336 lines
57 KiB
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336 lines
57 KiB
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<mods:title id="FFD2F9472C58A7327839A0330419A7E1">On the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species</mods:title>
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<taxonomicName id="4C084D54087F244CFF06FE19FE4E1256" ID-CoL="8VZ5G" authority="Maklin, 1880" authorityName="Maklin" authorityYear="1880" box="[151,504,510,537]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CEAC5087F244CFF06FE19FEFA1257" bold="true" box="[151,332,510,537]" italics="true" pageId="2" pageNumber="303">Mannerheimia</emphasis>
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Mäklin, 1880
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</emphasis>
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<paragraph id="8BB736D7087F244CFF06FDC4FED01273" blockId="2.[151,504,510,573]" box="[151,358,547,573]" pageId="2" pageNumber="303">
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(
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<figureCitation id="13332A52087F244CFF0EFDC4FEBD1273" box="[159,267,547,573]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1955,1979]" captionTargetBox="[187,1400,190,1920]" captionTargetId="figure-16@3.[173,1414,181,1931]" captionTargetPageId="3" captionText="FIGURES 1–9. Habitus of Mannerheimia: 1—M. afghanica (paratype), 2—M. arctica, 3—M. asiatica, 4—M. brevipennis, 5—M. japonica, 6—M. obscura, 7—M. pleshanovi (paratype), 8—M. yasudai, 9—M. zanettii (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5531374" httpUri="https://zenodo.org/record/5531374/files/figure.png" pageId="2" pageNumber="303">Figs. 1–9</figureCitation>
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,
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<figureCitation id="13332A52087F244CFE86FDC4FEE81273" box="[279,350,547,573]" captionStart-0="FIGURES 12–15" captionStart-1="FIGURES 16–25" captionStart-2="FIGURES 26–38" captionStartId-0="10.[151,264,1952,1976]" captionStartId-1="11.[151,264,1047,1071]" captionStartId-2="12.[151,264,1858,1882]" captionTargetBox-0="[167,1423,720,1921]" captionTargetBox-1="[169,1421,187,1017]" captionTargetBox-2="[172,1413,320,1826]" captionTargetId-0="figure-289@10.[151,1436,705,1929]" captionTargetId-1="figure-16@11.[163,1425,181,1023]" captionTargetId-2="figure-79@12.[160,1428,309,1835]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURES 12–15. Mouthparts of Mannerheimia arctica: 12—labrum (dorsal view), 13—mandibles (left and right, dorsal view), 14—maxilla (left, dorsal view), 15—labium and mentum (ventral view). Scale bar: 0.1 mm." captionText-1="FIGURES 16–25. Antenna (Figs. 16–19) and pronotum (Figs. 20–25) of Mannerheimia arctica (Figs. 16–17, 20–22), M. brevipennis (Figs. 18–19, 23–24) and M. pleshanovi (Fig. 25): 16, 18—right antenna of male, 17, 19—right antenna of female, 20—Irkutsk Area: Vitimskiy Nature Reserve, 21—Krasnoyarsk Territory: Potapovo, 22—Murmansk Area: Khibiny, 23—Krasnoyarsk Territory: Lebyazhya River, 24—China (Xinjiang), 25—Irkutsk Area: Khamar-Daban (paratype). Scale bar: 0.1 mm." captionText-2="FIGURES 26–38. Body parts of Mannerheimia arctica: 26—ventral aspect of head, 27—ventral aspect of prothorax, 28—ventral aspect of mesoventrite, 29—ventral aspect of metaventrite (without metanepisternum), 30—metendosternite (ventral view), 31—scutellum (without lateral structures, dorsal view), 32—ventral aspect of elytron, 33—hind tibia and tarsus (anterior view), 34—male tergite VIII (dorsal view), 35—male sternite VIII (ventral view), 36—female tergite VIII (dorsal view), 37—female sternite VIII (ventral view), 38—female genital segment (dorsal view). Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5531378" figureDoi-1="http://doi.org/10.5281/zenodo.5531380" figureDoi-2="http://doi.org/10.5281/zenodo.5531382" httpUri-0="https://zenodo.org/record/5531378/files/figure.png" httpUri-1="https://zenodo.org/record/5531380/files/figure.png" httpUri-2="https://zenodo.org/record/5531382/files/figure.png" pageId="2" pageNumber="303">12–38</figureCitation>
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)
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</paragraph>
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species:
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</emphasis>
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<taxonomicName id="4C084D54087F244CFEBDFD80FD371231" authority="Maklin, 1878" authorityName="Maklin" authorityYear="1878" box="[300,641,615,639]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="divergens">
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<emphasis id="B97CEAC5087F244CFEBDFD80FE451231" box="[300,499,615,639]" italics="true" pageId="2" pageNumber="303">Omalium divergens</emphasis>
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<bibRefCitation id="EF994B26087F244CFE69FD80FD371231" author="Maklin, F. G." box="[504,641,615,639]" pageId="2" pageNumber="303" pagination="15 - 32" refId="ref25649" refString="Maklin, F. G. (1878) Diagnoser ofver nagra nya siberiska insektarter. Ofversigt af Finska Vetenskaps-Societetens Forhandlingar, 19, 15 - 32." type="journal article" year="1878">Mäklin, 1878</bibRefCitation>
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<taxonomicName id="4C084D54087F244CFF06FD43FE5912F3" ID-CoL="8VZ5G" authority="Maklin, 1880 a: 80" authorityName="Maklin" authorityPageNumber="80" authorityYear="1880" box="[151,495,676,701]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CEAC5087F244CFF06FD43FE9A12F2" box="[151,300,676,700]" italics="true" pageId="2" pageNumber="303">Mannerheimia</emphasis>
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<bibRefCitation id="EF994B26087F244CFEA3FD43FE5912F3" author="Maklin, F. G." box="[306,495,676,701]" pageId="2" pageNumber="303" pagination="79 - 86" refId="ref25677" refString="Maklin, F. G. (1880 a) Ytterligare diagnoser ofver nagra nya sibiriska Coleopter-arter. Ofversigt af Finska Vetenskaps-Societetens Forhandlingar, 22, 79 - 86." type="journal article" year="1880">Mäklin, 1880a: 80</bibRefCitation>
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;
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<treatmentCitation id="0AA910C6087F244CFE6BFD43FD0E12F3" author="Maklin, F. G." box="[506,696,676,701]" page="39" pageId="2" pageNumber="303" year="1880">
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<bibRefCitation id="EF994B26087F244CFE6BFD43FD0E12F3" author="Maklin, F. G." box="[506,696,676,701]" pageId="2" pageNumber="303" pagination="1 - 48" refId="ref25707" refString="Maklin, F. G. (1880 b) Coleoptera insamlade under den Nordenskiold'ska expeditionen 1875 pa nagra oar vid Norges nordvest- kust, pa Novaja Semlja och on Waigatsch samt vid Jenissej i Sibirien. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 18 (4), 1 - 48." type="journal article" year="1880">Mäklin, 1880b: 39</bibRefCitation>
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,
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<treatmentCitation id="0AA910C6087F244CFD52FD43FC1112F2" author="Ganglbauer, L." box="[707,935,676,700]" page="725" pageId="2" pageNumber="303" year="1895">
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<bibRefCitation id="EF994B26087F244CFD52FD43FC1112F2" author="Ganglbauer, L." box="[707,935,676,700]" pageId="2" pageNumber="303" refId="ref24626" refString="Ganglbauer, L. (1895) Die Kafer von Mitteleuropa. 2. Familienreihe Staphylinoidea. Theil I. Staphylinidae, Pselaphidae. Carl Gerold's Sohn, Wien, 881 pp." type="book" year="1895">Ganglbauer 1895: 725</bibRefCitation>
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,
