treatments-xml/data/03/A1/87/03A187CAFFD5943B1FEAFF10965DFEDF.xml

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<mods:title id="D7B6F39D6A31348B54AEFB0B0CA75104">On the taxonomy and zoogeography of some West Palaearctic Quedius species, with a focus on the East Mediterranean and the species allied to Quedius umbrinus and Q. nivicola (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="8D61263F6BA8882188FBE58A0DF1884B">Assing, Volker</mods:namePart>
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<mods:date id="F24597991CA49E2F91AFCD124F1EBFB8">2018</mods:date>
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<paragraph id="8BB736DCFFD594381FEAFF109099FF57" blockId="13.[178,734,216,238]" box="[178,734,216,238]" pageId="13" pageNumber="162">
<heading id="D0FF81B0FFD594381FEAFF109099FF57" box="[178,734,216,238]" fontSize="8" level="2" pageId="13" pageNumber="162" reason="2">
<emphasis id="B97CEACEFFD594381FEAFF10907DFF57" bold="true" box="[178,570,216,238]" pageId="13" pageNumber="162">
<taxonomicName id="4C084D5FFFD594381FEAFF1093B7FF54" ID-CoL="4R37Z" authorityName="Baudi di Selve" authorityYear="1848" box="[178,496,216,238]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis" subGenus="Raphirus">
<emphasis id="B97CEACEFFD594381FEAFF10934EFF54" bold="true" box="[178,265,216,238]" italics="true" pageId="13" pageNumber="162">Quedius</emphasis>
(
<emphasis id="B97CEACEFFD594381E40FF10933EFF54" bold="true" box="[280,377,216,238]" italics="true" pageId="13" pageNumber="162">Raphirus</emphasis>
) 
<emphasis id="B97CEACEFFD594381ED0FF1093B7FF54" bold="true" box="[392,496,216,238]" italics="true" pageId="13" pageNumber="162">nemoralis</emphasis>
</taxonomicName>
<bibRefCitation id="EF994B2DFFD594381EAFFF11907DFF57" author="BAUDI DI SELVE F" box="[503,570,217,238]" pageId="13" pageNumber="162" pagination="113 - 148" refId="ref20032" refString="BAUDI DI SELVE F. (1848): Alcune specie nuove di stafilini. - Studi Entomologici 1 (2): 113 - 148." type="journal article" year="1848">BAUDI</bibRefCitation>
</emphasis>
DI 
<emphasis id="B97CEACEFFD594381D03FF109099FF57" bold="true" box="[603,734,216,238]" pageId="13" pageNumber="162">SELVE, 1848</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3126557FFD594381FEAFECC91EBFD75" pageId="13" pageNumber="162" type="reference_group">
<paragraph id="8BB736DCFFD594381FEAFECC9021FEA3" blockId="13.[178,1086,260,840]" box="[178,614,260,281]" pageId="13" pageNumber="162">
<treatmentCitationGroup id="AB1811F2FFD594381FEAFECC9021FEA3" box="[178,614,260,281]" pageId="13" pageNumber="162">
<treatmentCitation id="0AA910CDFFD594381FEAFECC9025FEA3" author="BAUDI DI SELVE F" box="[178,610,260,281]" page="131" pageId="13" pageNumber="162" year="1848">
<taxonomicName id="4C084D5FFFD594381FEAFECC9025FEA3" ID-CoL="4R37Z" authority="BAUDI DI SELVE, 1848: 131" authorityName="Baudi di Selve" authorityYear="1848" box="[178,610,260,281]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381FEAFECC9319FEA3" box="[178,350,260,281]" italics="true" pageId="13" pageNumber="162">Quedius nemoralis</emphasis>
<bibRefCitation id="EF994B2DFFD594381E3CFECC9025FEA3" author="BAUDI DI SELVE F" box="[356,610,260,281]" pageId="13" pageNumber="162" pagination="113 - 148" refId="ref20032" refString="BAUDI DI SELVE F. (1848): Alcune specie nuove di stafilini. - Studi Entomologici 1 (2): 113 - 148." type="journal article" year="1848">BAUDI DI SELVE, 1848: 131</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFD594381FEAFED690E7FE8E" blockId="13.[178,1086,260,840]" box="[178,672,286,308]" pageId="13" pageNumber="162">
<treatmentCitationGroup id="AB1811F2FFD594381FEAFED690E7FE8E" box="[178,672,286,308]" pageId="13" pageNumber="162">
<emphasis id="B97CEACEFFD594381FEAFED69322FE89" box="[178,357,286,307]" italics="true" pageId="13" pageNumber="162">
<taxonomicName id="4C084D5FFFD594381FEAFED692B9FE89" ID-CoL="8VS2M" box="[178,254,286,307]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
marginalis
</emphasis>
<treatmentCitation id="0AA910CDFFD594381E33FED69066FE8E" author="KRAATZ G" box="[363,545,286,308]" page="512" pageId="13" pageNumber="162" year="1857">
<bibRefCitation id="EF994B2DFFD594381E33FED69066FE8E" author="KRAATZ G" box="[363,545,286,308]" pageId="13" pageNumber="162" pagination="377 - 1080" refId="ref21018" refString="KRAATZ G. (1857): Naturgeschichte der Insecten Deutschlands. Erste Abtheilung Coleoptera. Zweiter Band. Lieferung 3 - 6. - Nicolai, Berlin: 377 - 1080." type="book chapter" year="1857">KRAATZ, 1857: 512</bibRefCitation>
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; preoccupied.
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</paragraph>
<paragraph id="8BB736DCFFD594381FEAFEF090FBFEF7" blockId="13.[178,1086,260,840]" box="[178,700,312,334]" pageId="13" pageNumber="162">
<treatmentCitationGroup id="AB1811F2FFD594381FEAFEF090FBFEF7" box="[178,700,312,334]" pageId="13" pageNumber="162">
<emphasis id="B97CEACEFFD594381FEAFEF0930CFEF4" box="[178,331,312,334]" italics="true" pageId="13" pageNumber="162">
<taxonomicName id="4C084D5FFFD594381FEAFEF092B9FEF7" ID-CoL="8VS2M" box="[178,254,312,333]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
mutatus
</emphasis>
<treatmentCitation id="0AA910CDFFD594381E09FEF09041FEF7" author="KRAATZ G" box="[337,518,312,334]" page="104" pageId="13" pageNumber="162" year="1868">
<bibRefCitation id="EF994B2DFFD594381E09FEF09041FEF7" author="KRAATZ G" box="[337,518,312,334]" pageId="13" pageNumber="162" pagination="104" refId="ref21086" refString="KRAATZ G. (1868): Abanderungen vergebener Namen. - Coleopterologische Hef te 4: 104." type="journal article" year="1868">KRAATZ, 1868: 104</bibRefCitation>
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; replacement name.
