160 lines
21 KiB
XML
160 lines
21 KiB
XML
<document id="C42BEFC6918B8339E9EA18BE5861D175" ID-DOI="10.11646/zootaxa.3674.1.1" ID-GBIF-Dataset="9c8e7b58-7cc0-407f-a8e9-d76311dcd290" ID-ISSN="1175-5326" ID-Zenodo-Dep="283954" ID-ZooBank="981F80ED-96D7-40C7-8A3C-677954416A2E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460635409297" checkinUser="plazi" docAuthor="Dupérré, Nadine" docDate="2013" docId="038D6700FFB85612118C00C0AF0AB26A" docLanguage="en" docName="zt03674p189.pdf" docOrigin="Zootaxa 3674 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Micronetinae" docType="treatment" docVersion="10" lastPageNumber="6" masterDocId="FFB41F78FFBC5617111B061BAE0BB136" masterDocTitle="Taxonomic revision of the spider genera Agyneta and Tennesseellum (Araneae, Linyphiidae) of North America north of Mexico with a study of the embolic division within Micronetinae sensu Saaristo & Tanasevitch 1996" masterLastPageNumber="189" masterPageNumber="1" pageNumber="5" updateTime="1698335287136" updateUser="plazi">
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<mods:title id="20B516D8F13A34797F18B0A62A7BD836">Taxonomic revision of the spider genera Agyneta and Tennesseellum (Araneae, Linyphiidae) of North America north of Mexico with a study of the embolic division within Micronetinae sensu Saaristo & Tanasevitch 1996</mods:title>
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<mods:namePart id="1ADF23FF3AB6A33B589C386E2BE9829F">Dupérré, Nadine</mods:namePart>
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<mods:title id="596B9E138CE4B1DF7551776D7E5E050D">Zootaxa</mods:title>
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<mods:date id="A0D6A9F0998C2BE021F2EFF41BAE42D7">2013</mods:date>
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<mods:number id="56960D545F1BF2257B6865668CD47D77">3674</mods:number>
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<treatment id="038D6700FFB85612118C00C0AF0AB26A" ID-DOI="http://doi.org/10.5281/zenodo.6162275" ID-GBIF-Taxon="119570642" ID-Zenodo-Dep="6162275" LSID="urn:lsid:plazi:treatment:038D6700FFB85612118C00C0AF0AB26A" httpUri="http://treatment.plazi.org/id/038D6700FFB85612118C00C0AF0AB26A" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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<subSubSection id="C33E859DFFB85612118C00C0AF0AB26A" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph id="8B9BD616FFB85613118C00C0AC3CB7C3" blockId="4.[151,567,1755,1781]" box="[151,567,1755,1781]" pageId="4" pageNumber="5">
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<heading id="D0D3617AFFB85613118C00C0AC3CB7C3" bold="true" box="[151,567,1755,1781]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
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<emphasis id="B9500A04FFB85613118C00C0AC3CB7C3" bold="true" box="[151,567,1755,1781]" pageId="4" pageNumber="5">
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SUBFAMILY
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<taxonomicName id="4C24AD95FFB85613105200C0AC3CB7C3" box="[329,567,1755,1781]" class="Arachnida" family="Linyphiidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Micronetinae">MICRONETINAE</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9BD616FFB85613118C0139AD61B690" blockId="4.[151,1436,1826,2031]" pageId="4" pageNumber="5">
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<bibRefCitation id="EFB5ABE7FFB85613118C0139AF14B60C" author="Hull" box="[151,287,1826,1850]" pageId="4" pageNumber="5" refString="Hull, J. E. (1920) The spider family Linyphiidae: an essay in taxonomy. Vasculum, 6, 7 - 11." type="journal article" year="1920">Hull (1920)</bibRefCitation>
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defined the subfamily
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<taxonomicName id="4C24AD95FFB85613132F0139ACC2B60C" box="[564,713,1826,1850]" class="Arachnida" family="Linyphiidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Micronetinae">Micronetinae</taxonomicName>
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by the absence of femoral and lateral spines, the absence of “auxiliary claws” and the presence of two, four or five promarginal teeth on the chelicerae. All these characters are very broad and can be found in large number of
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<taxonomicName id="4C24AD95FFB8561313A70171AD4CB6B4" box="[700,839,1898,1922]" class="Arachnida" family="Linyphiidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Linyphiidae</taxonomicName>
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.
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<bibRefCitation id="EFB5ABE7FFB8561312480171ADDCB6B4" author="Hull" box="[851,983,1898,1922]" pageId="4" pageNumber="5" refString="Hull, J. E. (1920) The spider family Linyphiidae: an essay in taxonomy. Vasculum, 6, 7 - 11." type="journal article" year="1920">Hull (1920)</bibRefCitation>
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included in the subfamily
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<taxonomicName id="4C24AD95FFB85613141C0171AB97B6B4" box="[1287,1436,1898,1922]" class="Arachnida" family="Linyphiidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Micronetinae">Micronetinae</taxonomicName>
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<taxonomicName id="4C24AD95FFB85613118C0195AEF8B690" box="[151,243,1934,1958]" class="Arachnida" family="Linyphiidae" genus="Agyneta" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B9500A04FFB85613118C0195AEF8B690" box="[151,243,1934,1958]" italics="true" pageId="4" pageNumber="5">Agyneta</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C24AD95FFB85613101B0194AF7FB690" box="[256,372,1935,1958]" class="Arachnida" family="Linyphiidae" genus="Microneta" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B9500A04FFB85613101B0194AF7FB690" box="[256,372,1935,1958]" italics="true" pageId="4" pageNumber="5">Microneta</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C24AD95FFB8561310990194AC79B690" authority="Rhabdoria" authorityName="Rhabdoria" box="[386,626,1934,1958]" class="Arachnida" family="Linyphiidae" genus="Meioneta" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B9500A04FFB8561310990194AFE7B690" box="[386,492,1935,1958]" italics="true" pageId="4" pageNumber="5">Meioneta</emphasis>
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,
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<emphasis id="B9500A04FFB8561310E10195AC79B690" box="[506,626,1934,1958]" italics="true" pageId="4" pageNumber="5">Rhabdoria</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C24AD95FFB85613139B0195ACC5B690" box="[640,718,1934,1958]" class="Arachnida" family="Linyphiidae" genus="Syedra" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B9500A04FFB85613139B0195ACC5B690" box="[640,718,1934,1958]" italics="true" pageId="4" pageNumber="5">Syedra</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C24AD95FFB85613121F0195AD68B690" box="[772,867,1934,1958]" class="Arachnida" family="Linyphiidae" genus="Aphileta" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B9500A04FFB85613121F0195AD68B690" box="[772,867,1934,1958]" italics="true" pageId="4" pageNumber="5">Aphileta</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8B9BD616FFB8561211D401A9ADC7B09A" blockId="4.[151,1436,1826,2031]" lastBlockId="5.[151,1436,151,860]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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<bibRefCitation id="EFB5ABE7FFB8561311D401A9AC23B6FC" author="Saaristo" box="[207,552,1970,1994]" pageId="4" pageNumber="5" refString="Saaristo, M. I. & Tanasevitch, A. V. (1996) Redelimitation of the subfamily Micronetinae Hull, 1920 and the genus Lepthyphantes Menge, 1866 with descriptions of some new genera (Aranei, Linyphiidae). Bericht des Naturwissens- Medizinischen Verein Innsbruck, 83, 163 - 186." type="journal article" year="1996">Saaristo & Tanasevitch (1996)</bibRefCitation>
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redelimited the subfamily
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<taxonomicName id="4C24AD95FFB85613124701A9ADFAB6FC" box="[860,1009,1970,1994]" class="Arachnida" family="Linyphiidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Micronetinae">Micronetinae</taxonomicName>
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in reference to the male palp and the conformation of the epigynum. They gave a general description of the male palp based on
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<taxonomicName id="4C24AD95FFB85612158301CDAF54B199" authority="Blackwall 1833" authorityName="Blackwall" authorityYear="1833" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" kingdom="Animalia" lastPageId="5" lastPageNumber="6" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="minutus">