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<treatmentCitation id="0AA910C6087F244CFC23FD42FBE512F2" author="Luze, G." box="[946,1107,676,700]" page="251" pageId="2" pageNumber="303" year="1905">
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<bibRefCitation id="EF994B26087F244CFC23FD42FBE512F2" author="Luze, G." box="[946,1107,676,700]" pageId="2" pageNumber="303" pagination="241 - 256" refId="ref25610" refString="Luze, G. (1905) Die palaarktischen Arten der Staphyliniden-Gattungen Deliphrum Er., Phyllodrepoidea Ganglb. und Mannerheimia Makl. Verhandlungen der koeniglich und keiserlich zoologisch-botanischen Gesellschaft in Wien, 55, 241 - 256." type="journal article" year="1905">Luze 1905: 251</bibRefCitation>
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</treatmentCitation>
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,
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<treatmentCitation id="0AA910C6087F244CFBCFFD42FB4312F3" author="Porta, A." box="[1118,1269,676,701]" page="24" pageId="2" pageNumber="303" year="1926">
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<bibRefCitation id="EF994B26087F244CFBCFFD42FB4312F3" author="Porta, A." box="[1118,1269,676,701]" pageId="2" pageNumber="303" refId="ref26180" refString="Porta, A. (1926) Fauna coleopterorum italica. II. Staphylinoidea. Stalilimento Tipografico Piacentino, Piacenza, 405 pp." type="book" year="1926">Porta 1926: 24</bibRefCitation>
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</treatmentCitation>
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,
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<treatmentCitation id="0AA910C6087F244CFB6EFD42FF591295" author="Cameron, M." page="149" pageId="2" pageNumber="303" year="1930">
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<bibRefCitation id="EF994B26087F244CFB6EFD42FF591295" author="Cameron, M." pageId="2" pageNumber="303" refId="ref24029" refString="Cameron, M. (1930) The fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. 1. Taylor and Francis, London, xvii + 471 pp." type="book" year="1930">Cameron 1930: 149</bibRefCitation>
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</treatmentCitation>
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,
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<treatmentCitation id="0AA910C6087F244CFF6BFD24FE261295" author="Palm, T." box="[250,400,707,731]" page="94" pageId="2" pageNumber="303" year="1948">
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<bibRefCitation id="EF994B26087F244CFF6BFD24FE261295" author="Palm, T." box="[250,400,707,731]" pageId="2" pageNumber="303" refId="ref25965" refString="Palm, T. (1948) Svensk Insekfauna. 9. Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Unterfam. Micropeplinae, Phloeocharinae, Olisthaerinae, Proteininae, Omaliinae. Almquist and Wiksells, Uppsala, 133 pp." type="book" year="1948">Palm 1948: 94</bibRefCitation>
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</treatmentCitation>
|
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,
|
||
<treatmentCitation id="0AA910C6087F244CFE0BFD24FDE61295" author="Zanetti, A." box="[410,592,707,731]" page="273" pageId="2" pageNumber="303" year="1987">
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<bibRefCitation id="EF994B26087F244CFE0BFD24FDE61295" author="Zanetti, A." box="[410,592,707,731]" pageId="2" pageNumber="303" pagination="1 - 472" refId="ref27488" refString="Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d'Italia. Vol. 25. Calderini, Bologna, pp. i - xii + 1 - 472." type="book chapter" year="1987">Zanetti 1987: 273</bibRefCitation>
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</treatmentCitation>
|
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,
|
||
<treatmentCitation id="0AA910C6087F244CFDCAFD24FCAB1295" author="Watanabe, Y." box="[603,797,707,731]" page="77" pageId="2" pageNumber="303" year="1990">
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<bibRefCitation id="EF994B26087F244CFDCAFD24FCAB1295" author="Watanabe, Y." box="[603,797,707,731]" pageId="2" pageNumber="303" pagination="59 - 391" refId="ref27421" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe 1990: 77</bibRefCitation>
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</treatmentCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C312655C087F244AFF06FCEEFB8C112B" lastPageId="4" lastPageNumber="305" pageId="2" pageNumber="303" type="description">
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<paragraph id="8BB736D7087F244CFF06FCEEFE9213B2" blockId="2.[151,1437,777,2027]" pageId="2" pageNumber="303">
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<emphasis id="B97CEAC5087F244CFF06FCEEFEF3136D" bold="true" box="[151,325,777,803]" pageId="2" pageNumber="303">Redescription.</emphasis>
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Body moderately wide (
|
||
<figureCitation id="13332A52087F244CFDF3FCEEFD5B136A" box="[610,749,777,804]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1955,1979]" captionTargetBox="[187,1400,190,1920]" captionTargetId="figure-16@3.[173,1414,181,1931]" captionTargetPageId="3" captionText="FIGURES 1–9. Habitus of Mannerheimia: 1—M. afghanica (paratype), 2—M. arctica, 3—M. asiatica, 4—M. brevipennis, 5—M. japonica, 6—M. obscura, 7—M. pleshanovi (paratype), 8—M. yasudai, 9—M. zanettii (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5531374" httpUri="https://zenodo.org/record/5531374/files/figure.png" pageId="2" pageNumber="303">Figs. 1, 5–9</figureCitation>
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) or elongate (
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<figureCitation id="13332A52087F244CFC1FFCEEFC4B136D" box="[910,1021,777,803]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1955,1979]" captionTargetBox="[187,1400,190,1920]" captionTargetId="figure-16@3.[173,1414,181,1931]" captionTargetPageId="3" captionText="FIGURES 1–9. Habitus of Mannerheimia: 1—M. afghanica (paratype), 2—M. arctica, 3—M. asiatica, 4—M. brevipennis, 5—M. japonica, 6—M. obscura, 7—M. pleshanovi (paratype), 8—M. yasudai, 9—M. zanettii (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5531374" httpUri="https://zenodo.org/record/5531374/files/figure.png" pageId="2" pageNumber="303">Figs. 2, 4</figureCitation>
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), slightly or strongly convex, shiny; length
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<quantity id="4CF09B32087F244CFF75FCCAFE3D1306" box="[228,395,813,840]" metricMagnitude="-3" metricUnit="m" metricValue="3.07" metricValueMax="4.24" metricValueMin="1.9" pageId="2" pageNumber="303" unit="mm" value="3.07" valueMax="4.24" valueMin="1.9">1.90–4.24 mm</quantity>
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. Coloration varying from yellow-brown to dark-brown, sometimes with paler pronotum and/ or elytra. Forebody with fine to coarse, dense to sparse punctation; punctation of abdomen fine or missing. Head and pronotum with fine to coarse microreticulation or without it; abdomen usually with distinct isodiametric microsculpture. Body glabrous, with sparse small setae on frontal portion of head and sometimes on pronotum and elytra or densely covered by short or moderately long setation (
|
||
<taxonomicName id="4C084D54087F244CFC94FC59FC331399" authorityName="Kastcheev" authorityYear="1999" box="[773,901,958,984]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="asiatica">
|
||
<emphasis id="B97CEAC5087F244CFC94FC59FCAA1399" box="[773,796,958,983]" italics="true" pageId="2" pageNumber="303">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087F244CFCBDFC59FC331399" box="[812,901,958,983]" italics="true" pageId="2" pageNumber="303">asiatica</emphasis>
|
||
</taxonomicName>
|
||
); abdominal tergites with or without very fine, sparse setae.
