pageId="2" pageNumber="431">Thelepus hemeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2680136FFE49D7AFE46F883FD80C24E" box="[502,584,1880,1901]" pageId="2" pageNumber="431" rank="species">
<emphasis id="B95BBC4DFFE49D7AFE46F883FD80C24E" bold="true" box="[502,584,1880,1901]" pageId="2" pageNumber="431">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype (NMNS7743-1): A, life animal; B, complete specimen in alcohol; CD, ventral views of anterior region; E, lateral view of left anterior body; F, latero-dorsal view of anterior body; G, close-up of branchial region, dorsal view; H, dorsal view of anterior body; I, lateral view of right anterior body; J, notochaetae, segment 28. Arrow and numerical numbers indicate eyespots and the number of body segment, respectively. Ga, Glandular area; Ll, Lower lip; Ul, Upper lip. Scale: A, 2.0 mm; B, 5.0 mm; CE, 1.0 mm; FI, 0.5 mm; J, 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C33533D4FFE59D7BFF77FF4CFDE8C6A3" pageId="3" pageNumber="432" type="discussion">
<paragraph id="8B90605FFFE59D7BFF77FF4CFC33C4D7" blockId="3.[151,1436,151,896]" pageId="3" pageNumber="432">
<materialsCitation id="3B476A02FFE59D7BFF77FF4CFC33C4D7" ID-GBIF-Occurrence="1440784456" country="China" county="Southeast Asia" location="Southeast Asia" municipality="Southern Japan" pageId="3" pageNumber="432" specimenCount="1" stateProvince="Taiwan)" typeStatus="holotype">
<emphasis id="B95BBC4DFFE59D7BFF77FF4CFEF3C593" bold="true" box="[199,315,151,176]" pageId="3" pageNumber="432">Remarks.</emphasis>
Of forty-eight valid species described for the genus, seventeen were reported from the East and Southeast Asia. These species are:
<taxonomicName id="4C2F1BDCFFE59D7BFD94FF67FC71C5F7" authority="Caullery, 1944" authorityName="Caullery" authorityYear="1944" box="[548,953,188,213]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="abyssorum">
<emphasis id="B95BBC4DFFE59D7BFD94FF67FCC2C5F6" box="[548,778,188,213]" italics="true" pageId="3" pageNumber="432">Thelepus abyssorum</emphasis>
<bibRefCitation id="EFBE1DAEFFE59D7BFCA3FF67FC71C5F7" author="Caullery" box="[787,953,188,213]" pageId="3" pageNumber="432" refString="Caullery, M. (1944) Polychetes Sedentaires de l'Expedition du Siboga: Ariciidae, Spionidae, Chaetopteridae, Chlorhaemidae, Opheliidae, Oweniidae, Sabellariidae, Sternaspidae, Amphictenidae, Ampharetidae, Terebellidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, XXIV (2 bis), 1 - 204." type="journal article" year="1944">Caullery, 1944</bibRefCitation>
</taxonomicName>
(
<typeStatus id="5494DEFDFFE59D7BFC79FF67FC33C5F7" box="[969,1019,188,212]" pageId="3" pageNumber="432">type</typeStatus>
locality: Southeast
<collectingCountry id="F33820CFFFE59D7BFB6CFF67FAEBC5F6" box="[1244,1315,188,213]" name="China" pageId="3" pageNumber="432">China</collectingCountry>
),
<taxonomicName id="4C2F1BDCFFE59D7BFA87FF67FE07C5DB" authority="Caullery, 1944" authorityName="Caullery" authorityYear="1944" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="angustitoris">
<emphasis id="B95BBC4DFFE59D7BFA87FF67FED5C5DB" italics="true" pageId="3" pageNumber="432">Thelepus angustitoris</emphasis>