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<emphasis id="B9500A04FFB85613158301CDAB97B6D8" box="[1176,1436,2006,2030]" italics="true" pageId="4" pageNumber="5">Lepthyphantes minutus</emphasis>
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(
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<bibRefCitation id="EFB5ABE7FFB956121185068CAF5DB199" author="Blackwall" box="[158,342,151,175]" pageId="5" pageNumber="6" refString="Blackwall, J. (1833) Characters of some undescribed genera and species of Araneidae. London and Edinburgh Philosophical Magazine and Journal of Science, (3) 3, 104 - 112, 187 - 197, 344 - 352, 436 - 443." type="journal article" year="1833">Blackwall 1833</bibRefCitation>
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)
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</taxonomicName>
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to represent the conformation found in the subfamily
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<taxonomicName id="4C24AD95FFB9561212FC068CAA8BB199" box="[999,1152,151,175]" class="Arachnida" family="Linyphiidae" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subFamily" subFamily="Micronetinae">Micronetinae</taxonomicName>
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.
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<bibRefCitation id="EFB5ABE7FFB956121594068CAF16B1E2" author="Saaristo" pageId="5" pageNumber="6" refString="Saaristo, M. I. & Tanasevitch, A. V. (1996) Redelimitation of the subfamily Micronetinae Hull, 1920 and the genus Lepthyphantes Menge, 1866 with descriptions of some new genera (Aranei, Linyphiidae). Bericht des Naturwissens- Medizinischen Verein Innsbruck, 83, 163 - 186." type="journal article" year="1996">Saaristo & Tanasevitch (1996: 170)</bibRefCitation>
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also mentioned the presence in the radix “at the site of the embolic base the sperm duct is dilated forming the so called Fickert’s gland”, and that “In certain cases this gland has been lost which sometimes is compensated by a secondary dilatation inside the embolus like for example in some
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<taxonomicName id="4C24AD95FFB95612155D071FAB2BB02A" authority="Hull, 1911" authorityName="Hull" authorityYear="1911" box="[1094,1312,260,284]" class="Arachnida" family="Linyphiidae" genus="Agyneta" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B9500A04FFB95612155D071FAAA9B02A" box="[1094,1186,260,284]" italics="true" pageId="5" pageNumber="6">Agyneta</emphasis>
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<bibRefCitation id="EFB5ABE7FFB9561215B2071FAB2BB02A" author="Hull" box="[1193,1312,260,284]" pageId="5" pageNumber="6" refString="Hull, J. E. (1911) Papers on spiders. Transactions of the Natural History Society of Northumberland (N. S.), 3 (3), 573 - 590." type="journal article" year="1911">Hull, 1911</bibRefCitation>
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</taxonomicName>
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species”. I found that not all
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<taxonomicName id="4C24AD95FFB95612107A0733AC73B009" authority="Saaristo" authorityName="Saaristo" box="[353,632,295,320]" class="Arachnida" family="Linyphiidae" genus="Agyneta" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B9500A04FFB95612107A0733AFB6B076" box="[353,445,296,320]" italics="true" pageId="5" pageNumber="6">Agyneta</emphasis>
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(
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<emphasis id="B9500A04FFB9561210D60732AC00B076" box="[461,523,297,320]" italics="true" pageId="5" pageNumber="6">sensu</emphasis>
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Saaristo)
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</taxonomicName>
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have a Fickert’s gland, but when present it is always in the embolus. It seems unlikely that this corresponds to a secondary dilation, and I would argue that the Fickert’s gland has been displaced to the embolus, since its position within the radix varies greatly and it has shifted so far as to be in the base of the embolus in
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<taxonomicName id="4C24AD95FFB956121081078FAD57B09A" authority="Blackwall 1841" authorityName="Blackwall" authorityYear="1841" box="[410,860,404,428]" class="Arachnida" family="Linyphiidae" genus="Poeciloneta" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="variegata">
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<emphasis id="B9500A04FFB956121081078FAC9AB09A" box="[410,657,404,428]" italics="true" pageId="5" pageNumber="6">Poeciloneta variegata</emphasis>
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(
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<bibRefCitation id="EFB5ABE7FFB9561213BB078FAD5FB09A" author="Blackwall" box="[672,852,404,428]" pageId="5" pageNumber="6" refString="Blackwall, J. (1841) The difference in the number of eyes with which spiders are provided proposed as the basis of their distribution into tribes; with descriptions of newly discovered species and the characters of a new family and three new genera of spiders. Transactions of the Linnean Society of London, 18, 601 - 670. http: // dx. doi. org / 10.1111 / j. 1095 - 8339.1838. tb 00210. x" type="journal article" year="1841">Blackwall 1841</bibRefCitation>
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)
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</taxonomicName>
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(
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<figureCitation id="131FCA93FFB956121270078FADCBB09A" box="[875,960,404,428]" captionStart="FIGURES 58 – 63" captionStartId="27.[151,264,1954,1977]" captionTargetBox="[176,1426,189,1929]" captionTargetId="figure@27.[173,1437,173,1934]" captionTargetPageId="27" captionText="FIGURES 58 – 63. Radical division, retrolateral view. 58. Bolyphantes luteolus (Blackwall 1852). 59. Poeciloneta variegata (Blackwall 1841). 60. Oreonetides vaginatus (Thorell 1872). 61. Arcuphantes fragilis Chamberlin & Ivie 1943. 62. Agnyphantes expunctus (O. Pickard-Cambridge 1875). 63. Megalepthyphantes nebulosus (Sundevall 1830)." httpUri="https://zenodo.org/record/283964/files/figure.png" pageId="5" pageNumber="6">Fig. 59</figureCitation>
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).
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</paragraph>
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<paragraph id="8B9BD616FFB9561211DC07ACAF0AB26A" blockId="5.[151,1436,151,860]" pageId="5" pageNumber="6">
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<bibRefCitation id="EFB5ABE7FFB9561211DC07ACAC53B0F9" author="Saaristo" box="[199,600,439,463]" pageId="5" pageNumber="6" refString="Saaristo, M. I. & Tanasevitch, A. V. (1996) Redelimitation of the subfamily Micronetinae Hull, 1920 and the genus Lepthyphantes Menge, 1866 with descriptions of some new genera (Aranei, Linyphiidae). Bericht des Naturwissens- Medizinischen Verein Innsbruck, 83, 163 - 186." type="journal article" year="1996">Saaristo & Tanasevitch (1996: 176)</bibRefCitation>
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gave a short description “Micronetids may be recognized by the complex
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<typeStatus id="549F68B4FFB95612118C07C6AECCB0C3" box="[151,199,477,501]" pageId="5" pageNumber="6">type</typeStatus>
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of embolic division as described above. Also the presence of the outward pointing suprategular apophysis just below the tip of the cymbium in the unexpanded palp is often diagnostic” and included 54 genera in the subfamily
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<taxonomicName id="4C24AD95FFB95612118C043FAF3BB30A" box="[151,304,548,572]" class="Arachnida" family="Linyphiidae" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subFamily" subFamily="Micronetinae">Micronetinae</taxonomicName>
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. In studying the complex embolic division, Saaristo & Tanasevitch overlooked some important characters such as the position of the embolus in relation to the radix and the position of the embolus proper (when present). They did mention that the embolus proper is situated on the retrolateral side of the embolus in
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<taxonomicName id="4C24AD95FFB95612118C048BAFDBB391" authority="Menge 1866" authorityName="Menge" authorityYear="1866" box="[151,464,655,680]" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B9500A04FFB95612118C048BAF31B39E" box="[151,314,656,680]" italics="true" pageId="5" pageNumber="6">Lepthyphantes</emphasis>
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Menge 1866
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</taxonomicName>
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, whereas it is on the prolateral side in
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<taxonomicName id="4C24AD95FFB956121261048BADDDB39E" box="[890,982,656,680]" class="Arachnida" family="Linyphiidae" genus="Agyneta" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B9500A04FFB956121261048BADDDB39E" box="[890,982,656,680]" italics="true" pageId="5" pageNumber="6">Agyneta</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C24AD95FFB956121517048AAA7DB39E" box="[1036,1142,657,680]" class="Arachnida" family="Linyphiidae" genus="Meioneta" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B9500A04FFB956121517048AAA7DB39E" box="[1036,1142,657,680]" italics="true" pageId="5" pageNumber="6">Meioneta</emphasis>
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</taxonomicName>
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, and that in
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<taxonomicName id="4C24AD95FFB9561215E4048AAB78B39E" box="[1279,1395,657,680]" class="Arachnida" family="Linyphiidae" genus="Microneta" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B9500A04FFB9561215E4048AAB78B39E" box="[1279,1395,657,680]" italics="true" pageId="5" pageNumber="6">Microneta</emphasis>
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</taxonomicName>
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the sperm duct just ends at the tip of the embolus without an embolus proper. I re-studied some of the genera included in Saaristo & Tanasevitch’s (1996) study, based on the
|
||
<typeStatus id="549F68B4FFB95612121D04C2AD3DB3C7" box="[774,822,729,753]" pageId="5" pageNumber="6">type</typeStatus>
|
||
species (except for
|
||
<taxonomicName id="4C24AD95FFB95612150104C2AB32B3D9" authority="Millidge 1988" authorityName="Millidge" authorityYear="1988" box="[1050,1337,727,752]" class="Arachnida" family="Linyphiidae" genus="Maorineta" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9500A04FFB95612150104C2AA9BB3C6" box="[1050,1168,729,752]" italics="true" pageId="5" pageNumber="6">Maorineta</emphasis>
|
||
<bibRefCitation id="EFB5ABE7FFB95612158304CCAB32B3D9" author="Millidge" box="[1176,1337,727,751]" pageId="5" pageNumber="6" refString="Millidge, A. F. (1988) The spiders of New Zealand: Part VI. Family Linyphiidae. Otago Museum Bulletin, 6, 35 - 67." type="journal article" year="1988">Millidge 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
, as I did not have access to the
|
||
<typeStatus id="549F68B4FFB9561210B904E6AFD9B223" box="[418,466,765,789]" pageId="5" pageNumber="6">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C24AD95FFB95612132C04E6ADBDB222" authority="Millidge 1988" authorityName="Millidge" authorityYear="1988" box="[567,950,764,788]" class="Arachnida" family="Linyphiidae" genus="Maorineta" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tibialis">
|
||
<emphasis id="B9500A04FFB95612132C04E6AD0EB222" box="[567,773,764,788]" italics="true" pageId="5" pageNumber="6">Maorineta tibialis</emphasis>
|
||
<bibRefCitation id="EFB5ABE7FFB95612121404E7ADBDB222" author="Millidge" box="[783,950,764,788]" pageId="5" pageNumber="6" refString="Millidge, A. F. (1988) The spiders of New Zealand: Part VI. Family Linyphiidae. Otago Museum Bulletin, 6, 35 - 67." type="journal article" year="1988">Millidge 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
). I examined the details of the male palp, particularly the embolus shape, morphology and its attachment to the radical division as well as the morphology of the radix.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |