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<document id="42C00875D70E101384F17AA9319B4E77" ID-DOI="10.11646/zootaxa.3948.3.5" ID-GBIF-Dataset="48fd46c1-a333-45ef-85fd-bb89451998da" ID-ISSN="1175-5326" ID-Zenodo-Dep="241857" ID-ZooBank="809AC895-779E-419D-8EBE-071F0ACCD72E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461044448529" checkinUser="plazi" docAuthor="Desutter-Grandcolas, Laure" docDate="2015" docId="038D8783FF8D3258FF7DFAB4FC8DFE92" docLanguage="en" docName="zt03948p496.pdf" docOrigin="Zootaxa 3948 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phasmagryllus Desutter-Grandcolas" docType="treatment" docVersion="9" lastPageNumber="476" masterDocId="FFB4FFFBFF953242FFEAFFA1FF9EFF8F" masterDocTitle="Phalangopsidae crickets from Tropical Africa (Orthoptera, Grylloidea), with descriptions of new taxa and an identification key for African genera" masterLastPageNumber="496" masterPageNumber="451" pageNumber="475" updateTime="1698576427866" updateUser="plazi">
<mods:mods id="633F895BE842163108A1CE421355EE67" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F5AFC1806FE2E409F54E41DFC73A6A8D">Phalangopsidae crickets from Tropical Africa (Orthoptera, Grylloidea), with descriptions of new taxa and an identification key for African genera</mods:title>
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<mods:name id="7324A777AEA7D064EB53E24B27E20EA7" type="personal">
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<mods:roleTerm id="608BCE99DCB942A2A92B1C6B42131926">Author</mods:roleTerm>
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<mods:namePart id="0CB11F0378CA6B55DF5351E25F5F4CC7">Desutter-Grandcolas, Laure</mods:namePart>
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<mods:typeOfResource id="6BA7900CFBDFD3C20B9C2A636D83F3B8">text</mods:typeOfResource>
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<mods:title id="912CDCBC4C501E76794B096869CC5184">Zootaxa</mods:title>
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<mods:date id="24D5F715536CDB1BFA581100DD5EB58B">2015</mods:date>
<mods:detail id="D0087AB2A6C7E6465E483F2D71C90EF7" type="volume">
<mods:number id="437334B51044815F9C5C8DC86FCCC1E6">3948</mods:number>
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<mods:identifier id="EABEC50989DEB9F3C8BA66E21058ACD9" type="DOI">10.11646/zootaxa.3948.3.5</mods:identifier>
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<treatment id="038D8783FF8D3258FF7DFAB4FC8DFE92" ID-DOI="http://doi.org/10.5281/zenodo.5687705" ID-GBIF-Taxon="119610265" ID-Zenodo-Dep="5687705" LSID="urn:lsid:plazi:treatment:038D8783FF8D3258FF7DFAB4FC8DFE92" httpUri="http://treatment.plazi.org/id/038D8783FF8D3258FF7DFAB4FC8DFE92" lastPageId="26" lastPageNumber="476" pageId="24" pageNumber="475">
<subSubSection id="C33E651EFF8D325AFF7DFAB4FE86FADF" pageId="24" pageNumber="475" type="nomenclature">
<paragraph id="8B9B3695FF8D325AFF7DFAB4FC91FAA0" blockId="24.[151,783,1301,1360]" box="[151,783,1301,1327]" pageId="24" pageNumber="475">
<heading id="D0D381F9FF8D325AFF7DFAB4FC91FAA0" bold="true" box="[151,783,1301,1327]" fontSize="11" level="1" pageId="24" pageNumber="475" reason="1">
<emphasis id="B950EA87FF8D325AFF7DFAB4FC91FAA0" bold="true" box="[151,783,1301,1327]" pageId="24" pageNumber="475">
Genus
<taxonomicName id="4C244D16FF8D325AFF04FAB4FD2FFAA0" ID-CoL="8NXG7" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[238,689,1301,1327]" class="Insecta" family="Phalangopsidae" genus="Phasmagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA87FF8D325AFF04FAB4FE3DFAA0" bold="true" box="[238,419,1301,1327]" italics="true" pageId="24" pageNumber="475">Phasmagryllus</emphasis>
Desutter-Grandcolas
</taxonomicName>
, n. gen.