<bibRefCitation id="EFBE1DAEFFE59D7BFE97FF04FE07C5DB" author="Caullery" box="[295,463,223,248]" pageId="3" pageNumber="432" refString="Caullery, M. (1944) Polychetes Sedentaires de l'Expedition du Siboga: Ariciidae, Spionidae, Chaetopteridae, Chlorhaemidae, Opheliidae, Oweniidae, Sabellariidae, Sternaspidae, Amphictenidae, Ampharetidae, Terebellidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, XXIV (2 bis), 1 - 204." type="journal article" year="1944">Caullery, 1944</bibRefCitation>
</taxonomicName>
(
<collectingCounty id="62F118D3FFE59D7BFE50FF04FD58C5DB" box="[480,656,223,248]" pageId="3" pageNumber="432">Southeast Asia</collectingCounty>
),
<taxonomicName id="4C2F1BDCFFE59D7BFD17FF04FB1CC5DB" authority="Pillai, 1965" authorityName="Pillai" authorityYear="1965" box="[679,1236,223,248]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="binakayensis">
<emphasis id="B95BBC4DFFE59D7BFD17FF04FC6FC5DB" box="[679,935,223,248]" italics="true" pageId="3" pageNumber="432">Thelepus binakayensis</emphasis>
Pillai, 1965 (
<collectingCountry id="F33820CFFFE59D7BFBF8FF04FB04C5DB" box="[1096,1228,223,248]" name="Philippines" pageId="3" pageNumber="432">Philippines</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C2F1BDCFFE59D7BFB52FF04FDF7C43E" authority="Caullery, 1944 (Malay Archipelago)" authorityName="Caullery, 1944 (Malay Archipelago" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="dubius">
<emphasis id="B95BBC4DFFE59D7BFB52FF04FA54C5DB" box="[1250,1436,223,248]" italics="true" pageId="3" pageNumber="432">Thelepus dubius</emphasis>
<bibRefCitation id="EFBE1DAEFFE59D7BFF27FEDFFE8AC43F" author="Caullery" box="[151,322,260,285]" pageId="3" pageNumber="432" refString="Caullery, M. (1944) Polychetes Sedentaires de l'Expedition du Siboga: Ariciidae, Spionidae, Chaetopteridae, Chlorhaemidae, Opheliidae, Oweniidae, Sabellariidae, Sternaspidae, Amphictenidae, Ampharetidae, Terebellidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, XXIV (2 bis), 1 - 204." type="journal article" year="1944">Caullery, 1944</bibRefCitation>
(Malay Archipelago)
</taxonomicName>
,
<taxonomicName id="4C2F1BDCFFE59D7BFDE0FEDFFBBCC43F" authority="Marenzeller, 1884" authorityName="Marenzeller" authorityYear="1884" box="[592,1140,260,285]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="japonicus">
<emphasis id="B95BBC4DFFE59D7BFDE0FEDFFCF8C43F" box="[592,816,260,285]" italics="true" pageId="3" pageNumber="432">Thelepus japonicus</emphasis>
Marenzeller, 1884 (
<collectingCountry id="F33820CFFFE59D7BFB96FEDFFBA3C43F" box="[1062,1131,260,284]" name="Japan" pageId="3" pageNumber="432">Japan</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C2F1BDCFFE59D7BFB36FEDFFE18C463" authority="(Grube, 1878) (Philippines)" authorityName="Grube" authorityYear="1878" baseAuthorityName="Grube" baseAuthorityYear="1878" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="leptoplocamus">
<emphasis id="B95BBC4DFFE59D7BFB36FEDFFA54C43E" box="[1158,1436,260,285]" italics="true" pageId="3" pageNumber="432">Thelepus leptoplocamus</emphasis>
(Grube, 1878) (
<collectingCountry id="F33820CFFFE59D7BFEF7FEFCFE01C463" box="[327,457,295,320]" name="Philippines" pageId="3" pageNumber="432">Philippines</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C2F1BDCFFE59D7BFE6FFEFCFBE4C463" authority="Hsueh &amp; Li, 2016" authorityName="Hsueh &amp; Li" authorityYear="2016" box="[479,1068,295,320]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="longtongensis">
<emphasis id="B95BBC4DFFE59D7BFE6FFEFCFD21C463" box="[479,745,295,320]" italics="true" pageId="3" pageNumber="432">Thelepus longtongensis</emphasis>
<bibRefCitation id="EFBE1DAEFFE59D7BFD40FEFCFC76C463" author="Hsueh" box="[752,958,295,320]" pageId="3" pageNumber="432" refString="Hsueh, P. - W. &amp; Li, K. - R. (2016) New species of Thelepodidae (Terebelliformia, Polychaeta) from Taiwan. Zootaxa, 4170 (3), 510 - 524." type="journal article" year="2016">Hsueh &amp; Li, 2016</bibRefCitation>
(
<collectingRegion id="49EBAEBDFFE59D7BFC7EFEFCFBE4C463" box="[974,1068,295,320]" country="China" name="Taiwan)" pageId="3" pageNumber="432">Taiwan)</collectingRegion>
</taxonomicName>
,
<taxonomicName id="4C2F1BDCFFE59D7BFB8BFEFCFE95C446" authority="(Caullery, 1944) (Malaysia)" authorityName="Caullery" authorityYear="1944" baseAuthorityName="Caullery" baseAuthorityYear="1944" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="malayensis">
<emphasis id="B95BBC4DFFE59D7BFB8BFEFCFAECC463" box="[1083,1316,295,320]" italics="true" pageId="3" pageNumber="432">Thelepus malayensis</emphasis>
(
<bibRefCitation id="EFBE1DAEFFE59D7BFA84FEFCFF18C447" author="Caullery" pageId="3" pageNumber="432" refString="Caullery, M. (1944) Polychetes Sedentaires de l'Expedition du Siboga: Ariciidae, Spionidae, Chaetopteridae, Chlorhaemidae, Opheliidae, Oweniidae, Sabellariidae, Sternaspidae, Amphictenidae, Ampharetidae, Terebellidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, XXIV (2 bis), 1 - 204." type="journal article" year="1944">Caullery, 1944</bibRefCitation>
) (
<collectingCountry id="F33820CFFFE59D7BFF59FE97FE9DC446" box="[233,341,332,357]" name="Malaysia" pageId="3" pageNumber="432">Malaysia</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C2F1BDCFFE59D7BFEDAFE97FCE8C446" authority="McIntosh, 1885" authorityName="McIntosh" authorityYear="1885" box="[362,800,332,357]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="marenzelleri">
<emphasis id="B95BBC4DFFE59D7BFEDAFE97FDADC446" box="[362,613,332,357]" italics="true" pageId="3" pageNumber="432">Thelepus marenzelleri</emphasis>
McIntosh, 1885
</taxonomicName>
(
<collectingMunicipality id="6BF4FA25FFE59D7BFC81FE97FC2DC447" box="[817,997,332,357]" pageId="3" pageNumber="432">Southern Japan</collectingMunicipality>
),
<taxonomicName id="4C2F1BDCFFE59D7BFC49FE97FF07C4AB" authority="Caullery, 1944" authorityName="Caullery" authorityYear="1944" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="microbranchiatus">
<emphasis id="B95BBC4DFFE59D7BFC49FE97FAE5C446" box="[1017,1325,332,357]" italics="true" pageId="3" pageNumber="432">Thelepus microbranchiatus</emphasis>
<bibRefCitation id="EFBE1DAEFFE59D7BFA86FE97FF07C4AB" author="Caullery" pageId="3" pageNumber="432" refString="Caullery, M. (1944) Polychetes Sedentaires de l'Expedition du Siboga: Ariciidae, Spionidae, Chaetopteridae, Chlorhaemidae, Opheliidae, Oweniidae, Sabellariidae, Sternaspidae, Amphictenidae, Ampharetidae, Terebellidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, XXIV (2 bis), 1 - 204." type="journal article" year="1944">Caullery, 1944</bibRefCitation>
</taxonomicName>
(