|
||
</paragraph>
|
||
<paragraph id="8BB736D7087F244CFF56FBE2FAE81581" blockId="2.[151,1437,777,2027]" pageId="2" pageNumber="303">
|
||
Head transverse, with slightly elevated supra-antennal prominences and middle portion; anteocellar foveae missing or present, short or elongate; mediobasal portion without longitudinal impression; postocular ridges missing, temples short, gradually narrowed toward neck or moderately long (
|
||
<taxonomicName id="4C084D54087F244CFC5FFBA9FBFB1429" authorityName="Kastcheev" authorityYear="1999" box="[974,1101,1102,1128]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="asiatica">
|
||
<emphasis id="B97CEAC5087F244CFC5FFBA9FC531429" box="[974,997,1102,1127]" italics="true" pageId="2" pageNumber="303">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087F244CFC65FBA9FBFB1429" box="[1012,1101,1102,1127]" italics="true" pageId="2" pageNumber="303">asiatica</emphasis>
|
||
</taxonomicName>
|
||
). Ocelli small, sometimes indistinct, or large and convex. Eyes convex, large or moderately small (
|
||
<taxonomicName id="4C084D54087F244CFC3CFB95FB9D14C5" authorityName="Kastcheev" authorityYear="1999" box="[941,1067,1138,1164]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="asiatica">
|
||
<emphasis id="B97CEAC5087F244CFC3CFB95FC7214C5" box="[941,964,1138,1163]" italics="true" pageId="2" pageNumber="303">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087F244CFC43FB95FB9D14C5" box="[978,1067,1138,1163]" italics="true" pageId="2" pageNumber="303">asiatica</emphasis>
|
||
</taxonomicName>
|
||
). Gular sutures widely separated at about level of posterior margins of eyes, slightly divergent posteriorly towards base of head (
|
||
<figureCitation id="13332A52087F244CFB48FB72FA9914FE" box="[1241,1327,1173,1200]" captionStart="FIGURES 26–38" captionStartId="12.[151,264,1858,1882]" captionTargetBox="[172,1413,320,1826]" captionTargetId="figure-79@12.[160,1428,309,1835]" captionTargetPageId="12" captionText="FIGURES 26–38. Body parts of Mannerheimia arctica: 26—ventral aspect of head, 27—ventral aspect of prothorax, 28—ventral aspect of mesoventrite, 29—ventral aspect of metaventrite (without metanepisternum), 30—metendosternite (ventral view), 31—scutellum (without lateral structures, dorsal view), 32—ventral aspect of elytron, 33—hind tibia and tarsus (anterior view), 34—male tergite VIII (dorsal view), 35—male sternite VIII (ventral view), 36—female tergite VIII (dorsal view), 37—female sternite VIII (ventral view), 38—female genital segment (dorsal view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531382" httpUri="https://zenodo.org/record/5531382/files/figure.png" pageId="2" pageNumber="303">Fig. 26</figureCitation>
|
||
). Labrum transverse, with widely rounded latero-apical portions, wide apical emargination and moderately long laterobasal projection (
|
||
<figureCitation id="13332A52087F244CFE88FB3AFEDB14B9" box="[281,365,1245,1271]" captionStart="FIGURES 12–15" captionStartId="10.[151,264,1952,1976]" captionTargetBox="[167,1423,720,1921]" captionTargetId="figure-289@10.[151,1436,705,1929]" captionTargetPageId="10" captionText="FIGURES 12–15. Mouthparts of Mannerheimia arctica: 12—labrum (dorsal view), 13—mandibles (left and right, dorsal view), 14—maxilla (left, dorsal view), 15—labium and mentum (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531378" httpUri="https://zenodo.org/record/5531378/files/figure.png" pageId="2" pageNumber="303">Fig. 12</figureCitation>
|
||
). Mandibles wide and subtriangular, each with moderately narrow and elongate apical portion, wide and large prostheca and molar area, with distinct subtriangular tooth in middle of right mandible (
|
||
<figureCitation id="13332A52087F244CFAA8FAE6FA391555" box="[1337,1423,1281,1307]" captionStart="FIGURES 12–15" captionStartId="10.[151,264,1952,1976]" captionTargetBox="[167,1423,720,1921]" captionTargetId="figure-289@10.[151,1436,705,1929]" captionTargetPageId="10" captionText="FIGURES 12–15. Mouthparts of Mannerheimia arctica: 12—labrum (dorsal view), 13—mandibles (left and right, dorsal view), 14—maxilla (left, dorsal view), 15—labium and mentum (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531378" httpUri="https://zenodo.org/record/5531378/files/figure.png" pageId="2" pageNumber="303">Fig. 13</figureCitation>
|
||
).