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B3695FF8D325AFF7DFA96FE86FADF" blockId="24.[151,783,1301,1360]" box="[151,280,1335,1360]" pageId="24" pageNumber="475">
(
<figureCitation id="131F2A10FF8D325AFF75FA96FE91FADF" box="[159,271,1335,1360]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStart-3="FIGURE 11" captionStartId-0="27.[151,250,1905,1927]" captionStartId-1="28.[151,250,1658,1680]" captionStartId-2="29.[151,250,1651,1673]" captionStartId-3="30.[151,250,1952,1974]" captionTargetBox-0="[204,1383,187,1884]" captionTargetBox-1="[153,1431,201,1629]" captionTargetBox-2="[152,1435,194,1619]" captionTargetBox-3="[227,1375,202,1930]" captionTargetId-0="figure@27.[202,1384,187,1884]" captionTargetId-1="figure@28.[151,1435,195,1637]" captionTargetId-2="figure@29.[151,1436,193,1629]" captionTargetId-3="figure@30.[211,1376,193,1931]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionTargetPageId-2="29" captionTargetPageId-3="30" captionText-0="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." captionText-1="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." captionText-2="FIGURE 10. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype, genitalia in dorsal (A), ventral (B) and lateral (C) views. Abbreviations see page 3." captionText-3="FIGURE 11. Phasmagryllus elegans Desutter-Grandcolas, n. sp., female allotype. A, dorsal view, scale 1 mm; B, lateral view; C, pronotum and FWs, dorsal view; D, ovipositor, lateral view." httpUri-0="https://zenodo.org/record/241865/files/figure.png" httpUri-1="https://zenodo.org/record/241866/files/figure.png" httpUri-2="https://zenodo.org/record/241867/files/figure.png" httpUri-3="https://zenodo.org/record/241868/files/figure.png" pageId="24" pageNumber="475">Figs 8–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33E651EFF8D325AFF7DFADEFCEBFA18" box="[151,885,1407,1432]" pageId="24" pageNumber="475" type="materials_examined">
<paragraph id="8B9B3695FF8D325AFF7DFADEFCEBFA18" blockId="24.[151,1437,1407,2009]" box="[151,885,1407,1432]" pageId="24" pageNumber="475">
<emphasis id="B950EA87FF8D325AFF7DFADEFEACFA17" bold="true" box="[151,306,1407,1432]" pageId="24" pageNumber="475">
<typeStatus id="549F8837FF8D325AFF7DFADEFF4CFA17" box="[151,210,1407,1432]" pageId="24" pageNumber="475">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C244D16FF8D325AFED3FADEFCB0FA17" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[313,814,1407,1432]" class="Insecta" family="Phalangopsidae" genus="Phasmagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="elegans" status="sp. nov.">
<emphasis id="B950EA87FF8D325AFED3FADEFDA0FA17" box="[313,574,1407,1432]" italics="true" pageId="24" pageNumber="475">Phasmagryllus elegans</emphasis>
Desutter-Grandcolas
</taxonomicName>
,
<taxonomicNameLabel id="A26357FCFF8D325AFCD3FA21FCEBFA18" box="[825,885,1408,1431]" pageId="24" pageNumber="475" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C33E651EFF8D325AFF2DFA05FE83FA50" pageId="24" pageNumber="475" type="etymology">
<paragraph id="8B9B3695FF8D325AFF2DFA05FE83FA50" blockId="24.[151,1437,1407,2009]" pageId="24" pageNumber="475">
<emphasis id="B950EA87FF8D325AFF2DFA05FED0FA32" bold="true" box="[199,334,1444,1469]" pageId="24" pageNumber="475">Etymology.</emphasis>
From
<emphasis id="B950EA87FF8D325AFE76FA05FE6DFA32" box="[412,499,1444,1469]" italics="true" pageId="24" pageNumber="475">phasma</emphasis>
, meaning ghost in Greek. Genus named after its delicate movements, and ghostly appearance.
</paragraph>
</subSubSection>
<subSubSection id="C33E651EFF8D325AFF2DFA4DFB9EF98A" box="[199,1024,1516,1541]" pageId="24" pageNumber="475" type="distribution">
<paragraph id="8B9B3695FF8D325AFF2DFA4DFB9EF98A" blockId="24.[151,1437,1407,2009]" box="[199,1024,1516,1541]" pageId="24" pageNumber="475">
<emphasis id="B950EA87FF8D325AFF2DFA4DFEFEF98A" bold="true" box="[199,352,1516,1541]" pageId="24" pageNumber="475">Distribution.</emphasis>
Known only from
<collectingCountry id="F3337605FF8D325AFDDDFA4DFD05F98A" box="[567,667,1516,1541]" name="Tanzania" pageId="24" pageNumber="475">Tanzania</collectingCountry>
(eastern Usambara mountains).
</paragraph>
</subSubSection>
<subSubSection id="C33E651EFF8D325BFF2DF9AEFC70FE93" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475" type="diagnosis">
<paragraph id="8B9B3695FF8D325BFF2DF9AEFC70FE93" blockId="24.[151,1437,1407,2009]" lastBlockId="25.[151,1437,151,2013]" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475">
<emphasis id="B950EA87FF8D325AFF2DF9AEFEDFF9A7" bold="true" box="[199,321,1551,1576]" pageId="24" pageNumber="475">Diagnosis.</emphasis>
Very small species with contrasted coloration (head orange, antennae bicolor with a wide white ring, legs and body dark brown:
<figureCitation id="131F2A10FF8D325AFDF3F995FDFAF9C3" box="[537,612,1588,1613]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="24" pageNumber="475">Figs 8</figureCitation>
,
<figureCitation id="131F2A10FF8D325AFD99F995FD0EF9C3" box="[627,656,1588,1612]" captionStart="FIGURE 11" captionStartId="30.[151,250,1952,1974]" captionTargetBox="[227,1375,202,1930]" captionTargetId="figure@30.[211,1376,193,1931]" captionTargetPageId="30" captionText="FIGURE 11. Phasmagryllus elegans Desutter-Grandcolas, n. sp., female allotype. A, dorsal view, scale 1 mm; B, lateral view; C, pronotum and FWs, dorsal view; D, ovipositor, lateral view." httpUri="https://zenodo.org/record/241868/files/figure.png" pageId="24" pageNumber="475">11</figureCitation>
). Head large, with small, not protruding eyes (
<figureCitation id="131F2A10FF8D325AFB5CF995FB62F9C3" box="[1206,1276,1588,1613]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="24" pageNumber="475">Fig. 8</figureCitation>
B), very wide fastigium (
<figureCitation id="131F2A10FF8D325AFEFAF9F6FECFF9FF" box="[272,337,1623,1648]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="24" pageNumber="475">Fig. 8</figureCitation>
B) and very elongate maxillary palpi (
<figureCitation id="131F2A10FF8D325AFD11F9F6FCDFF9FF" box="[763,833,1623,1648]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="24" pageNumber="475">Figs 8</figureCitation>
C, 9A). TI with a small, inner tympanum and no outer tympanum. TIII with three pairs of apical spurs, the upper ones the longest on both sides; four pairs of subapical spurs (
<figureCitation id="131F2A10FF8D325AFF0EF93EFEB7F937" box="[228,297,1695,1720]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="24" pageNumber="475">Fig. 8</figureCitation>
D), the first ones the smallest on both sides, the second ones the longest; these spurs alternate and almost aligned in only one row; serrulation sparse above spurs, almost lacking between spurs; all tarsomeres very long, the basitarsomeres III very elongate. FWs present in both males (
<figureCitation id="131F2A10FF8D325AFC50F946FC63F88F" box="[954,1021,1767,1792]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="24" pageNumber="475">Fig. 8</figureCitation>
) and females (
<figureCitation id="131F2A10FF8D325AFB4CF946FB66F88F" box="[1190,1272,1767,1792]" captionStart="FIGURE 11" captionStartId="30.[151,250,1952,1974]" captionTargetBox="[227,1375,202,1930]" captionTargetId="figure@30.[211,1376,193,1931]" captionTargetPageId="30" captionText="FIGURE 11. Phasmagryllus elegans Desutter-Grandcolas, n. sp., female allotype. A, dorsal view, scale 1 mm; B, lateral view; C, pronotum and FWs, dorsal view; D, ovipositor, lateral view." httpUri="https://zenodo.org/record/241868/files/figure.png" pageId="24" pageNumber="475">Fig. 11</figureCitation>
), not covering the whole abdomen, not widened in males.
<emphasis id="B950EA87FF8D325AFD6CF8ADFD5DF8AA" bold="true" box="[646,707,1804,1829]" pageId="24" pageNumber="475">Male</emphasis>
with a complete stridulum (
<figureCitation id="131F2A10FF8D325AFBEEF8ADFBD2F8AB" box="[1028,1100,1804,1829]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="24" pageNumber="475">Figs 8</figureCitation>
E, 9A): harp wider than long, crossed by few oblique parallel veins; mirror crossed by one transverse vein; a distinct transverse cell under the mirror; lateral field with few parallel veins (
<figureCitation id="131F2A10FF8D325AFD79F8F5FD46F8E3" box="[659,728,1876,1901]" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="24" pageNumber="475">Fig. 9</figureCitation>
C). Metanotum and tergites with possibly glandular structures (
<figureCitation id="131F2A10FF8D325AFF75F8D6FF74F81F" box="[159,234,1911,1936]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="24" pageNumber="475">Fig. 8</figureCitation>
F). Subgenital plate short and squared (
<figureCitation id="131F2A10FF8D325AFD38F8D6FCBEF81F" box="[722,800,1911,1936]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="24" pageNumber="475">Figs 8</figureCitation>
G, 9D–F). Male genitalia (
<figureCitation id="131F2A10FF8D325AFB83F8D6FB5CF81F" box="[1129,1218,1911,1936]" captionStart="FIGURE 10" captionStartId="29.[151,250,1651,1673]" captionTargetBox="[152,1435,194,1619]" captionTargetId="figure@29.[151,1436,193,1629]" captionTargetPageId="29" captionText="FIGURE 10. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype, genitalia in dorsal (A), ventral (B) and lateral (C) views. Abbreviations see page 3." httpUri="https://zenodo.org/record/241867/files/figure.png" pageId="24" pageNumber="475">Fig. 10</figureCitation>
) with asymetrical pseudepiphallic processes, each with a hollow spherical base, and each bifid distally, one of these going far beyond distal margin of subgenital plate (
<figureCitation id="131F2A10FF8D325AFDE7F861FDCFF857" box="[525,593,1984,2009]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="24" pageNumber="475">Fig. 8</figureCitation>
, arrows); pseudepiphallic parameres well-developed and partly sclerotized; short rami; ectophallic fold very short and membranous; ectophallic apodemes well-developed but short; endophallic sclerite short with a pair of lamellae like apodemes.
<emphasis id="B950EA87FF8C325BFC8EFF1DFC22FF5A" bold="true" box="[868,956,188,213]" pageId="25" pageNumber="476">Female</emphasis>
with short, slightly overlapping FWs (
<figureCitation id="131F2A10FF8C325BFA80FF1DFF3AFF77" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="25" pageNumber="476">Figs 9</figureCitation>
G, 11A, C), with few parallel, convex veins. Subgenital plate transverse (
<figureCitation id="131F2A10FF8C325BFC01FF7EFBAFFF77" box="[1003,1073,223,248]" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="25" pageNumber="476">
Fig.
<date id="FF9A1055FF8C325BFBCBFF41FBAFFF77" box="[1057,1073,224,248]" pageId="25" pageNumber="476">9</date>
</figureCitation>
I). Ovipositor flattened laterally, the apex smooth and not widened (
<figureCitation id="131F2A10FF8C325BFDF4FEA5FDF3FE93" box="[542,621,260,285]" captionStart="FIGURE 11" captionStartId="30.[151,250,1952,1974]" captionTargetBox="[227,1375,202,1930]" captionTargetId="figure@30.[211,1376,193,1931]" captionTargetPageId="30" captionText="FIGURE 11. Phasmagryllus elegans Desutter-Grandcolas, n. sp., female allotype. A, dorsal view, scale 1 mm; B, lateral view; C, pronotum and FWs, dorsal view; D, ovipositor, lateral view." httpUri="https://zenodo.org/record/241868/files/figure.png" pageId="25" pageNumber="476">Fig. 11</figureCitation>
D). Female genitalia as on
<figureCitation id="131F2A10FF8C325BFC71FEA5FC43FE93" box="[923,989,260,285]" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="25" pageNumber="476">Fig. 9</figureCitation>
J.
</paragraph>
</subSubSection>
<subSubSection id="C33E651EFF8C3258FF7DFE86FC8DFE92" lastPageId="26" lastPageNumber="477" pageId="25" pageNumber="476" type="description">
<paragraph id="8B9B3695FF8C325BFF7DFE86FEB7FECF" blockId="25.[151,1437,151,2013]" box="[151,297,295,320]" pageId="25" pageNumber="476">
<emphasis id="B950EA87FF8C325BFF7DFE86FEB7FECF" bold="true" box="[151,297,295,320]" pageId="25" pageNumber="476">Description.</emphasis>
</paragraph>
<paragraph id="8B9B3695FF8C325BFF2DFEECFB63FC0F" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="476">
Very small and delicate species (
<figureCitation id="131F2A10FF8C325BFDA2FEEDFD0CFEEB" box="[584,658,332,357]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Figs 8</figureCitation>
A, 11A, B), resembling the unrelated Guianese genus
<taxonomicName id="4C244D16FF8C325BFAFAFEEDFE5BFE07" authority="Desutter-Grandcolas, 1992" authorityName="Desutter-Grandcolas" authorityYear="1992" class="Insecta" family="Haglotettigoniidae" genus="Philippopsis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA87FF8C325BFAFAFEEDFA02FEEA" box="[1296,1436,332,357]" italics="true" pageId="25" pageNumber="476">Philippopsis</emphasis>
<bibRefCitation id="EFB54B64FF8C325BFF7DFECEFE5BFE07" author="Desutter-Grandcolas" box="[151,453,367,392]" pageId="25" pageNumber="476" refString="Desutter-Grandcolas, L. (1992) Les Phalangopsidae de Guyane francaise (Orthopteres, Grylloidea): systematique, elements de phylogenie et de biologie. Bulletin du Museum national d'Histoire naturelle, Paris, 14, 93 - 177." type="journal article" year="1992">
Desutter-Grandcolas,
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</bibRefCitation>
</taxonomicName>
in appearance and contrasted coloration, but still thinner. Very setose. Head very long and large, rounded. Vertex with many strong, widely separate setae. Fastigium very wide, not separate from the vertex (
<figureCitation id="131F2A10FF8C325BFF06FE16FEB1FE5F" box="[236,303,439,464]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Fig. 8</figureCitation>
B, C), but slanting. Ocelli small, widely apart (
<figureCitation id="131F2A10FF8C325BFCD4FE16FC1FFE5F" box="[830,897,439,464]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Fig. 8</figureCitation>
C); distance between the median and one lateral ocellus smaller than the distance between the lateral ocelli; median ocellus in apical position. Eyes small, not protruding, well apart from each other (
<figureCitation id="131F2A10FF8C325BFDB7FE5EFD3CFD97" box="[605,674,511,536]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Fig. 8</figureCitation>
B). Maxillary palpi very elongate for the subfamily (
<figureCitation id="131F2A10FF8C325BFB17FE5EFAD8FD97" box="[1277,1350,511,536]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Figs 8</figureCitation>
C, 9A); joint 3 well shorter than joint 4, which is smaller than joint 5; joint 5 concave dorsally, slightly widened toward apex, truncate obliquely over nearly half its length. Scapes slightly longer than wide (
<figureCitation id="131F2A10FF8C325BFBBEFDE6FB08FDEF" box="[1108,1174,583,608]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Fig. 8</figureCitation>
B). Pronotum (
<figureCitation id="131F2A10FF8C325BFAD4FDE6FAE1FDEF" box="[1342,1407,583,608]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Fig. 8</figureCitation>
B) transverse, rounded; DD anterior margin slightly concave, posterior margin more concave: lateral lobes squared. Legs very thin but not very elongate (
<figureCitation id="131F2A10FF8C325BFDA2FD2EFD11FD27" box="[584,655,655,680]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Figs 8</figureCitation>
A, 11A, B). TI with a inner tympanum, small, oval, obliterate; outer tympanum lacking; two ventral apical spurs. TII with two ventral apical spurs. TIII (
<figureCitation id="131F2A10FF8C325BFBBBFD15FB0DFD43" box="[1105,1171,692,717]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Fig. 8</figureCitation>
D) with three inner and three outer apical spurs, the ventral the smallest and the dorsal the longest on both sides; inner dorsal spur the longest, about as long as half basitarsomere III. TIII with four pairs of subapical spurs, alternate, but almost aligned on one row; subapical spurs all small, growing longer toward TIII apex, except for the first subapical spur, the smallest; serrulation very sparse, almost lacking between subapical spurs. Tarsomeres long, the basitarsomere III very elongate. Abdomen as collapsed distally, short with narrow tergites. Cerci not particularly long.
</paragraph>
<paragraph id="8B9B3695FF8C325BFF2DFC2DFADAFC2B" blockId="25.[151,1437,151,2013]" box="[199,1348,908,933]" pageId="25" pageNumber="476">
<emphasis id="B950EA87FF8C325BFF2DFC2DFED1FC2A" bold="true" box="[199,335,908,933]" pageId="25" pageNumber="476">Coloration.</emphasis>
Species dark brown with contrasted coloration (
<figureCitation id="131F2A10FF8C325BFC87FC2DFC2AFC2B" box="[877,948,908,933]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Figs 8</figureCitation>
B, 11A) on head, DD and antennae.
</paragraph>
<paragraph id="8B9B3695FF8C325BFF2DFC0EFDD0FB67" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="476">
<emphasis id="B950EA87FF8C325BFF2DFC0EFE95FC47" bold="true" box="[199,267,943,968]" pageId="25" pageNumber="476">Male.</emphasis>
Metanotum with a pair of high median buds, flattened apically (
<figureCitation id="131F2A10FF8C325BFC1DFC0EFBA2FC47" box="[1015,1084,943,968]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Fig. 8</figureCitation>
F). Tergites two to five clearly convex along their distal margin, with concave areas near their anterior margins. FWs covering about half abdomen only (
<figureCitation id="131F2A10FF8C325BFF32FC56FE87FB9F" box="[216,281,1015,1040]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Fig. 8</figureCitation>
A), somewhat thickened; cell limits on FW surface very strongly marked (
<figureCitation id="131F2A10FF8C325BFBB3FC56FB02FB9F" box="[1113,1180,1015,1040]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Fig. 8</figureCitation>
E); FW lateral margins almost parallel, the FWs not widened at the mirror level (
<figureCitation id="131F2A10FF8C325BFCFCFBBDFCC6FBBB" box="[790,856,1052,1077]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Fig. 8</figureCitation>
E). Stridulatory apparatus complete, with functional file (
<figureCitation id="131F2A10FF8C325BFF26FB9EFE8FFBD7" box="[204,273,1087,1112]" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="25" pageNumber="476">Fig. 9</figureCitation>
B): harp wider than long, crossed by few parallel, oblique veins; mirror crossed by one transverse vein; chords occupying a large part of the FWs, the chord 1 widely apart from chords 2 and 3; one transverse cell under the mirror; apical field almost lacking. Lateral field well developed, but shorter than the dorsal field (
<figureCitation id="131F2A10FF8C325BFAE9FB26FAD8FB2F" box="[1283,1350,1159,1184]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Fig. 8</figureCitation>
H); few parallel, longitudinal veins (
<figureCitation id="131F2A10FF8C325BFE3FFB0DFD84FB4B" box="[469,538,1196,1221]" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="25" pageNumber="476">Fig. 9</figureCitation>
C). HWs bud like. Supra anal plate without strong bristles (
<figureCitation id="131F2A10FF8C325BFB55FB0DFA9DFB4B" box="[1215,1283,1196,1221]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">
Fig.
<date id="FF9A1055FF8C325BFB1EFB0DFA9DFB4B" box="[1268,1283,1196,1220]" pageId="25" pageNumber="476">8</date>
</figureCitation>
I). Subgenital plate short, wide, truncate (
<figureCitation id="131F2A10FF8C325BFE22FB6EFD97FB67" box="[456,521,1231,1256]" captionStart="FIGURE 8" captionStartId="27.[151,250,1905,1927]" captionTargetBox="[204,1383,187,1884]" captionTargetId="figure@27.[202,1384,187,1884]" captionTargetPageId="27" captionText="FIGURE 8. Phasmagryllus elegans Desutter-Grandcolas, n. sp., male holotype. A, dorsal view, scale 1 mm; B, head and pronotum, dorsal view; C, face and maxillary palpi; D, hindtibia, dorsal view; E, right forewing, dorsal view; F, metanotum and tergite structures, dorsal view; G, subgenital plate, ventral view; H, subgenital plate and right forewing, lateral view; I, supra anal plate, dorsal view. Arrows point to the left pseudepiphallic distal process, which is extremely long." httpUri="https://zenodo.org/record/241865/files/figure.png" pageId="25" pageNumber="476">Fig. 8</figureCitation>
G, H).
</paragraph>
<paragraph id="8B9B3695FF8C325BFF2DFB55FA0AF9FA" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="476">
<emphasis id="B950EA87FF8C325BFF2DFB55FEE4FA82" bold="true" box="[199,378,1268,1293]" pageId="25" pageNumber="476">Male genitalia.</emphasis>
<figureCitation id="131F2A10FF8C325BFE68FB55FE59FA83" box="[386,455,1268,1293]" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="25" pageNumber="476">Fig. 9</figureCitation>
. Small, asymmetrical. Pseudepiphallic sclerite transverse, H-shaped, prolonged lateroventrally by a sclerotization facing the rami (
<figureCitation id="131F2A10FF8C325BFD74FAB6FD7AFABF" box="[670,740,1303,1328]" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="25" pageNumber="476">Fig. 9</figureCitation>
, v.ps.); connected to the rami by a thin, lateral sclerite; rami small and dejected ventrally. A pair of median rounded lobes, located distally to the pseudepiphallic sclerite; brisling with small spikes on inner face. A pair of disto-lateral huge pseudepiphallic structures, with a rounded and hollowed anterior part, and a bifid distal part; this bifid part with a short inner lobe and a spine-like outer lobe; left and right inner lobes facing each other; right spine-like lobe short, left one extremely elongate, prolonged far beyond subgenital distal margin (
<figureCitation id="131F2A10FF8C325BFDFFFA6DFDC0FA6B" box="[533,606,1484,1509]" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="25" pageNumber="476">Figs 9</figureCitation>
D, F, 10). Pseudepiphallic parameres located in central position between these pseudepiphallic structures, plicated, with a ribbon-like sclerotization. Ectophallic fold very short, triangular and membranous. Ectophallic apodemes quite long; arc complete. No developped ectophallic dorsal valves. Ventral valves extremely reduced. Endophallic sclerite 3-pronged distally. Endophallic apodemes with a pair of short lamellae. A pair of half-circled transverse sclerites between the endophallic sclerite and the tip of ectophallic fold.
</paragraph>
<paragraph id="8B9B3695FF8C325BFF2DF9DEFBA6F96F" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="476">
<emphasis id="B950EA87FF8C325BFF2DF9DEFEBBF917" bold="true" box="[199,293,1663,1688]" pageId="25" pageNumber="476">Female.</emphasis>
FWs very short and hardly overlapping (
<figureCitation id="131F2A10FF8C325BFD12F9DEFCD7F917" box="[760,841,1663,1688]" captionStart="FIGURE 11" captionStartId="30.[151,250,1952,1974]" captionTargetBox="[227,1375,202,1930]" captionTargetId="figure@30.[211,1376,193,1931]" captionTargetPageId="30" captionText="FIGURE 11. Phasmagryllus elegans Desutter-Grandcolas, n. sp., female allotype. A, dorsal view, scale 1 mm; B, lateral view; C, pronotum and FWs, dorsal view; D, ovipositor, lateral view." httpUri="https://zenodo.org/record/241868/files/figure.png" pageId="25" pageNumber="476">Fig. 11</figureCitation>
C); venation made of convex longitudinal veins only (
<figureCitation id="131F2A10FF8C325BFF75F905FF77F933" box="[159,233,1700,1725]" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="25" pageNumber="476">Fig. 9</figureCitation>
G); lateral field very narrow (
<figureCitation id="131F2A10FF8C325BFDBBF905FD05F933" box="[593,667,1700,1725]" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="25" pageNumber="476">Fig. 9</figureCitation>
H). Subgenital plate transverse, distal margin bisinuate (
<figureCitation id="131F2A10FF8C325BFAD0F905FA1AF933" box="[1338,1412,1700,1725]" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="25" pageNumber="476">
Fig.
<date id="FF9A1055FF8C325BFA9EF905FA1AF933" box="[1396,1412,1700,1724]" pageId="25" pageNumber="476">9</date>
</figureCitation>
I). Ovipositor flattened laterally; apex not widened and not ornemented (
<figureCitation id="131F2A10FF8C325BFC4EF966FC6DF96F" box="[932,1011,1735,1760]" captionStart="FIGURE 11" captionStartId="30.[151,250,1952,1974]" captionTargetBox="[227,1375,202,1930]" captionTargetId="figure@30.[211,1376,193,1931]" captionTargetPageId="30" captionText="FIGURE 11. Phasmagryllus elegans Desutter-Grandcolas, n. sp., female allotype. A, dorsal view, scale 1 mm; B, lateral view; C, pronotum and FWs, dorsal view; D, ovipositor, lateral view." httpUri="https://zenodo.org/record/241868/files/figure.png" pageId="25" pageNumber="476">Fig. 11</figureCitation>
B, D).
</paragraph>
<paragraph id="8B9B3695FF8C325BFF2DF94DFAB5F8A7" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="476">
<emphasis id="B950EA87FF8C325BFF2DF94DFE0AF88A" bold="true" box="[199,404,1772,1797]" pageId="25" pageNumber="476">Female genitalia.</emphasis>
In
<taxonomicName id="4C244D16FF8C325BFE51F94DFC2CF88A" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[443,946,1772,1797]" class="Insecta" family="Phalangopsidae" genus="Phasmagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="elegans" status="sp. nov.">
<emphasis id="B950EA87FF8C325BFE51F94DFD5EF88A" box="[443,704,1772,1797]" italics="true" pageId="25" pageNumber="476">Phasmagryllus elegans</emphasis>
Desutter-Grandcolas
</taxonomicName>
,
<taxonomicNameLabel id="A26357FCFF8C325BFC54F94CFC63F88B" box="[958,1021,1773,1796]" pageId="25" pageNumber="476" rank="species">n. sp.</taxonomicNameLabel>
, copulatory papilla having the shape of a circular sclerite, which membranous center is prolonged by a short and wide, plicated duct (
<figureCitation id="131F2A10FF8C325BFB27F8AEFA8EF8A7" box="[1229,1296,1807,1832]" captionStart="FIGURE 9" captionStartId="28.[151,250,1658,1680]" captionTargetBox="[153,1431,201,1629]" captionTargetId="figure@28.[151,1435,195,1637]" captionTargetPageId="28" captionText="FIGURE 9. Phasmagryllus elegans Desutter-Grandcolas, n. sp. A, maxillary palpus, female allotype; B, C, venation of dorsal and lateral fields of male holotype FWs; D-F, male holotype subgenital plate, ventral (D), right side (E) and left side (F), with projections of asymmetrical pseudepiphallic processes; G, H, venation of dorsal and lateral fields of female allotype right FW; I, female allotype subgenital plate, ventral view; J, female allotype copulatory papilla. Scale bars 1 mm." httpUri="https://zenodo.org/record/241866/files/figure.png" pageId="25" pageNumber="476">Fig. 9</figureCitation>
J).
</paragraph>
<paragraph id="8B9B3695FF8C3258FF2DF895FC8DFE92" blockId="25.[151,1437,151,2013]" lastBlockId="26.[151,1436,151,285]" lastPageId="26" lastPageNumber="477" pageId="25" pageNumber="476">
<emphasis id="B950EA87FF8C325BFF2DF895FD97F8C2" bold="true" box="[199,521,1844,1869]" pageId="25" pageNumber="476">Phylogenetic relationships.</emphasis>
By its morphology, and more specifically by its head shape (fastigium, ocelli), TIII shape, subapical spurs and serrulation, and male genitalia,
<taxonomicName id="4C244D16FF8C325BFCBFF8F6FB70F8FF" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[853,1262,1879,1904]" class="Insecta" family="Phalangopsidae" genus="Phasmagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA87FF8C325BFCBFF8F6FC63F8FF" box="[853,1021,1879,1904]" italics="true" pageId="25" pageNumber="476">Phasmagryllus</emphasis>
Desutter-Grandcolas
</taxonomicName>
, n. gen. is close to other African
<taxonomicName id="4C244D16FF8C325BFE80F8DDFE72F81A" box="[362,492,1916,1941]" class="Insecta" family="Phalangopsidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="subFamily" subFamily="Phaloriinae">Phaloriinae</taxonomicName>
, even though its body shape and way of life resemble other non phaloriine
<taxonomicName id="4C244D16FF8C325BFF7DF801FED9F836" box="[151,327,1952,1977]" class="Insecta" family="Phalangopsidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="family">Phalangopsidae</taxonomicName>
such as
<taxonomicName id="4C244D16FF8C325BFE5BF801FDA3F836" box="[433,573,1952,1977]" class="Insecta" family="Haglotettigoniidae" genus="Philippopsis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA87FF8C325BFE5BF801FDA3F836" box="[433,573,1952,1977]" italics="true" pageId="25" pageNumber="476">Philippopsis</emphasis>
</taxonomicName>
or
<taxonomicName id="4C244D16FF8C325BFD80F801FC33F837" authority="Serville, 1831" authorityName="Serville" authorityYear="1831" box="[618,941,1952,1977]" class="Insecta" family="Haglotettigoniidae" genus="Phalangopsis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA87FF8C325BFD80F801FC9DF836" box="[618,771,1952,1977]" italics="true" pageId="25" pageNumber="476">Phalangopsis</emphasis>
Serville,
<quantity id="4CDC9B70FF8C325BFC9FF801FC33F837" box="[885,941,1952,1976]" metricMagnitude="1" metricUnit="m" metricValue="4.65074" pageId="25" pageNumber="476" unit="in" value="1831.0">1831</quantity>
</taxonomicName>
in the Neotropical Region, or the African
<taxonomicName id="4C244D16FF8C325BFF7DF865FE7AF853" authority="Walker, 1871" authorityName="Walker" authorityYear="1871" box="[151,484,1988,2013]" class="Insecta" family="Haglotettigoniidae" genus="Phaeophilacris" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA87FF8C325BFF7DF865FEDCF852" box="[151,322,1988,2013]" italics="true" pageId="25" pageNumber="476">Phaeophilacris</emphasis>
Walker, 1871
</taxonomicName>
. These latter genera are straminicolous and cavicolous, foraging at night in the leaf litter, and hiding during the day in cavities at ground level such as burrows or hollow trees (
<bibRefCitation id="EFB54B64FF8F3258FB59FF36FF4CFF5B" author="Desutter-Grandcolas" pageId="26" pageNumber="477" refString="Desutter-Grandcolas, L. (1992) Les Phalangopsidae de Guyane francaise (Orthopteres, Grylloidea): systematique, elements de phylogenie et de biologie. Bulletin du Museum national d'Histoire naturelle, Paris, 14, 93 - 177." type="journal article" year="1992">Desutter-Grandcolas 1992</bibRefCitation>
,
<bibRefCitation id="EFB54B64FF8F3258FF0EFF1DFE80FF5A" author="Desutter-Grandcolas" box="[228,286,188,213]" pageId="26" pageNumber="477" refString="Desutter-Grandcolas, L. (1995) Toward the knowledge of the evolutionary biology of phalangopsid crickets (Orthoptera, Grylloidea, Phalangopsidae: data, questions and scenarios. Journal of Orthoptera Research, 4, 163 - 175. http: // dx. doi. org / 10.2307 / 3503472" type="journal article" year="1995">1995</bibRefCitation>
, pers. obs.). Convergence in body shape due to common habitat might have occurred between
<taxonomicName id="4C244D16FF8F3258FF7DFF7EFDA7FF77" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[151,569,223,248]" class="Insecta" family="Phalangopsidae" genus="Phasmagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA87FF8F3258FF7DFF7EFEA1FF77" box="[151,319,223,248]" italics="true" pageId="26" pageNumber="477">Phasmagryllus</emphasis>
Desutter-Grandcolas
</taxonomicName>
, n. gen. on one hand, and other &quot;truly&quot; long-legged crickets such as
<taxonomicName id="4C244D16FF8F3258FF7DFEA5FEBDFE92" box="[151,291,260,285]" class="Insecta" family="Haglotettigoniidae" genus="Philippopsis" kingdom="Animalia" order="Orthoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA87FF8F3258FF7DFEA5FEBDFE92" box="[151,291,260,285]" italics="true" pageId="26" pageNumber="477">Philippopsis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C244D16FF8F3258FEDBFEA5FE54FE92" box="[305,458,260,285]" class="Insecta" family="Haglotettigoniidae" genus="Phalangopsis" kingdom="Animalia" order="Orthoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA87FF8F3258FEDBFEA5FE54FE92" box="[305,458,260,285]" italics="true" pageId="26" pageNumber="477">Phalangopsis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C244D16FF8F3258FE3DFEA5FD1CFE92" box="[471,642,260,285]" class="Insecta" family="Haglotettigoniidae" genus="Phaeophilacris" kingdom="Animalia" order="Orthoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA87FF8F3258FE3DFEA5FD1CFE92" box="[471,642,260,285]" italics="true" pageId="26" pageNumber="477">Phaeophilacris</emphasis>
</taxonomicName>
and the like.
</paragraph>
</subSubSection>
</treatment>
</document>
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