<location id="8EF03684FFE59D7BFF58FEB4FE69C4AB" LSID="urn:lsid:plazi:treatment:0386D149FFE79D7BFF27FD5FFDE8C6A3:8EF03684FFE59D7BFF58FEB4FE69C4AB" box="[232,417,367,392]" country="China" county="Southeast Asia" municipality="Southern Japan" name="Southeast Asia" pageId="3" pageNumber="432" stateProvince="Taiwan)">Southeast Asia</location>
),
<taxonomicName id="4C2F1BDCFFE59D7BFE0EFEB4FC95C4AB" authority="Hutchings, 1990" authorityName="Hutchings" authorityYear="1990" box="[446,861,367,392]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="opimus">
<emphasis id="B95BBC4DFFE59D7BFE0EFEB4FD4EC4AB" box="[446,646,367,392]" italics="true" pageId="3" pageNumber="432">Thelepus opimus</emphasis>
Hutchings, 1990
</taxonomicName>
(
<collectingCountry id="F33820CFFFE59D7BFCDCFEABFBC8C4AB" box="[876,1024,368,392]" name="Hong Kong" pageId="3" pageNumber="432">Hong Kong</collectingCountry>
),
<taxonomicName id="4C2F1BDCFFE59D7BFBAEFEB4FEE3C48E" authority="(Grube, 1878) (Philippines)" authorityName="Grube" authorityYear="1878" baseAuthorityName="Grube" baseAuthorityYear="1878" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="parcus">
<emphasis id="B95BBC4DFFE59D7BFBAEFEB4FB28C4AB" box="[1054,1248,367,392]" italics="true" pageId="3" pageNumber="432">Thelepus parcus</emphasis>
(Grube, 1878) (
<collectingCountry id="F33820CFFFE59D7BFF2EFE4FFEEAC48E" box="[158,290,404,429]" name="Philippines" pageId="3" pageNumber="432">Philippines</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C2F1BDCFFE59D7BFE8BFE4FFC55C48E" authority="(Grube, 1878) (Philippines)" authorityName="Grube" authorityYear="1878" baseAuthorityName="Grube" baseAuthorityYear="1878" box="[315,925,404,429]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="paucibranchis">
<emphasis id="B95BBC4DFFE59D7BFE8BFE4FFD84C48E" box="[315,588,404,429]" italics="true" pageId="3" pageNumber="432">Thelepus paucibranchis</emphasis>
(Grube, 1878) (
<collectingCountry id="F33820CFFFE59D7BFCA1FE4FFC5DC48E" box="[785,917,404,429]" name="Philippines" pageId="3" pageNumber="432">Philippines</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C2F1BDCFFE59D7BFC1DFE4FFA80C48F" authority="Hutchings, 1990" authorityName="Hutchings" authorityYear="1990" box="[941,1352,404,429]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="pulvinus">
<emphasis id="B95BBC4DFFE59D7BFC1DFE4FFBB5C48E" box="[941,1149,404,429]" italics="true" pageId="3" pageNumber="432">Thelepus pulvinus</emphasis>
Hutchings, 1990
</taxonomicName>
(
<collectingCountry id="F33820CFFFE59D7BFAECFE4FFF11C4F3" name="Hong Kong" pageId="3" pageNumber="432">Hong Kong</collectingCountry>
),
<taxonomicName id="4C2F1BDCFFE59D7BFF40FE6CFCC5C4F3" authority="(Grube, 1878) (Philippines)" authorityName="Grube" authorityYear="1878" baseAuthorityName="Grube" baseAuthorityYear="1878" box="[240,781,439,464]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="robustus">
<emphasis id="B95BBC4DFFE59D7BFF40FE6CFE08C4F3" box="[240,448,439,464]" italics="true" pageId="3" pageNumber="432">Thelepus robustus</emphasis>
(Grube, 1878) (
<collectingCountry id="F33820CFFFE59D7BFD31FE6CFCCDC4F3" box="[641,773,439,464]" name="Philippines" pageId="3" pageNumber="432">Philippines</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C2F1BDCFFE59D7BFCACFE6CFAACC4F3" authority="Hsueh &amp; Li, 2016" authorityName="Hsueh &amp; Li" authorityYear="2016" box="[796,1380,439,464]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="taiwanensis">
<emphasis id="B95BBC4DFFE59D7BFCACFE6CFBD8C4F3" box="[796,1040,439,464]" italics="true" pageId="3" pageNumber="432">Thelepus taiwanensis</emphasis>
<bibRefCitation id="EFBE1DAEFFE59D7BFBABFE6CFB3AC4F3" author="Hsueh" box="[1051,1266,439,464]" pageId="3" pageNumber="432" refString="Hsueh, P. - W. &amp; Li, K. - R. (2016) New species of Thelepodidae (Terebelliformia, Polychaeta) from Taiwan. Zootaxa, 4170 (3), 510 - 524." type="journal article" year="2016">Hsueh &amp; Li, 2016</bibRefCitation>
(
<collectingRegion id="49EBAEBDFFE59D7BFAB5FE6CFAACC4F3" box="[1285,1380,439,464]" country="China" name="Taiwan)" pageId="3" pageNumber="432">Taiwan)</collectingRegion>
</taxonomicName>
, and
<taxonomicName id="4C2F1BDCFFE59D7BFF27FE07FDB6C4D7" authority="Okuda, 1936" authorityName="Okuda" authorityYear="1936" box="[151,638,476,501]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="toyamaensis">
<emphasis id="B95BBC4DFFE59D7BFF27FE07FE46C4D7" box="[151,398,476,501]" italics="true" pageId="3" pageNumber="432">Thelepus toyamaensis</emphasis>
<bibRefCitation id="EFBE1DAEFFE59D7BFE24FE07FDEDC4D6" author="Okuda" box="[404,549,476,501]" pageId="3" pageNumber="432" refString="Okuda, S. (1936) Polychaetous annelids from Toyama Bay and its adjacent waters. I. Polychaeta Sedentaria. Bulletin of the Biogeographical Society of Japan, 6 (14), 147 - 157." type="journal article" year="1936">Okuda, 1936</bibRefCitation>
(
<collectingCountry id="F33820CFFFE59D7BFD83FE07FDBEC4D7" box="[563,630,476,500]" name="Japan" pageId="3" pageNumber="432">Japan</collectingCountry>
)
</taxonomicName>
(
<bibRefCitation id="EFBE1DAEFFE59D7BFD3DFE07FC22C4D7" author="Hsueh" box="[653,1002,476,501]" pageId="3" pageNumber="432" refString="Hsueh, P. - W. &amp; Li, K. - R. (2016) New species of Thelepodidae (Terebelliformia, Polychaeta) from Taiwan. Zootaxa, 4170 (3), 510 - 524." type="journal article" year="2016">
<location id="8EF03684FFE59D7BFD3DFE07FD1EC4D6" LSID="urn:lsid:plazi:treatment:0386D149FFE79D7BFF27FD5FFDE8C6A3:8EF03684FFE59D7BFD3DFE07FD1EC4D6" box="[653,726,476,501]" country="China" county="Southeast Asia" municipality="Southern Japan" name="Hsueh" pageId="3" pageNumber="432" stateProvince="Taiwan)">Hsueh</location>
&amp;
<location id="8EF03684FFE59D7BFD4AFE07FCDBC4D6" LSID="urn:lsid:plazi:treatment:0386D149FFE79D7BFF27FD5FFDE8C6A3:8EF03684FFE59D7BFD4AFE07FCDBC4D6" box="[762,787,476,501]" country="China" county="Southeast Asia" municipality="Southern Japan" name="Li" pageId="3" pageNumber="432" stateProvince="Taiwan)">Li</location>
2016: 519,
<location id="8EF03684FFE59D7BFC28FE07FC1EC4D6" LSID="urn:lsid:plazi:treatment:0386D149FFE79D7BFF27FD5FFDE8C6A3:8EF03684FFE59D7BFC28FE07FC1EC4D6" box="[920,982,476,501]" country="China" county="Southeast Asia" municipality="Southern Japan" name="Table" pageId="3" pageNumber="432" stateProvince="Taiwan)">Table</location>
1
</bibRefCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B90605FFFE59D7BFF77FDDBFDE8C6A3" blockId="3.[151,1436,151,896]" pageId="3" pageNumber="432">
Among these species, only
<taxonomicName id="4C2F1BDCFFE59D7BFE4AFDDAFDA8C73B" box="[506,608,511,536]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="dubius">
<emphasis id="B95BBC4DFFE59D7BFE4AFDDAFDC0C73B" box="[506,520,513,536]" italics="true" pageId="3" pageNumber="432">T</emphasis>
.
<emphasis id="B95BBC4DFFE59D7BFDA5FE24FDA8C73B" box="[533,608,511,536]" italics="true" pageId="3" pageNumber="432">dubius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2F1BDCFFE59D7BFDDEFDDAFD13C73B" box="[622,731,512,536]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="opimus">
<emphasis id="B95BBC4DFFE59D7BFDDEFDDAFDB3C73B" box="[622,635,513,536]" italics="true" pageId="3" pageNumber="432">T</emphasis>
.
<emphasis id="B95BBC4DFFE59D7BFD3AFDDAFD13C73B" box="[650,731,513,536]" italics="true" pageId="3" pageNumber="432">opimus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2F1BDCFFE59D7BFCA8FDDAFC76C73B" box="[792,958,512,536]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="toyamaensis">
<emphasis id="B95BBC4DFFE59D7BFCA8FDDAFCEDC73B" box="[792,805,513,536]" italics="true" pageId="3" pageNumber="432">T</emphasis>
.
<emphasis id="B95BBC4DFFE59D7BFC84FDDAFC76C73B" box="[820,958,513,536]" italics="true" pageId="3" pageNumber="432">toyamaensis</emphasis>
</taxonomicName>
have similar ranges of branchial filaments and the presence of eyespots (
<bibRefCitation id="EFBE1DAEFFE59D7BFE42FDFFFC95C71F" author="Hsueh" box="[498,861,548,573]" pageId="3" pageNumber="432" refString="Hsueh, P. - W. &amp; Li, K. - R. (2016) New species of Thelepodidae (Terebelliformia, Polychaeta) from Taiwan. Zootaxa, 4170 (3), 510 - 524." type="journal article" year="2016">Hsueh &amp; Li 2016: 519, Table 1</bibRefCitation>
) as in
<taxonomicName id="4C2F1BDCFFE59D7BFC02FDFFFB54C71E" box="[946,1180,548,573]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="hemeiensis" status="sp. nov.">
<emphasis id="B95BBC4DFFE59D7BFC02FDFFFB54C71E" box="[946,1180,548,573]" italics="true" pageId="3" pageNumber="432">Thelepus hemeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2680136FFE59D7BFB15FDFFFAC9C71E" box="[1189,1281,548,573]" pageId="3" pageNumber="432" rank="species">
<emphasis id="B95BBC4DFFE59D7BFB15FDFFFAC9C71E" bold="true" box="[1189,1281,548,573]" pageId="3" pageNumber="432">sp. nov.</emphasis>
</taxonomicNameLabel>
. However,
<taxonomicName id="4C2F1BDCFFE59D7BFA37FDFEFF2AC743" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="dubius">
<emphasis id="B95BBC4DFFE59D7BFA37FDFEFA5CC71F" box="[1415,1428,549,572]" italics="true" pageId="3" pageNumber="432">T</emphasis>
.
<emphasis id="B95BBC4DFFE59D7BFF27FD9CFF2AC743" box="[151,226,583,608]" italics="true" pageId="3" pageNumber="432">dubius</emphasis>
</taxonomicName>
has only 29 pairs of notopodia and 60 abdominal segments (
<bibRefCitation id="EFBE1DAEFFE59D7BFC12FD9CFB74C743" author="Caullery" box="[930,1212,583,608]" pageId="3" pageNumber="432" refString="Caullery, M. (1944) Polychetes Sedentaires de l'Expedition du Siboga: Ariciidae, Spionidae, Chaetopteridae, Chlorhaemidae, Opheliidae, Oweniidae, Sabellariidae, Sternaspidae, Amphictenidae, Ampharetidae, Terebellidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, XXIV (2 bis), 1 - 204." type="journal article" year="1944">Caullery 1944: 170171</bibRefCitation>
), whereas
<taxonomicName id="4C2F1BDCFFE59D7BFA87FD9CFEDAC7A6" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="hemeiensis" status="sp. nov.">
<emphasis id="B95BBC4DFFE59D7BFA87FD9CFEDAC7A6" italics="true" pageId="3" pageNumber="432">Thelepus hemeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2680136FFE59D7BFEAEFDB7FEB5C7A6" box="[286,381,620,645]" pageId="3" pageNumber="432" rank="species">
<emphasis id="B95BBC4DFFE59D7BFEAEFDB7FEB5C7A6" bold="true" box="[286,381,620,645]" pageId="3" pageNumber="432">sp. nov.</emphasis>
</taxonomicNameLabel>
has 37 pairs of notopodia and 80 abdominal segments.
<taxonomicName id="4C2F1BDCFFE59D7BFB91FDB7FB2CC7A7" box="[1057,1252,620,645]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="opimus">
<emphasis id="B95BBC4DFFE59D7BFB91FDB7FB2CC7A7" box="[1057,1252,620,645]" italics="true" pageId="3" pageNumber="432">Thelepus opimus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C2F1BDCFFE59D7BFA37FDB6FEDAC78B" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="hemeiensis" status="sp. nov.">
<emphasis id="B95BBC4DFFE59D7BFA37FDB6FA5CC7A7" box="[1415,1428,621,644]" italics="true" pageId="3" pageNumber="432">T</emphasis>
.
<emphasis id="B95BBC4DFFE59D7BFF27FD54FEDAC78B" box="[151,274,655,680]" italics="true" pageId="3" pageNumber="432">hemeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2680136FFE59D7BFEAAFD54FEBDC78B" box="[282,373,655,680]" pageId="3" pageNumber="432" rank="species">
<emphasis id="B95BBC4DFFE59D7BFEAAFD54FEBDC78B" bold="true" box="[282,373,655,680]" pageId="3" pageNumber="432">sp. nov.</emphasis>
</taxonomicNameLabel>
by having many more pairs of notopodia (&gt;55 vs. 37) and different uncini dental formula on anterior and posterior tori (MF:2:23:Ɗ/MF:23:17:05 vs. MF:2:1 throughout) (
<bibRefCitation id="EFBE1DAEFFE59D7BFB85FD6FFEFDC7D3" author="Hsueh" pageId="3" pageNumber="432" refString="Hsueh, P. - W. &amp; Li, K. - R. (2016) New species of Thelepodidae (Terebelliformia, Polychaeta) from Taiwan. Zootaxa, 4170 (3), 510 - 524." type="journal article" year="2016">Hsueh &amp; Li 2016: 519, Table 1; present study</bibRefCitation>
).
<taxonomicName id="4C2F1BDCFFE59D7BFEFDFD0CFD80C7D3" box="[333,584,727,752]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="toyamaensis">
<emphasis id="B95BBC4DFFE59D7BFEFDFD0CFD80C7D3" box="[333,584,727,752]" italics="true" pageId="3" pageNumber="432">Thelepus toyamaensis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C2F1BDCFFE59D7BFC23FD02FBE7C7D3" box="[915,1071,727,752]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="432" phylum="Annelida" rank="species" species="hemeiensis" status="sp. nov.">
<emphasis id="B95BBC4DFFE59D7BFC23FD02FC69C7D3" box="[915,929,729,752]" italics="true" pageId="3" pageNumber="432">T</emphasis>
.
<emphasis id="B95BBC4DFFE59D7BFC04FD0CFBE7C7D3" box="[948,1071,727,752]" italics="true" pageId="3" pageNumber="432">hemeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2680136FFE59D7BFB8BFD0CFB51C7D3" box="[1083,1177,727,752]" pageId="3" pageNumber="432" rank="species">
<emphasis id="B95BBC4DFFE59D7BFB8BFD0CFB51C7D3" bold="true" box="[1083,1177,727,752]" pageId="3" pageNumber="432">sp. nov.</emphasis>
</taxonomicNameLabel>
by having very short branchial filaments (vs. moderate elongate branchial filaments), dorsal body surface with longitudinal furrows (vs. horizontally corrugated), about 30 segments with glandular ventral surface (vs. about 10 segments with glandular ventral surface), and one row of secondary teeth above main fang (vs. two rows of secondary teeth above main fang) (
<bibRefCitation id="EFBE1DAEFFE59D7BFF50FCBCFDDCC6A3" author="Okuda" box="[224,532,871,896]" pageId="3" pageNumber="432" refString="Okuda, S. (1936) Polychaetous annelids from Toyama Bay and its adjacent waters. I. Polychaeta Sedentaria. Bulletin of the Biogeographical Society of Japan, 6 (14), 147 - 157." type="journal article" year="1936">Okuda 1936; present study</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>