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<document id="A505ADC6FE9F9A3CDA43B47FB37D38CD" ID-CLB-Dataset="46921" ID-DOI="10.5281/zenodo.199982" ID-GBIF-Dataset="558b2fa1-879c-44e4-956d-9abd9c9a249a" ID-ISSN="1175-5326" ID-Zenodo-Dep="199982" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460351134285" checkinUser="plazi" docAuthor="Wu, Min &amp; Gao, Linhui" docDate="2010" docId="03A187F4FFB6FFD79E8007127B05FA68" docLanguage="en" docName="zt02725p027.pdf" docOrigin="Zootaxa 2725" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pupopsis subtorquilla Wu &amp; Gao, 2010, sp. nov." docType="treatment" docVersion="10" lastPageNumber="20" masterDocId="FF98FF8CFFA5FFC49E170677796BFFDC" masterDocTitle="A review of the genus Pupopsis Gredler, 1898 (Gastropoda: Stylommatophora: Enidae), with the descriptions of eight new species from China" masterLastPageNumber="27" masterPageNumber="1" pageNumber="20" updateTime="1698293683227" updateUser="ExternalLinkService">
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<mods:title id="BB9170894E9697C884FFDC2D3325A6DE">A review of the genus Pupopsis Gredler, 1898 (Gastropoda: Stylommatophora: Enidae), with the descriptions of eight new species from China</mods:title>
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<mods:namePart id="40E48C41AE500B3CF5B247D1976B7831">Wu, Min</mods:namePart>
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<mods:namePart id="E768153ED561DEEE5C9A55F8082218EB">Gao, Linhui</mods:namePart>
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<mods:date id="56C4CE9FCBD5BE4DCCFFE034938DE276">2010</mods:date>
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<mods:number id="C96E0E2F843E1FD7DB1E6C15F9A77F6F">2725</mods:number>
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<mods:identifier id="ABFA4608DD922558569A1A822D52F523" type="CLB-Dataset">46921</mods:identifier>
<mods:identifier id="8596B03768306D497E9FEA158063ED60" type="DOI">10.5281/zenodo.199982</mods:identifier>
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<treatment id="03A187F4FFB6FFD79E8007127B05FA68" ID-DOI="http://doi.org/10.5281/zenodo.5625400" ID-GBIF-Taxon="119407881" ID-Zenodo-Dep="5625400" LSID="urn:lsid:plazi:treatment:03A187F4FFB6FFD79E8007127B05FA68" httpUri="http://treatment.plazi.org/id/03A187F4FFB6FFD79E8007127B05FA68" lastPageNumber="20" pageId="19" pageNumber="20">
<subSubSection id="C3126569FFB6FFD79E8007127B46FEA3" box="[151,557,357,383]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8BB736E2FFB6FFD79E8007127B46FEA3" blockId="19.[151,557,357,415]" box="[151,557,357,383]" pageId="19" pageNumber="20">
<heading id="D0FF818EFFB6FFD79E8007127B46FEA3" bold="true" box="[151,557,357,383]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<emphasis id="B97CEAF0FFB6FFD79E8007127B46FEA3" bold="true" box="[151,557,357,383]" pageId="19" pageNumber="20">
11.
<taxonomicName id="4C084D61FFB6FFD79ED6071278A3FEA3" ID-CoL="7SX89" box="[193,456,357,383]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="subtorquilla" status="sp. nov.">
<emphasis id="B97CEAF0FFB6FFD79ED6071278A3FEA3" bold="true" box="[193,456,357,383]" italics="true" pageId="19" pageNumber="20">Pupopsis subtorquilla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F578BFFB6FFD79FD807127B46FEA3" box="[463,557,357,383]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3126569FFB6FFD79E8007F07888FE43" box="[151,483,391,415]" pageId="19" pageNumber="20" type="description">
<paragraph id="8BB736E2FFB6FFD79E8007F07888FE43" blockId="19.[151,557,357,415]" box="[151,483,391,415]" pageId="19" pageNumber="20">
<figureCitation id="13332A67FFB6FFD79E8007F07988FE43" box="[151,227,391,415]" captionStart="FIGURE 1" captionStartId="4.[151,255,1432,1456]" captionTargetBox="[151,1430,346,1400]" captionTargetId="figure@4.[151,1436,303,1411]" captionTargetPageId="4" captionText="FIGURE 1. Distribution map of the genus Pupopsis. Red dot, Pupopsis dissociabilis (Sturany, 1900); blue dot, P. retrodens (Martens, 1879); blue square, P. maoxian sp. nov.; red square, P. rhodostoma sp. nov.; green dot, P. subpupopsis sp. nov.; green square, P. yengiawat sp. nov. Two grey dots each with two or five species as indicated in the boxes on the map. The species P. paraplesius (Sturany, 1900), P. polystreptus (Sturany, 1900) and P. zilchi sp. nov. are not mapped due to no precise locality information available." httpUri="https://zenodo.org/record/199983/files/figure.png" pageId="19" pageNumber="20">Figs. 1</figureCitation>
,
<date id="FFB61022FFB6FFD79EE607F07861FE43" box="[241,266,391,415]" pageId="19" pageNumber="20">
<figureCitation id="13332A67FFB6FFD79EE607F07994FE43" box="[241,255,391,415]" captionStart="FIGURE 3" captionStartId="6.[151,255,1784,1808]" captionTargetBox="[172,1419,194,1772]" captionTargetId="figure@6.[172,1420,194,1774]" captionTargetPageId="6" captionText="FIGURE 3. Apertural view of Pupopsis species. A, Pupopsis pupopsis (Gredler, 1898), SMF- 42076; B, Pupopsis retrodens (Martens, 1879), HBUMM- 06460 specimen 1; C-D, Pupopsis torquilla (Möllendorff, 1901). C, lectotype, SMF- 42082; D, HBUMM- 05584 specimen 1; E, Pupopsis hendan sp. nov., holotype, HBUMM- 05592 specimen 2; F, Pupopsis maoxian sp. nov., holotype, HBUMM- 01023 specimen 1; G, Pupopsis rhodostoma sp. nov., holotype, HBUMM- 05872 specimen 2; H, Pupopsis subpupopsis sp. nov., holotype, HBUMM- 05676 specimen 1; I, Pupopsis subtorquilla sp. nov. A, holotype, HBUMM- 05429 specimen 2; J, Pupopsis yengiawat sp. nov., holotype, HBUMM- 04439 specimen 1; K, Pupopsis yuxu sp. nov., Holotype, HBUMM- 06089 specimen 3; L, Pupopsis zilchi sp. nov., holotype, SMF- 334777." httpUri="https://zenodo.org/record/199985/files/figure.png" pageId="19" pageNumber="20">3</figureCitation>
I
</date>
, 11A11C;
<tableCitation id="C68A0359FFB6FFD79F8707F07888FE43" box="[400,483,391,415]" captionStart="TABLE 2" captionStartId="3.[151,239,617,641]" captionTargetBox="[151,1435,700,1395]" captionTargetPageId="3" captionText="TABLE 2. Pupopsis species and their conchological measurements (data from Sturany, 1900; Möllendorff, 1901; Kobelt, 1902 and this study; *, calculated from the published data; /, data unavailable)." httpUri="http://table.plazi.org/id/DF77666AFFA6FFC79E80041E7D5DFD7C" pageId="19" pageNumber="20" tableUuid="DF77666AFFA6FFC79E80041E7D5DFD7C">Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3126569FFB6FFD79E8007A77B32FCF0" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BB736E2FFB6FFD79E8007A77D14FD38" blockId="19.[151,1437,463,1460]" pageId="19" pageNumber="20">
<emphasis id="B97CEAF0FFB6FFD79E8007A7782CFE34" bold="true" box="[151,327,463,488]" pageId="19" pageNumber="20">
<typeStatus id="54B38840FFB6FFD79E8007A779B9FE34" box="[151,210,464,488]" pageId="19" pageNumber="20">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54B38840FFB6FFD79F5807B878D7FE3B" box="[335,444,463,487]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
: HBUMM-05429specimen 2 (fms), Yuxushan Hill,
<quantity id="4CF09B07FFB6FFD79A0C07B87D0FFE3B" box="[1051,1124,463,487]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="km" value="10.0">10 km</quantity>
north of town of Wenxian, site No. 0 45, 1070 m a.s.l., 32°57ʹ35.5ʹ N, 104°40ʹ41.6ʹ E, Wenxian County, S. Gansu,
<collectingCountry id="F31F7672FFB6FFD79A7707837DCDFDD0" box="[1120,1190,500,524]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
,
<date id="FFB61022FFB6FFD79AA607837C2AFDD0" box="[1201,1345,500,524]" pageId="19" pageNumber="20" value="2006-09-27">2006IX27</date>
, leg. M. Wu, J. Liu, W. Zheng &amp; L. Gao.
<typeStatus id="54B38840FFB6FFD79C1F046F7B12FDEC" box="[520,633,536,560]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
: HBUMM-05429specimens 1, 313, (12 fms), same collection data as
<typeStatus id="54B38840FFB6FFD79E80044B7996FD88" box="[151,253,572,596]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
; HBUMM-05545specimens 13, (3 fms),
<quantity id="4CF09B07FFB6FFD79CF9044B7A5CFD88" box="[750,823,572,596]" metricMagnitude="2" metricUnit="m" metricValue="8.29" pageId="19" pageNumber="20" unit="m" value="829.0">829 m</quantity>
a.s.l., 32°51ʹ44.7ʹ N, 104°50ʹ48.2ʹ E,
<taxonomicName id="4C084D61FFB6FFD79AF3044B7C50FD88" box="[1252,1339,572,596]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="17" phylum="Mollusca" rank="species" species="hendan">Hendan</taxonomicName>
Village, along national highway no. 212, S. Gansu,
<collectingCountry id="F31F7672FFB6FFD79C6004287BD6FDAB" box="[631,701,607,631]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
,
<date id="FFB61022FFB6FFD79CD004287A3CFDAB" box="[711,855,607,631]" pageId="19" pageNumber="20" value="2006-09-29">2006IX29</date>
, leg. M. Wu, J. Liu, W. Zheng &amp; L. Gao; HBUMM- 0 0 349 (1 fms), Bikou,
<quantity id="4CF09B07FFB6FFD79F8B04F3788FFD40" box="[412,484,644,668]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="19" pageNumber="20" unit="m" value="720.0">720 m</quantity>
a.s.l., 32°48ʹ, 105°12ʹE, Wenxian County, Gansu,
<collectingCountry id="F31F7672FFB6FFD79A3404F37D02FD40" box="[1059,1129,644,668]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
,
<date id="FFB61022FFB6FFD79A6104F37C6DFD40" box="[1142,1286,644,668]" pageId="19" pageNumber="20" value="2004-04-18">2004IV18</date>
, leg. M. Wu; HBUMM-00314specimens 14 (4 fms), Wenxian, Shangdanpu,
<quantity id="4CF09B07FFB6FFD79D9004D07AB4FD63" box="[903,991,679,703]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="19" pageNumber="20" unit="m" value="1450.0">1450 m</quantity>
a.s.l., 33°0ʹN, 104°36ʹE,
<date id="FFB61022FFB6FFD79B1004D07CF3FD63" box="[1287,1432,679,703]" pageId="19" pageNumber="20" value="2004-04-22">2004IV22</date>
, leg. M. Wu; HBUMM-00337specimens 13 (3 fms), Wenxian,
<date id="FFB61022FFB6FFD79D7C04BB7A85FD38" box="[875,1006,716,740]" pageId="19" pageNumber="20" value="2004-04-07">2004IV7</date>
, leg. M. Wu.
</paragraph>
<paragraph id="8BB736E2FFB6FFD79EDE04877D1AFCDB" blockId="19.[151,1437,463,1460]" box="[201,1137,751,776]" pageId="19" pageNumber="20">
<emphasis id="B97CEAF0FFB6FFD79EDE0487780EFCD4" bold="true" box="[201,357,751,776]" pageId="19" pageNumber="20">
<typeStatus id="54B38840FFB6FFD79EDE04877869FCD4" box="[201,258,752,776]" pageId="19" pageNumber="20">Type</typeStatus>
locality.
</emphasis>
Yuxushan Hill,
<quantity id="4CF09B07FFB6FFD79C0A04987B0FFCDB" box="[541,612,751,775]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="km" value="10.0">10 km</quantity>
north of Wenxian, Wenxian County, S. Gansu.
</paragraph>
<paragraph id="8BB736E2FFB6FFD79EDE05647B32FCF0" blockId="19.[151,1437,463,1460]" box="[201,601,787,813]" pageId="19" pageNumber="20">
<emphasis id="B97CEAF0FFB6FFD79EDE056478D9FCF0" bold="true" box="[201,434,787,812]" pageId="19" pageNumber="20">Material examined.</emphasis>
<typeStatus id="54B38840FFB6FFD79FAE05627884FCF1" box="[441,495,789,813]" pageId="19" pageNumber="20">Type</typeStatus>
material.
</paragraph>
</subSubSection>
<subSubSection id="C3126569FFB6FFD79EDE05407BEAFC93" box="[201,641,823,848]" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8BB736E2FFB6FFD79EDE05407BEAFC93" blockId="19.[151,1437,463,1460]" box="[201,641,823,848]" pageId="19" pageNumber="20">
<emphasis id="B97CEAF0FFB6FFD79EDE05407809FC8C" bold="true" box="[201,354,823,848]" pageId="19" pageNumber="20">Distribution.</emphasis>
Wenxian County, Gansu.