|
||
<collectingRegion id="49CCF835087F244CFF06FAC1FF40150E" box="[151,246,1318,1344]" country="Myanmar" name="Chin" pageId="2" pageNumber="303">Mentum</collectingRegion>
|
||
small, with two long latero-apical setae; labium wide, with short preapical labial palpomere, about as long as broad or distinctly transverse, apical palpomere, more than twice as long and about as broad as preceeding segment (
|
||
<figureCitation id="13332A52087F244CFF4EFA8AFE8515C6" box="[223,307,1389,1416]" captionStart="FIGURES 12–15" captionStartId="10.[151,264,1952,1976]" captionTargetBox="[167,1423,720,1921]" captionTargetId="figure-289@10.[151,1436,705,1929]" captionTargetPageId="10" captionText="FIGURES 12–15. Mouthparts of Mannerheimia arctica: 12—labrum (dorsal view), 13—mandibles (left and right, dorsal view), 14—maxilla (left, dorsal view), 15—labium and mentum (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531378" httpUri="https://zenodo.org/record/5531378/files/figure.png" pageId="2" pageNumber="303">Fig. 15</figureCitation>
|
||
). Maxilla as in
|
||
<figureCitation id="13332A52087F244CFE72FA8AFD8015C9" box="[483,566,1389,1415]" captionStart="FIGURES 12–15" captionStartId="10.[151,264,1952,1976]" captionTargetBox="[167,1423,720,1921]" captionTargetId="figure-289@10.[151,1436,705,1929]" captionTargetPageId="10" captionText="FIGURES 12–15. Mouthparts of Mannerheimia arctica: 12—labrum (dorsal view), 13—mandibles (left and right, dorsal view), 14—maxilla (left, dorsal view), 15—labium and mentum (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531378" httpUri="https://zenodo.org/record/5531378/files/figure.png" pageId="2" pageNumber="303">Fig. 14</figureCitation>
|
||
; preapical segment of maxillary palpus long, more than twice as long as, and slightly broader than preceding segment. Antenna short, reaching posterior margin of pronotum or apical third of elytra when reclined, with transverse (
|
||
<figureCitation id="13332A52087F244CFDD2FA52FD7C1581" box="[579,714,1461,1487]" captionStart="FIGURES 16–25" captionStartId="11.[151,264,1047,1071]" captionTargetBox="[169,1421,187,1017]" captionTargetId="figure-16@11.[163,1425,181,1023]" captionTargetPageId="11" captionText="FIGURES 16–25. Antenna (Figs. 16–19) and pronotum (Figs. 20–25) of Mannerheimia arctica (Figs. 16–17, 20–22), M. brevipennis (Figs. 18–19, 23–24) and M. pleshanovi (Fig. 25): 16, 18—right antenna of male, 17, 19—right antenna of female, 20—Irkutsk Area: Vitimskiy Nature Reserve, 21—Krasnoyarsk Territory: Potapovo, 22—Murmansk Area: Khibiny, 23—Krasnoyarsk Territory: Lebyazhya River, 24—China (Xinjiang), 25—Irkutsk Area: Khamar-Daban (paratype). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531380" httpUri="https://zenodo.org/record/5531380/files/figure.png" pageId="2" pageNumber="303">Figs. 18–19</figureCitation>
|
||
) or elongate (
|
||
<taxonomicName id="4C084D54087F244CFCF7FA51FC6C1581" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[870,986,1462,1488]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="arctica">
|
||
<emphasis id="B97CEAC5087F244CFCF7FA51FCCB1581" box="[870,893,1462,1487]" italics="true" pageId="2" pageNumber="303">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087F244CFC1DFA51FC6C1581" box="[908,986,1462,1487]" italics="true" pageId="2" pageNumber="303">arctica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="13332A52087F244CFC79FA52FBDB159E" box="[1000,1133,1461,1488]" captionStart="FIGURES 16–25" captionStartId="11.[151,264,1047,1071]" captionTargetBox="[169,1421,187,1017]" captionTargetId="figure-16@11.[163,1425,181,1023]" captionTargetPageId="11" captionText="FIGURES 16–25. Antenna (Figs. 16–19) and pronotum (Figs. 20–25) of Mannerheimia arctica (Figs. 16–17, 20–22), M. brevipennis (Figs. 18–19, 23–24) and M. pleshanovi (Fig. 25): 16, 18—right antenna of male, 17, 19—right antenna of female, 20—Irkutsk Area: Vitimskiy Nature Reserve, 21—Krasnoyarsk Territory: Potapovo, 22—Murmansk Area: Khibiny, 23—Krasnoyarsk Territory: Lebyazhya River, 24—China (Xinjiang), 25—Irkutsk Area: Khamar-Daban (paratype). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531380" httpUri="https://zenodo.org/record/5531380/files/figure.png" pageId="2" pageNumber="303">Figs. 16–17</figureCitation>
|
||
) antennomeres 4–10.
|
||
</paragraph>
|
||
<paragraph id="8BB736D7087F244CFF56FA3DFE331685" blockId="2.[151,1437,777,2027]" pageId="2" pageNumber="303">
|
||
Pronotum transverse (
|
||
<figureCitation id="13332A52087F244CFE2AFA3EFDF215BA" box="[443,580,1497,1524]" captionStart="FIGURES 16–25" captionStartId="11.[151,264,1047,1071]" captionTargetBox="[169,1421,187,1017]" captionTargetId="figure-16@11.[163,1425,181,1023]" captionTargetPageId="11" captionText="FIGURES 16–25. Antenna (Figs. 16–19) and pronotum (Figs. 20–25) of Mannerheimia arctica (Figs. 16–17, 20–22), M. brevipennis (Figs. 18–19, 23–24) and M. pleshanovi (Fig. 25): 16, 18—right antenna of male, 17, 19—right antenna of female, 20—Irkutsk Area: Vitimskiy Nature Reserve, 21—Krasnoyarsk Territory: Potapovo, 22—Murmansk Area: Khibiny, 23—Krasnoyarsk Territory: Lebyazhya River, 24—China (Xinjiang), 25—Irkutsk Area: Khamar-Daban (paratype). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531380" httpUri="https://zenodo.org/record/5531380/files/figure.png" pageId="2" pageNumber="303">Figs. 20–25</figureCitation>
|
||
), widely rounded toward latero-apical portions, sometimes slightly protruded anteriad, and slightly or distinctly narrowed toward obtuse or widely rounded posterior angles, with or without laterobasal impressions; lateral portions narrow, slightly explanate. Prothorax with elongate, narrow intercoxal process and strongly protruded, long prosternal processes (
|
||
<figureCitation id="13332A52087F244CFD43F9A2FC911611" box="[722,807,1605,1631]" captionStart="FIGURES 26–38" captionStartId="12.[151,264,1858,1882]" captionTargetBox="[172,1413,320,1826]" captionTargetId="figure-79@12.[160,1428,309,1835]" captionTargetPageId="12" captionText="FIGURES 26–38. Body parts of Mannerheimia arctica: 26—ventral aspect of head, 27—ventral aspect of prothorax, 28—ventral aspect of mesoventrite, 29—ventral aspect of metaventrite (without metanepisternum), 30—metendosternite (ventral view), 31—scutellum (without lateral structures, dorsal view), 32—ventral aspect of elytron, 33—hind tibia and tarsus (anterior view), 34—male tergite VIII (dorsal view), 35—male sternite VIII (ventral view), 36—female tergite VIII (dorsal view), 37—female sternite VIII (ventral view), 38—female genital segment (dorsal view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531382" httpUri="https://zenodo.org/record/5531382/files/figure.png" pageId="2" pageNumber="303">Fig. 27</figureCitation>
|
||
). Mesoventrite transverse, with long intercoxal process, reaching about middle of mesocoxae (
|
||
<figureCitation id="13332A52087F244CFDDEF98EFD1016CD" box="[591,678,1641,1667]" captionStart="FIGURES 26–38" captionStartId="12.[151,264,1858,1882]" captionTargetBox="[172,1413,320,1826]" captionTargetId="figure-79@12.[160,1428,309,1835]" captionTargetPageId="12" captionText="FIGURES 26–38. Body parts of Mannerheimia arctica: 26—ventral aspect of head, 27—ventral aspect of prothorax, 28—ventral aspect of mesoventrite, 29—ventral aspect of metaventrite (without metanepisternum), 30—metendosternite (ventral view), 31—scutellum (without lateral structures, dorsal view), 32—ventral aspect of elytron, 33—hind tibia and tarsus (anterior view), 34—male tergite VIII (dorsal view), 35—male sternite VIII (ventral view), 36—female tergite VIII (dorsal view), 37—female sternite VIII (ventral view), 38—female genital segment (dorsal view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531382" httpUri="https://zenodo.org/record/5531382/files/figure.png" pageId="2" pageNumber="303">Fig. 28</figureCitation>
|
||
). Scutellum wide, with rounded apex (
|
||
<figureCitation id="13332A52087F244CFBF4F98EFB0D16CD" box="[1125,1211,1641,1667]" captionStart="FIGURES 26–38" captionStartId="12.[151,264,1858,1882]" captionTargetBox="[172,1413,320,1826]" captionTargetId="figure-79@12.[160,1428,309,1835]" captionTargetPageId="12" captionText="FIGURES 26–38. Body parts of Mannerheimia arctica: 26—ventral aspect of head, 27—ventral aspect of prothorax, 28—ventral aspect of mesoventrite, 29—ventral aspect of metaventrite (without metanepisternum), 30—metendosternite (ventral view), 31—scutellum (without lateral structures, dorsal view), 32—ventral aspect of elytron, 33—hind tibia and tarsus (anterior view), 34—male tergite VIII (dorsal view), 35—male sternite VIII (ventral view), 36—female tergite VIII (dorsal view), 37—female sternite VIII (ventral view), 38—female genital segment (dorsal view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531382" httpUri="https://zenodo.org/record/5531382/files/figure.png" pageId="2" pageNumber="303">Fig. 31</figureCitation>
|
||
). Metaventrite with wide and deep intercoxal cavities and moderately short, subtriangular intercoxal process, almost reaching mesosternal process (
|
||
<figureCitation id="13332A52087F244CFEB5F956FECE1685" box="[292,376,1713,1739]" captionStart="FIGURES 26–38" captionStartId="12.[151,264,1858,1882]" captionTargetBox="[172,1413,320,1826]" captionTargetId="figure-79@12.[160,1428,309,1835]" captionTargetPageId="12" captionText="FIGURES 26–38. Body parts of Mannerheimia arctica: 26—ventral aspect of head, 27—ventral aspect of prothorax, 28—ventral aspect of mesoventrite, 29—ventral aspect of metaventrite (without metanepisternum), 30—metendosternite (ventral view), 31—scutellum (without lateral structures, dorsal view), 32—ventral aspect of elytron, 33—hind tibia and tarsus (anterior view), 34—male tergite VIII (dorsal view), 35—male sternite VIII (ventral view), 36—female tergite VIII (dorsal view), 37—female sternite VIII (ventral view), 38—female genital segment (dorsal view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531382" httpUri="https://zenodo.org/record/5531382/files/figure.png" pageId="2" pageNumber="303">Fig. 29</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB736D7087F244CFF56F932FB4C175A" blockId="2.[151,1437,777,2027]" pageId="2" pageNumber="303">
|
||
Elytra short or moderately long, gradually or strongly widened apicad (
|
||
<figureCitation id="13332A52087F244CFC79F932FB8A16A1" box="[1000,1084,1749,1775]" captionStart="FIGURES 26–38" captionStartId="12.[151,264,1858,1882]" captionTargetBox="[172,1413,320,1826]" captionTargetId="figure-79@12.[160,1428,309,1835]" captionTargetPageId="12" captionText="FIGURES 26–38. Body parts of Mannerheimia arctica: 26—ventral aspect of head, 27—ventral aspect of prothorax, 28—ventral aspect of mesoventrite, 29—ventral aspect of metaventrite (without metanepisternum), 30—metendosternite (ventral view), 31—scutellum (without lateral structures, dorsal view), 32—ventral aspect of elytron, 33—hind tibia and tarsus (anterior view), 34—male tergite VIII (dorsal view), 35—male sternite VIII (ventral view), 36—female tergite VIII (dorsal view), 37—female sternite VIII (ventral view), 38—female genital segment (dorsal view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531382" httpUri="https://zenodo.org/record/5531382/files/figure.png" pageId="2" pageNumber="303">Fig. 32</figureCitation>
|
||
); lateral portions narrowly flattened, with bordered margins. Wings fully developed or reduced (
|
||
<taxonomicName id="4C084D54087F244CFCE9F91DFC41175D" authorityName="Kastcheev" authorityYear="1999" box="[888,1015,1786,1812]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="asiatica">
|
||
<emphasis id="B97CEAC5087F244CFCE9F91DFC39175D" box="[888,911,1786,1811]" italics="true" pageId="2" pageNumber="303">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087F244CFC0FF91DFC41175D" box="[926,1015,1786,1811]" italics="true" pageId="2" pageNumber="303">asiatica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C084D54087F244CFB95F91DFB15175D" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[1028,1187,1785,1812]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="pleshanovi" status="sp. nov.">
|
||
<emphasis id="B97CEAC5087F244CFB95F91DFBAD175D" box="[1028,1051,1786,1811]" italics="true" pageId="2" pageNumber="303">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087F244CFBBBF91EFB15175D" box="[1066,1187,1785,1811]" italics="true" pageId="2" pageNumber="303">pleshanovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97CEAC5087F244CFB3BF91EFB5C175D" bold="true" box="[1194,1258,1785,1811]" pageId="2" pageNumber="303">
|
||
<taxonomicNameLabel id="A24F57BE087F244CFB3BF91EFB5C175D" box="[1194,1258,1785,1811]" pageId="2" pageNumber="303" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB736D7087F244CFF56F8F9FE591731" blockId="2.[151,1437,777,2027]" pageId="2" pageNumber="303">
|
||
Legs long and slender; tibiae covered with moderately sparse and long setae, these setae stouter on meso- and metatibiae; metatarsus about twice as long as metatibia (
|
||
<figureCitation id="13332A52087F244CFC83F8A6FCD01715" box="[786,870,1857,1883]" captionStart="FIGURES 26–38" captionStartId="12.[151,264,1858,1882]" captionTargetBox="[172,1413,320,1826]" captionTargetId="figure-79@12.[160,1428,309,1835]" captionTargetPageId="12" captionText="FIGURES 26–38. Body parts of Mannerheimia arctica: 26—ventral aspect of head, 27—ventral aspect of prothorax, 28—ventral aspect of mesoventrite, 29—ventral aspect of metaventrite (without metanepisternum), 30—metendosternite (ventral view), 31—scutellum (without lateral structures, dorsal view), 32—ventral aspect of elytron, 33—hind tibia and tarsus (anterior view), 34—male tergite VIII (dorsal view), 35—male sternite VIII (ventral view), 36—female tergite VIII (dorsal view), 37—female sternite VIII (ventral view), 38—female genital segment (dorsal view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531382" httpUri="https://zenodo.org/record/5531382/files/figure.png" pageId="2" pageNumber="303">Fig. 33</figureCitation>
|
||
); apical metatarsomere about twice as long as two preceding tarsomeres together.
|
||
</paragraph>
|
||
<paragraph id="8BB736D7087F244CFF56F86EFEEF17A5" blockId="2.[151,1437,777,2027]" pageId="2" pageNumber="303">Abdomen convex, short or strongly extended, with or without two transverse wing-folding spots in middle of tergite V, with narrow palisade fringe on apical margin of tergite VII in winged species, without palisade fringe in wingless species.</paragraph>
|
||
<caption id="DF77665F087E244DFF06F844FA3D1791" ID-DOI="http://doi.org/10.5281/zenodo.5531374" ID-Zenodo-Dep="5531374" httpUri="https://zenodo.org/record/5531374/files/figure.png" pageId="3" pageNumber="304" startId="3.[151,264,1955,1979]" targetBox="[187,1400,190,1920]" targetPageId="3">
|
||
<paragraph id="8BB736D7087E244DFF06F844FA3D1791" blockId="3.[151,1436,1955,2015]" pageId="3" pageNumber="304">
|
||
<emphasis id="B97CEAC5087E244DFF06F844FE8B17F5" bold="true" box="[151,317,1955,1979]" pageId="3" pageNumber="304">FIGURES 1–9.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C084D54087E244DFE2BF844FDF917F5" authorityName="Maklin" authorityYear="1880" box="[442,591,1955,1979]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAC5087E244DFE2BF844FDF917F5" box="[442,591,1955,1979]" italics="true" pageId="3" pageNumber="304">Mannerheimia</emphasis>
|
||
</taxonomicName>
|
||
: 1—
|
||
<taxonomicName id="4C084D54087E244DFD14F843FCB817F5" authorityName="Coiffait" authorityYear="1982" box="[645,782,1955,1979]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="afghanica">
|
||
<emphasis id="B97CEAC5087E244DFD14F843FD2C17F5" box="[645,666,1956,1979]" italics="true" pageId="3" pageNumber="304">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087E244DFD38F844FCB817F5" box="[681,782,1955,1979]" italics="true" pageId="3" pageNumber="304">afghanica</emphasis>
|
||
</taxonomicName>
|
||
(paratype), 2—
|
||
<taxonomicName id="4C084D54087E244DFC25F843FBA917F5" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[948,1055,1956,1979]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="arctica">
|
||
<emphasis id="B97CEAC5087E244DFC25F843FC7F17F5" box="[948,969,1956,1979]" italics="true" pageId="3" pageNumber="304">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087E244DFC49F843FBA917F5" box="[984,1055,1956,1979]" italics="true" pageId="3" pageNumber="304">arctica</emphasis>
|
||
</taxonomicName>
|
||
, 3—
|
||
<taxonomicName id="4C084D54087E244DFBC2F843FB7117F5" authorityName="Kastcheev" authorityYear="1999" box="[1107,1223,1956,1979]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="asiatica">
|
||
<emphasis id="B97CEAC5087E244DFBC2F843FBDE17F5" box="[1107,1128,1956,1979]" italics="true" pageId="3" pageNumber="304">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087E244DFBE6F843FB7117F5" box="[1143,1223,1956,1979]" italics="true" pageId="3" pageNumber="304">asiatica</emphasis>
|
||
</taxonomicName>
|
||
, 4—
|
||
<taxonomicName id="4C084D54087E244DFB6CF843FA2017F5" authorityName="Motschulsky" authorityYear="1860" box="[1277,1430,1955,1979]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="brevipennis">
|
||
<emphasis id="B97CEAC5087E244DFB6CF843FAA417F5" box="[1277,1298,1956,1979]" italics="true" pageId="3" pageNumber="304">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087E244DFAB0F844FA2017F5" box="[1313,1430,1955,1979]" italics="true" pageId="3" pageNumber="304">brevipennis</emphasis>
|
||
</taxonomicName>
|
||
, 5—
|
||
<taxonomicName id="4C084D54087E244DFF2CF82FFE8E1791" authorityName="Watanabe" authorityYear="1990" box="[189,312,1992,2015]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="B97CEAC5087E244DFF2CF82FFF641791" box="[189,210,1992,2015]" italics="true" pageId="3" pageNumber="304">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087E244DFF4EF82FFE8E1791" box="[223,312,1992,2015]" italics="true" pageId="3" pageNumber="304">japonica</emphasis>
|
||
</taxonomicName>
|
||
, 6—
|
||
<taxonomicName id="4C084D54087E244DFEFBF82FFE681791" authorityName="Coiffait" authorityYear="1982" box="[362,478,1991,2015]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="obscura">
|
||
<emphasis id="B97CEAC5087E244DFEFBF82FFEC91791" box="[362,383,1992,2015]" italics="true" pageId="3" pageNumber="304">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087E244DFE1DF820FE681791" box="[396,478,1991,2015]" italics="true" pageId="3" pageNumber="304">obscura</emphasis>
|
||
</taxonomicName>
|
||
, 7—
|
||
<taxonomicName id="4C084D54087E244DFD81F82FFD161791" authorityName="Shavrin" authorityYear="2021" box="[528,672,1991,2015]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="pleshanovi">
|
||
<emphasis id="B97CEAC5087E244DFD81F82FFD931791" box="[528,549,1992,2015]" italics="true" pageId="3" pageNumber="304">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087E244DFDA3F820FD161791" box="[562,672,1991,2015]" italics="true" pageId="3" pageNumber="304">pleshanovi</emphasis>
|
||
</taxonomicName>
|
||
(paratype), 8—
|
||
<taxonomicName id="4C084D54087E244DFCD0F82FFC041791" authorityName="Watanabe" authorityYear="1990" box="[833,946,1991,2015]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="yasudai">
|
||
<emphasis id="B97CEAC5087E244DFCD0F82FFCE01791" box="[833,854,1992,2015]" italics="true" pageId="3" pageNumber="304">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087E244DFCF2F820FC041791" box="[867,946,1991,2015]" italics="true" pageId="3" pageNumber="304">yasudai</emphasis>
|
||
</taxonomicName>
|
||
, 9—
|
||
<taxonomicName id="4C084D54087E244DFC75F82FFBE71791" authorityName="Shavrin" authorityYear="2021" box="[996,1105,1992,2015]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="zanettii">
|
||
<emphasis id="B97CEAC5087E244DFC75F82FFC4F1791" box="[996,1017,1992,2015]" italics="true" pageId="3" pageNumber="304">M</emphasis>
|
||
.
