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<document id="022ADD2B351E49E0E4D6A31A1BEE1498" ID-CLB-Dataset="20513" ID-DOI="10.15553/c2018v732a1" ID-GBIF-Dataset="025e1235-99de-4ef9-8849-de54ad29a779" ID-ISSN="2235-3658" ID-Zenodo-Dep="5724514" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1637763397625" checkinUser="carolina" docAuthor="Vorontsova, Maria S." docDate="2018" docId="03A187BAFFCDFF9CFFCFFD4E193E5309" docLanguage="en" docName="Candollea.73.2.143-186.pdf" docOrigin="Candollea 73 (2)" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.3:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="3" docTitle="Panicum humbertii A. Camus" docType="treatment" docVersion="10" lastPageNumber="162" masterDocId="FF98FFC2FFDEFF8FFFBEFF821D085607" masterDocTitle="Revision of the group previously known as Panicum L. (Poaceae: Panicoideae) in Madagascar" masterLastPageNumber="186" masterPageNumber="143" pageNumber="162" updateTime="1699470019059" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="5A306F1D3BE440C4B2E4C840260A50C3">Revision of the group previously known as Panicum L. (Poaceae: Panicoideae) in Madagascar</mods:title>
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<mods:namePart id="75128DCAA11DCB96D47E4E259AB7A2C0">Vorontsova, Maria S.</mods:namePart>
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<mods:date id="8765141A593446F34A1AC8DC7B86760D">2018</mods:date>
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<mods:number id="7D844A51772713CDA8C0B847108FB276">2018-11-05</mods:number>
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<treatment id="03A187BAFFCDFF9CFFCFFD4E193E5309" ID-DOI="http://doi.org/10.5281/zenodo.6312729" ID-GBIF-Taxon="193580205" ID-Zenodo-Dep="6312729" LSID="urn:lsid:plazi:treatment:03A187BAFFCDFF9CFFCFFD4E193E5309" httpUri="http://treatment.plazi.org/id/03A187BAFFCDFF9CFFCFFD4E193E5309" lastPageNumber="162" pageId="19" pageNumber="162">
<subSubSection id="C3126527FFCDFF9CFFCFFD4E1C0A5503" pageId="19" pageNumber="162" type="nomenclature">
<paragraph id="8BB736ACFFCDFF9CFFCFFD4E1C0A5503" blockId="19.[113,764,716,772]" pageId="19" pageNumber="162">
13.
<treatmentCitation id="0AA910BDFFCDFF9CFF29FD4E1DF65503" author="A. Camus" journal="Bull. Soc. Bot. France" page="620" pageId="19" pageNumber="162" score="67.94127085788433" volume="72" year="1925">
<taxonomicName id="4C084D2FFFCDFF9CFF29FD4E1CDB54E3" ID-CoL="4CDZK" authority="A. Camus" authorityName="A. Camus" box="[151,467,716,740]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="19" pageNumber="162" phylum="Tracheophyta" rank="species" species="humbertii">
<emphasis id="B97CEABEFFCDFF9CFF29FD4E1C6A54E3" bold="true" box="[151,354,716,740]" italics="true" pageId="19" pageNumber="162">Panicum humbertii</emphasis>
A. Camus
</taxonomicName>
in
<bibCitation id="5E223EC0FFCDFF9CFE4AFD4E1DF65503" journal="Bull. Soc. Bot. France" page="620" pageId="19" pageNumber="162" score="67.94127085788433" volume="72" year="1925">
Bull. Soc. Bot.
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72: 620. 1925
</bibCitation>
</treatmentCitation>
.
</paragraph>
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<paragraph id="8BB736ACFFCDFF9CFF29FC951FF65569" blockId="19.[151,766,791,879]" pageId="19" pageNumber="162">
<materialsCitation id="3B603CF1FFCDFF9CFF29FC951CDF5568" ID-GBIF-Occurrence="3472816879" collectingDate="1909-04" collectionCode="P" collectorName="Perrier de la Bathie" country="Madagascar" location="Massif de Manongarivo" municipality="Massif de Manongarivo" pageId="19" pageNumber="162" specimenCode="P00450267" specimenCount="1" stateProvince="Prov. Antsiranana" typeStatus="lectotype">
<emphasis id="B97CEABEFFCDFF9CFF29FC951C145528" bold="true" box="[151,284,791,815]" pageId="19" pageNumber="162">
<typeStatus id="54B3880EFFCDFF9CFF29FC951C145528" box="[151,284,791,815]" pageId="19" pageNumber="162" type="lectotype">Lectotypus</typeStatus>
</emphasis>
(designated here):
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<collectingCountry id="F31F763CFFCDFF9CFDB4FC951FA65528" box="[522,686,791,815]" name="Madagascar" pageId="19" pageNumber="162">MADAGASCAR</collectingCountry>
.
<collectingRegion id="49CCF84EFFCDFF9CFD7EFC951C155548" pageId="19" pageNumber="162">Prov. Antsiranana</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BD3ACD6FFCDFF9CFE98FCB51F165548" box="[294,542,823,847]" country="Madagascar" name="Massif de Manongarivo" pageId="19" pageNumber="162" stateProvince="Antsiranana">Massif de Manongarivo</collectingMunicipality>
,
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<collectingDate id="EFF2E984FFCDFF9CFD99FCB51F715548" box="[551,633,823,847]" pageId="19" pageNumber="162" value="1909-04">IV.1909</collectingDate>
</date>
,
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<collectorName id="26FD537AFFCDFF9CFD3DFCBA1DD15568" pageId="19" pageNumber="162">Perrier de la Bâthie</collectorName>
11088
</emphasis>
(
<collectionCode id="ED19AE69FFCDFF9CFE94FCD51C325569" box="[298,314,855,878]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBAE9ED7FFCDFF9CFEF4FCD51CC95568" box="[330,449,855,879]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P00450267</specimenCode>
]!;
</materialsCitation>
<materialsCitation id="3B603CF1FFCDFF9CFE60FCD51FFF556A" ID-GBIF-Occurrence="3472816351" box="[478,759,853,879]" collectionCode="P" collectorName="Perrier de la Bathie" country="Madagascar" pageId="19" pageNumber="162" specimenCode="P00450268" specimenCount="1" stateProvince="Prov." typeStatus="isolectotype">
<typeStatus id="54B3880EFFCDFF9CFE60FCD51F345568" box="[478,572,855,879]" pageId="19" pageNumber="162" type="isolectotype">isolectotype</typeStatus>
:
<collectionCode id="ED19AE69FFCDFF9CFDFBFCD51F5D5569" box="[581,597,855,878]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBAE9ED7FFCDFF9CFDDBFCD51FD55568" box="[613,733,855,879]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P00450268</specimenCode>
]!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3126527FFCDFF9CFF29FC001FF652C3" pageId="19" pageNumber="162" type="reference_group">
<paragraph id="8BB736ACFFCDFF9CFF29FC001F74521E" blockId="19.[151,763,898,1050]" pageId="19" pageNumber="162">
<treatmentCitationGroup id="AB181182FFCDFF9CFF29FC001F74521E" pageId="19" pageNumber="162">
<emphasis id="B97CEABEFFCDFF9CFF29FC001DAC559E" box="[151,164,898,921]" italics="true" pageId="19" pageNumber="162">=</emphasis>
<materialsCitation id="3B603CF1FFCDFF9CFF03FC001C5C521D" ID-GBIF-Occurrence="3472816462" collectingDate="1933-08" collectionCode="P" collectorName="Perrier de la Bathie" country="Madagascar" location="Montagne d'Ambre" municipality="Montagne d'Ambre" pageId="19" pageNumber="162" specimenCode="P00450256" specimenCount="1" stateProvince="Prov. Antsiranana">
<treatmentCitation id="0AA910BDFFCDFF9CFF03FC001C4555BE" author="A. Camus" journal="Bull. Soc. Bot. France" page="40" pageId="19" pageNumber="162" score="67.94127085788433" volume="94" year="1947">
<taxonomicName id="4C084D2FFFCDFF9CFF03FC001CE6559D" ID-CoL="6TN32" authority="A. Camus" authorityName="A. Camus" box="[189,494,898,922]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="19" pageNumber="162" phylum="Tracheophyta" rank="species" species="muscicola">
<originalDetermination id="A8168E50FFCDFF9CFF03FC001CE6559D" box="[189,494,898,922]" pageId="19" pageNumber="162">
<emphasis id="B97CEABEFFCDFF9CFF03FC001C73559D" box="[189,379,898,922]" italics="true" pageId="19" pageNumber="162">Panicum muscicola</emphasis>
A. Camus
</originalDetermination>
</taxonomicName>
in
<bibCitation id="5E223EC0FFCDFF9CFDACFC001C3655BD" journal="Bull. Soc. Bot. France" page="40" pageId="19" pageNumber="162" score="67.94127085788433" volume="94" year="1947">Bull. Soc. Bot. France 94: 40. 1947</bibCitation>
[
</treatmentCitation>
as
<emphasis id="B97CEABEFFCDFF9CFED3FC201CEA55BD" box="[365,482,930,954]" italics="true" pageId="19" pageNumber="162">muscicolum</emphasis>
].
<emphasis id="B97CEABEFFCDFF9CFE46FC201F7D55BD" bold="true" box="[504,629,930,954]" pageId="19" pageNumber="162">Lectotypus</emphasis>
(designated here):
<emphasis id="B97CEABEFFCDFF9CFEBAFC431F8A55DE" bold="true" box="[260,642,961,986]" pageId="19" pageNumber="162">
<collectingCountry id="F31F763CFFCDFF9CFEBAFC431CA855DE" box="[260,416,961,985]" name="Madagascar" pageId="19" pageNumber="162">MADAGASCAR</collectingCountry>
.
<collectingRegion id="49CCF84EFFCDFF9CFE0EFC401F7555DE" box="[432,637,961,986]" pageId="19" pageNumber="162">Prov. Antsiranana</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BD3ACD6FFCDFF9CFD36FC401C2B55FD" country="Madagascar" name="Montagne d'Ambre" pageId="19" pageNumber="162" stateProvince="Antsiranana">Montagne d’Ambre</collectingMunicipality>
,
<date id="FFB6106CFFCDFF9CFE8FFC601CAA55FD" box="[305,418,994,1018]" pageId="19" pageNumber="162" value="1933-08">
<collectingDate id="EFF2E984FFCDFF9CFE8FFC601CAA55FD" box="[305,418,994,1018]" pageId="19" pageNumber="162" value="1933-08">VIII.1933</collectingDate>
</date>
,
<emphasis id="B97CEABEFFCDFF9CFE0FFC601FDF55FD" box="[433,727,994,1018]" italics="true" pageId="19" pageNumber="162">
<collectorName id="26FD537AFFCDFF9CFE0FFC601F8355FD" box="[433,651,994,1018]" pageId="19" pageNumber="162">Perrier de la Bâthie</collectorName>
19300
</emphasis>
(
<collectionCode id="ED19AE69FFCDFF9CFD55FC601FF355FE" box="[747,763,994,1017]" pageId="19" pageNumber="162">P</collectionCode>
[
<specimenCode id="DBAE9ED7FFCDFF9CFF78FB831C36521E" box="[198,318,1025,1049]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P00450256</specimenCode>
]!;
</materialsCitation>
<materialsCitation id="3B603CF1FFCDFF9CFEE5FB801F74521E" ID-GBIF-Occurrence="3472816754" box="[347,636,1024,1050]" collectionCode="P" pageId="19" pageNumber="162" specimenCode="P02263028" specimenCount="1" typeStatus="isolectotype">
<typeStatus id="54B3880EFFCDFF9CFEE5FB801CB1521D" box="[347,441,1026,1050]" pageId="19" pageNumber="162" type="isolectotype">isolectotype</typeStatus>
:
<collectionCode id="ED19AE69FFCDFF9CFE7DFB801CDB521E" box="[451,467,1026,1049]" pageId="19" pageNumber="162">P</collectionCode>
[
<specimenCode id="DBAE9ED7FFCDFF9CFE5AFB831F54521E" box="[484,604,1025,1049]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P02263028</specimenCode>
]!).
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736ACFFCDFF9CFF29FBAF1FF652C3" blockId="19.[151,766,1068,1220]" pageId="19" pageNumber="162">
<treatmentCitationGroup id="AB181182FFCDFF9CFF29FBAF1FF652C3" pageId="19" pageNumber="162">
<emphasis id="B97CEABEFFCDFF9CFF29FBAF1DAC5243" box="[151,164,1069,1092]" italics="true" pageId="19" pageNumber="162">=</emphasis>
<materialsCitation id="3B603CF1FFCDFF9CFF03FBAF1C5B52C3" ID-GBIF-Occurrence="3472816783" collectingDate="1923-05" collectionCode="P" collectorName="Perrier de la Bathie" country="Madagascar" location="Montagne des Francais" municipality="Montagne des Francais" pageId="19" pageNumber="162" specimenCode="P00450281" specimenCount="1" stateProvince="Prov. Antsiranana">
<treatmentCitation id="0AA910BDFFCDFF9CFF03FBAF1C485263" author="A. Camus" journal="Bull. Soc. Bot. France" page="39" pageId="19" pageNumber="162" score="67.94127085788433" volume="94" year="1947">
<taxonomicName id="4C084D2FFFCDFF9CFF03FBAF1CEF5243" ID-CoL="4CD83" authority="A. Camus" authorityName="A. Camus" box="[189,487,1068,1092]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="19" pageNumber="162" phylum="Tracheophyta" rank="species" species="calcicola">
<originalDetermination id="A8168E50FFCDFF9CFF03FBAF1CEF5243" box="[189,487,1068,1092]" pageId="19" pageNumber="162">
<emphasis id="B97CEABEFFCDFF9CFF03FBAF1C7A5243" box="[189,370,1068,1092]" italics="true" pageId="19" pageNumber="162">Panicum calcicola</emphasis>
A. Camus
</originalDetermination>
</taxonomicName>
in
<bibCitation id="5E223EC0FFCDFF9CFDB2FBAE1C485263" journal="Bull. Soc. Bot. France" page="39" pageId="19" pageNumber="162" score="67.94127085788433" volume="94" year="1947">Bull. Soc. Bot. France 94: 39. 1947</bibCitation>
</treatmentCitation>
[as
<emphasis id="B97CEABEFFCDFF9CFECFFBCE1CD35263" box="[369,475,1100,1124]" italics="true" pageId="19" pageNumber="162">calcicolum</emphasis>
].
<emphasis id="B97CEABEFFCDFF9CFE4CFBCE1F795263" bold="true" box="[498,625,1100,1124]" pageId="19" pageNumber="162">Lectotypus</emphasis>
(designated here):
<emphasis id="B97CEABEFFCDFF9CFEBAFBEE1F8A5283" bold="true" box="[260,642,1132,1156]" pageId="19" pageNumber="162">
<collectingCountry id="F31F763CFFCDFF9CFEBAFBEE1CA85283" box="[260,416,1132,1156]" name="Madagascar" pageId="19" pageNumber="162">MADAGASCAR</collectingCountry>
.
<collectingRegion id="49CCF84EFFCDFF9CFE0EFBEE1F755283" box="[432,637,1132,1156]" pageId="19" pageNumber="162">Prov. Antsiranana</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BD3ACD6FFCDFF9CFD36FBEF1C4452A3" country="Madagascar" name="Montagne des Francais" pageId="19" pageNumber="162" stateProvince="Antsiranana">Montagne des Français</collectingMunicipality>
,
<date id="FFB6106CFFCDFF9CFEE4FB0E1CAF52A3" box="[346,423,1164,1188]" pageId="19" pageNumber="162" value="1923-05">
<collectingDate id="EFF2E984FFCDFF9CFEE4FB0E1CAF52A3" box="[346,423,1164,1188]" pageId="19" pageNumber="162" value="1923-05">V.1923</collectingDate>
</date>
,
<emphasis id="B97CEABEFFCDFF9CFE0BFB0F1FD052A3" box="[437,728,1164,1188]" italics="true" pageId="19" pageNumber="162">
<collectorName id="26FD537AFFCDFF9CFE0BFB0F1F8552A3" box="[437,653,1164,1188]" pageId="19" pageNumber="162">Perrier de la Bâthie</collectorName>
16209
</emphasis>
(
<collectionCode id="ED19AE69FFCDFF9CFD55FB0F1FF352A3" box="[747,763,1165,1188]" pageId="19" pageNumber="162">P</collectionCode>
[
<specimenCode id="DBAE9ED7FFCDFF9CFF79FB2E1C3552C3" box="[199,317,1196,1220]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P00450281</specimenCode>
]!;
</materialsCitation>
<materialsCitation id="3B603CF1FFCDFF9CFEE7FB2E1FF652C3" ID-GBIF-Occurrence="3472817089" box="[345,766,1195,1220]" collectionCode="P" pageId="19" pageNumber="162" specimenCode="P00450279, P00450280" specimenCount="1" typeStatus="isolectotype">
<typeStatus id="54B3880EFFCDFF9CFEE7FB2E1CBF52C3" box="[345,439,1196,1220]" pageId="19" pageNumber="162" type="isolectotype">isolectotype</typeStatus>
:
<collectionCode id="ED19AE69FFCDFF9CFE7FFB2F1CD952C3" box="[449,465,1197,1220]" pageId="19" pageNumber="162">P</collectionCode>
[
<specimenCode id="DBAE9ED7FFCDFF9CFE5FFB2E1F5152C3" box="[481,601,1196,1220]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P00450279</specimenCode>
,
<specimenCode id="DBAE9ED7FFCDFF9CFDDAFB2E1FD652C3" box="[612,734,1196,1220]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P00450280</specimenCode>
]!).
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3126527FFCDFF9CFF29FB6E1DB55163" pageId="19" pageNumber="162" type="description">
<paragraph id="8BB736ACFFCDFF9CFF29FB6E1DB55163" blockId="19.[114,764,1260,1892]" pageId="19" pageNumber="162">
Scandent perennial woody at base, rooting at nodes, to
<quantity id="4CF09B49FFCDFF9CFFCCFA8E1D935323" box="[114,155,1292,1316]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="162" unit="m" value="2.0">2 m</quantity>
long, the culms glabrous, the nodes dark and glabrous.
<emphasis id="B97CEABEFFCDFF9CFFCCFAAE1DE15343" box="[114,233,1324,1348]" italics="true" pageId="19" pageNumber="162">Leaf sheaths</emphasis>
glabrous, with bulbous-based hairs on the margins.
<emphasis id="B97CEABEFFCDFF9CFFCCFACE1DBA5363" box="[114,178,1356,1380]" italics="true" pageId="19" pageNumber="162">Ligule</emphasis>
membranous, obtuse.
<emphasis id="B97CEABEFFCDFF9CFE27FACE1F0E5363" box="[409,518,1356,1380]" italics="true" pageId="19" pageNumber="162">Leaf blades</emphasis>
lanceolate, often asymmetric at base, flat, membranous, 4– 15 ×
<quantity id="4CF09B49FFCDFF9CFD81FAEE1FAE5383" box="[575,678,1387,1412]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="19" pageNumber="162" unit="cm" value="1.75" valueMax="2.5" valueMin="1.0">1 –2.5 cm</quantity>
, drying green-brown, cross veins visible when dry, glabrous to sparsely pubescent on both sides, sometimes with bulbous based cilia on the lower part of the margin.
<emphasis id="B97CEABEFFCDFF9CFE53FA4E1F4A53E3" box="[493,578,1484,1508]" italics="true" pageId="19" pageNumber="162">Panicles</emphasis>
terminal, partly exserted, (5–)
<quantity id="4CF09B49FFCDFF9CFEBAFA6E1C665003" box="[260,366,1515,1540]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="19" pageNumber="162" unit="cm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 cm</quantity>
long, effuse, the branches dehiscent, appressed when young and becoming broadly divergent at maturity, usually with a dense cover of bulbous-based white cilia several times longer than the spikelet.
<emphasis id="B97CEABEFFCDFF9CFDF2F9CE1FAD5063" box="[588,677,1612,1636]" italics="true" pageId="19" pageNumber="162">Spikelets</emphasis>
elliptic, apically acute,
<quantity id="4CF09B49FFCDFF9CFEA6F9EE1C815083" box="[280,393,1643,1668]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="19" pageNumber="162" unit="mm" value="1.2" valueMax="1.4" valueMin="1.0">1 – 1.4 mm</quantity>
long, brown, never gaping open.
<emphasis id="B97CEABEFFCDFF9CFFCCF90F1DFF50A3" box="[114,247,1676,1700]" italics="true" pageId="19" pageNumber="162">Lower glume</emphasis>
c. 2/3 as long as the spikelet, membranous, acute, 3-veined, with small prickle hairs.
<emphasis id="B97CEABEFFCDFF9CFE65F92F1F5350C3" box="[475,603,1708,1732]" italics="true" pageId="19" pageNumber="162">Upper glume</emphasis>
usually as long as the spikelet, sometimes 2/3–3/4 as long, herbaceous, (3–)5- veined, with small prickle hairs, dehiscing early.
<emphasis id="B97CEABEFFCDFF9CFDC3F96F1FF35103" box="[637,763,1772,1796]" italics="true" pageId="19" pageNumber="162">Lower floret</emphasis>
barren, without a significant palea.
<emphasis id="B97CEABEFFCDFF9CFE52F88F1F715123" box="[492,633,1804,1828]" italics="true" pageId="19" pageNumber="162">Lower lemma</emphasis>
herbaceous, 5-veined, with small prickle hairs.
<emphasis id="B97CEABEFFCDFF9CFE67F8AF1F685143" box="[473,608,1836,1860]" italics="true" pageId="19" pageNumber="162">Upper lemma</emphasis>
smooth, shiny, brown.
</paragraph>
</subSubSection>
<subSubSection id="C3126527FFCDFF9CFCF9FE9519745788" pageId="19" pageNumber="162" type="distribution">
<paragraph id="8BB736ACFFCDFF9CFCF9FE9519745788" blockId="19.[801,1452,279,399]" pageId="19" pageNumber="162">
<emphasis id="B97CEABEFFCDFF9CFCF9FE9519335728" box="[839,1083,279,303]" italics="true" pageId="19" pageNumber="162">Distribution and ecology</emphasis>
. – Endemic to northern
<collectingCountry id="F31F763CFFCDFF9CFAFFFE951E485749" name="Madagascar" pageId="19" pageNumber="162">Madagascar</collectingCountry>
seasonally dry forest on limestone and gneiss,
<quantity id="4CF09B49FFCDFF9CFA99FEB518A45749" box="[1319,1452,310,335]" metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="12.0" metricValueMin="2.5" pageId="19" pageNumber="162" unit="m" value="725.0" valueMax="1200.0" valueMin="250.0">250–1200 m</quantity>
(
<figureCitation id="13332A29FFCDFF9CFC95FED51E755768" box="[811,893,343,367]" captionStart="Fig" captionStartId="17.[113,141,1193,1212]" captionTargetBox="[133,1420,284,1164]" captionTargetId="figure-428@17.[128,784,207,1210]" captionTargetPageId="17" captionText="Fig. 8. – Distribution maps. A. Panicum cupressifolium A. Camus (stars), P. danguyi A. Camus (triangles), P. dregeanum Nees (circles) and P. flacourtii A. Camus (squares); B. Panicum humbertii A. Camus (stars), P. humile Nees ex Steud. (triangles), P. ibitense A. Camus (circles) and P. inconspicuum Voronts. (squares)." figureDoi="http://doi.org/10.5281/zenodo.5724530" httpUri="https://zenodo.org/record/5724530/files/figure.png" pageId="19" pageNumber="162">Fig. 8B</figureCitation>
). Seems to be rare; an expedition by the author in 2015 failed to locate this species.
</paragraph>
</subSubSection>
<subSubSection id="C3126527FFCDFF9CFCF9FE3A181755A8" pageId="19" pageNumber="162" type="discussion">
<paragraph id="8BB736ACFFCDFF9CFCF9FE3A19165568" blockId="19.[801,1452,439,943]" pageId="19" pageNumber="162">
<emphasis id="B97CEABEFFCDFF9CFCF9FE3A1E7657C8" box="[839,894,440,463]" italics="true" pageId="19" pageNumber="162">Notes</emphasis>
. – This species has dehiscent inflorescence branches, a rare character in grasses. Panicle branches are appressed to the axis on emergence. As the inflorescence ages the upper branches of the inflorescence dehisce, leaving broadly divergent lower inflorescence branches. Hirsute inflorescence branches can form a tangled mass
<emphasis id="B97CEABEFFCDFF9CFB94FDD418A3546A" box="[1066,1451,598,623]" italics="true" pageId="19" pageNumber="162">(Wohlhauser &amp; Andriamalaza 60285)</emphasis>
which may attach itself to passing animals. Spikelets are similar to
<taxonomicName id="4C084D2FFFCDFF9CFC33FD15182A54A8" authority="(Lam.) Clayton" authorityName="Clayton" baseAuthorityName="Lam." box="[909,1314,661,687]" class="Liliopsida" family="Poaceae" genus="Cyrtococcum" kingdom="Plantae" order="Poales" pageId="19" pageNumber="162" phylum="Tracheophyta" rank="species" species="multinode">
<emphasis id="B97CEABEFFCDFF9CFC33FD15197B54A8" box="[909,1139,663,687]" italics="true" pageId="19" pageNumber="162">Cyrtococcum multinode</emphasis>
(Lam.) Clayton
</taxonomicName>
but not dorsally compressed, the spikelets are single, and panicle branches almost always have long white hairs.
<taxonomicName id="4C084D2FFFCDFF9CFB02FD5A187654E8" box="[1212,1406,727,751]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="19" pageNumber="162" phylum="Tracheophyta" rank="species" species="muscicola">
<emphasis id="B97CEABEFFCDFF9CFB02FD5A187654E8" box="[1212,1406,727,751]" italics="true" pageId="19" pageNumber="162">Panicum muscicola</emphasis>
</taxonomicName>
is a variant with no trichomes in the inflorescence. Inflorescences of
<taxonomicName id="4C084D2FFFCDFF9CFC83FC9A1EBD5528" box="[829,949,791,815]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="19" pageNumber="162" phylum="Tracheophyta" rank="species" species="humbertii">
<emphasis id="B97CEABEFFCDFF9CFC83FC9A1EBD5528" box="[829,949,791,815]" italics="true" pageId="19" pageNumber="162">P. humbertii</emphasis>
</taxonomicName>
are at least
<quantity id="4CF09B49FFCDFF9CFB91FC9519665529" box="[1071,1134,791,815]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="162" unit="cm" value="10.0">10 cm</quantity>
long; a single collection
<emphasis id="B97CEABEFFCDFF9CFAD1FC9A1E6B5548" italics="true" pageId="19" pageNumber="162">Bosser 20382</emphasis>
has inflorescences c.
<quantity id="4CF09B49FFCDFF9CFBFBFCB5197F5549" box="[1093,1143,823,847]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="162" unit="cm" value="5.0">5 cm</quantity>
long, likely associated with a particularly dry habitat.
</paragraph>
<paragraph id="8BB736ACFFCDFF9CFCF9FCF5181755A8" blockId="19.[801,1452,439,943]" pageId="19" pageNumber="162">
The
<typeStatus id="54B3880EFFCDFF9CFCC9FCF51EEB5588" box="[887,995,887,911]" pageId="19" pageNumber="162" type="lectotype">lectotypes</typeStatus>
of
<taxonomicName id="4C084D2FFFCDFF9CFBB6FCFA198E5588" box="[1032,1158,887,911]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="19" pageNumber="162" phylum="Tracheophyta" rank="species" species="humbertii">
<emphasis id="B97CEABEFFCDFF9CFBB6FCFA198E5588" box="[1032,1158,887,911]" italics="true" pageId="19" pageNumber="162">P. humbertii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C084D2FFFCDFF9CFB2CFCFA18035588" box="[1170,1291,887,911]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="19" pageNumber="162" phylum="Tracheophyta" rank="species" species="muscicola">
<emphasis id="B97CEABEFFCDFF9CFB2CFCFA18035588" box="[1170,1291,887,911]" italics="true" pageId="19" pageNumber="162">P. muscicola</emphasis>
</taxonomicName>
and
<taxonomicName id="4C084D2FFFCDFF9CFAFFFCFA18A45588" box="[1345,1452,887,911]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="19" pageNumber="162" phylum="Tracheophyta" rank="species" species="calcicola">
<emphasis id="B97CEABEFFCDFF9CFAFFFCFA18A45588" box="[1345,1452,887,911]" italics="true" pageId="19" pageNumber="162">P. calcicola</emphasis>
</taxonomicName>
are chosen for their superior flowering material.
</paragraph>
</subSubSection>
<subSubSection id="C3126527FFCDFF9CFCF9FC57193E5309" pageId="19" pageNumber="162" type="materials_examined">
<paragraph id="8BB736ACFFCDFF9CFCF9FC57193E5309" blockId="19.[801,1452,981,1294]" pageId="19" pageNumber="162">
<emphasis id="B97CEABEFFCDFF9CFCF9FC57195855EF" box="[839,1104,981,1000]" italics="true" pageId="19" pageNumber="162">Additional material examined</emphasis>
. –
<materialsCitation id="3B603CF1FFCDFF9CFBCEFC5718705205" ID-GBIF-Occurrence="3472816317" collectingDate="1953-07" collectionCode="P, TAN" collectorName="Bosser" country="Madagascar" location="Montagne d'Ambre" pageId="19" pageNumber="162" specimenCode="P02608857" specimenCount="1" stateProvince="Prov. Antsiranana">
<emphasis id="B97CEABEFFCDFF9CFBCEFC5718A355EF" bold="true" box="[1136,1451,981,1000]" pageId="19" pageNumber="162">
<collectingCountry id="F31F763CFFCDFF9CFBCEFC5719FB55EF" box="[1136,1267,981,1000]" name="Madagascar" pageId="19" pageNumber="162">MADAGASCAR</collectingCountry>
.
<collectingRegion id="49CCF84EFFCDFF9CFB42FC5718AF55EF" box="[1276,1447,981,1000]" pageId="19" pageNumber="162">Prov. Antsiranana</collectingRegion>
:
</emphasis>
<location id="8ED76077FFCDFF9CFC9FFC721EDE5204" LSID="urn:lsid:plazi:treatment:03A187BAFFCDFF9CFFCFFD4E193E5309:8ED76077FFCDFF9CFC9FFC721EDE5204" box="[801,982,1008,1027]" country="Madagascar" name="Montagne d'Ambre" pageId="19" pageNumber="162" stateProvince="Prov. Antsiranana">Montagne d’Ambre</location>
,
<date id="FFB6106CFFCDFF9CFC5FFC7219275204" box="[993,1071,1008,1027]" pageId="19" pageNumber="162" value="1953-07">
<collectingDate id="EFF2E984FFCDFF9CFC5FFC7219275204" box="[993,1071,1008,1027]" pageId="19" pageNumber="162" value="1953-07">VII.1953</collectingDate>
</date>
,
<emphasis id="B97CEABEFFCDFF9CFB84FC7219A85204" box="[1082,1184,1008,1027]" italics="true" pageId="19" pageNumber="162">
<collectorName id="26FD537AFFCDFF9CFB84FC7219665205" box="[1082,1134,1008,1026]" pageId="19" pageNumber="162">Bosser</collectorName>
5342
</emphasis>
(
<collectionCode id="ED19AE69FFCDFF9CFB0EFC7219B55204" box="[1200,1213,1008,1027]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBAE9ED7FFCDFF9CFB72FC7218275204" box="[1228,1327,1008,1027]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P02608857</specimenCode>
],
<collectionCode id="ED19AE69FFCDFF9CFAFCFC7218795204" box="[1346,1393,1008,1027]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="19" pageNumber="162" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B603CF1FFCDFF9CFA3BFC72199F521B" ID-GBIF-Occurrence="3472816343" collectingDate="1953-07" collectionCode="P" collectorName="Bosser" country="Madagascar" location="Montagne d'Ambre" pageId="19" pageNumber="162" specimenCode="P02263027" specimenCount="1" stateProvince="Prov.">
<emphasis id="B97CEABEFFCDFF9CFA3BFC721E36521A" italics="true" pageId="19" pageNumber="162">ibid. loco</emphasis>
,
<date id="FFB6106CFFCDFF9CFCF6FB881E9C521A" box="[840,916,1034,1053]" pageId="19" pageNumber="162" value="1953-07">
<collectingDate id="EFF2E984FFCDFF9CFCF6FB881E9C521A" box="[840,916,1034,1053]" pageId="19" pageNumber="162" value="1953-07">VII.1953</collectingDate>
</date>
,
<emphasis id="B97CEABEFFCDFF9CFC20FB891EF75219" box="[926,1023,1035,1054]" italics="true" pageId="19" pageNumber="162">
<collectorName id="26FD537AFFCDFF9CFC20FB891ED8521A" box="[926,976,1035,1053]" pageId="19" pageNumber="162">Bosser</collectorName>
5379
</emphasis>
(
<collectionCode id="ED19AE69FFCDFF9CFBB2FB8919115219" box="[1036,1049,1035,1054]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBAE9ED7FFCDFF9CFB99FB881980521A" box="[1063,1160,1034,1053]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P02263027</specimenCode>
])
</materialsCitation>
;
<materialsCitation id="3B603CF1FFCDFF9CFB1CFB8819575230" ID-GBIF-Occurrence="3472816949" collectingDate="1953-07" collectionCode="K, P" collectorName="Bosser" country="Madagascar" location="Montagne d'Ambre" pageId="19" pageNumber="162" specimenCode="K000805468, P02222308" specimenCount="1" stateProvince="Prov.">
<emphasis id="B97CEABEFFCDFF9CFB1CFB8819E2521A" box="[1186,1258,1034,1053]" italics="true" pageId="19" pageNumber="162">ibid. loco</emphasis>
,
<date id="FFB6106CFFCDFF9CFB4AFB881848521A" box="[1268,1344,1034,1053]" pageId="19" pageNumber="162" value="1953-07">
<collectingDate id="EFF2E984FFCDFF9CFB4AFB881848521A" box="[1268,1344,1034,1053]" pageId="19" pageNumber="162" value="1953-07">VII.1953</collectingDate>
</date>
,
<emphasis id="B97CEABEFFCDFF9CFAF4FB8918A35219" box="[1354,1451,1035,1054]" italics="true" pageId="19" pageNumber="162">
<collectorName id="26FD537AFFCDFF9CFAF4FB891874521A" box="[1354,1404,1035,1053]" pageId="19" pageNumber="162">Bosser</collectorName>
5543
</emphasis>
(
<collectionCode id="ED19AE69FFCDFF9CFC97FBA41E30523E" box="[809,824,1062,1081]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="19" pageNumber="162" type="Herbarium">K</collectionCode>
[
<specimenCode id="DBAE9ED7FFCDFF9CFCF6FBA71EB3523F" box="[840,955,1061,1080]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="19" pageNumber="162" type="Herbarium">K000805468</specimenCode>
],
<collectionCode id="ED19AE69FFCDFF9CFC71FBA41ED5523E" box="[975,989,1062,1081]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBAE9ED7FFCDFF9CFC55FBA71958523F" box="[1003,1104,1061,1080]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P02222308</specimenCode>
])
</materialsCitation>
;
<materialsCitation id="3B603CF1FFCDFF9CFBD2FBA419355255" ID-GBIF-Occurrence="3472817255" collectingDate="1970-04-13" collectionCode="P" collectorName="Bosser" country="Madagascar" location="montagne des Francais" pageId="19" pageNumber="162" specimenCode="P06795924" specimenCount="1" stateProvince="Prov.">
<location id="8ED76077FFCDFF9CFBD2FBA41837523F" LSID="urn:lsid:plazi:treatment:03A187BAFFCDFF9CFFCFFD4E193E5309:8ED76077FFCDFF9CFBD2FBA41837523F" box="[1132,1343,1061,1081]" country="Madagascar" name="montagne des Francais" pageId="19" pageNumber="162" stateProvince="Prov.">montagne des Francais</location>
,
<date id="FFB6106CFFCDFF9CFAF7FBA718AF523F" box="[1353,1447,1061,1080]" pageId="19" pageNumber="162" value="1970-04-13">
<collectingDate id="EFF2E984FFCDFF9CFAF7FBA718AF523F" box="[1353,1447,1061,1080]" pageId="19" pageNumber="162" value="1970-04-13">13.IV.1970</collectingDate>
</date>
,
<emphasis id="B97CEABEFFCDFF9CFC9FFBC21E9E5254" box="[801,918,1088,1107]" italics="true" pageId="19" pageNumber="162">
<collectorName id="26FD537AFFCDFF9CFC9FFBC21E5F5255" box="[801,855,1088,1106]" pageId="19" pageNumber="162">Bosser</collectorName>
20162
</emphasis>
(
<collectionCode id="ED19AE69FFCDFF9CFC19FBC21EBC5254" box="[935,948,1088,1107]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBAE9ED7FFCDFF9CFC78FBC219245254" box="[966,1068,1088,1107]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P06795924</specimenCode>
])
</materialsCitation>
;
<materialsCitation id="3B603CF1FFCDFF9CFBF5FBC21E9F526B" ID-GBIF-Occurrence="3472816964" collectingDate="1970-04-13" collectionCode="P" collectorName="Bosser" country="Madagascar" location="Francais" pageId="19" pageNumber="162" specimenCode="P06795925" specimenCount="1" stateProvince="Prov.">
<emphasis id="B97CEABEFFCDFF9CFBF5FBC219935254" box="[1099,1179,1088,1107]" italics="true" pageId="19" pageNumber="162">ibid. loco</emphasis>
,
<date id="FFB6106CFFCDFF9CFB17FBC218045254" box="[1193,1292,1088,1107]" pageId="19" pageNumber="162" value="1970-04-13">
<collectingDate id="EFF2E984FFCDFF9CFB17FBC218045254" box="[1193,1292,1088,1107]" pageId="19" pageNumber="162" value="1970-04-13">13.IV.1970</collectingDate>
</date>
,
<emphasis id="B97CEABEFFCDFF9CFAA7FBC218865254" box="[1305,1422,1088,1107]" italics="true" pageId="19" pageNumber="162">
<collectorName id="26FD537AFFCDFF9CFAA7FBC218475255" box="[1305,1359,1088,1106]" pageId="19" pageNumber="162">Bosser</collectorName>
20169
</emphasis>
(
<collectionCode id="ED19AE69FFCDFF9CFA20FBC218A35254" box="[1438,1451,1088,1107]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBAE9ED7FFCDFF9CFC97FBD81E80526A" box="[809,904,1114,1133]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P06795925</specimenCode>
])
</materialsCitation>
;
<materialsCitation id="3B603CF1FFCDFF9CFC1DFBD818AE526B" ID-GBIF-Occurrence="3472816834" box="[931,1446,1113,1134]" collectingDate="1970-06" collectionCode="P" collectorName="Bosser" country="Madagascar" location="foret de Sahafary" pageId="19" pageNumber="162" specimenCode="P02222320" specimenCount="1" stateProvince="Prov.">
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<collectorName id="26FD537AFFCDFF9CFAE5FB291E6252DF" pageId="19" pageNumber="162">Perrier de la Bâthie</collectorName>
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<collectorName id="26FD537AFFCDFF9CFB7BFB62184152F4" box="[1221,1353,1248,1267]" pageId="19" pageNumber="162">Andriamalaza</collectorName>
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[
<specimenCode id="DBAE9ED7FFCDFF9CFC7EFB79192B5309" box="[960,1059,1275,1294]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="162" type="Herbarium">P04430800</specimenCode>
])
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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