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</paragraph>
<paragraph id="8BB736DCFFD594381FEAFE9A902CFEDD" blockId="13.[178,1086,260,840]" box="[178,619,338,360]" pageId="13" pageNumber="162">
<emphasis id="B97CEACEFFD594381FEAFE9A931EFEDD" box="[178,345,338,359]" italics="true" pageId="13" pageNumber="162">
<taxonomicName id="4C084D5FFFD594381FEAFE9A92B9FEDD" ID-CoL="8VS2M" box="[178,254,338,359]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
ovaliceps
</emphasis>
MULSANT &amp; REY, 1876: 744.
</paragraph>
<paragraph id="8BB736DCFFD594381FEAFEA4902DFE3B" blockId="13.[178,1086,260,840]" box="[178,618,364,386]" pageId="13" pageNumber="162">
<treatmentCitationGroup id="AB1811F2FFD594381FEAFEA4902DFE3B" box="[178,618,364,386]" pageId="13" pageNumber="162">
<emphasis id="B97CEACEFFD594381FEAFEA49311FE3B" box="[178,342,364,385]" italics="true" pageId="13" pageNumber="162">
<taxonomicName id="4C084D5FFFD594381FEAFEA492B9FE3B" ID-CoL="8VS2M" box="[178,254,364,385]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
safaensis
</emphasis>
<treatmentCitation id="0AA910CDFFD594381E04FEA593B3FE3B" author="FAGEL G" box="[348,500,364,386]" page="8" pageId="13" pageNumber="162" year="1968">
<bibRefCitation id="EF994B2DFFD594381E04FEA593B3FE3B" author="FAGEL G" box="[348,500,364,386]" pageId="13" pageNumber="162" pagination="1 - 14" refId="ref20541" refString="FAGEL G. (1968 b): Contribution a la connaissance des Staphylinidae CIII. - Le complexe des Quedius oblitteratus ER. - nemoralis BAUDI. - Bulletin Institut Royale des Sciences Naturelles de Belgique 44 (24): 1 - 14." type="journal article" year="1968">FAGEL, 1968b: 8</bibRefCitation>
</treatmentCitation>
ff.; 
<emphasis id="B97CEACEFFD594381D43FEA4902DFE3B" bold="true" box="[539,618,364,385]" pageId="13" pageNumber="162">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFD594381FEAFE4E9085FE26" blockId="13.[178,1086,260,840]" box="[178,706,390,412]" pageId="13" pageNumber="162">
<treatmentCitationGroup id="AB1811F2FFD594381FEAFE4E9085FE26" box="[178,706,390,412]" pageId="13" pageNumber="162">
<emphasis id="B97CEACEFFD594381FEAFE4E93ECFE26" box="[178,427,390,412]" italics="true" pageId="13" pageNumber="162">
<taxonomicName id="4C084D5FFFD594381FEAFE4E92B9FE21" ID-CoL="8VS2M" box="[178,254,390,411]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
safaensis ormanus
</emphasis>
<treatmentCitation id="0AA910CDFFD594381EE9FE4F9013FE26" author="FAGEL G" box="[433,596,391,412]" page="129" pageId="13" pageNumber="162" year="1971">
<bibRefCitation id="EF994B2DFFD594381EE9FE4F9013FE26" author="FAGEL G" box="[433,596,391,412]" pageId="13" pageNumber="162" pagination="119 - 141" refId="ref20674" refString="FAGEL G. (1971): Contribution a la connaissance des Staphylinidae CXIII. - Remarques sur la faune de l'Uludagh, en Anatolie egeenne. - Bulletin et Annales de la Societe Royale d'Entomologie de Belgique 107: 119 - 141." type="journal article" year="1971">
FAGEL, 1971: 
<specimenCount id="9D0EFD55FFD594381D6AFE4F9013FE26" box="[562,596,391,412]" pageId="13" pageNumber="162" type="generic">129</specimenCount>
</bibRefCitation>
</treatmentCitation>
f.; 
<emphasis id="B97CEACEFFD594381D2BFE4F9085FE26" bold="true" box="[627,706,391,412]" pageId="13" pageNumber="162">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFD594381FEAFE6890E7FE0C" blockId="13.[178,1086,260,840]" box="[178,672,416,438]" pageId="13" pageNumber="162">
<treatmentCitationGroup id="AB1811F2FFD594381FEAFE6890E7FE0C" box="[178,672,416,438]" pageId="13" pageNumber="162">
<emphasis id="B97CEACEFFD594381FEAFE6893DEFE0C" box="[178,409,416,438]" italics="true" pageId="13" pageNumber="162">
<taxonomicName id="4C084D5FFFD594381FEAFE689319FE0F" ID-CoL="4R37Z" authorityName="Baudi di Selve" authorityYear="1848" box="[178,350,416,437]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">Quedius nemoralis</taxonomicName>
erinci
</emphasis>
<treatmentCitation id="0AA910CDFFD594381EC6FE699000FE0C" author="KORGE H" box="[414,583,416,438]" page="55" pageId="13" pageNumber="162" year="1971">
<bibRefCitation id="EF994B2DFFD594381EC6FE699000FE0C" author="KORGE H" box="[414,583,416,438]" pageId="13" pageNumber="162" pagination="1 - 68" refId="ref20958" refString="KORGE H. (1971 a): Beitrage zur Kenntnis der Koleopterenfauna Kleinasiens. - Annotationes Zoologicae et Botanicae 67: 1 - 68." type="journal article" year="1971">KORGE, 1971a: 55</bibRefCitation>
</treatmentCitation>
; 
<emphasis id="B97CEACEFFD594381D09FE6990E7FE0C" bold="true" box="[593,672,417,438]" pageId="13" pageNumber="162">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFD594381FEAFE0891EBFD75" blockId="13.[178,1086,260,840]" pageId="13" pageNumber="162">
T y p e m a t e r i a l e x a m i n e d: 