|
||
<emphasis id="B97CEAC5087E244DFB97F82FFBE71791" box="[1030,1105,1992,2015]" italics="true" pageId="3" pageNumber="304">zanettii</emphasis>
|
||
</taxonomicName>
|
||
(paratype). Scale bar: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB736D70879244AFF56FF70FA8A1153" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="305">
|
||
Male. First four protarsomeres widened. Aedeagus suboval or elongate, with narrow or widely rounded apex; parameres slightly or distinctly exceeding apex of median lobe or shorter, slightly or significantly widened apically; middle portion of median lobe with distinct transverse sclerotized structure (
|
||
<figureCitation id="13332A520879244AFC6BFF38FB3510B7" box="[1018,1155,223,249]" captionStart="FIGURES 42–47" captionStartId="15.[151,264,1523,1547]" captionTargetBox="[163,1417,178,1506]" captionTargetId="figure-16@15.[151,1436,166,1507]" captionTargetPageId="15" captionText="FIGURES 42–47. Aedeagus of Mannerheimia arctica (42: Buryatia: Gitara Lake, 43–44: Murmansk Area: Khibiny, 45: Tymen Area: Sob` St., 46: Yamal: Shchuchye, 47: Kemerovo Area: Belogorsk): 42–43, 45–47—parameral view, 44—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531388" httpUri="https://zenodo.org/record/5531388/files/figure.png" pageId="4" pageNumber="305">Figs. 42, 47</figureCitation>
|
||
,
|
||
<figureCitation id="13332A520879244AFB01FF38FB6F10B7" box="[1168,1241,222,249]" captionStart="FIGURES 48–56" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[286,1301,192,1910]" captionTargetId="figure-16@17.[274,1314,181,1919]" captionTargetPageId="17" captionText="FIGURES 48–56.Aedeagus of Mannerheimia brevipennis (48–49: Kazakhstan:Aksu, 50: Krasnoyarsk Territory:Turukhanskiy District, 51: Buryatia: Okinskiy District, 52: Yamal: Sopkey; 53: Kyrgyzstan:Ala Too; 54: Yakutia: Spasskiy; 55: Iran: Esfahan; 56: China: Xinjiang): 48, 50–56—parameral view, 49—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531390" httpUri="https://zenodo.org/record/5531390/files/figure.png" pageId="4" pageNumber="305">48, 50</figureCitation>
|
||
,
|
||
<figureCitation id="13332A520879244AFB77FF39FA8710B7" box="[1254,1329,222,249]" captionStart="FIGURES 57–62" captionStartId="23.[151,264,1913,1937]" captionTargetBox="[158,1429,187,1881]" captionTargetId="figure-16@23.[151,1436,181,1889]" captionTargetPageId="23" captionText="FIGURES 57–62. Aedeagus of Mannerheimia asiatica (Figs. 57–58), M. japonica (Figs. 59–60), M. obscura (Figs. 61–62): 57, 59, 61—parameral view, 58, 60, 62—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531394" httpUri="https://zenodo.org/record/5531394/files/figure.png" pageId="4" pageNumber="305">57, 59</figureCitation>
|
||
); internal sac long, narrow or wide, usually with two long fields of elongate thorns (
|
||
<figureCitation id="13332A520879244AFC45FEE4FBEA1153" box="[980,1116,259,285]" captionStart="FIGURES 42–47" captionStartId="15.[151,264,1523,1547]" captionTargetBox="[163,1417,178,1506]" captionTargetId="figure-16@15.[151,1436,166,1507]" captionTargetPageId="15" captionText="FIGURES 42–47. Aedeagus of Mannerheimia arctica (42: Buryatia: Gitara Lake, 43–44: Murmansk Area: Khibiny, 45: Tymen Area: Sob` St., 46: Yamal: Shchuchye, 47: Kemerovo Area: Belogorsk): 42–43, 45–47—parameral view, 44—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531388" httpUri="https://zenodo.org/record/5531388/files/figure.png" pageId="4" pageNumber="305">Figs. 42–43</figureCitation>
|
||
,
|
||
<figureCitation id="13332A520879244AFBF6FEE4FB331153" box="[1127,1157,259,285]" captionStart="FIGURES 48–56" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[286,1301,192,1910]" captionTargetId="figure-16@17.[274,1314,181,1919]" captionTargetPageId="17" captionText="FIGURES 48–56.Aedeagus of Mannerheimia brevipennis (48–49: Kazakhstan:Aksu, 50: Krasnoyarsk Territory:Turukhanskiy District, 51: Buryatia: Okinskiy District, 52: Yamal: Sopkey; 53: Kyrgyzstan:Ala Too; 54: Yakutia: Spasskiy; 55: Iran: Esfahan; 56: China: Xinjiang): 48, 50–56—parameral view, 49—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531390" httpUri="https://zenodo.org/record/5531390/files/figure.png" pageId="4" pageNumber="305">48</figureCitation>
|
||
,
|
||
<figureCitation id="13332A520879244AFB00FEE5FB6F1153" box="[1169,1241,258,285]" captionStart="FIGURES 57–62" captionStartId="23.[151,264,1913,1937]" captionTargetBox="[158,1429,187,1881]" captionTargetId="figure-16@23.[151,1436,181,1889]" captionTargetPageId="23" captionText="FIGURES 57–62. Aedeagus of Mannerheimia asiatica (Figs. 57–58), M. japonica (Figs. 59–60), M. obscura (Figs. 61–62): 57, 59, 61—parameral view, 58, 60, 62—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531394" httpUri="https://zenodo.org/record/5531394/files/figure.png" pageId="4" pageNumber="305">57, 61</figureCitation>
|
||
,
|
||
<figureCitation id="13332A520879244AFB74FEE4FAB51153" box="[1253,1283,259,285]" captionStart="FIGURES 66–68" captionStartId="26.[151,264,1705,1729]" captionTargetBox="[158,1428,891,1676]" captionTargetId="figure-309@26.[151,1436,885,1681]" captionTargetPageId="26" captionText="FIGURES 66–68. Aedeagus of Mannerheimia pleshanovi (paratypes): 66, 68—parameral view, 67—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531398" httpUri="https://zenodo.org/record/5531398/files/figure.png" pageId="4" pageNumber="305">66</figureCitation>
|
||
,
|
||
<figureCitation id="13332A520879244AFA9EFEE4FA991153" box="[1295,1327,259,285]" captionStart="FIGURES 70–71" captionStartId="28.[151,264,1990,2014]" captionTargetBox="[162,1426,892,1959]" captionTargetId="figure-283@28.[151,1436,885,1966]" captionTargetPageId="28" captionText="FIGURES 70–71. Aedeagus of Mannerheimia zanettii (paratype): 70—parameral view, 71—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531402" httpUri="https://zenodo.org/record/5531402/files/figure.png" pageId="4" pageNumber="305">70</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB736D70879244AFF56FEC0FB8C112B" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="305">
|
||
Female. First four protarsomeres narrow. Genital segment with narrow short sternite IX, wide and long gonocoxites, and with narrow, elongate styli, each with a very long apical seta (
|
||
<figureCitation id="13332A520879244AFC48FEACFB9B112B" box="[985,1069,331,357]" captionStart="FIGURES 26–38" captionStartId="12.[151,264,1858,1882]" captionTargetBox="[172,1413,320,1826]" captionTargetId="figure-79@12.[160,1428,309,1835]" captionTargetPageId="12" captionText="FIGURES 26–38. Body parts of Mannerheimia arctica: 26—ventral aspect of head, 27—ventral aspect of prothorax, 28—ventral aspect of mesoventrite, 29—ventral aspect of metaventrite (without metanepisternum), 30—metendosternite (ventral view), 31—scutellum (without lateral structures, dorsal view), 32—ventral aspect of elytron, 33—hind tibia and tarsus (anterior view), 34—male tergite VIII (dorsal view), 35—male sternite VIII (ventral view), 36—female tergite VIII (dorsal view), 37—female sternite VIII (ventral view), 38—female genital segment (dorsal view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531382" httpUri="https://zenodo.org/record/5531382/files/figure.png" pageId="4" pageNumber="305">Fig. 38</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C312655C0879244AFF56FE89FAC11283" pageId="4" pageNumber="305" type="diagnosis">
|
||
<paragraph id="8BB736D70879244AFF56FE89FE04122F" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="305">
|
||
<emphasis id="B97CEAC50879244AFF56FE89FE7911C6" bold="true" box="[199,463,366,392]" pageId="4" pageNumber="305">Differential diagnosis.</emphasis>
|
||
Regarding the shape of the smaller and less pubescent antennomere 5 and the general shape of the aedeagus,
|
||
<taxonomicName id="4C084D540879244AFE0AFE74FD8911E3" authorityName="Maklin" authorityYear="1880" box="[411,575,403,429]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAC50879244AFE0AFE74FD8911E3" box="[411,575,403,429]" italics="true" pageId="4" pageNumber="305">Mannerheimia</emphasis>
|
||
</taxonomicName>
|
||
is similar to the monotypic genus
|
||
<taxonomicName id="4C084D540879244AFC55FE74FAE611E3" authority="Ganglbauer, 1895" authorityName="Ganglbauer" authorityYear="1895" box="[964,1360,402,429]" class="Insecta" family="Staphylinidae" genus="Phyllodrepoidea" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAC50879244AFC55FE74FBC811E3" box="[964,1150,403,429]" italics="true" pageId="4" pageNumber="305">Phyllodrepoidea</emphasis>
|
||
<bibRefCitation id="EF994B260879244AFB14FE74FAE611E3" author="Ganglbauer, L." box="[1157,1360,402,429]" pageId="4" pageNumber="305" refId="ref24626" refString="Ganglbauer, L. (1895) Die Kafer von Mitteleuropa. 2. Familienreihe Staphylinoidea. Theil I. Staphylinidae, Pselaphidae. Carl Gerold's Sohn, Wien, 881 pp." type="book" year="1895">Ganglbauer, 1895</bibRefCitation>
|
||
</taxonomicName>
|
||
. It differs from
|
||
<taxonomicName id="4C084D540879244AFE99FE50FE74119F" authorityName="Ganglbauer" authorityYear="1895" box="[264,450,439,465]" class="Insecta" family="Staphylinidae" genus="Phyllodrepoidea" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAC50879244AFE99FE50FE74119F" box="[264,450,439,465]" italics="true" pageId="4" pageNumber="305">Phyllodrepoidea</emphasis>
|
||
</taxonomicName>
|
||
by a less regular arrangement of the elytral punctation and narrower apical portions of the parameres. Based on the general shape of the body, these two genera are somewhat similar to species of the
|
||
<taxonomicName id="4C084D540879244AFF06FE18FEA41257" box="[151,274,511,537]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAC50879244AFF06FE18FEA41257" box="[151,274,511,537]" italics="true" pageId="4" pageNumber="305">Anthobium</emphasis>
|
||
</taxonomicName>
|
||
group of genera (e.g.
|
||
<bibRefCitation id="EF994B260879244AFD9BFE18FC911257" author="Shavrin, A. V. & Smetana, A." box="[522,807,511,537]" pageId="4" pageNumber="305" pagination="1 - 39" refId="ref26967" refString="Shavrin, A. V. & Smetana, A. (2017) A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups. Zootaxa, 4365 (1), 1 - 39. https: // doi. org / 10.11646 / zootaxa. 4365.1.1" type="journal article" year="2017">Shavrin & Smetana 2017</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF994B260879244AFCA3FE18FCD81257" author="Shavrin, A. V. & Smetana, A." box="[818,878,511,537]" pageId="4" pageNumber="305" pagination="576 - 586" refId="ref27032" refString="Shavrin, A. V. & Smetana, A. (2020) Anthobiomorphus, a new genus of Anthophagini Thomson, 1859 (Coleoptera: Staphylinidae: Omaliinae), with description of two new species from China, India and Nepal. Zootaxa, 4755 (3), 576 - 586. https: // doi. org / 10.11646 / zootaxa. 4755.3.9" type="journal article" year="2020">2020</bibRefCitation>
|
||
), from which they can be distinguished by the absence of postocular ridges on the head and lateral pits on the pronotum, greater distance between the gular sutures, and broader gonocoxites.
|
||
</paragraph>
|
||
<paragraph id="8BB736D70879244AFF56FD8DFAC11283" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="305">
|
||
<emphasis id="B97CEAC50879244AFF56FD8DFEFD12CA" bold="true" box="[199,331,618,644]" pageId="4" pageNumber="305">Bionomics.</emphasis>
|
||
For most species of
|
||
<taxonomicName id="4C084D540879244AFDA1FD8CFD6212CB" authorityName="Maklin" authorityYear="1880" box="[560,724,619,645]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAC50879244AFDA1FD8CFD6212CB" box="[560,724,619,645]" italics="true" pageId="4" pageNumber="305">Mannerheimia</emphasis>
|
||
</taxonomicName>
|
||
ecological data are unknown. Members of the genus may be regarded as hygromesophilous. They inhabit mixed forests from low to high elevations and can be found in wet habitats such as mosses, litter, soil, various debris, etc.; some species have been found near snowfields and glaciers.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |