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<mods:title id="E571D1B62EF15B9C4DBA1F4988B58229">A review of the genus Scaphidium Oliver (Coleoptera, Staphylinidae, Scaphidiinae) from Tianmushan, East China</mods:title>
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<mods:namePart id="0F3A2A5D81BD641BF5829A9F19D00FDE">He, Wen-Jia</mods:namePart>
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Key to
<taxonomicName id="4C244D0EA672FFF78F8EC48BFDB1977F" ID-CoL="8VS34" box="[240,385,1021,1047]" class="Insecta" family="Staphylinidae" genus="Scaphidium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA9FA672FFF78F8EC48BFDB1977F" bold="true" box="[240,385,1021,1047]" italics="true" pageId="1" pageNumber="56">Scaphidium</emphasis>
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species of Tianmushan
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<keyLead id="30D595B8A672FFF78FE9C33EF9A5970A" box="[151,1429,1096,1122]" pageId="1" pageNumber="56">1 Pronotum and elytra entirely black ............................................................................................................ 2</keyLead>
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<keyLead id="30D595B8A672FFF78FE9C306F9A597E2" box="[151,1429,1136,1162]" pageId="1" pageNumber="56">- At least elytra with spots or fascia ............................................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D02E28A672FFF78FE9C3EEF9A59572" pageId="1" pageNumber="56">
<paragraph id="8B9B368DA672FFF78FE9C3EEF9A4963A" blockId="1.[151,1436,1096,2002]" pageId="1" pageNumber="56">
<keyLead id="30D595B8A672FFF78FE9C3EEF9A4963A" pageId="1" pageNumber="56">
2 BL
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. Last antennal segment symmetrical. Legs black. Male profemur curved, without microsculpture; protibia slightly expanded from basal 1/5 to median portion, gradually narrowed to apical 1/6 and then forming obtuse low protuberance subapically. (see
<figureCitation id="131F2A08A672FFF78CE7C39EFFDE966A" box="[921,1006,1256,1282]" captionStart="FIGURES 9 17" captionStartId="5.[151,269,942,966]" captionTargetBox="[153,1415,278,941]" captionTargetId="figure@5.[151,1436,248,944]" captionTargetPageId="5" captionText="FIGURES 9 17. Front (9 12) and hind (13) leg and antenna (14 17) of Scaphidium. 9, 14 — S. wuyongxiangi sp. nov.; 10, 15 — S. comes; 11, 16 — S. sinense; 12, 13, 17 — S. stigmatinotum. Scale bars 1 mm." httpUri="https://zenodo.org/record/184482/files/figure.png" pageId="1" pageNumber="56">
Fig.
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in this paper;
<figureCitation id="131F2A08A672FFF78BE1C39EF8D5966A" box="[1183,1253,1256,1282]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1405,1429]" captionTargetBox="[172,1403,199,1362]" captionTargetId="figure@3.[151,1436,172,1381]" captionTargetPageId="3" captionText="FIGURES 1 8. Habitus (1 4) and metaventrite (5 8) of Scaphidium. 1, 5 — S. wuyongxingi sp. nov.; 2, 3, 7 — S. sinense (Baishanzu (2), Tianmushan (3 )); 4, 8 — S. stigmatinotum; 6 — S. comes. Scale bars 1 mm." httpUri="https://zenodo.org/record/184481/files/figure.png" pageId="1" pageNumber="56">
Fig.
<quantity id="4CDC9B68A672FFF78BA9C39EF8D5966A" box="[1239,1253,1256,1282]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="1" pageNumber="56" unit="in" value="6.0">6</quantity>
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in
<bibRefCitation id="EFB54B7CA672FFF78A71C39EF9BF966A" author="Lobl" box="[1295,1423,1256,1282]" pageId="1" pageNumber="62" refString="Lobl, I. (1968) Decription of Scaphidium comes sp. n. and notes on some other Palaearctic species of the genus Scaphidium (Coleoptera, Scaphididae). Acta entomologica Bohemoslovaca, 65, 386 - 390." type="journal article" year="1968">Löbl 1968</bibRefCitation>
). Aedeagus
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long (see
<figureCitation id="131F2A08A672FFF78D54C266FEAA9642" box="[554,666,1296,1322]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1405,1429]" captionTargetBox="[172,1403,199,1362]" captionTargetId="figure@3.[151,1436,172,1381]" captionTargetPageId="3" captionText="FIGURES 1 8. Habitus (1 4) and metaventrite (5 8) of Scaphidium. 1, 5 — S. wuyongxingi sp. nov.; 2, 3, 7 — S. sinense (Baishanzu (2), Tianmushan (3 )); 4, 8 — S. stigmatinotum; 6 — S. comes. Scale bars 1 mm." httpUri="https://zenodo.org/record/184481/files/figure.png" pageId="1" pageNumber="56">Figs. 1, 2</figureCitation>
in
<bibRefCitation id="EFB54B7CA672FFF78DBCC266FF709642" author="Lobl" box="[706,832,1296,1322]" pageId="1" pageNumber="62" refString="Lobl, I. (1968) Decription of Scaphidium comes sp. n. and notes on some other Palaearctic species of the genus Scaphidium (Coleoptera, Scaphididae). Acta entomologica Bohemoslovaca, 65, 386 - 390." type="journal article" year="1968">Löbl 1968</bibRefCitation>
;
<figureCitation id="131F2A08A672FFF78C33C266FFA39642" box="[845,915,1296,1322]" captionStart="FIGURES 9 17" captionStartId="5.[151,269,942,966]" captionTargetBox="[153,1415,278,941]" captionTargetId="figure@5.[151,1436,248,944]" captionTargetPageId="5" captionText="FIGURES 9 17. Front (9 12) and hind (13) leg and antenna (14 17) of Scaphidium. 9, 14 — S. wuyongxiangi sp. nov.; 10, 15 — S. comes; 11, 16 — S. sinense; 12, 13, 17 — S. stigmatinotum. Scale bars 1 mm." httpUri="https://zenodo.org/record/184482/files/figure.png" pageId="1" pageNumber="56">
Fig.
<quantity id="4CDC9B68A672FFF78CFAC266FFA39642" box="[900,915,1296,1322]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="1" pageNumber="56" unit="in" value="9.0">9</quantity>
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in
<bibRefCitation id="EFB54B7CA672FFF78CC5C266F8589642" author="He" box="[955,1128,1296,1322]" pageId="1" pageNumber="62" refString="He, W. - J., Tang, L. &amp; Li, L. - Zh. (2008) Notes on the genus Scaphidium Olivier of China with description of a new species (Coleoptera: Staphylinidae: Scaphidiinae). The Entomological Review of Japan, 62 (2), 177 - 182." type="journal article" year="2008">He et al. 2008</bibRefCitation>
).
<collectingCountry id="F333761DA672FFF78B03C266F8F39642" box="[1149,1219,1296,1322]" name="China" pageId="1" pageNumber="56">China</collectingCountry>
(Zhejiang),
<collectingCountry id="F333761DA672FFF78A26C266FD3E963A" name="North Korea" pageId="1" pageNumber="56">North Korea</collectingCountry>
.....................................................................................................................................
<taxonomicName id="4C244D0EA672FFF78B91C24EF9A4963A" ID-CoL="79TR9" authority="Lobl" authorityName="Lobl" box="[1263,1428,1336,1362]" class="Insecta" family="Staphylinidae" genus="Scaphidium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="comes">
<emphasis id="B950EA9FA672FFF78B91C24EF9669639" box="[1263,1366,1336,1361]" italics="true" pageId="1" pageNumber="56">S. comes</emphasis>
Löbl
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<keyLead id="30D595B8A672FFF78FE9C216F9A59572" pageId="1" pageNumber="56">
- BL 6.0
<quantity id="4CDC9B68A672FFF78E57C216FDB79612" box="[297,391,1376,1402]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="56" unit="mm" value="9.5">9.5 mm</quantity>
. Last antennal segment asymmetrical. Meso- and metafemora with wide reddish fasciae on median portion. Male profemur slightly expanded with distinct protuberances and mesh-like microsculpture; protibia not expanded, with distinct protuberances subapically. Aedeagus
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long (see Figs. 39, 55,
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<bibRefCitation id="EFB54B7CA672FFF78ED1C2AEFE1F969A" author="Lobl" box="[431,559,1496,1522]" pageId="1" pageNumber="62" refString="Lobl, I. (1992) The Scaphidiidae (Coleoptera) of the Nepal Himalaya. Revue Suisse de Zoologie, 99 (3), 471 - 627." type="journal article" year="1992">Löbl 1992</bibRefCitation>
).
<collectingCountry id="F333761DA672FFF78D3BC2AEFEBB969A" box="[581,651,1496,1522]" name="China" pageId="1" pageNumber="56">China</collectingCountry>
(widely distributed in the south),
<collectingCountry id="F333761DA672FFF78B59C2AEF856969A" box="[1063,1126,1496,1522]" name="India" pageId="1" pageNumber="56">India</collectingCountry>
,
<collectingCountry id="F333761DA672FFF78B0AC2AEF88D969A" box="[1140,1213,1496,1522]" name="Nepal" pageId="1" pageNumber="56">Nepal</collectingCountry>
,
<collectingCountry id="F333761DA672FFF78BB5C2AEF92E969A" box="[1227,1310,1496,1522]" name="Myanmar" pageId="1" pageNumber="56">Burma</collectingCountry>
,
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,
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,
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,
<collectingCountry id="F333761DA672FFF78EF6C176FDDE9572" box="[392,494,1536,1562]" name="Vietnam" pageId="1" pageNumber="56">Vietnam</collectingCountry>
,
<collectingCountry id="F333761DA672FFF78E84C176FE5B9572" box="[506,619,1536,1562]" name="Indonesia" pageId="1" pageNumber="56">Indonesia</collectingCountry>
.................................................................................
<taxonomicName id="4C244D0EA672FFF78BB0C176F9A59572" ID-CoL="6Y5TR" authority="Gestro" authorityName="Gestro" box="[1230,1429,1536,1562]" class="Insecta" family="Staphylinidae" genus="Scaphidium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="grande">
<emphasis id="B950EA9FA672FFF78BB0C176F90F9571" box="[1230,1343,1536,1561]" italics="true" pageId="1" pageNumber="56">S. grande</emphasis>
Gestro
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<keyLead id="30D595B8A672FFF78FE9C15EF9A5952A" box="[151,1429,1576,1602]" pageId="1" pageNumber="56">3 Pronotum reddish brown, with spots or fascia ........................................................................................... 4</keyLead>
</paragraph>
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<keyLead id="30D595B8A672FFF78FE9C126F9A59502" box="[151,1429,1616,1642]" pageId="1" pageNumber="56">- Pronotum entirely black ............................................................................................................................. 5</keyLead>
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<paragraph id="8B9B368DA672FFF78FE9C10EF9A49432" blockId="1.[151,1436,1096,2002]" pageId="1" pageNumber="56">
<keyLead id="30D595B8A672FFF78FE9C10EF9A49432" pageId="1" pageNumber="56">
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4 ..... Basal portion of pronotum with black strip along antebasal puncture row; each elytron without round black spot delimited by basal fascia near scutellum (see
<figureCitation id="131F2A08A672FFF78C26C1D6FFF595D2" box="[856,965,1696,1722]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1405,1429]" captionTargetBox="[172,1403,199,1362]" captionTargetId="figure@3.[151,1436,172,1381]" captionTargetPageId="3" captionText="FIGURES 1 8. Habitus (1 4) and metaventrite (5 8) of Scaphidium. 1, 5 — S. wuyongxingi sp. nov.; 2, 3, 7 — S. sinense (Baishanzu (2), Tianmushan (3 )); 4, 8 — S. stigmatinotum; 6 — S. comes. Scale bars 1 mm." httpUri="https://zenodo.org/record/184481/files/figure.png" pageId="1" pageNumber="56">Figs. 2, 3</figureCitation>
in this paper;
<figureCitation id="131F2A08A672FFF78B12C1D6F88F95D2" box="[1132,1215,1696,1722]" captionStart="FIGURES 9 17" captionStartId="5.[151,269,942,966]" captionTargetBox="[153,1415,278,941]" captionTargetId="figure@5.[151,1436,248,944]" captionTargetPageId="5" captionText="FIGURES 9 17. Front (9 12) and hind (13) leg and antenna (14 17) of Scaphidium. 9, 14 — S. wuyongxiangi sp. nov.; 10, 15 — S. comes; 11, 16 — S. sinense; 12, 13, 17 — S. stigmatinotum. Scale bars 1 mm." httpUri="https://zenodo.org/record/184482/files/figure.png" pageId="1" pageNumber="56">
Fig.
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in
<bibRefCitation id="EFB54B7CA672FFF78B9AC1D6F95195D2" author="Lobl" box="[1252,1377,1696,1722]" pageId="1" pageNumber="62" refString="Lobl, I. (1992) The Scaphidiidae (Coleoptera) of the Nepal Himalaya. Revue Suisse de Zoologie, 99 (3), 471 - 627." type="journal article" year="1992">Löbl 1992</bibRefCitation>
;
<figureCitation id="131F2A08A672FFF78A13C1D6FCD4958A" captionStart="FIGURES 18 23" captionStartId="6.[151,269,1538,1562]" captionTargetBox="[184,1385,310,1486]" captionTargetId="figure@6.[151,1436,276,1530]" captionTargetPageId="6" captionText="FIGURES 18 23. Aedeagus (18 20; dorsal view) and details of internal sac (21 23) of Scaphidium. 18, 21 — S. wuyongxaingi sp. nov.; 19, 22 — S. sinense; 20, 23 — S. stigmatinotum. Scale bars 0.25 mm." httpUri="https://zenodo.org/record/184483/files/figure.png" pageId="1" pageNumber="56">
Fig.
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</figureCitation>
in
<bibRefCitation id="EFB54B7CA672FFF78E73C1BEFDBC958A" author="Lobl" box="[269,396,1736,1762]" pageId="1" pageNumber="62" refString="Lobl, I. (1999) A review of the Scaphidiinae (Coleoptera: Staphylinidae) of the People's Republic of the China, I. Revue Suisse de Zoologie, 106 (3), 691 - 744." type="journal article" year="1999">Löbl 1999</bibRefCitation>
). Male profemora gradually expanded from base to middle and then narrowed apically, forming two low ridges in median portion. Protibiae straight, not expanded, with low protuberance subapically. (see
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Fig.
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in this paper) Aedeagus
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long. (see
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in this paper). BL
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long.
<collectingCountry id="F333761DA672FFF78E77C036FD7F9432" box="[265,335,1856,1882]" name="China" pageId="1" pageNumber="56">China</collectingCountry>
(Fujian, Zhejiang) ...............................................................................................
<taxonomicName id="4C244D0EA672FFF78B8AC036F9A49432" authority="Pic" authorityName="Pic" box="[1268,1428,1856,1882]" class="Insecta" family="Staphylinidae" genus="Scaphidum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="B950EA9FA672FFF78B8AC036F9579431" box="[1268,1383,1856,1881]" italics="true" pageId="1" pageNumber="56">S. sinense</emphasis>
Pic
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- Basal portion of pronotum without black strip; each elytron with a round black spot delimited by basal fascia near scutellum (see
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Fig.