<emphasis id="B97CEACEFFD594381D3CFE08909CFE6F" box="[612,731,448,469]" italics="true" pageId="13" pageNumber="162">Q. safaensis</emphasis>
: 
<typeStatus id="54B3887EFFD594381DB7FE08910CFE6F" box="[751,843,448,469]" pageId="13" pageNumber="162" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D0EFD55FFD594381C04FE0891EFFE6F" box="[860,936,448,469]" pageId="13" pageNumber="162" type="generic">31 exs.</specimenCount>
: &quot; 
<collectingCountry id="F31F764CFFD594381C98FE08964DFE6F" box="[960,1034,448,469]" name="Lebanon" pageId="13" pageNumber="162">LIBAN</collectingCountry>
: Ain Zhalta, Jabal el Jaidi, cédraie, 
<quantity id="4CF09B39FFD594381E9DFE1E904BFE56" box="[453,524,470,492]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="13" pageNumber="162" unit="m" value="1750.0">1750 m</quantity>
, 
<date id="FFB6101CFFD594381D4DFE1E901FFE56" box="[533,600,470,492]" pageId="13" pageNumber="162" value="1966-05">V.1966</date>
G. Fagel / G. Fagel det., 
<emphasis id="B97CEACEFFD594381C67FE1E91D5FE51" box="[831,914,470,491]" italics="true" pageId="13" pageNumber="162">safaensis</emphasis>
nov. sp. / 
<taxonomicName id="4C084D5FFFD594381CA9FE1E93E6FDB9" ID-CoL="4R37Z" authority="Baudi di Selve" authorityName="Baudi di Selve" authorityYear="1848" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381CA9FE1E934AFDB8" italics="true" pageId="13" pageNumber="162">Quedius nemoralis</emphasis>
Baudi di Selve
</taxonomicName>
, det. V. 
<bibRefCitation id="EF994B2DFFD594381EACFE25902DFDB8" author="ASSING V &amp; SCHULKE M &amp; BRACHAT V" box="[500,618,493,515]" pageId="13" pageNumber="162" pagination="1" refId="ref19985" refString="ASSING V., SCHULKE M., BRACHAT V. &amp; H. MEYBOHM (in press): On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). - Contributions to En tomology 68 (1) (2018)." type="journal article" year="2018">Assing 2018</bibRefCitation>
&quot; (IRSNB); 
<specimenCount id="9D0EFD55FFD594381D82FE259167FDB8" box="[730,800,493,515]" pageId="13" pageNumber="162" type="generic">26 exs.</specimenCount>
: &quot; 
<collectingCountry id="F31F764CFFD594381C6AFE25913BFDB8" box="[818,892,493,514]" name="Lebanon" pageId="13" pageNumber="162">LIBAN</collectingCountry>
: Nabeh Safa, humus sous cistes [or &quot;bois marécageux&quot; or &quot;mousses gorges d'eau&quot;], 
<quantity id="4CF09B39FFD594381DA9FDCC917EFDA3" box="[753,825,516,537]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="162" unit="m" value="1000.0">1000 m</quantity>
, 
<date id="FFB6101CFFD594381C1AFDCB91C2FDA3" box="[834,901,515,537]" pageId="13" pageNumber="162" value="1966-05">V.1966</date>
G. Fagel / G. Fagel det., 
<emphasis id="B97CEACEFFD594381FB8FDD29374FD95" box="[224,307,538,559]" italics="true" pageId="13" pageNumber="162">safaensis</emphasis>
n. sp. / 
<taxonomicName id="4C084D5FFFD594381ED9FDD29086FD8A" ID-CoL="4R37Z" authority="Baudi di Selve" authorityName="Baudi di Selve" authorityYear="1848" box="[385,705,538,560]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381ED9FDD29068FD95" box="[385,559,538,559]" italics="true" pageId="13" pageNumber="162">Quedius nemoralis</emphasis>
Baudi di Selve
</taxonomicName>
, det. V. 
<bibRefCitation id="EF994B2DFFD594381C49FDD291C0FD95" author="ASSING V &amp; SCHULKE M &amp; BRACHAT V" box="[785,903,538,560]" pageId="13" pageNumber="162" pagination="1" refId="ref19985" refString="ASSING V., SCHULKE M., BRACHAT V. &amp; H. MEYBOHM (in press): On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). - Contributions to En tomology 68 (1) (2018)." type="journal article" year="2018">Assing 2018</bibRefCitation>
&quot; (IRSNB); 
<specimenCount id="9D0EFD55FFD594381CADFDD2967DFD8A" box="[1013,1082,538,560]" pageId="13" pageNumber="162" type="generic">16 exs.</specimenCount>
: &quot; 
<collectingCountry id="F31F764CFFD594381FE2FDF99344FDFC" box="[186,259,561,582]" name="Lebanon" pageId="13" pageNumber="162">LIBAN</collectingCountry>
, Maasser ech Chouf, cédraie, 
<quantity id="4CF09B39FFD594381D43FDF99023FDFC" box="[539,612,561,583]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="13" pageNumber="162" unit="m" value="1850.0">1850 m</quantity>
, 
<date id="FFB6101CFFD594381D36FDF990F5FDFD" box="[622,690,561,583]" pageId="13" pageNumber="162" value="1966-05">V.1966</date>
G. Fagel / G. Fagel det., 
<emphasis id="B97CEACEFFD594381CFBFDF991B1FDFC" box="[931,1014,561,582]" italics="true" pageId="13" pageNumber="162">safaensis</emphasis>
n. sp. / 
<taxonomicName id="4C084D5FFFD594381FEAFD8F93B0FDE7" ID-CoL="4R37Z" authority="Baudi di Selve" authorityName="Baudi di Selve" authorityYear="1848" box="[178,503,583,605]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381FEAFD8F9327FDE6" box="[178,352,583,604]" italics="true" pageId="13" pageNumber="162">Quedius nemoralis</emphasis>
Baudi di Selve
</taxonomicName>
, det. V. 
<bibRefCitation id="EF994B2DFFD594381D14FD8F9084FDE7" author="ASSING V &amp; SCHULKE M &amp; BRACHAT V" box="[588,707,583,605]" pageId="13" pageNumber="162" pagination="1" refId="ref19985" refString="ASSING V., SCHULKE M., BRACHAT V. &amp; H. MEYBOHM (in press): On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). - Contributions to En tomology 68 (1) (2018)." type="journal article" year="2018">Assing 2018</bibRefCitation>
&quot; (IRSNB); 
<specimenCount id="9D0EFD55FFD594381C6CFD809128FDE7" box="[820,879,584,605]" pageId="13" pageNumber="162" type="generic">2 exs.</specimenCount>
: &quot; 
<collectingCountry id="F31F764CFFD594381CDAFD80918BFDE7" box="[898,972,584,605]" name="Lebanon" pageId="13" pageNumber="162">LIBAN</collectingCountry>
: El Barouk, bois de Lab Houl, 
<quantity id="4CF09B39FFD594381E3AFD9693ECFDCE" box="[354,427,606,628]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="162" unit="m" value="1700.0">1700 m</quantity>
, 
<date id="FFB6101CFFD594381EEEFD9693BEFDCE" box="[438,505,606,628]" pageId="13" pageNumber="162" value="1966-05">V.1966</date>
G. Fagel / G. Fagel det., 
<emphasis id="B97CEACEFFD594381DB5FD969107FDC9" box="[749,832,606,627]" italics="true" pageId="13" pageNumber="162">safaensis</emphasis>
n. sp. / 
<taxonomicName id="4C084D5FFFD594381CD7FD96937AFD31" ID-CoL="4R37Z" authority="Baudi di Selve" authorityName="Baudi di Selve" authorityYear="1848" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381CD7FD96967AFDC9" box="[911,1085,606,627]" italics="true" pageId="13" pageNumber="162">Quedius nemoralis</emphasis>
Baudi di Selve
</taxonomicName>
, det. V. 
<bibRefCitation id="EF994B2DFFD594381ED7FDBD9043FD30" author="ASSING V &amp; SCHULKE M &amp; BRACHAT V" box="[399,516,629,651]" pageId="13" pageNumber="162" pagination="1" refId="ref19985" refString="ASSING V., SCHULKE M., BRACHAT V. &amp; H. MEYBOHM (in press): On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). - Contributions to En tomology 68 (1) (2018)." type="journal article" year="2018">Assing 2018</bibRefCitation>
&quot; (IRSNB); 
<specimenCount id="9D0EFD55FFD594381D2BFDBD90E9FD30" box="[627,686,629,650]" pageId="13" pageNumber="162" type="generic">3 exs.</specimenCount>
: &quot; 
<collectingCountry id="F31F764CFFD594381D98FDBD914DFD30" box="[704,778,629,650]" name="Lebanon" pageId="13" pageNumber="162">LIBAN</collectingCountry>
: Ain Dara, Nahr Jesâyer, 
<quantity id="4CF09B39FFD594381CA4FDBD967DFD30" box="[1020,1082,629,650]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="162" unit="m" value="900.0">900 m</quantity>
, 
<date id="FFB6101CFFD594381FEAFD4392B2FD1B" box="[178,245,651,673]" pageId="13" pageNumber="162" value="1966-05">V.1966</date>
G. Fagel / G. Fagel det., 
<emphasis id="B97CEACEFFD594381E84FD439068FD1A" box="[476,559,651,672]" italics="true" pageId="13" pageNumber="162">safaensis</emphasis>
n. sp. / 
<taxonomicName id="4C084D5FFFD594381D20FD4391F5FD1B" ID-CoL="4R37Z" authority="Baudi di Selve" authorityName="Baudi di Selve" authorityYear="1848" box="[632,946,651,673]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381D20FD439163FD1A" box="[632,804,651,672]" italics="true" pageId="13" pageNumber="162">Quedius nemoralis</emphasis>
Baudi di Selve
</taxonomicName>
, det. V. 
<bibRefCitation id="EF994B2DFFD594381CA6FD4392A6FD0D" author="ASSING V &amp; SCHULKE M &amp; BRACHAT V" pageId="13" pageNumber="162" pagination="1" refId="ref19985" refString="ASSING V., SCHULKE M., BRACHAT V. &amp; H. MEYBOHM (in press): On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). - Contributions to En tomology 68 (1) (2018)." type="journal article" year="2018">Assing 2018</bibRefCitation>
&quot; (IRSNB); 
<specimenCount id="9D0EFD55FFD594381E0FFD6A93E6FD02" box="[343,417,674,696]" pageId="13" pageNumber="162" type="generic">17 exs.</specimenCount>
: &quot; 
<collectingCountry id="F31F764CFFD594381EEEFD6A93B7FD02" box="[438,496,674,696]" name="Lebanon" pageId="13" pageNumber="162">Liban</collectingCountry>
: Kartaba, 
<quantity id="4CF09B39FFD594381D0FFD6A909FFD02" box="[599,728,674,696]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="13" pageNumber="162" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">1200-1400 m</quantity>
, 
<date id="FFB6101CFFD594381DBEFD6A916EFD02" box="[742,809,674,696]" pageId="13" pageNumber="162" value="1964-05">V.1964</date>
- G. Fagel / G. Fagel det., 
<emphasis id="B97CEACEFFD594381FEAFD719342FD74" box="[178,261,697,718]" italics="true" pageId="13" pageNumber="162">safaensis</emphasis>
n. sp. / 
<taxonomicName id="4C084D5FFFD594381E15FD7190C0FD75" ID-CoL="4R37Z" authority="Baudi di Selve" authorityName="Baudi di Selve" authorityYear="1848" box="[333,647,697,719]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381E15FD7193BEFD74" box="[333,505,697,718]" italics="true" pageId="13" pageNumber="162">Quedius nemoralis</emphasis>
Baudi di Selve
</taxonomicName>
, det. V. 
<bibRefCitation id="EF994B2DFFD594381D8AFD719101FD74" author="ASSING V &amp; SCHULKE M &amp; BRACHAT V" box="[722,838,697,719]" pageId="13" pageNumber="162" pagination="1" refId="ref19985" refString="ASSING V., SCHULKE M., BRACHAT V. &amp; H. MEYBOHM (in press): On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). - Contributions to En tomology 68 (1) (2018)." type="journal article" year="2018">Assing 2018</bibRefCitation>
&quot; (IRSNB).
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFD5943B1FEAFD10965DFEDF" lastPageId="14" lastPageNumber="163" pageId="13" pageNumber="162" type="description">
<paragraph id="8BB736DCFFD594381FEAFD109114FCF2" blockId="13.[178,1086,260,840]" pageId="13" pageNumber="162">
<emphasis id="B97CEACEFFD594381FEAFD1093C3FD57" box="[178,388,727,749]" italics="true" pageId="13" pageNumber="162">Q. safaensis ormanus</emphasis>
: 
<materialsCitation id="3B603C81FFD594381ECFFD10914AFCA1" ID-GBIF-Occurrence="2846796434" collectingDate="1970-06" collectionCode="IRSNB" collectorName="G. Fagel &amp; R. I. Sc." determinerName="G. Fagel" elevation="1850" pageId="13" pageNumber="162" specimenCount="32" specimenCount-female="14" specimenCount-male="18" typeStatus="paratype">
<typeStatus id="54B3887EFFD594381ECFFD1093B4FD57" box="[407,499,728,749]" pageId="13" pageNumber="162" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D0EFD55FFD594381D5AFD10907AFD57" box="[514,573,728,749]" pageId="13" pageNumber="162" type="male">18♂♂</specimenCount>
, 
<specimenCount id="9D0EFD55FFD594381D16FD1090C1FD4A" box="[590,646,728,752]" pageId="13" pageNumber="162" type="female">14♀♀</specimenCount>
: &quot;Anatolie occ., Uludagh, 
<quantity id="4CF09B39FFD594381CCEFD109651FD57" box="[918,1046,728,749]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="13" pageNumber="162" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">
<elevation id="0025D1EFFFD594381CCEFD109651FD57" box="[918,1046,728,749]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="13" pageNumber="162" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">1800-1900 m</elevation>
</quantity>
[or 
<quantity id="4CF09B39FFD594381FEAFD2692BDFCB9" box="[178,250,749,771]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="162" unit="m" value="2000.0">
<elevation id="0025D1EFFFD594381FEAFD2692BDFCB9" box="[178,250,749,771]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="162" unit="m" value="2000.0">2000 m</elevation>
</quantity>
], 
<date id="FFB6101CFFD594381E54FD269310FCB9" box="[268,343,750,771]" pageId="13" pageNumber="162" value="1970-06">
<collectingDate id="EFF2E9F4FFD594381E54FD269310FCB9" box="[268,343,750,771]" pageId="13" pageNumber="162" value="1970-06">VI.1970</collectingDate>
</date>
<collectorName id="26FD530AFFD594381E07FD2693F7FCBE" box="[351,432,750,772]" pageId="13" pageNumber="162">G. Fagel</collectorName>
/ 
<determinerName id="7E6BDA18FFD594381E9EFD269050FCBE" box="[454,535,750,772]" pageId="13" pageNumber="162">G. Fagel</determinerName>
det., 
<emphasis id="B97CEACEFFD594381D16FD2690BEFCBE" box="[590,761,750,772]" italics="true" pageId="13" pageNumber="162">safaensis ormanus</emphasis>
nov. / 
<typeStatus id="54B3887EFFD594381C67FD2791C9FCBE" box="[831,910,751,772]" pageId="13" pageNumber="162" type="paratype">Paratype</typeStatus>
/ 
<collectorName id="26FD530AFFD594381CFDFD2791B2FCB9" box="[933,1013,750,772]" pageId="13" pageNumber="162">R. I. Sc.</collectorName>
N. B. / 
<taxonomicName id="4C084D5FFFD594381FEAFCCD93ABFCA1" authority="Baudi di Selve" authorityName="Baudi di Selve" authorityYear="1848" box="[178,492,773,795]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381FEAFCCD9319FCA0" box="[178,350,773,794]" italics="true" pageId="13" pageNumber="162">Quedius nemoralis</emphasis>
Baudi di Selve
</taxonomicName>
, det. V. 
<bibRefCitation id="EF994B2DFFD594381D60FCCD90ECFCA0" author="ASSING V &amp; SCHULKE M &amp; BRACHAT V" box="[568,683,773,795]" pageId="13" pageNumber="162" pagination="1" refId="ref19985" refString="ASSING V., SCHULKE M., BRACHAT V. &amp; H. MEYBOHM (in press): On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). - Contributions to En tomology 68 (1) (2018)." type="journal article" year="2018">Assing 2018</bibRefCitation>
&quot; (
<collectionCode id="ED19AE19FFD594381DE7FCCD9140FCA1" box="[703,775,773,795]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="162">IRSNB</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B603C81FFD594381C4FFCCD9109FCF2" ID-GBIF-Occurrence="2846796345" collectingDate="1967-05" collectionCode="IRSNB" collectorName="G. Fagel &amp; R. I. Sc." determinerName="G. Fagel" elevation="1475" pageId="13" pageNumber="162" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D0EFD55FFD594381C4FFCCD9173FCA0" box="[791,820,773,794]" pageId="13" pageNumber="162" type="male">1♂</specimenCount>
, 
<specimenCount id="9D0EFD55FFD594381C66FCCD911CFCA7" box="[830,859,773,797]" pageId="13" pageNumber="162" type="female">1♀</specimenCount>
: &quot;Anatolie occid., Abant Dagh. 
<quantity id="4CF09B39FFD594381FB7FCD3932BFC8B" box="[239,364,795,817]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.55" metricValueMin="1.4" pageId="13" pageNumber="162" unit="m" value="1475.0" valueMax="1550.0" valueMin="1400.0">
<elevation id="0025D1EFFFD594381FB7FCD3932BFC8B" box="[239,364,795,817]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.55" metricValueMin="1.4" pageId="13" pageNumber="162" unit="m" value="1475.0" valueMax="1550.0" valueMin="1400.0">1400-1550 m</elevation>
</quantity>
, 
<date id="FFB6101CFFD594381E2EFCD393FEFC8B" box="[374,441,795,817]" pageId="13" pageNumber="162" value="1967-05">
<collectingDate id="EFF2E9F4FFD594381E2EFCD393FEFC8B" box="[374,441,795,817]" pageId="13" pageNumber="162" value="1967-05">V.1967</collectingDate>
</date>
<collectorName id="26FD530AFFD594381E98FCD49049FC8B" box="[448,526,795,817]" pageId="13" pageNumber="162">G. Fagel</collectorName>
/ 
<determinerName id="7E6BDA18FFD594381D7AFCD49036FC8B" box="[546,625,795,817]" pageId="13" pageNumber="162">G. Fagel</determinerName>
det., 
<emphasis id="B97CEACEFFD594381DFDFCD3910AFC8B" box="[677,845,795,817]" italics="true" pageId="13" pageNumber="162">safaensis ormanus</emphasis>
nov. / 
<typeStatus id="54B3887EFFD594381CD7FCD49199FC8B" box="[911,990,796,817]" pageId="13" pageNumber="162" type="paratype">Paratype</typeStatus>
/ 
<collectorName id="26FD530AFFD594381CAAFCD49679FC8B" box="[1010,1086,796,817]" pageId="13" pageNumber="162">R. I. Sc.</collectorName>
N. B. / 
<taxonomicName id="4C084D5FFFD594381FACFCFA9069FCF2" authority="Baudi di Selve" authorityName="Baudi di Selve" authorityYear="1848" box="[244,558,818,840]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381FACFCFA93E7FCFD" box="[244,416,818,839]" italics="true" pageId="13" pageNumber="162">Quedius nemoralis</emphasis>
Baudi di Selve
</taxonomicName>
, det. V. 
<bibRefCitation id="EF994B2DFFD594381D21FCFA90AAFCFD" author="ASSING V &amp; SCHULKE M &amp; BRACHAT V" box="[633,749,818,840]" pageId="13" pageNumber="162" pagination="1" refId="ref19985" refString="ASSING V., SCHULKE M., BRACHAT V. &amp; H. MEYBOHM (in press): On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). - Contributions to En tomology 68 (1) (2018)." type="journal article" year="2018">Assing 2018</bibRefCitation>
&quot; (
<collectionCode id="ED19AE19FFD594381C59FCFA910EFCF2" box="[769,841,818,840]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="162">IRSNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB736DCFFD594381FEAFC9E9100FC7E" blockId="13.[178,1085,854,964]" pageId="13" pageNumber="162">
C o m m e n t: The original description of 
<taxonomicName id="4C084D5FFFD594381DD6FC9F9151FCD1" authorityName="Baudi di Selve" authorityYear="1848" box="[654,790,854,876]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381DD6FC9F9151FCD1" box="[654,790,854,876]" italics="true" pageId="13" pageNumber="162">Q. nemoralis</emphasis>
</taxonomicName>
is based on an unspecified number of 
<typeStatus id="54B3887EFFD594381E46FCBC9335FC33" box="[286,370,884,905]" pageId="13" pageNumber="162" type="syntype">syntypes</typeStatus>
from 
<collectingRegion id="49CCF83EFFD594381EF7FCBC904AFC30" box="[431,525,884,906]" country="Italy" name="Piemonte" pageId="13" pageNumber="162">Piemonte</collectingRegion>
, North 
<collectingCountry id="F31F764CFFD594381D0EFCBC90C5FC30" box="[598,642,884,906]" name="Italy" pageId="13" pageNumber="162">Italy</collectingCountry>
(&quot;in provincia Montisferrati (
<collectingRegion id="49CCF83EFFD594381CFAFCBC9668FC30" box="[930,1071,884,906]" country="Italy" name="Piemonte" pageId="13" pageNumber="162">Pedemontium</collectingRegion>
)&quot; (
<bibRefCitation id="EF994B2DFFD594381FE2FC59937CFC1D" author="BAUDI DI SELVE F" box="[186,315,913,935]" pageId="13" pageNumber="162" pagination="113 - 148" refId="ref20032" refString="BAUDI DI SELVE F. (1848): Alcune specie nuove di stafilini. - Studi Entomologici 1 (2): 113 - 148." type="journal article" year="1848">BAUDI 1848</bibRefCitation>
). 
<emphasis id="B97CEACEFFD594381E15FC599051FC1C" box="[333,534,913,934]" italics="true" pageId="13" pageNumber="162">
<taxonomicName id="4C084D5FFFD594381E15FC5993E6FC1C" box="[333,417,913,934]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
marginalis
</emphasis>
, 
<emphasis id="B97CEACEFFD594381D7CFC5990D4FC1C" box="[548,659,913,934]" italics="true" pageId="13" pageNumber="162">Q. mutatus</emphasis>
, and 
<emphasis id="B97CEACEFFD594381D95FC59910AFC1C" box="[717,845,913,934]" italics="true" pageId="13" pageNumber="162">Q. ovaliceps</emphasis>
had been synonymized with 
<taxonomicName id="4C084D5FFFD594381FBDFC67932EFC79" authorityName="Baudi di Selve" authorityYear="1848" box="[229,361,942,964]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381FBDFC67932EFC79" box="[229,361,942,964]" italics="true" pageId="13" pageNumber="162">Q. nemoralis</emphasis>
</taxonomicName>
prior to the present paper (see 
<bibRefCitation id="EF994B2DFFD594381DFCFC67917BFC7E" author="HERMAN L" box="[676,828,942,964]" pageId="13" pageNumber="162" refId="ref20779" refString="HERMAN L. H. (2001): Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Volumes I-VII. - Bulletin of the America n Museum of Natural History 265: 4218 pp." type="book" year="2001">HERMAN 2001</bibRefCitation>
).
</paragraph>
<paragraph id="8BB736DCFFD594381FEAFC1C9036FB9E" blockId="13.[178,1085,980,1060]" pageId="13" pageNumber="162">
<emphasis id="B97CEACEFFD594381FEAFC1C9328FC53" box="[178,367,980,1001]" italics="true" pageId="13" pageNumber="162">
<taxonomicName id="4C084D5FFFD594381FEAFC1C9341FC53" box="[178,262,980,1001]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
safaensis
</emphasis>
was originally described based on a male 
<typeStatus id="54B3887EFFD594381C1DFC1C91D9FC50" box="[837,926,980,1002]" pageId="13" pageNumber="162" type="holotype">holotype</typeStatus>
and numerous 
<typeStatus id="54B3887EFFD594381FEAFC399354FBBC" box="[178,275,1009,1030]" pageId="13" pageNumber="162" type="paratype">paratypes</typeStatus>
from &quot; 
<collectingCountry id="F31F764CFFD594381E3BFC3993E4FBBD" box="[355,419,1009,1031]" name="Lebanon" pageId="13" pageNumber="162">Liban</collectingCountry>
: Nabeh Safa&quot; and on additional 
<typeStatus id="54B3887EFFD594381C5FFC39912FFBBC" box="[775,872,1009,1030]" pageId="13" pageNumber="162" type="paratype">paratypes</typeStatus>
from &quot;Maasser ech Chouf&quot; and from &quot;Kartaba&quot; (
<bibRefCitation id="EF994B2DFFD594381E8FFBC69021FB9E" author="FAGEL G" box="[471,614,1038,1060]" pageId="13" pageNumber="162" pagination="1 - 14" refId="ref20541" refString="FAGEL G. (1968 b): Contribution a la connaissance des Staphylinidae CIII. - Le complexe des Quedius oblitteratus ER. - nemoralis BAUDI. - Bulletin Institut Royale des Sciences Naturelles de Belgique 44 (24): 1 - 14." type="journal article" year="1968">FAGEL 1968b</bibRefCitation>
).
</paragraph>
<paragraph id="8BB736DCFFD594381FEAFBFC91A7FB66" blockId="13.[178,1085,1076,1244]" pageId="13" pageNumber="162">
The original description of 
<emphasis id="B97CEACEFFD594381E82FBFC90F8FBF3" box="[474,703,1076,1097]" italics="true" pageId="13" pageNumber="162">Q. safaensis ormanus</emphasis>
is based on a 
<typeStatus id="54B3887EFFD594381C3EFBFC91F8FBF0" box="[870,959,1076,1098]" pageId="13" pageNumber="162" type="holotype">holotype</typeStatus>
and twelve 
<typeStatus id="54B3887EFFD594381FEAFB999354FBDC" box="[178,275,1105,1126]" pageId="13" pageNumber="162" type="paratype">paratypes</typeStatus>
from &quot;Anatolie occidentale: vilayet de 
<collectingRegion id="49CCF83EFFD594381DFEFB9990A4FBDC" box="[678,739,1105,1126]" country="Turkey" name="Bursa" pageId="13" pageNumber="162">Bursa</collectingRegion>
, Uludagh, 
<quantity id="4CF09B39FFD594381C0AFB99919BFBDC" box="[850,988,1105,1126]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="13" pageNumber="162" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">1800-1900 m</quantity>
&quot; and two 
<typeStatus id="54B3887EFFD594381FEAFBA79354FB3E" box="[178,275,1135,1156]" pageId="13" pageNumber="162" type="paratype">paratypes</typeStatus>
from &quot;vilayet de 
<collectingRegion id="49CCF83EFFD594381E96FBA69045FB3E" box="[462,514,1134,1156]" country="Turkey" name="Bolu" pageId="13" pageNumber="162">Bolu</collectingRegion>
, Abant Dagh, 
<quantity id="4CF09B39FFD594381DC2FBA69161FB39" box="[666,806,1134,1156]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="13" pageNumber="162" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400-1500 m</quantity>
&quot; (
<bibRefCitation id="EF994B2DFFD594381C67FBA69186FB3E" author="FAGEL G" box="[831,961,1134,1156]" pageId="13" pageNumber="162" pagination="119 - 141" refId="ref20674" refString="FAGEL G. (1971): Contribution a la connaissance des Staphylinidae CXIII. - Remarques sur la faune de l'Uludagh, en Anatolie egeenne. - Bulletin et Annales de la Societe Royale d'Entomologie de Belgique 107: 119 - 141." type="journal article" year="1971">FAGEL 1971</bibRefCitation>
). Strangely, as many as 
<specimenCount id="9D0EFD55FFD594381E75FB4493FFFB18" box="[301,440,1164,1186]" pageId="13" pageNumber="162" type="generic">27 specimens</specimenCount>
labelled as 
<typeStatus id="54B3887EFFD594381D6DFB4490D1FB1B" box="[565,662,1164,1185]" pageId="13" pageNumber="162" type="paratype">paratypes</typeStatus>
from the type locality were found in the Fagel collection at the IRSNB. The pr 
<emphasis id="B97CEACEFFD594381D1DFB61900BFB04" box="[581,588,1193,1214]" italics="true" pageId="13" pageNumber="162">i</emphasis>
nciple difference indicated in the description as distinguishing 
<emphasis id="B97CEACEFFD594381E10FB0F9387FB66" box="[328,448,1223,1244]" italics="true" pageId="13" pageNumber="162">Q. ormanus</emphasis>
from 
<emphasis id="B97CEACEFFD594381EA5FB0F903EFB61" box="[509,633,1222,1244]" italics="true" pageId="13" pageNumber="162">Q. safaensis</emphasis>
and 
<taxonomicName id="4C084D5FFFD594381DF2FB0F9169FB61" authorityName="Baudi di Selve" authorityYear="1848" box="[682,814,1222,1244]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381DF2FB0F9169FB61" box="[682,814,1222,1244]" italics="true" pageId="13" pageNumber="162">Q. nemoralis</emphasis>
</taxonomicName>
is the coloration.
</paragraph>
<paragraph id="8BB736DCFFD594381FEAFB249191FA08" blockId="13.[178,1085,1260,1458]" pageId="13" pageNumber="162">
According to the original description of 
<emphasis id="B97CEACEFFD594381D2CFB249100FABB" box="[628,839,1260,1281]" italics="true" pageId="13" pageNumber="162">
<taxonomicName id="4C084D5FFFD594381D2CFB2490B9FABB" authorityName="Baudi di Selve" authorityYear="1848" box="[628,766,1260,1281]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">Q. nemoralis</taxonomicName>
erinci
</emphasis>
, which is based on a 
<typeStatus id="54B3887EFFD594381FEAFAC1934CFAA5" box="[178,267,1289,1311]" pageId="13" pageNumber="162" type="holotype">holotype</typeStatus>
and six 
<typeStatus id="54B3887EFFD594381E32FAC1938CFAA4" box="[362,459,1289,1310]" pageId="13" pageNumber="162" type="paratype">paratypes</typeStatus>
from &quot;Bosburun-Gebirge im Pisidischen Taurus&quot;, this subspecies is distinguished from the nominal subspecies by slight differences in head shape, coloration, and the size of the aedeagus (
<bibRefCitation id="EF994B2DFFD594381D3AFA8C90B1FAE0" author="KORGE H" box="[610,758,1348,1370]" pageId="13" pageNumber="162" pagination="1 - 68" refId="ref20958" refString="KORGE H. (1971 a): Beitrage zur Kenntnis der Koleopterenfauna Kleinasiens. - Annotationes Zoologicae et Botanicae 67: 1 - 68." type="journal article" year="1971">KORGE 1971a</bibRefCitation>
). A study of abundant material from various localities both in South and North 
<collectingCountry id="F31F764CFFD594381DE4FAA99151FACD" box="[700,790,1377,1399]" name="Turkey" pageId="13" pageNumber="162">Anatolia</collectingCountry>
, however, revealed that the Turkish populations are within the range of intraspecific variation of 
<taxonomicName id="4C084D5FFFD594381CF4FAB79671FA29" authorityName="Baudi di Selve" authorityYear="1848" box="[940,1078,1406,1428]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381CF4FAB79671FA29" box="[940,1078,1406,1428]" italics="true" pageId="13" pageNumber="162">Q. nemoralis</emphasis>
</taxonomicName>
. Moreover, a zoogeographically plausible distribution pattern was not observed.
</paragraph>
<paragraph id="8BB736DCFFD5943B1FEAFA09965DFEDF" blockId="13.[178,1085,1473,1583]" lastBlockId="14.[178,1086,219,358]" lastPageId="14" lastPageNumber="163" pageId="13" pageNumber="162">
The known range of 
<taxonomicName id="4C084D5FFFD594381EDFFA099000FA6C" authorityName="Baudi di Selve" authorityYear="1848" box="[391,583,1473,1494]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="nemoralis">
<emphasis id="B97CEACEFFD594381EDFFA099000FA6C" box="[391,583,1473,1494]" italics="true" pageId="13" pageNumber="162">Quedius nemoralis</emphasis>
</taxonomicName>
ranges from the Iberian Peninsula across Central and South Europe eastwards to the Caucasus region, 
<collectingCountry id="F31F764CFFD594381D96FA16915DFA4E" box="[718,794,1502,1524]" name="Turkey" pageId="13" pageNumber="162">Turkey</collectingCountry>
, 
<collectingCountry id="F31F764CFFD594381C7CFA169137FA49" box="[804,880,1502,1523]" name="Cyprus" pageId="13" pageNumber="162">Cyprus</collectingCountry>
, and 
<collectingCountry id="F31F764CFFD594381CFEFA179188FA4E" box="[934,975,1503,1524]" name="Iran" pageId="13" pageNumber="162">Iran</collectingCountry>
(SCHÜLKE &amp; SMETANA 2015). Additional material from numerous localities in 
<collectingCountry id="F31F764CFFD594381CFCFA3491A7F9A8" box="[932,992,1532,1554]" name="Spain" pageId="13" pageNumber="162">Spain</collectingCountry>
, 
<collectingCountry id="F31F764CFFD594381CAAFA34967DF9AB" box="[1010,1082,1532,1553]" name="France" pageId="13" pageNumber="162">France</collectingCountry>
, 
<collectingCountry id="F31F764CFFD594381FEAF9D19354F994" box="[178,275,1561,1582]" name="Germany" pageId="13" pageNumber="162">Germany</collectingCountry>
, 
<collectingCountry id="F31F764CFFD594381E7CF9D19336F995" box="[292,369,1561,1583]" name="Austria" pageId="13" pageNumber="162">Austria</collectingCountry>
, 
<collectingCountry id="F31F764CFFD594381EDAF9D193F4F995" box="[386,435,1561,1583]" name="Italy" pageId="13" pageNumber="162">Italy</collectingCountry>
, 
<collectingCountry id="F31F764CFFD594381E9DF9D19048F995" box="[453,527,1561,1583]" name="Croatia" pageId="13" pageNumber="162">Croatia</collectingCountry>
, 
<collectingCountry id="F31F764CFFD594381D7BF9D19021F995" box="[547,614,1561,1583]" name="Serbia" pageId="13" pageNumber="162">Serbia</collectingCountry>
, 
<collectingCountry id="F31F764CFFD594381D2FF9D190B2F994" box="[631,757,1561,1582]" name="Serbia" pageId="13" pageNumber="162">Montenegro</collectingCountry>
, 
<collectingCountry id="F31F764CFFD594381C5FF9D1911DF995" box="[775,858,1561,1583]" name="Albania" pageId="13" pageNumber="162">Albania</collectingCountry>
, 
<collectingRegion id="49CCF83EFFD594381C32F9D1919AF995" box="[874,989,1561,1583]" country="Greece" name="Dytiki Makedonia" pageId="13" pageNumber="162">Macedonia</collectingRegion>
, 
<collectingCountry id="F31F764CFFD594381CB7F9D1967EF994" box="[1007,1081,1561,1582]" name="Greece" pageId="13" pageNumber="162">Greece</collectingCountry>
, 
<collectingCountry id="F31F764CFFD6943B1FEAFF1392B9FF4B" box="[178,254,219,241]" name="Turkey" pageId="14" pageNumber="163">Turkey</collectingCountry>
, and 
<collectingCountry id="F31F764CFFD6943B1E63FF1393D4FF4B" box="[315,403,219,241]" name="Lebanon" pageId="14" pageNumber="163">Lebanon</collectingCountry>
was examined. Based on personal observations it is one of the most common 
<taxonomicName id="4C084D5FFFD6943B1E11FF3093E1FEB7" box="[329,422,248,269]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="167" phylum="Arthropoda" rank="subGenus" subGenus="Raphirus">
<emphasis id="B97CEACEFFD6943B1E11FF3093E1FEB7" box="[329,422,248,269]" italics="true" pageId="14" pageNumber="163">Raphirus</emphasis>
</taxonomicName>
species in the East Mediterranean, and can be found in leaf litter and debris in various shrub and forest habitats, often in large numbers. It is subject to considerable intraspecific variation of coloration (sometimes even within populations). The variability of the aedeagus is moderate, i.e., not as pronounced as in 
<taxonomicName id="4C084D5FFFD6943B1CCDFE989653FEDF" box="[917,1044,336,357]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="163" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFD6943B1CCDFE989653FEDF" box="[917,1044,336,357]" italics="true" pageId="14" pageNumber="163">Q. umbrinus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>