</paragraph>
</subSubSection>
<subSubSection id="C3126569FFB6FFD79EDE052B7884FC4B" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8BB736E2FFB6FFD79EDE052B7884FC4B" blockId="19.[151,1437,463,1460]" pageId="19" pageNumber="20">
<emphasis id="B97CEAF0FFB6FFD79EDE052B7824FCA8" bold="true" box="[201,335,860,884]" pageId="19" pageNumber="20">Etymology.</emphasis>
The new species is named for its congener,
<taxonomicName id="4C084D61FFB6FFD79D52052A7AA8FCA8" box="[837,963,860,884]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="torquilla">
<emphasis id="B97CEAF0FFB6FFD79D52052A7A38FCA8" box="[837,851,861,884]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="B97CEAF0FFB6FFD79D75052B7AA8FCA8" box="[866,963,860,884]" italics="true" pageId="19" pageNumber="20">torquilla</emphasis>
</taxonomicName>
, to which it is taxonomically closest. The epithet is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3126569FFB6FFD79EDE05D478FDFA9B" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BB736E2FFB6FFD79EDE05D47AB6FC03" blockId="19.[151,1437,463,1460]" pageId="19" pageNumber="20">
<emphasis id="B97CEAF0FFB6FFD79EDE05D47828FC60" bold="true" box="[201,323,931,956]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
Shell with 6.5007.500 whorls. Height 7.08.4, diam. maj. 3.54.5. Parietal tooth present. Columellar tooth prominent and expanded inward. Palatal tooth ridge-like.
</paragraph>
<paragraph id="8BB736E2FFB6FFD79EDE059C78FDFA9B" blockId="19.[151,1437,463,1460]" pageId="19" pageNumber="20">
<emphasis id="B97CEAF0FFB6FFD79EDE059C7860FBD8" bold="true" box="[201,267,1003,1028]" pageId="19" pageNumber="20">Shell.</emphasis>
Bulbous-ovate; apex not acuminate; dextral; thick-shelled; solid; opaque; glossy; with 6.5007.025 7.500 whorls. Whorls never spirally grooved; somewhat convex. Embryonic shell smooth; not polished; with 1.2501.5542.500 whorls. Postnuclear whorls wrinkled. Suture simple. Last whorl gradually ascending toward aperture; rounded at periphery. Shell 7.07.88.4 high; 3.54.0
<quantity id="4CF09B07FFB6FFD79D6E02207AD0FBB3" box="[889,955,1111,1135]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" pageId="19" pageNumber="20" unit="in" value="4.5">4.5 in</quantity>
diam. maj. Height/diam. maj. ratio 1.79 1.972.30. Aperture subovate; almost vertical; its insertions separated; 2.52.93.4 high, 2.32.62.8 wide. Ratio of shell height to aperture height 2.272.713.16. Palatal margin rounded. Peristome expanded, sharp. Palatal tooth ridge-like, extending through nearly the entire body whorl. Parietal callus distinct. Angular tubercle and parietal tooth present. Columellar margin reflexed; with one tooth which is hardly visible in apertural view but extends inward along the columella. Umbilicus a narrow slit. Shell, including upper whorls, uniformly light corneousbrown; aperture white.
</paragraph>
</subSubSection>
<subSubSection id="C3126569FFB6FFD79EDE03237B05FA68" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8BB736E2FFB6FFD79EDE03237B05FA68" blockId="19.[151,1437,463,1460]" pageId="19" pageNumber="20">
<emphasis id="B97CEAF0FFB6FFD79EDE03237856FAB0" bold="true" box="[201,317,1364,1388]" pageId="19" pageNumber="20">Remarks.</emphasis>
The new species is close to
<taxonomicName id="4C084D61FFB6FFD79C9A03227A67FAB0" box="[653,780,1364,1388]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="torquilla">
<emphasis id="B97CEAF0FFB6FFD79C9A03227BF0FAB0" box="[653,667,1365,1388]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="B97CEAF0FFB6FFD79CBC03237A67FAB0" box="[683,780,1364,1388]" italics="true" pageId="19" pageNumber="20">torquilla</emphasis>
</taxonomicName>
due to similar size and the development of the apertural armature. However, the shell of the new species is much more swollen, with much fewer whorls and the columellar tooth is almost invisible in apertural view.
</paragraph>
</subSubSection>
</treatment>
</document>