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in
<bibRefCitation id="EFB54B7CA672FFF78D0CC0E6FF1394C2" author="He" box="[626,803,1936,1962]" pageId="1" pageNumber="62" refString="He, W. - J., Tang, L. &amp; Li, L. - Zh. (2008) Notes on the genus Scaphidium Olivier of China with description of a new species (Coleoptera: Staphylinidae: Scaphidiinae). The Entomological Review of Japan, 62 (2), 177 - 182." type="journal article" year="2008">
He
<emphasis id="B950EA9FA672FFF78DE3C0E6FEE394C1" box="[669,723,1936,1961]" italics="true" pageId="1" pageNumber="56">et al</emphasis>
. 2008
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). Male profemora with ventral side cornuted in basal third, followed by ridge. Protibiae straight, not expanded, without subapical protuberance (see
<figureCitation id="131F2A08A672FFF78A51C0CEF94694BA" box="[1327,1398,1976,2002]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1405,1429]" captionTargetBox="[172,1403,199,1362]" captionTargetId="figure@3.[151,1436,172,1381]" captionTargetPageId="3" captionText="FIGURES 1 8. Habitus (1 4) and metaventrite (5 8) of Scaphidium. 1, 5 — S. wuyongxingi sp. nov.; 2, 3, 7 — S. sinense (Baishanzu (2), Tianmushan (3 )); 4, 8 — S. stigmatinotum; 6 — S. comes. Scale bars 1 mm." httpUri="https://zenodo.org/record/184481/files/figure.png" pageId="1" pageNumber="56">Fig. 5</figureCitation>
, in
</keyLead>
</paragraph>
<paragraph id="8B9B368DA671FFF48FB9C7EEF9A492C9" blockId="2.[151,1437,152,658]" pageId="2" pageNumber="57">
<bibRefCitation id="EFB54B7CA671FFF48FB9C7EEFD4693DA" author="He" box="[199,374,152,178]" pageId="2" pageNumber="62" refString="He, W. - J., Tang, L. &amp; Li, L. - Zh. (2008) Notes on the genus Scaphidium Olivier of China with description of a new species (Coleoptera: Staphylinidae: Scaphidiinae). The Entomological Review of Japan, 62 (2), 177 - 182." type="journal article" year="2008">He et al., 2008</bibRefCitation>
). Aedeagus (
<figureCitation id="131F2A08A671FFF48D73C7EEFE4A93DA" box="[525,634,152,178]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1405,1429]" captionTargetBox="[172,1403,199,1362]" captionTargetId="figure@3.[151,1436,172,1381]" captionTargetPageId="3" captionText="FIGURES 1 8. Habitus (1 4) and metaventrite (5 8) of Scaphidium. 1, 5 — S. wuyongxingi sp. nov.; 2, 3, 7 — S. sinense (Baishanzu (2), Tianmushan (3 )); 4, 8 — S. stigmatinotum; 6 — S. comes. Scale bars 1 mm." httpUri="https://zenodo.org/record/184481/files/figure.png" pageId="2" pageNumber="57">Figs. 7, 8</figureCitation>
in
<bibRefCitation id="EFB54B7CA671FFF48DE0C7EEFF7593DA" author="He" box="[670,837,152,178]" pageId="2" pageNumber="62" refString="He, W. - J., Tang, L. &amp; Li, L. - Zh. (2008) Notes on the genus Scaphidium Olivier of China with description of a new species (Coleoptera: Staphylinidae: Scaphidiinae). The Entomological Review of Japan, 62 (2), 177 - 182." type="journal article" year="2008">
He
<emphasis id="B950EA9FA671FFF48DB9C7EEFECA93D9" box="[711,762,152,177]" italics="true" pageId="2" pageNumber="57">et al</emphasis>
. 2008
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)
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long. BL 5.0
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.
<collectingCountry id="F333761DA671FFF48A70C7EEF96493DA" box="[1294,1364,152,178]" name="China" pageId="2" pageNumber="57">China</collectingCountry>
(Zhejiang, Anhui) ................................................................................................
<taxonomicName id="4C244D0EA671FFF48B40C7B6F9A493B2" ID-CoL="79TR5" authority="He, Tang &amp; Li" authorityName="He, Tang &amp; Li" box="[1086,1428,192,218]" class="Insecta" family="Staphylinidae" genus="Scaphidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="biwenxuani">
<emphasis id="B950EA9FA671FFF48B40C7B6F8D393B1" box="[1086,1251,192,217]" italics="true" pageId="2" pageNumber="57">S. biwenxuani</emphasis>
He, Tang &amp; Li
</taxonomicName>
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5 Pronotum with antebasal row of punctures impressed. Subbasal fascia on each elytron yellowish, small, of simple shape (see
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Fig.
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in this paper). Male: profemora expanded gradually, with two widest points. Protibiae sinuate, with angular protuberance at widest point, narrowed toward apex, followed by ridge. (see
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Fig.
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in this paper). Aedeagus (see
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Fig.
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in this paper) 2.00 mm long. BL
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(Zhejiang, Anhui) .............................................................................
<taxonomicName id="4C244D0EA671FFF48CCAC6FEF91B92CA" authority="He, Tang &amp; Li" authorityName="He, Tang &amp; Li" box="[948,1323,392,418]" class="Insecta" family="Staphylinidae" genus="Scaphidum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="wuyongxiangi" status="sp. nov.">
<emphasis id="B950EA9FA671FFF48CCAC6FEF84692C9" box="[948,1142,392,417]" italics="true" pageId="2" pageNumber="57">S. wuyongxiangi</emphasis>
He, Tang &amp; Li
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,
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<taxonomicNameLabel id="A26357E4A671FFF48A49C6F1F9A492C9" box="[1335,1428,391,417]" pageId="2" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
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</keyLead>
</paragraph>
<paragraph id="8B9B368DA671FFF48FE9C6C6F9A5929A" blockId="2.[151,1437,152,658]" pageId="2" pageNumber="57">
<keyLead id="30D595B8A671FFF48FE9C6C6F9A5929A" pageId="2" pageNumber="57">- Pronotum with antebasal row of punctures not impressed. Basal fascia on each elytron reddish, large, of complex shape ........................................................................................................................................... 6</keyLead>
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6 Basal fascia of each elytron encircling only one black spot, near humerus. (see
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Fig.
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in this paper)
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(Yunnan, Zhejiang, Jiangsu, Fujian, Hunan) ...........................................................
<taxonomicName id="4C244D0EA671FFF48BEFC55EF9A4912A" authority="Lobl" authorityName="Lobl" box="[1169,1428,552,578]" class="Insecta" family="Staphylinidae" genus="Scaphidum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="stigmatinotum">
<emphasis id="B950EA9FA671FFF48BEFC55EF9659129" box="[1169,1365,552,577]" italics="true" pageId="2" pageNumber="57">S. stigmatinotum</emphasis>
Löbl
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- Basal fascia of each elytron encircling two black spots, one on humerus and another near scutellum. (see Fig.
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<bibRefCitation id="EFB54B7CA671FFF48E41C50EFD8A91FA" author="Lobl" box="[319,442,632,658]" pageId="2" pageNumber="62" refString="Lobl, I. (1999) A review of the Scaphidiinae (Coleoptera: Staphylinidae) of the People's Republic of the China, I. Revue Suisse de Zoologie, 106 (3), 691 - 744." type="journal article" year="1999">Löbl 1999</bibRefCitation>
)
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(Fujian, Anhui) ................................................................
<taxonomicName id="4C244D0EA671FFF48BBBC50EF9A491FA" ID-CoL="6Y5SC" authority="Pic" authorityName="Pic" box="[1221,1428,632,658]" class="Insecta" family="Staphylinidae" genus="Scaphidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="B950EA9FA671FFF48BBBC50EF95791F9" box="[1221,1383,632,657]" italics="true" pageId="2" pageNumber="57">S. klapperichi</emphasis>
Pic
</taxonomicName>
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</subSubSection>
</treatment>
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