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<document id="8742F9F6A5DB77C0EB536C2B6300EFFF" ID-CLB-Dataset="148081" ID-DOI="10.11646/zootaxa.5188.6.2" ID-GBIF-Dataset="ecb5de27-f5fe-4b16-903b-633548186b0a" ID-ISSN="1175-5326" ID-Zenodo-Dep="7103618" ID-ZooBank="BF01B6B4-3415-41A1-86A0-F4B187541A55" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1663829136880" checkinUser="plazi" docAuthor="Brooks, Scott E. &amp; Cumming, Jeffrey M." docDate="2022" docId="038D87B8C813200AEBC3E2A29FA8AF13" docLanguage="en" docName="zootaxa.5186.6.2.pdf" docOrigin="Zootaxa 5188 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chimerothalassius Shamshev &amp; Grootaert 2003" docType="treatment" docVersion="7" lastPageNumber="524" masterDocId="FFB4FFC0C8102009EB54E27D9F48AB33" masterDocTitle="New Australasian Parathalassiinae (Diptera: Dolichopodidae sensu lato)" masterLastPageNumber="543" masterPageNumber="521" pageNumber="524" updateTime="1699383858559" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C9DABB0EAA245CF4C320451CB7E833D5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BFC5667622177B9C019ACA731138F929">
<mods:title id="DB598AD6C148DE9010AEB5AD5071FE46">New Australasian Parathalassiinae (Diptera: Dolichopodidae sensu lato)</mods:title>
</mods:titleInfo>
<mods:name id="95608816E23FBA812479E4D080544609" type="personal">
<mods:role id="143DAAA0CF11B18EA4BDC5AED15F060A">
<mods:roleTerm id="727770FD4307DDCC7F4FB7A4F91B88E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="986A1162B3DD82DF9167F90799F17D6A">Brooks, Scott E.</mods:namePart>
<mods:nameIdentifier id="9317AE057180621B6FEA62CD9D64BAB1" type="ORCID">0000-0001-7915-1941</mods:nameIdentifier>
<mods:affiliation id="D065E85DF33DEFD4F91B4A16F6F7B880">Diptera Unit, Canadian National Collection of Insects, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, Ontario, K 1 A 0 C 6, CANADA &amp; scott. brooks @ agr. gc. ca; https: // orcid. org / 0000 - 0001 - 7915 - 1941</mods:affiliation>
<mods:nameIdentifier id="AF48AC9CB9D613C4444D5CA01A2BDA65" type="email">scott.brooks@agr.gc.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="6097200505F5B1B6B67B32548A403361" type="personal">
<mods:role id="2B8D95F442D7F2D63CE49B744EB3972E">
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<mods:namePart id="A0839C8DEF31437EB40C82CC31A0FA2E">Cumming, Jeffrey M.</mods:namePart>
<mods:nameIdentifier id="92078459606EBDB3A019BE36D47AD6D4" type="ORCID">0000-0002-3619-3700</mods:nameIdentifier>
<mods:affiliation id="CAA6F90C5D84CFEA8C10A10B7D1F650A">Diptera Unit, Canadian National Collection of Insects, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, Ontario, K 1 A 0 C 6, CANADA &amp; jeff. cumming @ agr. gc. ca; https: // orcid. org / 0000 - 0002 - 3619 - 3700</mods:affiliation>
<mods:nameIdentifier id="5C32C5AA2D7E88293BBED33FC6A72D68" type="email">jeff.cumming@agr.gc.ca</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="3337136E402FA09A447F8AC0EB097A1D">text</mods:typeOfResource>
<mods:relatedItem id="067E928EA6627A2B1ECD7C19A022257B" type="host">
<mods:titleInfo id="234A958985DFB02A57E660890A11C368">
<mods:title id="D99D8E0E333CDA8058E31836BC032C0C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B1F663E00BEA5F89C2F8CC81F4F62D2F">
<mods:date id="845B0F4162F8A99AF2A33BACC2158F27">2022</mods:date>
<mods:detail id="FEA7B9D3D368AFAED60B844F74B0A951" type="pubDate">
<mods:number id="266069FA132B83E6E4D05C62E5EC796A">2022-09-22</mods:number>
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<mods:detail id="E899E61BF20F85B7B2099CBC56B0904A" type="volume">
<mods:number id="D89D0E588834E90443476F5B567B2DCE">5188</mods:number>
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<mods:detail id="D470649CB7DB7E7F11B73BE4001394F9" type="issue">
<mods:number id="6458DC1B0E9F0CFE5CDDA30D0739F89C">6</mods:number>
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<mods:classification id="F7E5280C3BE6FC78DD1517557EDAB352">journal article</mods:classification>
<mods:identifier id="C8D171739D8F1E5658CCCCEEC36BDB7D" type="CLB-Dataset">148081</mods:identifier>
<mods:identifier id="94695BD186FF401F5F913C58EE65AFD6" type="DOI">10.11646/zootaxa.5188.6.2</mods:identifier>
<mods:identifier id="C7448D6A110610CE27A0AFA0C558B10A" type="GBIF-Dataset">ecb5de27-f5fe-4b16-903b-633548186b0a</mods:identifier>
<mods:identifier id="786985B23F9A6AFB56DCDB2904458256" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="50479F6FC64117FB4AD5FA8D29D1340B" type="Zenodo-Dep">7103618</mods:identifier>
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<treatment id="038D87B8C813200AEBC3E2A29FA8AF13" ID-DOI="http://doi.org/10.5281/zenodo.7103614" ID-GBIF-Taxon="201105645" ID-Zenodo-Dep="7103614" LSID="urn:lsid:plazi:treatment:038D87B8C813200AEBC3E2A29FA8AF13" httpUri="http://treatment.plazi.org/id/038D87B8C813200AEBC3E2A29FA8AF13" lastPageNumber="524" pageId="3" pageNumber="524">
<subSubSection id="C33E6525C813200AEBC3E2A29DECABCA" box="[151,676,223,250]" pageId="3" pageNumber="524" type="nomenclature">
<paragraph id="8B9B36AEC813200AEBC3E2A29DECABCA" blockId="3.[151,676,223,250]" box="[151,676,223,250]" pageId="3" pageNumber="524">
<heading id="D0D381C2C813200AEBC3E2A29DECABCA" bold="true" box="[151,676,223,250]" fontSize="11" level="1" pageId="3" pageNumber="524" reason="1">
<taxonomicName id="4C244D2DC813200AEBC3E2A29DECABCA" ID-CoL="3N6L" authority="Shamshev &amp; Grootaert" authorityName="Shamshev &amp; Grootaert" authorityYear="2003" box="[151,676,223,250]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="B950EABCC813200AEBC3E2A29DECABCA" bold="true" box="[151,676,223,250]" pageId="3" pageNumber="524">
<emphasis id="B950EABCC813200AEBC3E2A29E32ABC9" bold="true" box="[151,378,223,250]" italics="true" pageId="3" pageNumber="524">Chimerothalassius</emphasis>
Shamshev &amp; Grootaert
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33E6525C813200AEBC3E3639D8BAA05" box="[151,707,286,310]" pageId="3" pageNumber="524" type="reference_group">
<paragraph id="8B9B36AEC813200AEBC3E3639D8BAA05" blockId="3.[151,1436,286,341]" box="[151,707,286,310]" pageId="3" pageNumber="524">
<treatmentCitationGroup id="AB341180C813200AEBC3E3639D8BAA05" box="[151,707,286,310]" pageId="3" pageNumber="524">
<treatmentCitation id="0A8510BFC813200AEBC3E3639D88AA05" author="Shamshev, I. V. &amp; Grootaert, P." box="[151,704,286,310]" page="131" pageId="3" pageNumber="524" year="2002">
<taxonomicName id="4C244D2DC813200AEBC3E3639D88AA05" ID-CoL="3N6L" authority="Shamshev &amp; Grootaert, 2002: 131" authorityName="Shamshev &amp; Grootaert" authorityPageNumber="131" authorityYear="2003" box="[151,704,286,310]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="B950EABCC813200AEBC3E3639E1CAA05" box="[151,340,286,310]" italics="true" pageId="3" pageNumber="524">Chimerothalassius</emphasis>
<bibRefCitation id="EFB54B5FC813200AEA08E3639D88AA05" author="Shamshev, I. V. &amp; Grootaert, P." box="[348,704,286,310]" pageId="3" pageNumber="524" pagination="129 - 144" refId="ref15415" refString="Shamshev, I. V. &amp; Grootaert, P. (2002) A new genus of Microphorinae (Diptera Empidoidea) from New Zealand. Belgian Journal of Entomology, 4, 129 - 144." type="journal article" year="2002">Shamshev &amp; Grootaert, 2002: 131</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33E6525C813200AE99EE3629EF0AA66" pageId="3" pageNumber="524" type="type_taxon">
<paragraph id="8B9B36AEC813200AE99EE3629EF0AA66" blockId="3.[151,1436,286,341]" pageId="3" pageNumber="524">
<typeStatus id="549F880CC813200AE99EE3629DB4AA04" box="[714,764,287,311]" pageId="3" pageNumber="524">Type</typeStatus>
species:
<taxonomicName id="4C244D2DC813200AE80FE3639AD0AA05" ID-CoL="5Y42J" authority="Shamshev &amp; Grootaert, 2002" authorityName="Shamshev &amp; Grootaert" authorityYear="2003" box="[859,1432,286,310]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="ismayi">
<emphasis id="B950EABCC813200AE80FE3639B2AAA05" box="[859,1122,286,310]" italics="true" pageId="3" pageNumber="524">Chimerothalassius ismayi</emphasis>
<bibRefCitation id="EFB54B5FC813200AEF3EE3639AD0AA05" author="Shamshev, I. V. &amp; Grootaert, P." box="[1130,1432,286,310]" pageId="3" pageNumber="524" pagination="129 - 144" refId="ref15415" refString="Shamshev, I. V. &amp; Grootaert, P. (2002) A new genus of Microphorinae (Diptera Empidoidea) from New Zealand. Belgian Journal of Entomology, 4, 129 - 144." type="journal article" year="2002">Shamshev &amp; Grootaert, 2002</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C33E6525C813200AEBC3E3009DAAA85F" pageId="3" pageNumber="524" type="diagnosis">
<paragraph id="8B9B36AEC813200AEBC3E3009DAAA85F" blockId="3.[151,1437,381,1056]" pageId="3" pageNumber="524">
<emphasis id="B950EABCC813200AEBC3E3009E5AAAA4" bold="true" box="[151,274,381,407]" pageId="3" pageNumber="524">Diagnosis.</emphasis>
The genus
<taxonomicName id="4C244D2DC813200AEAC0E3039D2DAAAB" authorityName="Shamshev &amp; Grootaert" authorityYear="2003" box="[404,613,382,408]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="B950EABCC813200AEAC0E3039D2DAAAB" box="[404,613,382,408]" italics="true" pageId="3" pageNumber="524">Chimerothalassius</emphasis>
</taxonomicName>
is distinguished from other parathalassiine genera by the following characters: head with gena scarcely projected below eye (
<figureCitation id="131F2A2BC813200AE9A9E3DF9CCAAA8F" box="[765,898,418,444]" captionStart="FIGURES 12–15" captionStartId="4.[151,264,1842,1866]" captionTargetBox="[151,1436,788,1809]" captionTargetId="figure-325@4.[151,1436,777,1819]" captionTargetPageId="4" captionText="FIGURES 12–15. Chimerothalassius ismayi Shamshev &amp; Grootaert and C. marshalli sp. nov. 12. Male of C. ismayi Shamshev &amp; Grootaert from Port Levy, New Zealand (debu00102042), left lateral view. 13. Female of C. ismayi Shamshev &amp; Grootaert from Cable Bay, New Zealand (USNM ENT 00085568), head and foreleg. 14. Male holotype of C. marshalli sp. nov., left lateral view of head and thorax. 15. Male holotype of C. marshalli sp. nov., dorsal view of head and thorax. Abbreviation: plp—palpus." figureDoi="http://doi.org/10.5281/zenodo.7103630" httpUri="https://zenodo.org/record/7103630/files/figure.png" pageId="3" pageNumber="524">Figs 12, 14</figureCitation>
,
<figureCitation id="131F2A2BC813200AE8C4E3DF9CE7AA8F" box="[912,943,418,444]" captionStart="FIGURES 26–30" captionStartId="11.[151,264,1882,1906]" captionTargetBox="[151,1436,496,1852]" captionTargetId="figure-146@11.[151,1436,489,1859]" captionTargetPageId="11" captionText="FIGURES 26–30. Male holotype of Chimerothalassius sinclairi sp. nov. (CNC1155798). 26. Habitus. 27. Head, lateral view. 28. Head, anterior view. 29. Abdomen and terminalia, dorsal view. 30. Abdomen and terminalia, right lateral view. Photos taken prior to removal, dissection and illustration of terminalia. Abbreviation: plp—palpus." figureDoi="http://doi.org/10.5281/zenodo.7103640" httpUri="https://zenodo.org/record/7103640/files/figure.png" pageId="3" pageNumber="524">27</figureCitation>
), mouthparts directed ventrally with fleshy labellum (
<emphasis id="B950EABCC813200AEA5DE3BA9E79AAD3" box="[265,305,455,480]" italics="true" pageId="3" pageNumber="524">e.g.</emphasis>
,
<figureCitation id="131F2A2BC813200AEA6BE3BB9EDCAAD3" box="[319,404,454,480]" captionStart="FIGURES 12–15" captionStartId="4.[151,264,1842,1866]" captionTargetBox="[151,1436,788,1809]" captionTargetId="figure-325@4.[151,1436,777,1819]" captionTargetPageId="4" captionText="FIGURES 12–15. Chimerothalassius ismayi Shamshev &amp; Grootaert and C. marshalli sp. nov. 12. Male of C. ismayi Shamshev &amp; Grootaert from Port Levy, New Zealand (debu00102042), left lateral view. 13. Female of C. ismayi Shamshev &amp; Grootaert from Cable Bay, New Zealand (USNM ENT 00085568), head and foreleg. 14. Male holotype of C. marshalli sp. nov., left lateral view of head and thorax. 15. Male holotype of C. marshalli sp. nov., dorsal view of head and thorax. Abbreviation: plp—palpus." figureDoi="http://doi.org/10.5281/zenodo.7103630" httpUri="https://zenodo.org/record/7103630/files/figure.png" pageId="3" pageNumber="524">Figs 14</figureCitation>
,
<figureCitation id="131F2A2BC813200AEACBE3BB9EF6AAD3" box="[415,446,454,480]" captionStart="FIGURES 26–30" captionStartId="11.[151,264,1882,1906]" captionTargetBox="[151,1436,496,1852]" captionTargetId="figure-146@11.[151,1436,489,1859]" captionTargetPageId="11" captionText="FIGURES 26–30. Male holotype of Chimerothalassius sinclairi sp. nov. (CNC1155798). 26. Habitus. 27. Head, lateral view. 28. Head, anterior view. 29. Abdomen and terminalia, dorsal view. 30. Abdomen and terminalia, right lateral view. Photos taken prior to removal, dissection and illustration of terminalia. Abbreviation: plp—palpus." figureDoi="http://doi.org/10.5281/zenodo.7103640" httpUri="https://zenodo.org/record/7103640/files/figure.png" pageId="3" pageNumber="524">27</figureCitation>
), palpus elongate and narrow (
<figureCitation id="131F2A2BC813200AE847E3BB9CDBAAD3" box="[787,915,454,480]" captionStart="FIGURES 12–15" captionStartId="4.[151,264,1842,1866]" captionTargetBox="[151,1436,788,1809]" captionTargetId="figure-325@4.[151,1436,777,1819]" captionTargetPageId="4" captionText="FIGURES 12–15. Chimerothalassius ismayi Shamshev &amp; Grootaert and C. marshalli sp. nov. 12. Male of C. ismayi Shamshev &amp; Grootaert from Port Levy, New Zealand (debu00102042), left lateral view. 13. Female of C. ismayi Shamshev &amp; Grootaert from Cable Bay, New Zealand (USNM ENT 00085568), head and foreleg. 14. Male holotype of C. marshalli sp. nov., left lateral view of head and thorax. 15. Male holotype of C. marshalli sp. nov., dorsal view of head and thorax. Abbreviation: plp—palpus." figureDoi="http://doi.org/10.5281/zenodo.7103630" httpUri="https://zenodo.org/record/7103630/files/figure.png" pageId="3" pageNumber="524">Figs 12, 14</figureCitation>
), abruptly capitate apically and narrow basally (
<figureCitation id="131F2A2BC813200AEBCBE3979FBDA937" box="[159,245,490,516]" captionStart="FIGURES 19–22" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[219,1369,1001,1909]" captionTargetId="figure-389@8.[219,1369,993,1914]" captionTargetPageId="8" captionText="FIGURES 19–22. Chimerothalassius riparius sp. nov. 19. Head of male holotype (CNC1155797), anterior view. 20. Head of male holotype, showing left antenna in medial view. 21. Abdomen of female paratype, lateral view. 22. Right wing of male holotype, dorsal view. Abbreviations: cerc—cercus; M1, M4—medial veins; plp—palpus; R1, R2+3, R 4+5 —radial veins; tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.7103634" httpUri="https://zenodo.org/record/7103634/files/figure.png" pageId="3" pageNumber="524">Figs 19</figureCitation>
,
<figureCitation id="131F2A2BC813200AEA54E3979E00A937" box="[256,328,490,516]" captionStart="FIGURES 26–30" captionStartId="11.[151,264,1882,1906]" captionTargetBox="[151,1436,496,1852]" captionTargetId="figure-146@11.[151,1436,489,1859]" captionTargetPageId="11" captionText="FIGURES 26–30. Male holotype of Chimerothalassius sinclairi sp. nov. (CNC1155798). 26. Habitus. 27. Head, lateral view. 28. Head, anterior view. 29. Abdomen and terminalia, dorsal view. 30. Abdomen and terminalia, right lateral view. Photos taken prior to removal, dissection and illustration of terminalia. Abbreviation: plp—palpus." figureDoi="http://doi.org/10.5281/zenodo.7103640" httpUri="https://zenodo.org/record/7103640/files/figure.png" pageId="3" pageNumber="524">27, 28</figureCitation>
), or broadly subtriangular; thorax with prosternum fused to proepisternum forming precoxal bridge, scutellum with 1 pair of strong dorsally directed setae near apex (
<emphasis id="B950EABCC813200AE822E0729CD6A91B" box="[886,926,527,552]" italics="true" pageId="3" pageNumber="524">e.g.</emphasis>
,
<figureCitation id="131F2A2BC813200AE8F8E0739CB7A91B" box="[940,1023,526,552]" captionStart="FIGURES 12–15" captionStartId="4.[151,264,1842,1866]" captionTargetBox="[151,1436,788,1809]" captionTargetId="figure-325@4.[151,1436,777,1819]" captionTargetPageId="4" captionText="FIGURES 12–15. Chimerothalassius ismayi Shamshev &amp; Grootaert and C. marshalli sp. nov. 12. Male of C. ismayi Shamshev &amp; Grootaert from Port Levy, New Zealand (debu00102042), left lateral view. 13. Female of C. ismayi Shamshev &amp; Grootaert from Cable Bay, New Zealand (USNM ENT 00085568), head and foreleg. 14. Male holotype of C. marshalli sp. nov., left lateral view of head and thorax. 15. Male holotype of C. marshalli sp. nov., dorsal view of head and thorax. Abbreviation: plp—palpus." figureDoi="http://doi.org/10.5281/zenodo.7103630" httpUri="https://zenodo.org/record/7103630/files/figure.png" pageId="3" pageNumber="524">Fig. 14</figureCitation>
); legs with fore coxa lacking field of short stout spinose setae on anterior surface, tarsomere 5 of each leg with medial apical projection, although sometimes weakly developed; wing (
<figureCitation id="131F2A2BC813200AEAADE02B9D02A943" box="[505,586,598,624]" captionStart="FIGURES 19–22" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[219,1369,1001,1909]" captionTargetId="figure-389@8.[219,1369,993,1914]" captionTargetPageId="8" captionText="FIGURES 19–22. Chimerothalassius riparius sp. nov. 19. Head of male holotype (CNC1155797), anterior view. 20. Head of male holotype, showing left antenna in medial view. 21. Abdomen of female paratype, lateral view. 22. Right wing of male holotype, dorsal view. Abbreviations: cerc—cercus; M1, M4—medial veins; plp—palpus; R1, R2+3, R 4+5 —radial veins; tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.7103634" httpUri="https://zenodo.org/record/7103634/files/figure.png" pageId="3" pageNumber="524">Fig. 22</figureCitation>
) with R
<subScript id="17A034EBC813200AE9F0E0159DE4A945" attach="left" box="[676,684,616,630]" fontSize="6" pageId="3" pageNumber="524">1</subScript>
reaching costa before middle of wing, crossvein bm-m complete or incomplete, cell dm absent without veins M
<subScript id="17A034EBC813200AE92FE0F19DCBA9A9" attach="left" box="[635,643,652,666]" fontSize="6" pageId="3" pageNumber="524">2</subScript>
and dm-m, CuA rounded, cell cua convex apically, CuA+CuP absent or vestigial, anal lobe not developed; male terminalia with hypopygium small (
<figureCitation id="131F2A2BC813200AE8B0E0E39BC5A98B" box="[996,1165,670,696]" captionStart="FIGURES 26–30" captionStartId="11.[151,264,1882,1906]" captionTargetBox="[151,1436,496,1852]" captionTargetId="figure-146@11.[151,1436,489,1859]" captionTargetPageId="11" captionText="FIGURES 26–30. Male holotype of Chimerothalassius sinclairi sp. nov. (CNC1155798). 26. Habitus. 27. Head, lateral view. 28. Head, anterior view. 29. Abdomen and terminalia, dorsal view. 30. Abdomen and terminalia, right lateral view. Photos taken prior to removal, dissection and illustration of terminalia. Abbreviation: plp—palpus." figureDoi="http://doi.org/10.5281/zenodo.7103640" httpUri="https://zenodo.org/record/7103640/files/figure.png" pageId="3" pageNumber="524">Figs 26, 29, 30</figureCitation>
), right epandrial lamella usually with dorsally directed ventral process (
<figureCitation id="131F2A2BC813200AE9FAE0BF9C4FA9EF" box="[686,775,706,732]" captionStart="FIGURES 16–18" captionStartId="5.[151,264,1581,1605]" captionTargetBox="[225,1367,706,1548]" captionTargetId="figure-181@5.[205,1382,700,1557]" captionTargetPageId="5" captionText="FIGURES 16–18. Hypopygium of Chimerothalassius marshalli sp. nov., holotype (debu00102042). 16. Hypopygium, left lateral view. 17. Hypopygium, right lateral view. 18. Cerci and hypoproct, dorsal view.Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103632" httpUri="https://zenodo.org/record/7103632/files/figure.png" pageId="3" pageNumber="524">Figs 17</figureCitation>
,
<figureCitation id="131F2A2BC813200AE841E0BF9C7BA9EF" box="[789,819,706,732]" captionStart="FIGURES 23–25" captionStartId="9.[151,264,1559,1583]" captionTargetBox="[151,1426,641,1526]" captionTargetId="figure-221@9.[151,1436,632,1536]" captionTargetPageId="9" captionText="FIGURES 23–25. Hypopygium of Chimerothalassius riparius sp. nov., holotype (CNC1155797). 23. Hypopygium, left lateral view. 24. Hypopygium, right lateral view. 25. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103638" httpUri="https://zenodo.org/record/7103638/files/figure.png" pageId="3" pageNumber="524">24</figureCitation>
,
<figureCitation id="131F2A2BC813200AE815E0BF9C28A9EF" box="[833,864,706,732]" captionStart="FIGURES 31–33" captionStartId="12.[151,264,1148,1172]" captionTargetBox="[155,1432,196,1113]" captionTargetId="figure-19@12.[151,1436,181,1125]" captionTargetPageId="12" captionText="FIGURES 31–33. Hypopygium of Chimerothalassius sinclairi sp. nov., holotype (CNC1155798). 31. Hypopygium, left lateral view. 32. Hypopygium, right lateral view. 33. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103644" httpUri="https://zenodo.org/record/7103644/files/figure.png" pageId="3" pageNumber="524">32</figureCitation>
), cerci symmetrical (or nearly so) and moderately short, hypoproct projected (
<figureCitation id="131F2A2BC813200AEA84E09B9D60A833" box="[464,552,742,768]" captionStart="FIGURES 16–18" captionStartId="5.[151,264,1581,1605]" captionTargetBox="[225,1367,706,1548]" captionTargetId="figure-181@5.[205,1382,700,1557]" captionTargetPageId="5" captionText="FIGURES 16–18. Hypopygium of Chimerothalassius marshalli sp. nov., holotype (debu00102042). 16. Hypopygium, left lateral view. 17. Hypopygium, right lateral view. 18. Cerci and hypoproct, dorsal view.Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103632" httpUri="https://zenodo.org/record/7103632/files/figure.png" pageId="3" pageNumber="524">Figs 18</figureCitation>
,
<figureCitation id="131F2A2BC813200AE961E09B9D1BA832" box="[565,595,742,769]" captionStart="FIGURES 23–25" captionStartId="9.[151,264,1559,1583]" captionTargetBox="[151,1426,641,1526]" captionTargetId="figure-221@9.[151,1436,632,1536]" captionTargetPageId="9" captionText="FIGURES 23–25. Hypopygium of Chimerothalassius riparius sp. nov., holotype (CNC1155797). 23. Hypopygium, left lateral view. 24. Hypopygium, right lateral view. 25. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103638" httpUri="https://zenodo.org/record/7103638/files/figure.png" pageId="3" pageNumber="524">25</figureCitation>
,
<figureCitation id="131F2A2BC813200AE934E09B9D37A833" box="[608,639,742,768]" captionStart="FIGURES 31–33" captionStartId="12.[151,264,1148,1172]" captionTargetBox="[155,1432,196,1113]" captionTargetId="figure-19@12.[151,1436,181,1125]" captionTargetPageId="12" captionText="FIGURES 31–33. Hypopygium of Chimerothalassius sinclairi sp. nov., holotype (CNC1155798). 31. Hypopygium, left lateral view. 32. Hypopygium, right lateral view. 33. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103644" httpUri="https://zenodo.org/record/7103644/files/figure.png" pageId="3" pageNumber="524">33</figureCitation>
); female abdomen with apical segments retracted into segment 5 (
<figureCitation id="131F2A2BC813200AEE3BE09B9FFDA817" captionStart="FIGURES 19–22" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[219,1369,1001,1909]" captionTargetId="figure-389@8.[219,1369,993,1914]" captionTargetPageId="8" captionText="FIGURES 19–22. Chimerothalassius riparius sp. nov. 19. Head of male holotype (CNC1155797), anterior view. 20. Head of male holotype, showing left antenna in medial view. 21. Abdomen of female paratype, lateral view. 22. Right wing of male holotype, dorsal view. Abbreviations: cerc—cercus; M1, M4—medial veins; plp—palpus; R1, R2+3, R 4+5 —radial veins; tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.7103634" httpUri="https://zenodo.org/record/7103634/files/figure.png" pageId="3" pageNumber="524">Fig. 21</figureCitation>
), terminalia with syntergite 9+10 undivided and bearing acanthophorous setae and cercus narrowly rounded apically with prominent apical or preapical seta, or syntergite 9+10 divided and bearing acanthophorous spines and cercus pointed apically without apical seta (
<figureCitation id="131F2A2BC813200AE9D6E12F9D9DA85F" box="[642,725,850,876]" captionStart="FIGURES 19–22" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[219,1369,1001,1909]" captionTargetId="figure-389@8.[219,1369,993,1914]" captionTargetPageId="8" captionText="FIGURES 19–22. Chimerothalassius riparius sp. nov. 19. Head of male holotype (CNC1155797), anterior view. 20. Head of male holotype, showing left antenna in medial view. 21. Abdomen of female paratype, lateral view. 22. Right wing of male holotype, dorsal view. Abbreviations: cerc—cercus; M1, M4—medial veins; plp—palpus; R1, R2+3, R 4+5 —radial veins; tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.7103634" httpUri="https://zenodo.org/record/7103634/files/figure.png" pageId="3" pageNumber="524">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E6525C813200AEB93E10B9FA8AF13" pageId="3" pageNumber="524" type="discussion">
<paragraph id="8B9B36AEC813200AEB93E10B9FA8AF13" blockId="3.[151,1437,381,1056]" pageId="3" pageNumber="524">
<emphasis id="B950EABCC813200AEB93E10B9E73A8A3" bold="true" box="[199,315,886,912]" pageId="3" pageNumber="524">Remarks.</emphasis>
Species of
<taxonomicName id="4C244D2DC813200AEAEBE10B9DD8A8A3" authorityName="Shamshev &amp; Grootaert" authorityYear="2003" box="[447,656,886,912]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="B950EABCC813200AEAEBE10B9DD8A8A3" box="[447,656,886,912]" italics="true" pageId="3" pageNumber="524">Chimerothalassius</emphasis>
</taxonomicName>
are known from the Caribbean,
<collectingCountry id="F333763EC813200AEF54E10B9B35A8A3" box="[1024,1149,886,912]" name="Costa Rica" pageId="3" pageNumber="524">Costa Rica</collectingCountry>
,
<collectingCountry id="F333763EC813200AEFDDE10B9A68A8A3" box="[1161,1312,886,912]" name="New Zealand" pageId="3" pageNumber="524">New Zealand</collectingCountry>
(
<figureCitation id="131F2A2BC813200AEE7BE10B9ADCA8A3" box="[1327,1428,886,912]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–7" captionStartId-0="1.[151,264,1747,1771]" captionStartId-1="2.[151,264,785,809]" captionTargetBox-0="[151,1436,497,1719]" captionTargetBox-1="[166,1422,188,754]" captionTargetId-0="figure-163@1.[151,1436,489,1724]" captionTargetId-1="figure-19@2.[166,1422,181,761]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1–4. Known geographical distribution and collection localities of Microphorella and Chimerothalassius from Australia and New Zealand. 1. Distribution of M. bungle sp. nov., M. iota Colless and M. viticula sp. nov. 2. Govetts Leap,Australia, type locality of M. viticula sp. nov. (photo by Brad Sinclair). 3. Distribution of C. ismayi Shamshev &amp; Grootaert, C. marshalli sp. nov., Chimerothalassius sp. 1 (female from Takapuna Beach) and “Chimerothalassius” sp. 2 (female from Three Kings Is.). 4. Birdlings Flat, New Zealand, type locality of C. ismayi Shamshev &amp; Grootaert (photo by Rebecca Le Grice)." captionText-1="FIGURES 5–7. Port Levy, New Zealand, type locality of Chimerothalassius marshalli sp. nov. (photos by Morgane Merien)." figureDoi-0="http://doi.org/10.5281/zenodo.7103620" figureDoi-1="http://doi.org/10.5281/zenodo.7103624" httpUri-0="https://zenodo.org/record/7103620/files/figure.png" httpUri-1="https://zenodo.org/record/7103624/files/figure.png" pageId="3" pageNumber="524">Figs 3–7</figureCitation>
) (
<bibRefCitation id="EFB54B5FC813200AEBCBE1E79E98A887" author="Brooks, S. E. &amp; Cumming, J. M." box="[159,464,922,948]" pageId="3" pageNumber="524" pagination="511 - 523" refId="ref14887" refString="Brooks, S. E. &amp; Cumming, J. M. (2018) New species of Chimerothalassius Shamshev &amp; Grootaert (Diptera: Dolichopodidae: Parathalassiinae) from the West Indies and Costa Rica. Zootaxa, 4387 (3), 511 - 523. https: // doi. org / 10.11646 / zootaxa. 4387.3.6" type="journal article" year="2018">Brooks &amp; Cumming 2018</bibRefCitation>
) and now
<collectingCountry id="F333763EC813200AE91BE1E79C48A887" box="[591,768,922,948]" name="New Caledonia" pageId="3" pageNumber="524">New Caledonia</collectingCountry>
(
<figureCitation id="131F2A2BC813200AE845E1E79CCEA887" box="[785,902,922,948]" captionStart="FIGURES 8–11" captionStartId="2.[151,264,1908,1932]" captionTargetBox="[167,1420,842,1878]" captionTargetId="figure-39@2.[167,1420,833,1884]" captionTargetPageId="2" captionText="FIGURES 8–11. Known geographical distribution and collection localities of Chimerothalassius and Eothalassius from New Caledonia. 8. Distribution of C. riparius sp. nov., C. sinclairi sp. nov. and Eothalassius sp. 9–10. Rivère Bleue, type locality of C. riparius sp. nov. and C. sinclairi sp. nov. (photos by Brad Sinclair). 11. Plage de Poé, collection locality of Eothalassius sp. (photo by Brad Sinclair)." figureDoi="http://doi.org/10.5281/zenodo.7103626" httpUri="https://zenodo.org/record/7103626/files/figure.png" pageId="3" pageNumber="524">Figs 8–10</figureCitation>
) (
<bibRefCitation id="EFB54B5FC813200AE8F4E1E79B9AA887" author="Cumming, J. M. &amp; Brooks, S. E." box="[928,1234,922,948]" pageId="3" pageNumber="524" pagination="111 - 129" refId="ref15062" refString="Cumming, J. M. &amp; Brooks, S. E. (2019) Phylogenetic analysis and preliminary classification of the Parathalassiinae (Empidoidea: Dolichopodidae sensu lato). Zootaxa, 4648 (1), 111 - 129. https: // doi. org / 10.11646 / zootaxa. 4648.1.5" type="journal article" year="2019">Cumming &amp; Brooks 2019</bibRefCitation>
, as “Undescribed genus [
<collectingCountry id="F333763EC813200AEBB3E1C39EE9A8EB" box="[231,417,958,984]" name="New Caledonia" pageId="3" pageNumber="524">New Caledonia</collectingCountry>
]”). The genus is found on rocky, stony or sandy habitats of coastal beaches (
<figureCitation id="131F2A2BC813200AEFABE1C39A2CA8EB" box="[1279,1380,958,984]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–7" captionStartId-0="1.[151,264,1747,1771]" captionStartId-1="2.[151,264,785,809]" captionTargetBox-0="[151,1436,497,1719]" captionTargetBox-1="[166,1422,188,754]" captionTargetId-0="figure-163@1.[151,1436,489,1724]" captionTargetId-1="figure-19@2.[166,1422,181,761]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1–4. Known geographical distribution and collection localities of Microphorella and Chimerothalassius from Australia and New Zealand. 1. Distribution of M. bungle sp. nov., M. iota Colless and M. viticula sp. nov. 2. Govetts Leap,Australia, type locality of M. viticula sp. nov. (photo by Brad Sinclair). 3. Distribution of C. ismayi Shamshev &amp; Grootaert, C. marshalli sp. nov., Chimerothalassius sp. 1 (female from Takapuna Beach) and “Chimerothalassius” sp. 2 (female from Three Kings Is.). 4. Birdlings Flat, New Zealand, type locality of C. ismayi Shamshev &amp; Grootaert (photo by Rebecca Le Grice)." captionText-1="FIGURES 5–7. Port Levy, New Zealand, type locality of Chimerothalassius marshalli sp. nov. (photos by Morgane Merien)." figureDoi-0="http://doi.org/10.5281/zenodo.7103620" figureDoi-1="http://doi.org/10.5281/zenodo.7103624" httpUri-0="https://zenodo.org/record/7103620/files/figure.png" httpUri-1="https://zenodo.org/record/7103624/files/figure.png" pageId="3" pageNumber="524">Figs 4–7</figureCitation>
) and emerged rocks in rivers (
<figureCitation id="131F2A2BC813200AEAF9E19F9D56A8CF" box="[429,542,994,1020]" captionStart="FIGURES 8–11" captionStartId="2.[151,264,1908,1932]" captionTargetBox="[167,1420,842,1878]" captionTargetId="figure-39@2.[167,1420,833,1884]" captionTargetPageId="2" captionText="FIGURES 8–11. Known geographical distribution and collection localities of Chimerothalassius and Eothalassius from New Caledonia. 8. Distribution of C. riparius sp. nov., C. sinclairi sp. nov. and Eothalassius sp. 9–10. Rivère Bleue, type locality of C. riparius sp. nov. and C. sinclairi sp. nov. (photos by Brad Sinclair). 11. Plage de Poé, collection locality of Eothalassius sp. (photo by Brad Sinclair)." figureDoi="http://doi.org/10.5281/zenodo.7103626" httpUri="https://zenodo.org/record/7103626/files/figure.png" pageId="3" pageNumber="524">Figs 9, 10</figureCitation>
) (
<bibRefCitation id="EFB54B5FC813200AE961E19F9C33A8CF" author="Shamshev, I. V. &amp; Grootaert, P." box="[565,891,994,1020]" pageId="3" pageNumber="524" pagination="129 - 144" refId="ref15415" refString="Shamshev, I. V. &amp; Grootaert, P. (2002) A new genus of Microphorinae (Diptera Empidoidea) from New Zealand. Belgian Journal of Entomology, 4, 129 - 144." type="journal article" year="2002">Shamshev &amp; Grootaert 2002</bibRefCitation>
;
<bibRefCitation id="EFB54B5FC813200AE8D2E19F9BE4A8CF" author="Brooks, S. E. &amp; Cumming, J. M." box="[902,1196,994,1020]" pageId="3" pageNumber="524" pagination="511 - 523" refId="ref14887" refString="Brooks, S. E. &amp; Cumming, J. M. (2018) New species of Chimerothalassius Shamshev &amp; Grootaert (Diptera: Dolichopodidae: Parathalassiinae) from the West Indies and Costa Rica. Zootaxa, 4387 (3), 511 - 523. https: // doi. org / 10.11646 / zootaxa. 4387.3.6" type="journal article" year="2018">Brooks &amp; Cumming 2018</bibRefCitation>
;
<bibRefCitation id="EFB54B5FC813200AEFECE19F9F9CAF13" author="Cumming, J. M. &amp; Brooks, S. E." pageId="3" pageNumber="524" pagination="111 - 129" refId="ref15062" refString="Cumming, J. M. &amp; Brooks, S. E. (2019) Phylogenetic analysis and preliminary classification of the Parathalassiinae (Empidoidea: Dolichopodidae sensu lato). Zootaxa, 4648 (1), 111 - 129. https: // doi. org / 10.11646 / zootaxa. 4648.1.5" type="journal article" year="2019">Cumming &amp; Brooks 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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