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<document id="C7B14AE140CCD4229C2D9C65F31B0E79" ID-DOI="10.11646/zootaxa.3949.1.4" ID-GBIF-Dataset="60b71b57-6ebd-42e4-9d9a-3689579bab1b" ID-ISSN="1175-5326" ID-Zenodo-Dep="241799" ID-ZooBank="22266A34-5663-4180-91F3-814AA12D8ABE" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461045206382" checkinUser="plazi" docAuthor="Ariyama, Hiroyuki" docDate="2015" docId="038D5362FF9DFFC2FF7DF901FB5DF92B" docLanguage="en" docName="zt03949p110.pdf" docOrigin="Zootaxa 3949 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psammogammarus lobatus Ariyama, 2015, sp. nov." docType="treatment" docVersion="10" lastPageNumber="103" masterDocId="FFB42B1AFF9AFFCEFFEAFF86FF81FFCB" masterDocTitle="Three new species of the Eriopisa group (Crustacea: Amphipoda: Eriopisidae) from Japan, with the description of a new genus" masterLastPageNumber="110" masterPageNumber="91" pageNumber="98" updateTime="1698576547859" updateUser="plazi">
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<mods:title id="267948D7E337AB54681A9067F0302342">Three new species of the Eriopisa group (Crustacea: Amphipoda: Eriopisidae) from Japan, with the description of a new genus</mods:title>
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<mods:name id="F37CED68CC1815B1C74589BC2DA723AC" type="personal">
<mods:role id="56DC1FB4EE0D6F2FD61551570E28FBA2">
<mods:roleTerm id="AE0A24F0FACD20806A46576AF3EFE1EB">Author</mods:roleTerm>
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<mods:namePart id="945853F1B3B3D15E19328549C619F987">Ariyama, Hiroyuki</mods:namePart>
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<mods:title id="067B7E709658FA39DE46E8C6E688D1C7">Zootaxa</mods:title>
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<mods:part id="2B121887F2E997F9481A3C1367A1A871">
<mods:date id="F4EF6E722EAD46870542D85DE9453564">2015</mods:date>
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<mods:number id="F78225C9EFD3467EAA9D727FABAE98EC">3949</mods:number>
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<mods:identifier id="12C3D2C2907D923E2B05D040381975A9" type="DOI">10.11646/zootaxa.3949.1.4</mods:identifier>
<mods:identifier id="974AA933E2BDE28BDF347C73C1DCDBA5" type="GBIF-Dataset">60b71b57-6ebd-42e4-9d9a-3689579bab1b</mods:identifier>
<mods:identifier id="7CFAC1A4D64CCBCF924A58D069728E49" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C0BE9A54AD796273B6882004F972842B" type="Zenodo-Dep">241799</mods:identifier>
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<treatment id="038D5362FF9DFFC2FF7DF901FB5DF92B" ID-DOI="http://doi.org/10.5281/zenodo.6108629" ID-GBIF-Taxon="119610335" ID-Zenodo-Dep="6108629" LSID="urn:lsid:plazi:treatment:038D5362FF9DFFC2FF7DF901FB5DF92B" httpUri="http://treatment.plazi.org/id/038D5362FF9DFFC2FF7DF901FB5DF92B" lastPageId="12" lastPageNumber="103" pageId="7" pageNumber="98">
<subSubSection id="C33EB1FFFF9DFFC9FF7DF901FEB8F908" pageId="7" pageNumber="98" type="nomenclature">
<paragraph id="8B9BE274FF9DFFC9FF7DF901FDC5F969" blockId="7.[151,580,1671,1731]" box="[151,580,1671,1698]" pageId="7" pageNumber="98">
<heading id="D0D35518FF9DFFC9FF7DF901FDC5F969" bold="true" box="[151,580,1671,1698]" fontSize="11" level="1" pageId="7" pageNumber="98" reason="1">
<emphasis id="B9503E66FF9DFFC9FF7DF901FDC5F969" bold="true" box="[151,580,1671,1698]" pageId="7" pageNumber="98">
<taxonomicName id="4C2499F7FF9DFFC9FF7DF901FE5EF96A" box="[151,479,1671,1697]" class="Malacostraca" family="Melitidae" genus="Psammogammarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="lobatus" status="sp. nov.">
<emphasis id="B9503E66FF9DFFC9FF7DF901FE5EF96A" bold="true" box="[151,479,1671,1697]" italics="true" pageId="7" pageNumber="98">Psammogammarus lobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A263831DFF9DFFC9FE0CF90EFDC5F969" box="[486,580,1672,1698]" pageId="7" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9BE274FF9DFFC9FF7DF92CFEB8F908" blockId="7.[151,580,1671,1731]" box="[151,313,1706,1731]" pageId="7" pageNumber="98">
(
<figureCitation id="131FFEF1FF9DFFC9FF75F92CFF63F908" box="[159,226,1706,1731]" captionStart="FIGURE 1" captionStartId="2.[151,250,978,1000]" captionTargetBox="[350,1240,276,943]" captionTargetId="figure@2.[338,1249,267,956]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the collecting sites. A B, Victoriopisa wadai sp. nov.; C, Paraflagitopisa excavata gen. et sp. nov. and Psammogammarus lobatus sp. nov." httpUri="https://zenodo.org/record/241800/files/figure.png" pageId="7" pageNumber="98">Figs 1</figureCitation>
C, 69)
</paragraph>
</subSubSection>
<subSubSection id="C33EB1FFFF9DFFC9FF7DF974FDA5F8BC" pageId="7" pageNumber="98" type="materials_examined">
<paragraph id="8B9BE274FF9DFFC9FF7DF974FDA5F8BC" blockId="7.[151,1437,1778,2019]" pageId="7" pageNumber="98">
<emphasis id="B9503E66FF9DFFC9FF7DF974FE02F8C0" bold="true" box="[151,387,1778,1803]" pageId="7" pageNumber="98">Material examined.</emphasis>
<typeStatus id="549F5CD6FF9DFFC9FE66F974FE78F8C0" box="[396,505,1778,1803]" pageId="7" pageNumber="98" type="holotype">Holotype</typeStatus>
: male (OMNH-Ar-9717), 4.0 mm, mouth of Higashi River, Misaki Town, Osaka Prefecture,
<geoCoordinate id="EE1084B3FF9DFFC9FEFDF891FE1FF8E4" box="[279,414,1815,1839]" direction="north" orientation="latitude" pageId="7" pageNumber="98" precision="15" value="34.32222">34°19'20&quot;N</geoCoordinate>
, 135°07'09&quot; (
<figureCitation id="131FFEF1FF9DFFC9FDDFF890FDF5F8E4" box="[565,628,1814,1839]" captionStart="FIGURE 1" captionStartId="2.[151,250,978,1000]" captionTargetBox="[350,1240,276,943]" captionTargetId="figure@2.[338,1249,267,956]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the collecting sites. A B, Victoriopisa wadai sp. nov.; C, Paraflagitopisa excavata gen. et sp. nov. and Psammogammarus lobatus sp. nov." httpUri="https://zenodo.org/record/241800/files/figure.png" pageId="7" pageNumber="98">Fig. 1</figureCitation>
C), sandy tidal flat, reverse surface of large stone,
<date id="FF9AC4B4FF9DFFC9FB4DF891FABFF8E4" box="[1191,1342,1815,1839]" pageId="7" pageNumber="98" value="1991-05-14">14 May 1991</date>
, coll. H. Ariyama.
<typeStatus id="549F5CD6FF9DFFC9FEEDF8BDFEF8F898" box="[263,377,1851,1875]" pageId="7" pageNumber="98" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D2229FDFF9DFFC9FE69F8BDFE4EF898" box="[387,463,1850,1875]" pageId="7" pageNumber="98" type="male">1 male</specimenCount>
(OMNH-Ar-9718),
<quantity id="4CDC4F91FF9DFFC9FD5FF8BDFC8CF899" box="[693,781,1851,1875]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="7" pageNumber="98" unit="mm" value="4.4">4.4 mm</quantity>
, and
<specimenCount id="9D2229FDFF9DFFC9FCA3F8BDFC37F898" box="[841,950,1850,1875]" pageId="7" pageNumber="98" type="female">2 females</specimenCount>
(OMNH-Ar-9719, 9720),
<quantity id="4CDC4F91FF9DFFC9FB08F8BDFABBF899" box="[1250,1338,1851,1875]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="98" unit="mm" value="4.2">4.2 mm</quantity>
,
<quantity id="4CDC4F91FF9DFFC9FAACF8BDFA1CF899" box="[1350,1437,1851,1875]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="98" unit="mm" value="3.7">3.7 mm</quantity>
(ovigerous), same data as
<typeStatus id="549F5CD6FF9DFFC9FE50F8D8FDA1F8BC" box="[442,544,1886,1911]" pageId="7" pageNumber="98" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C33EB1FFFF9DFFC2FF2DF804FD89FA56" lastPageId="12" lastPageNumber="103" pageId="7" pageNumber="98" type="description">
<paragraph id="8B9BE274FF9DFFC9FF2DF804FC0AF828" blockId="7.[151,1437,1778,2019]" pageId="7" pageNumber="98">
<emphasis id="B9503E66FF9DFFC9FF2DF804FED8F850" bold="true" box="[199,345,1922,1947]" pageId="7" pageNumber="98">Description.</emphasis>
Male [
<typeStatus id="549F5CD6FF9DFFC9FE42F804FD91F850" box="[424,528,1922,1947]" pageId="7" pageNumber="98" type="holotype">holotype</typeStatus>
, 4.0 mm (OMNH-Ar-9717)].
<emphasis id="B9503E66FF9DFFC9FC8EF804FC1FF850" box="[868,926,1922,1947]" italics="true" pageId="7" pageNumber="98">Body</emphasis>
(
<figureCitation id="131FFEF1FF9DFFC9FC44F804FC72F850" box="[942,1011,1922,1947]" captionStart="FIGURE 6" captionStartId="8.[151,250,958,980]" captionTargetBox="[160,1429,201,929]" captionTargetId="figure@8.[151,1435,193,937]" captionTargetPageId="8" captionText="FIGURE 6. Psammogammarus lobatus sp. nov. Holotype, male (OMNH-Ar- 9717), 4.0 mm. Habitus, lateral view. Scale: 1 mm." httpUri="https://zenodo.org/record/241805/files/figure.png" pageId="7" pageNumber="98">Fig. 6</figureCitation>
) slender, depressed laterally, smooth; pereonites and pleonites without dorsal setae, urosomites free.
<emphasis id="B9503E66FF9DFFC9FC88F820FC1EF874" box="[866,927,1958,1983]" italics="true" pageId="7" pageNumber="98">Head</emphasis>
longer than pereonite 1, rostrum small, eyes absent; lateral cephalic lobe rounded, anteroventral sinus indistinct.
</paragraph>
<caption id="DF5BB2FCFF92FFC6FF7DFC38FF47FC38" httpUri="https://zenodo.org/record/241805/files/figure.png" pageId="8" pageNumber="99" targetBox="[160,1429,201,929]" targetPageId="8">
<paragraph id="8B9BE274FF92FFC6FF7DFC38FF47FC38" blockId="8.[151,1435,958,1011]" pageId="8" pageNumber="99">
<emphasis id="B9503E66FF92FFC6FF7DFC38FE94FC18" bold="true" box="[151,277,958,980]" pageId="8" pageNumber="99">FIGURE 6.</emphasis>
<taxonomicName id="4C2499F7FF92FFC6FEF7FC39FDB2FC1F" box="[285,563,958,980]" class="Malacostraca" family="Melitidae" genus="Psammogammarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="99" phylum="Arthropoda" rank="species" species="lobatus" status="sp. nov.">
<emphasis id="B9503E66FF92FFC6FEF7FC39FDB2FC1F" box="[285,563,958,980]" italics="true" pageId="8" pageNumber="99">Psammogammarus lobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A263831DFF92FFC6FDD0FC39FD0CFC1F" box="[570,653,959,980]" pageId="8" pageNumber="99" rank="species">
<emphasis id="B9503E66FF92FFC6FDD0FC39FD0CFC1F" bold="true" box="[570,653,959,980]" pageId="8" pageNumber="99">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype, male (OMNH-Ar-9717), 4.0 mm. Habitus, lateral view. Scale: 1 mm.
</paragraph>
</caption>
<paragraph id="8B9BE274FF92FFC6FF2DFB9DFE4CFB08" blockId="8.[151,1437,1050,2011]" pageId="8" pageNumber="99">
<emphasis id="B9503E66FF92FFC6FF2DFB9DFEBEFBF8" box="[199,319,1051,1075]" italics="true" pageId="8" pageNumber="99">Antenna 1</emphasis>
(
<figureCitation id="131FFEF1FF92FFC6FEBEFB9CFE1DFBF8" box="[340,412,1050,1075]" captionStart="FIGURE 7" captionStartId="9.[151,250,1755,1777]" captionTargetBox="[167,1419,208,1721]" captionTargetId="figure@9.[151,1435,193,1733]" captionTargetPageId="9" captionText="FIGURE 7. Psammogammarus lobatus sp. nov. All but C, holotype, male (OMNH-Ar- 9717), 4.0 mm; C, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, lateral view; B, left antenna 2, lateral view; C, upper lip, anterior view; D, left mandible, lateral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, lateral view; E, right mandible, medial view; E 1, incisor, lacinia mobilis and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, outer plate and palp of left maxilla 1, dorsal view; G 1, inner plate of left maxilla 1, dorsal view; G 2, distal part of left maxilla 1, dorsal view, normal setae omitted; H, right maxilla 2, ventral view; I, right maxilliped, dorsal view; I 1, distal part of inner plate of right maxilliped, dorsal view, normal setae omitted; I 2, distal part of outer plate of right maxilliped, dorsal view, normal setae omitted. Scale: A, 1 mm; A 1, B, 0.5 mm; C I, G 1, 0.25 mm; D 1, E 1, G 2, I 1 2, 0.13 mm." httpUri="https://zenodo.org/record/241806/files/figure.png" pageId="8" pageNumber="99">Fig. 7</figureCitation>
A, A1) long, with ratio of lengths of peduncular articles 13 1:0.7:0.3, article 1 with ventrodistal robust seta; accessory flagellum short, 2-articulated, each article with distal setae; primary flagellum with 11 articles, articles 310 with 1, 2, 1, 1, 1, 0, 1, 1 aesthetascs at each distal end, respectively.
<emphasis id="B9503E66FF92FFC6FB06FBE5FADEFBB0" box="[1260,1375,1123,1147]" italics="true" pageId="8" pageNumber="99">Antenna 2</emphasis>
(
<figureCitation id="131FFEF1FF92FFC6FA84FBE4FF24FB54" captionStart="FIGURE 7" captionStartId="9.[151,250,1755,1777]" captionTargetBox="[167,1419,208,1721]" captionTargetId="figure@9.[151,1435,193,1733]" captionTargetPageId="9" captionText="FIGURE 7. Psammogammarus lobatus sp. nov. All but C, holotype, male (OMNH-Ar- 9717), 4.0 mm; C, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, lateral view; B, left antenna 2, lateral view; C, upper lip, anterior view; D, left mandible, lateral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, lateral view; E, right mandible, medial view; E 1, incisor, lacinia mobilis and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, outer plate and palp of left maxilla 1, dorsal view; G 1, inner plate of left maxilla 1, dorsal view; G 2, distal part of left maxilla 1, dorsal view, normal setae omitted; H, right maxilla 2, ventral view; I, right maxilliped, dorsal view; I 1, distal part of inner plate of right maxilliped, dorsal view, normal setae omitted; I 2, distal part of outer plate of right maxilliped, dorsal view, normal setae omitted. Scale: A, 1 mm; A 1, B, 0.5 mm; C I, G 1, 0.25 mm; D 1, E 1, G 2, I 1 2, 0.13 mm." httpUri="https://zenodo.org/record/241806/files/figure.png" pageId="8" pageNumber="99">Fig. 7</figureCitation>
B) short, slender, with ratio of lengths of peduncular articles 35 1:1.5:1.3; flagellum short, with 6 articles, longer than last peduncular article.
</paragraph>
<paragraph id="8B9BE274FF92FFC6FF2DFB48FC8AF9E1" blockId="8.[151,1437,1050,2011]" pageId="8" pageNumber="99">
<emphasis id="B9503E66FF92FFC6FF2DFB48FEB2FB2C" box="[199,307,1230,1255]" italics="true" pageId="8" pageNumber="99">Mandible</emphasis>
(
<figureCitation id="131FFEF1FF92FFC6FEAFFB48FE0DFB2D" box="[325,396,1230,1255]" captionStart="FIGURE 7" captionStartId="9.[151,250,1755,1777]" captionTargetBox="[167,1419,208,1721]" captionTargetId="figure@9.[151,1435,193,1733]" captionTargetPageId="9" captionText="FIGURE 7. Psammogammarus lobatus sp. nov. All but C, holotype, male (OMNH-Ar- 9717), 4.0 mm; C, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, lateral view; B, left antenna 2, lateral view; C, upper lip, anterior view; D, left mandible, lateral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, lateral view; E, right mandible, medial view; E 1, incisor, lacinia mobilis and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, outer plate and palp of left maxilla 1, dorsal view; G 1, inner plate of left maxilla 1, dorsal view; G 2, distal part of left maxilla 1, dorsal view, normal setae omitted; H, right maxilla 2, ventral view; I, right maxilliped, dorsal view; I 1, distal part of inner plate of right maxilliped, dorsal view, normal setae omitted; I 2, distal part of outer plate of right maxilliped, dorsal view, normal setae omitted. Scale: A, 1 mm; A 1, B, 0.5 mm; C I, G 1, 0.25 mm; D 1, E 1, G 2, I 1 2, 0.13 mm." httpUri="https://zenodo.org/record/241806/files/figure.png" pageId="8" pageNumber="99">Fig. 7</figureCitation>
D, D1, E, E1), each incisor bearing 5 cusps, left lacinia mobilis parallelogramatic, distal margin minutely serrated, right lacinia mobilis with 3 cusps, each accessory blade
<quantity id="4CDC4F91FF92FFC6FBDDFB74FBE3FAC0" box="[1079,1122,1266,1291]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="8" pageNumber="99" unit="in" value="6.0">6 in</quantity>
number; molar strong; palp article length ratio 1:2.2:
<quantity id="4CDC4F91FF92FFC6FE44FA91FE74FAE4" box="[430,501,1302,1327]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="8" pageNumber="99" unit="in" value="1.2">1.2 in</quantity>
left (probably regenerated) and 1:2.5:
<quantity id="4CDC4F91FF92FFC6FC4FFA90FC6DFAE4" box="[933,1004,1302,1327]" metricMagnitude="-2" metricUnit="m" metricValue="4.826" pageId="8" pageNumber="99" unit="in" value="1.9">1.9 in</quantity>
right, article 2 with 4 setae ventrally, article 3 with 3 and 5 setae in left and right, respectively.
<emphasis id="B9503E66FF92FFC6FCF0FABDFC07FA98" box="[794,902,1338,1363]" italics="true" pageId="8" pageNumber="99">Lower lip</emphasis>
(
<figureCitation id="131FFEF1FF92FFC6FC7FFABCFC59FA98" box="[917,984,1338,1363]" captionStart="FIGURE 7" captionStartId="9.[151,250,1755,1777]" captionTargetBox="[167,1419,208,1721]" captionTargetId="figure@9.[151,1435,193,1733]" captionTargetPageId="9" captionText="FIGURE 7. Psammogammarus lobatus sp. nov. All but C, holotype, male (OMNH-Ar- 9717), 4.0 mm; C, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, lateral view; B, left antenna 2, lateral view; C, upper lip, anterior view; D, left mandible, lateral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, lateral view; E, right mandible, medial view; E 1, incisor, lacinia mobilis and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, outer plate and palp of left maxilla 1, dorsal view; G 1, inner plate of left maxilla 1, dorsal view; G 2, distal part of left maxilla 1, dorsal view, normal setae omitted; H, right maxilla 2, ventral view; I, right maxilliped, dorsal view; I 1, distal part of inner plate of right maxilliped, dorsal view, normal setae omitted; I 2, distal part of outer plate of right maxilliped, dorsal view, normal setae omitted. Scale: A, 1 mm; A 1, B, 0.5 mm; C I, G 1, 0.25 mm; D 1, E 1, G 2, I 1 2, 0.13 mm." httpUri="https://zenodo.org/record/241806/files/figure.png" pageId="8" pageNumber="99">Fig. 7</figureCitation>
F) with small inner lobes, distal margins of outer lobes setose.
<emphasis id="B9503E66FF92FFC6FE7EFAD8FD82FABC" box="[404,515,1374,1399]" italics="true" pageId="8" pageNumber="99">Maxilla 1</emphasis>
(
<figureCitation id="131FFEF1FF92FFC6FDFFFAD8FDD8FABC" box="[533,601,1374,1399]" captionStart="FIGURE 7" captionStartId="9.[151,250,1755,1777]" captionTargetBox="[167,1419,208,1721]" captionTargetId="figure@9.[151,1435,193,1733]" captionTargetPageId="9" captionText="FIGURE 7. Psammogammarus lobatus sp. nov. All but C, holotype, male (OMNH-Ar- 9717), 4.0 mm; C, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, lateral view; B, left antenna 2, lateral view; C, upper lip, anterior view; D, left mandible, lateral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, lateral view; E, right mandible, medial view; E 1, incisor, lacinia mobilis and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, outer plate and palp of left maxilla 1, dorsal view; G 1, inner plate of left maxilla 1, dorsal view; G 2, distal part of left maxilla 1, dorsal view, normal setae omitted; H, right maxilla 2, ventral view; I, right maxilliped, dorsal view; I 1, distal part of inner plate of right maxilliped, dorsal view, normal setae omitted; I 2, distal part of outer plate of right maxilliped, dorsal view, normal setae omitted. Scale: A, 1 mm; A 1, B, 0.5 mm; C I, G 1, 0.25 mm; D 1, E 1, G 2, I 1 2, 0.13 mm." httpUri="https://zenodo.org/record/241806/files/figure.png" pageId="8" pageNumber="99">Fig. 7</figureCitation>
G, G1
<emphasis id="B9503E66FF92FFC6FD77FAD9FD2AFABD" box="[669,683,1375,1398]" italics="true" pageId="8" pageNumber="99"></emphasis>
2), inner plate roundish triangular, with several distal setae; outer plate with 9 robust setae apically; palp 2-articulated, tip of article 2 projected in middle, with 3 robust setae.
<emphasis id="B9503E66FF92FFC6FF7DFA20FE85FA74" box="[151,260,1446,1471]" italics="true" pageId="8" pageNumber="99">Maxilla 2</emphasis>
(
<figureCitation id="131FFEF1FF92FFC6FEFFFA20FED8FA74" box="[277,345,1446,1471]" captionStart="FIGURE 7" captionStartId="9.[151,250,1755,1777]" captionTargetBox="[167,1419,208,1721]" captionTargetId="figure@9.[151,1435,193,1733]" captionTargetPageId="9" captionText="FIGURE 7. Psammogammarus lobatus sp. nov. All but C, holotype, male (OMNH-Ar- 9717), 4.0 mm; C, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, lateral view; B, left antenna 2, lateral view; C, upper lip, anterior view; D, left mandible, lateral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, lateral view; E, right mandible, medial view; E 1, incisor, lacinia mobilis and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, outer plate and palp of left maxilla 1, dorsal view; G 1, inner plate of left maxilla 1, dorsal view; G 2, distal part of left maxilla 1, dorsal view, normal setae omitted; H, right maxilla 2, ventral view; I, right maxilliped, dorsal view; I 1, distal part of inner plate of right maxilliped, dorsal view, normal setae omitted; I 2, distal part of outer plate of right maxilliped, dorsal view, normal setae omitted. Scale: A, 1 mm; A 1, B, 0.5 mm; C I, G 1, 0.25 mm; D 1, E 1, G 2, I 1 2, 0.13 mm." httpUri="https://zenodo.org/record/241806/files/figure.png" pageId="8" pageNumber="99">Fig. 7</figureCitation>
H), inner plate as wide as outer, dorsal surface of inner plate with sparse setae, terminal ends of both plates setose.
<emphasis id="B9503E66FF92FFC6FE99FA4CFE6CFA28" box="[371,493,1482,1507]" italics="true" pageId="8" pageNumber="99">Maxilliped</emphasis>
(
<figureCitation id="131FFEF1FF92FFC6FE14FA4CFDC5FA29" box="[510,580,1482,1507]" captionStart="FIGURE 7" captionStartId="9.[151,250,1755,1777]" captionTargetBox="[167,1419,208,1721]" captionTargetId="figure@9.[151,1435,193,1733]" captionTargetPageId="9" captionText="FIGURE 7. Psammogammarus lobatus sp. nov. All but C, holotype, male (OMNH-Ar- 9717), 4.0 mm; C, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, lateral view; B, left antenna 2, lateral view; C, upper lip, anterior view; D, left mandible, lateral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, lateral view; E, right mandible, medial view; E 1, incisor, lacinia mobilis and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, outer plate and palp of left maxilla 1, dorsal view; G 1, inner plate of left maxilla 1, dorsal view; G 2, distal part of left maxilla 1, dorsal view, normal setae omitted; H, right maxilla 2, ventral view; I, right maxilliped, dorsal view; I 1, distal part of inner plate of right maxilliped, dorsal view, normal setae omitted; I 2, distal part of outer plate of right maxilliped, dorsal view, normal setae omitted. Scale: A, 1 mm; A 1, B, 0.5 mm; C I, G 1, 0.25 mm; D 1, E 1, G 2, I 1 2, 0.13 mm." httpUri="https://zenodo.org/record/241806/files/figure.png" pageId="8" pageNumber="99">
Fig.
<date id="FF9AC4B4FF92FFC6FDDFFA4DFDC5FA29" box="[565,580,1483,1506]" pageId="8" pageNumber="99">7</date>
</figureCitation>
I,
<date id="FF9AC4B4FF92FFC6FDB7FA4DFD12FA28" box="[605,659,1483,1507]" pageId="8" pageNumber="99">
I1
<emphasis id="B9503E66FF92FFC6FD9EFA4DFD03FA29" box="[628,642,1483,1506]" italics="true" pageId="8" pageNumber="99"></emphasis>
2
</date>
), inner plate truncate, distal end bearing 4 short robust setae; outer plate produced distomedially, with 3 long robust setae distally; palp 4-articulated, article 4 with short nail, dorsomedial surface of article 4 covered with thin setae.
</paragraph>
<paragraph id="8B9BE274FF92FFC6FF2DF9B0FB0DF8A4" blockId="8.[151,1437,1050,2011]" pageId="8" pageNumber="99">
<emphasis id="B9503E66FF92FFC6FF2DF9B0FEDFF984" box="[199,350,1590,1615]" italics="true" pageId="8" pageNumber="99">Gnathopod 1</emphasis>
(
<figureCitation id="131FFEF1FF92FFC6FE98F9B0FE39F984" box="[370,440,1590,1615]" captionStart="FIGURE 8" captionStartId="10.[151,250,1952,1974]" captionTargetBox="[152,1433,227,1951]" captionTargetId="figure@10.[151,1436,161,1981]" captionTargetPageId="10" captionText="FIGURE 8. Psammogammarus lobatus sp. nov. Holotype, male (OMNH-Ar- 9717), 4.0 mm. A B, left gnathopods 1 2, lateral views; A 1, B 1, palms of left gnathopods 1 2, lateral views, normal setae omitted; C G, left pereopods 3 7, lateral views; E 1, F 1, G 1, posterior parts of carpi of left pereopods 5 7, lateral views. Scales: A G, 0.5 mm; A 1, B 1, E 1, F 1, G 1, 0.25 mm." httpUri="https://zenodo.org/record/241807/files/figure.png" pageId="8" pageNumber="99">Fig. 8</figureCitation>
A, A1) small, subchelate; coxa short, trapezoidal; basis widened in distal two thirds, posterior margin with 2 long setae; posterior margin of ischium with long seta; posterior margin of merus with long setae, medial surface covered with fine setae; posterior margin of carpus setose; propodus about 1.3 times as long as carpus, palm defined by 2 medial robust setae, bearing 9 minute medial robust setae; dactylus long, curved.
<emphasis id="B9503E66FF92FFC6FF7DF940FEACF914" box="[151,301,1734,1759]" italics="true" pageId="8" pageNumber="99">Gnathopod 2</emphasis>
(
<figureCitation id="131FFEF1FF92FFC6FEAAF940FE07F914" box="[320,390,1734,1759]" captionStart="FIGURE 8" captionStartId="10.[151,250,1952,1974]" captionTargetBox="[152,1433,227,1951]" captionTargetId="figure@10.[151,1436,161,1981]" captionTargetPageId="10" captionText="FIGURE 8. Psammogammarus lobatus sp. nov. Holotype, male (OMNH-Ar- 9717), 4.0 mm. A B, left gnathopods 1 2, lateral views; A 1, B 1, palms of left gnathopods 1 2, lateral views, normal setae omitted; C G, left pereopods 3 7, lateral views; E 1, F 1, G 1, posterior parts of carpi of left pereopods 5 7, lateral views. Scales: A G, 0.5 mm; A 1, B 1, E 1, F 1, G 1, 0.25 mm." httpUri="https://zenodo.org/record/241807/files/figure.png" pageId="8" pageNumber="99">Fig. 8</figureCitation>
B, B1) large, subchelate; coxa short, rectangular, gill curved posteriorly; basis widened in distal two thirds, posterior margin with 3 long setae; carpus lobed, posterior margin setose; propodus dilated, about 3.1 times as long as carpus, posterior margin with several bundle of setae; palm oblique, palmer margin composed of 2 lobes, 1 excavation and 2 large medial robust setae, first lobe with 2 lateral and 3 medial small robust setae, second lobe with 1 lateral and 2 medial small robust setae; dactylus stout, strongly curved.
</paragraph>
<paragraph id="8B9BE274FF92FFC2FF2DF8FCFF6DFDF7" blockId="8.[151,1437,1050,2011]" lastBlockId="12.[151,1437,151,1760]" lastPageId="12" lastPageNumber="103" pageId="8" pageNumber="99">
<emphasis id="B9503E66FF92FFC6FF2DF8FCFEF1F858" box="[199,368,1914,1939]" italics="true" pageId="8" pageNumber="99">Pereopods 34</emphasis>
(
<figureCitation id="131FFEF1FF92FFC6FE6BF8FCFE42F858" box="[385,451,1914,1939]" captionStart="FIGURE 8" captionStartId="10.[151,250,1952,1974]" captionTargetBox="[152,1433,227,1951]" captionTargetId="figure@10.[151,1436,161,1981]" captionTargetPageId="10" captionText="FIGURE 8. Psammogammarus lobatus sp. nov. Holotype, male (OMNH-Ar- 9717), 4.0 mm. A B, left gnathopods 1 2, lateral views; A 1, B 1, palms of left gnathopods 1 2, lateral views, normal setae omitted; C G, left pereopods 3 7, lateral views; E 1, F 1, G 1, posterior parts of carpi of left pereopods 5 7, lateral views. Scales: A G, 0.5 mm; A 1, B 1, E 1, F 1, G 1, 0.25 mm." httpUri="https://zenodo.org/record/241807/files/figure.png" pageId="8" pageNumber="99">Fig. 8</figureCitation>
C
<emphasis id="B9503E66FF92FFC6FE3DF8FDFE64F859" box="[471,485,1915,1938]" italics="true" pageId="8" pageNumber="99"></emphasis>
D) similar, slender; coxae short, trapezoidal, gills large, oval; bases linear, posterior margins with a few long setae; meri slender, widened distally; dactyli narrow, almost straight.
<emphasis id="B9503E66FF92FFC6FB58F818FAB3F87D" box="[1202,1330,1950,1975]" italics="true" pageId="8" pageNumber="99">Pereopod 5</emphasis>
(
<figureCitation id="131FFEF1FF92FFC6FAA8F818FA02F87C" box="[1346,1411,1950,1975]" captionStart="FIGURE 8" captionStartId="10.[151,250,1952,1974]" captionTargetBox="[152,1433,227,1951]" captionTargetId="figure@10.[151,1436,161,1981]" captionTargetPageId="10" captionText="FIGURE 8. Psammogammarus lobatus sp. nov. Holotype, male (OMNH-Ar- 9717), 4.0 mm. A B, left gnathopods 1 2, lateral views; A 1, B 1, palms of left gnathopods 1 2, lateral views, normal setae omitted; C G, left pereopods 3 7, lateral views; E 1, F 1, G 1, posterior parts of carpi of left pereopods 5 7, lateral views. Scales: A G, 0.5 mm; A 1, B 1, E 1, F 1, G 1, 0.25 mm." httpUri="https://zenodo.org/record/241807/files/figure.png" pageId="8" pageNumber="99">Fig. 8</figureCitation>
E, E1), posterior lobe of coxa about 0.6 times as long as anterior lobe; basis rectangular, posterodistal corner projected distally; posterodistal corner of merus with 2 robust setae; anterior margin and anterodistal and posterodistal corners of carpus with 1, 3 and 3 robust setae, respectively; anterior margin and anterodistal corner of propodus each bearing robust seta; dactylus narrow.
<emphasis id="B9503E66FF96FFC2FD9AFF59FD71FF33" box="[624,752,223,248]" italics="true" pageId="12" pageNumber="103">Pereopod 6</emphasis>
(
<figureCitation id="131FFEF1FF96FFC2FD15FF59FCC1FF33" box="[767,832,223,248]" captionStart="FIGURE 8" captionStartId="10.[151,250,1952,1974]" captionTargetBox="[152,1433,227,1951]" captionTargetId="figure@10.[151,1436,161,1981]" captionTargetPageId="10" captionText="FIGURE 8. Psammogammarus lobatus sp. nov. Holotype, male (OMNH-Ar- 9717), 4.0 mm. A B, left gnathopods 1 2, lateral views; A 1, B 1, palms of left gnathopods 1 2, lateral views, normal setae omitted; C G, left pereopods 3 7, lateral views; E 1, F 1, G 1, posterior parts of carpi of left pereopods 5 7, lateral views. Scales: A G, 0.5 mm; A 1, B 1, E 1, F 1, G 1, 0.25 mm." httpUri="https://zenodo.org/record/241807/files/figure.png" pageId="12" pageNumber="103">Fig. 8</figureCitation>
F, F1) about 1.5 times as long as pereopod 5; posterior lobe of coxa 85 % length of anterior lobe; basis rectangular, posterodistal corner projected distally like pereopod 5; posterior margin and posterodistal corner of merus with 1 and 2 robust setae, respectively; anterior margin and anterodistal and posterodistal corners of carpus with 2, 3 and 3 robust setae, respectively; anterior margin and anterodistal and posterodistal corners of propodus each bearing robust seta; dactylus narrow.
<emphasis id="B9503E66FF96FFC2FB5AFEE9FAB0FE4C" box="[1200,1329,367,392]" italics="true" pageId="12" pageNumber="103">Pereopod 7</emphasis>
(
<figureCitation id="131FFEF1FF96FFC2FAABFEE9FA03FE43" box="[1345,1410,367,392]" captionStart="FIGURE 8" captionStartId="10.[151,250,1952,1974]" captionTargetBox="[152,1433,227,1951]" captionTargetId="figure@10.[151,1436,161,1981]" captionTargetPageId="10" captionText="FIGURE 8. Psammogammarus lobatus sp. nov. Holotype, male (OMNH-Ar- 9717), 4.0 mm. A B, left gnathopods 1 2, lateral views; A 1, B 1, palms of left gnathopods 1 2, lateral views, normal setae omitted; C G, left pereopods 3 7, lateral views; E 1, F 1, G 1, posterior parts of carpi of left pereopods 5 7, lateral views. Scales: A G, 0.5 mm; A 1, B 1, E 1, F 1, G 1, 0.25 mm." httpUri="https://zenodo.org/record/241807/files/figure.png" pageId="12" pageNumber="103">Fig. 8</figureCitation>
G, G1) about 1.2 times as long as pereopod 6; coxa short triangular, not lobed; basis rectangular, posterodistal corner projected distally like pereopods 56; posterior margin and anterodistal and posterodistal corners of merus with 1, 1 and 2 robust setae, respectively; anterior margin and anterodistal and posterodistal corners of carpus with 2, 3 and 2 robust setae, respectively; anterior margin and anterodistal corner of propodus each bearing robust seta; dactylus narrow.
</paragraph>
<caption id="DF5BB2FCFF93FFC7FF7DF95DFE09F82D" httpUri="https://zenodo.org/record/241806/files/figure.png" pageId="9" pageNumber="100" targetBox="[167,1419,208,1721]" targetPageId="9">
<paragraph id="8B9BE274FF93FFC7FF7DF95DFE09F82D" blockId="9.[151,1436,1755,2022]" pageId="9" pageNumber="100">
<emphasis id="B9503E66FF93FFC7FF7DF95DFE94F93B" bold="true" box="[151,277,1755,1777]" pageId="9" pageNumber="100">FIGURE 7.</emphasis>
<taxonomicName id="4C2499F7FF93FFC7FEF7F95AFDB3F93A" box="[285,562,1755,1777]" class="Malacostraca" family="Melitidae" genus="Psammogammarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="lobatus" status="sp. nov.">
<emphasis id="B9503E66FF93FFC7FEF7F95AFDB3F93A" box="[285,562,1755,1777]" italics="true" pageId="9" pageNumber="100">Psammogammarus lobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A263831DFF93FFC7FDD1F95DFD0CF93B" box="[571,653,1755,1776]" pageId="9" pageNumber="100" rank="species">
<emphasis id="B9503E66FF93FFC7FDD1F95DFD0CF93B" bold="true" box="[571,653,1755,1776]" pageId="9" pageNumber="100">sp. nov.</emphasis>
</taxonomicNameLabel>
All but C, holotype, male (OMNH-Ar-9717), 4.0 mm; C, paratype, female (OMNH-Ar-9719), 4.2 mm. A, left antenna 1, lateral view; A1, accessory flagellum of left antenna 1, lateral view; B, left antenna 2, lateral view; C, upper lip, anterior view; D, left mandible, lateral view; D1, incisor, lacinia mobilis and accessory blades of left mandible, lateral view; E, right mandible, medial view; E1, incisor, lacinia mobilis and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, outer plate and palp of left maxilla 1, dorsal view; G1, inner plate of left maxilla 1, dorsal view; G2, distal part of left maxilla 1, dorsal view, normal setae omitted; H, right maxilla 2, ventral view; I, right maxilliped, dorsal view; I1, distal part of inner plate of right maxilliped, dorsal view, normal setae omitted; I2, distal part of outer plate of right maxilliped, dorsal view, normal setae omitted. Scale: A, 1 mm; A1, B, 0.5 mm; CI, G1, 0.25 mm; D1, E1, G2, I12, 0.13 mm.
</paragraph>
</caption>
<caption id="DF5BB2FCFF90FFC4FF7DF826FADFF83F" httpUri="https://zenodo.org/record/241807/files/figure.png" pageId="10" pageNumber="101" targetBox="[152,1433,227,1951]" targetPageId="10">
<paragraph id="8B9BE274FF90FFC4FF7DF826FADFF83F" blockId="10.[151,1436,1952,2036]" pageId="10" pageNumber="101">
<emphasis id="B9503E66FF90FFC4FF7DF826FE93F87E" bold="true" box="[151,274,1952,1974]" pageId="10" pageNumber="101">FIGURE 8.</emphasis>
<taxonomicName id="4C2499F7FF90FFC4FEF2F824FDAAF87D" box="[280,555,1952,1975]" class="Malacostraca" family="Melitidae" genus="Psammogammarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="lobatus" status="sp. nov.">
<emphasis id="B9503E66FF90FFC4FEF2F824FDAAF87D" box="[280,555,1952,1975]" italics="true" pageId="10" pageNumber="101">Psammogammarus lobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A263831DFF90FFC4FDDBF827FD00F87D" box="[561,641,1953,1974]" pageId="10" pageNumber="101" rank="species">
<emphasis id="B9503E66FF90FFC4FDDBF827FD00F87D" bold="true" box="[561,641,1953,1974]" pageId="10" pageNumber="101">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype, male (OMNH-Ar-9717), 4.0 mm. AB, left gnathopods 12, lateral views; A1, B1, palms of left gnathopods 12, lateral views, normal setae omitted; CG, left pereopods 37, lateral views; E1, F1, G1, posterior parts of carpi of left pereopods 57, lateral views. Scales: AG, 0.5 mm; A1, B1, E1, F1, G1, 0.25 mm.
</paragraph>
</caption>
<caption id="DF5BB2FCFF91FFC5FF7DF8E4FBCEF839" httpUri="https://zenodo.org/record/241808/files/figure.png" pageId="11" pageNumber="102" targetBox="[184,1375,201,1888]" targetPageId="11">
<paragraph id="8B9BE274FF91FFC5FF7DF8E4FBCEF839" blockId="11.[151,1436,1890,2034]" pageId="11" pageNumber="102">
<emphasis id="B9503E66FF91FFC5FF7DF8E4FE96F8BC" bold="true" box="[151,279,1890,1912]" pageId="11" pageNumber="102">FIGURE 9.</emphasis>
<taxonomicName id="4C2499F7FF91FFC5FEC8F8E5FDBBF8B3" box="[290,570,1890,1912]" class="Malacostraca" family="Melitidae" genus="Psammogammarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="lobatus" status="sp. nov.">
<emphasis id="B9503E66FF91FFC5FEC8F8E5FDBBF8B3" box="[290,570,1890,1912]" italics="true" pageId="11" pageNumber="102">Psammogammarus lobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A263831DFF91FFC5FDAEF8E4FD18F8BC" box="[580,665,1890,1911]" pageId="11" pageNumber="102" rank="species">
<emphasis id="B9503E66FF91FFC5FDAEF8E4FD18F8BC" bold="true" box="[580,665,1890,1911]" pageId="11" pageNumber="102">sp. nov.</emphasis>
</taxonomicNameLabel>
AH, holotype, male (OMNH-Ar-9717), 4.0 mm; IJ, paratype, female (OMNH-Ar-9719), 4.2 mm. A, right pleonal epimera 13, lateral view; B, left pleopod 1, posterior view; C, left pleopod 2, anterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsolateral view; F, left uropod 2, dorsomedial view; G, left uropod 3, dorsal view; H, telson, dorsal view; IJ, left gnathopods 12, lateral views; I1, J1, palms of left gnathopods 12, lateral views, normal setae omitted. Scale: G, 0.75 mm; AF, IJ, 0.5 mm; H, I1, J1, 0.25 mm.
</paragraph>
</caption>
<paragraph id="8B9BE274FF96FFC2FF2DFDC1FEDAFB0F" blockId="12.[151,1437,151,1760]" pageId="12" pageNumber="103">
<emphasis id="B9503E66FF96FFC2FF2DFDC1FE07FDAB" box="[199,390,583,608]" italics="true" pageId="12" pageNumber="103">Pleonal epimera</emphasis>
(
<figureCitation id="131FFEF1FF96FFC2FE73FDC1FE61FDAB" box="[409,480,583,608]" captionStart="FIGURE 9" captionStartId="11.[151,250,1890,1912]" captionTargetBox="[184,1375,201,1888]" captionTargetId="figure@11.[171,1416,149,1913]" captionTargetPageId="11" captionText="FIGURE 9. Psammogammarus lobatus sp. nov. A H, holotype, male (OMNH-Ar- 9717), 4.0 mm; I J, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, posterior view; C, left pleopod 2, anterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsolateral view; F, left uropod 2, dorsomedial view; G, left uropod 3, dorsal view; H, telson, dorsal view; I J, left gnathopods 1 2, lateral views; I 1, J 1, palms of left gnathopods 1 2, lateral views, normal setae omitted. Scale: G, 0.75 mm; A F, I J, 0.5 mm; H, I 1, J 1, 0.25 mm." httpUri="https://zenodo.org/record/241808/files/figure.png" pageId="12" pageNumber="103">Fig. 9</figureCitation>
A), each posteroventral corner quadrate, each posterior margin with 2 short setae (epimeron 3 probably lost 1 seta), ventral margin of epimeron 3 bearing short seta, posteroventral corners of epimera 2
<emphasis id="B9503E66FF96FFC2FEE1FD17FE98FD63" box="[267,281,657,680]" italics="true" pageId="12" pageNumber="103"></emphasis>
3 minutely projected.
<emphasis id="B9503E66FF96FFC2FDF6FD09FD3BFD63" box="[540,698,655,680]" italics="true" pageId="12" pageNumber="103">Pleopods 13</emphasis>
(
<figureCitation id="131FFEF1FF96FFC2FD25FD09FC97FD63" box="[719,790,655,680]" captionStart="FIGURE 9" captionStartId="11.[151,250,1890,1912]" captionTargetBox="[184,1375,201,1888]" captionTargetId="figure@11.[171,1416,149,1913]" captionTargetPageId="11" captionText="FIGURE 9. Psammogammarus lobatus sp. nov. A H, holotype, male (OMNH-Ar- 9717), 4.0 mm; I J, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, posterior view; C, left pleopod 2, anterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsolateral view; F, left uropod 2, dorsomedial view; G, left uropod 3, dorsal view; H, telson, dorsal view; I J, left gnathopods 1 2, lateral views; I 1, J 1, palms of left gnathopods 1 2, lateral views, normal setae omitted. Scale: G, 0.75 mm; A F, I J, 0.5 mm; H, I 1, J 1, 0.25 mm." httpUri="https://zenodo.org/record/241808/files/figure.png" pageId="12" pageNumber="103">Fig. 9</figureCitation>
B
<emphasis id="B9503E66FF96FFC2FCC2FD17FCB7FD63" box="[808,822,657,680]" italics="true" pageId="12" pageNumber="103"></emphasis>
D), pleopod 3 shortest; peduncles long, each with 2 coupling hooks and distal projection; inner rami longer than outer, inner and outer rami with 7 and 8 articles in pleopod 1, 6 and 6 articles in pleopod 2, and 4 and 5 articles in pleopod 3, respectively.
<emphasis id="B9503E66FF96FFC2FB52FD51FAA9FD3B" box="[1208,1320,727,752]" italics="true" pageId="12" pageNumber="103">Uropod 1</emphasis>
(
<figureCitation id="131FFEF1FF96FFC2FAD1FD51FA03FD3B" box="[1339,1410,727,752]" captionStart="FIGURE 9" captionStartId="11.[151,250,1890,1912]" captionTargetBox="[184,1375,201,1888]" captionTargetId="figure@11.[171,1416,149,1913]" captionTargetPageId="11" captionText="FIGURE 9. Psammogammarus lobatus sp. nov. A H, holotype, male (OMNH-Ar- 9717), 4.0 mm; I J, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, posterior view; C, left pleopod 2, anterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsolateral view; F, left uropod 2, dorsomedial view; G, left uropod 3, dorsal view; H, telson, dorsal view; I J, left gnathopods 1 2, lateral views; I 1, J 1, palms of left gnathopods 1 2, lateral views, normal setae omitted. Scale: G, 0.75 mm; A F, I J, 0.5 mm; H, I 1, J 1, 0.25 mm." httpUri="https://zenodo.org/record/241808/files/figure.png" pageId="12" pageNumber="103">Fig. 9</figureCitation>
E) biramous, long; peduncle bearing 1 ventrofacial and 1 dorsolateral robust setae, distomedial and distolateral corners with 2 (1 long and 1 short) and 2 robust setae, respectively; inner ramus almost same length as peduncle, with 5 distal robust setae; outer ramus 83 % length of peduncle, with 1 lateral and 5 distal robust setae.
<emphasis id="B9503E66FF96FFC2FADAFCC2FA1AFC97" box="[1328,1435,836,861]" italics="true" pageId="12" pageNumber="103">Uropod 2</emphasis>
(
<figureCitation id="131FFEF1FF96FFC2FF75FCE1FF63FC4B" box="[159,226,871,896]" captionStart="FIGURE 9" captionStartId="11.[151,250,1890,1912]" captionTargetBox="[184,1375,201,1888]" captionTargetId="figure@11.[171,1416,149,1913]" captionTargetPageId="11" captionText="FIGURE 9. Psammogammarus lobatus sp. nov. A H, holotype, male (OMNH-Ar- 9717), 4.0 mm; I J, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, posterior view; C, left pleopod 2, anterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsolateral view; F, left uropod 2, dorsomedial view; G, left uropod 3, dorsal view; H, telson, dorsal view; I J, left gnathopods 1 2, lateral views; I 1, J 1, palms of left gnathopods 1 2, lateral views, normal setae omitted. Scale: G, 0.75 mm; A F, I J, 0.5 mm; H, I 1, J 1, 0.25 mm." httpUri="https://zenodo.org/record/241808/files/figure.png" pageId="12" pageNumber="103">Fig. 9</figureCitation>
F) biramous, short, 65 % as long as uropod 1; peduncle short, bearing dorsolateral robust seta, distomedial and distolateral corners with 2 and 1 robust setae, respectively; inner ramus about 1.1 times as long as peduncle, with 1 medial, 1 lateral and 5 distal robust setae; outer ramus 94 % length of peduncle, with 1 lateral and 4 distal robust setae.
<emphasis id="B9503E66FF96FFC2FEC1FC52FE17FC27" box="[299,406,980,1005]" italics="true" pageId="12" pageNumber="103">Uropod 3</emphasis>
(
<figureCitation id="131FFEF1FF96FFC2FE4CFC52FE68FC27" box="[422,489,980,1005]" captionStart="FIGURE 9" captionStartId="11.[151,250,1890,1912]" captionTargetBox="[184,1375,201,1888]" captionTargetId="figure@11.[171,1416,149,1913]" captionTargetPageId="11" captionText="FIGURE 9. Psammogammarus lobatus sp. nov. A H, holotype, male (OMNH-Ar- 9717), 4.0 mm; I J, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, posterior view; C, left pleopod 2, anterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsolateral view; F, left uropod 2, dorsomedial view; G, left uropod 3, dorsal view; H, telson, dorsal view; I J, left gnathopods 1 2, lateral views; I 1, J 1, palms of left gnathopods 1 2, lateral views, normal setae omitted. Scale: G, 0.75 mm; A F, I J, 0.5 mm; H, I 1, J 1, 0.25 mm." httpUri="https://zenodo.org/record/241808/files/figure.png" pageId="12" pageNumber="103">Fig. 9</figureCitation>
G) very large, about 2.8 times as long as uropod 1, biramous; peduncle short, with 3 ventral robust setae; inner ramus very short, scale-like, 60 % length of peduncle, with seta on tip; outer ramus long, about 6.0 times as long as peduncle; article 1 sparsely setose on medial and lateral margins, distomedial and distolateral corners each with 3 robust setae, article 2 about 1.4 times as long as article 1, sparsely setose on medial and lateral margins, distal margin with many long setae.
<emphasis id="B9503E66FF96FFC2FCD2FBE2FCFFFBB6" box="[824,894,1124,1149]" italics="true" pageId="12" pageNumber="103">Telson</emphasis>
(
<figureCitation id="131FFEF1FF96FFC2FC79FBE2FC5BFBB7" box="[915,986,1124,1149]" captionStart="FIGURE 9" captionStartId="11.[151,250,1890,1912]" captionTargetBox="[184,1375,201,1888]" captionTargetId="figure@11.[171,1416,149,1913]" captionTargetPageId="11" captionText="FIGURE 9. Psammogammarus lobatus sp. nov. A H, holotype, male (OMNH-Ar- 9717), 4.0 mm; I J, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, posterior view; C, left pleopod 2, anterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsolateral view; F, left uropod 2, dorsomedial view; G, left uropod 3, dorsal view; H, telson, dorsal view; I J, left gnathopods 1 2, lateral views; I 1, J 1, palms of left gnathopods 1 2, lateral views, normal setae omitted. Scale: G, 0.75 mm; A F, I J, 0.5 mm; H, I 1, J 1, 0.25 mm." httpUri="https://zenodo.org/record/241808/files/figure.png" pageId="12" pageNumber="103">Fig. 9</figureCitation>
H), breadth subequal to length, deeply incised; each lateral margin with 2 robust setae; distal margins bearing 12 robust setae, dorsal surface with 2 pairs of plumose setae.
</paragraph>
<paragraph id="8B9BE274FF96FFC2FF2DFB49FD89FA56" blockId="12.[151,1437,151,1760]" pageId="12" pageNumber="103">
Female [
<typeStatus id="549F5CD6FF96FFC2FEDAFB57FE17FB22" box="[304,406,1233,1257]" pageId="12" pageNumber="103" type="paratype">paratype</typeStatus>
,
<quantity id="4CDC4F91FF96FFC2FE41FB56FD8AFB23" box="[427,523,1232,1256]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="103" unit="mm" value="4.2">4.2 mm</quantity>
(OMNH-Ar-9719)]. Generally similar to male except for gnathopods and oostegites.
<emphasis id="B9503E66FF96FFC2FEFEFB72FE26FAC7" box="[276,423,1268,1293]" italics="true" pageId="12" pageNumber="103">Gnathopod 1</emphasis>
(
<figureCitation id="131FFEF1FF96FFC2FE5DFB72FE7BFAC7" box="[439,506,1268,1293]" captionStart="FIGURE 9" captionStartId="11.[151,250,1890,1912]" captionTargetBox="[184,1375,201,1888]" captionTargetId="figure@11.[171,1416,149,1913]" captionTargetPageId="11" captionText="FIGURE 9. Psammogammarus lobatus sp. nov. A H, holotype, male (OMNH-Ar- 9717), 4.0 mm; I J, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, posterior view; C, left pleopod 2, anterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsolateral view; F, left uropod 2, dorsomedial view; G, left uropod 3, dorsal view; H, telson, dorsal view; I J, left gnathopods 1 2, lateral views; I 1, J 1, palms of left gnathopods 1 2, lateral views, normal setae omitted. Scale: G, 0.75 mm; A F, I J, 0.5 mm; H, I 1, J 1, 0.25 mm." httpUri="https://zenodo.org/record/241808/files/figure.png" pageId="12" pageNumber="103">
Fig.
<date id="FF9AC4B4FF96FFC2FE01FB72FE7BFAC7" box="[491,506,1268,1292]" pageId="12" pageNumber="103">9</date>
</figureCitation>
I,
<date id="FF9AC4B4FF96FFC2FDFAFB72FDA9FAC7" box="[528,552,1268,1292]" pageId="12" pageNumber="103">I1</date>
), carpus wider, propodus longer than that of male; propodus about 1.4 times as long as carpus, palm defined by 2 medial robust setae, bearing 11 minute medial robust setae.
<emphasis id="B9503E66FF96FFC2FB20FA91FADFFAFB" box="[1226,1374,1303,1328]" italics="true" pageId="12" pageNumber="103">Gnathopod 2</emphasis>
(
<figureCitation id="131FFEF1FF96FFC2FA84FA91FF27FA9F" captionStart="FIGURE 9" captionStartId="11.[151,250,1890,1912]" captionTargetBox="[184,1375,201,1888]" captionTargetId="figure@11.[171,1416,149,1913]" captionTargetPageId="11" captionText="FIGURE 9. Psammogammarus lobatus sp. nov. A H, holotype, male (OMNH-Ar- 9717), 4.0 mm; I J, paratype, female (OMNH-Ar- 9719), 4.2 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, posterior view; C, left pleopod 2, anterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsolateral view; F, left uropod 2, dorsomedial view; G, left uropod 3, dorsal view; H, telson, dorsal view; I J, left gnathopods 1 2, lateral views; I 1, J 1, palms of left gnathopods 1 2, lateral views, normal setae omitted. Scale: G, 0.75 mm; A F, I J, 0.5 mm; H, I 1, J 1, 0.25 mm." httpUri="https://zenodo.org/record/241808/files/figure.png" pageId="12" pageNumber="103">Fig. 9</figureCitation>
J, J1) smaller than that of male; coxa roundish rhomboidal; carpus longer; propodus shorter, about 1.9 times as long as carpus, palm without excavation, defined by 2 medial robust setae, bearing 8 minute medial robust setae.
<emphasis id="B9503E66FF96FFC2FF7DFA03FE8CFA57" box="[151,269,1413,1436]" italics="true" pageId="12" pageNumber="103">Oostegites</emphasis>
present on coxae 25.
</paragraph>
</subSubSection>
<subSubSection id="C33EB1FFFF96FFC2FF2DFA21FB7BFA0B" box="[199,1274,1447,1472]" pageId="12" pageNumber="103" type="etymology">
<paragraph id="8B9BE274FF96FFC2FF2DFA21FB7BFA0B" blockId="12.[151,1437,151,1760]" box="[199,1274,1447,1472]" pageId="12" pageNumber="103">
<emphasis id="B9503E66FF96FFC2FF2DFA21FECFFA0B" bold="true" box="[199,334,1447,1472]" pageId="12" pageNumber="103">Etymology.</emphasis>
From the Latin
<taxonomicName id="4C2499F7FF96FFC2FDE9FA21FDD7FA0B" box="[515,598,1447,1472]" class="Malacostraca" family="Melitidae" genus="Psammogammarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="103" phylum="Arthropoda" rank="species" species="lobatus">
<emphasis id="B9503E66FF96FFC2FDE9FA21FDD7FA0B" box="[515,598,1447,1472]" italics="true" pageId="12" pageNumber="103">lobatus</emphasis>
</taxonomicName>
(= lobate), referring to the shape of bases of pereopods 57.
</paragraph>
</subSubSection>
<subSubSection id="C33EB1FFFF96FFC2FF2DFA4AFB5DF92B" pageId="12" pageNumber="103" type="discussion">
<paragraph id="8B9BE274FF96FFC2FF2DFA4AFDADF976" blockId="12.[151,1437,151,1760]" pageId="12" pageNumber="103">
<emphasis id="B9503E66FF96FFC2FF2DFA4AFEBAFA2E" bold="true" box="[199,315,1484,1509]" pageId="12" pageNumber="103">Remarks.</emphasis>
<taxonomicName id="4C2499F7FF96FFC2FEA8FA4BFDF2FA2E" box="[322,627,1484,1509]" class="Malacostraca" family="Melitidae" genus="Psammogammarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="103" phylum="Arthropoda" rank="species" species="lobatus" status="sp. nov.">
<emphasis id="B9503E66FF96FFC2FEA8FA4BFDF2FA2E" box="[322,627,1484,1509]" italics="true" pageId="12" pageNumber="103">Psammogammarus lobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A263831DFF96FFC2FD90FA4AFD52FA2E" box="[634,723,1484,1509]" pageId="12" pageNumber="103" rank="species">
<emphasis id="B9503E66FF96FFC2FD90FA4AFD52FA2E" bold="true" box="[634,723,1484,1509]" pageId="12" pageNumber="103">sp. nov.</emphasis>
</taxonomicNameLabel>
is characterized by (1) male gnathopod 2 with excavated palm, (2) posterodistally projected bases of pereopods 57, (3) quadrate posteroventral corner of pleonal epimeron 3, (4) short inner ramus of uropod 3, and (5) article 2 of uropod 3 outer ramus longer than article 1. Although 15 species have been recorded so far in the genus
<taxonomicName id="4C2499F7FF96FFC2FD9EF9BFFBC0F99B" authority="Jaume et al. 2013" authorityName="Jaume et al." authorityYear="2013" box="[628,1089,1591,1616]" class="Malacostraca" family="Melitidae" genus="Psammogammarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B9503E66FF96FFC2FD9EF9BFFCCBF99B" box="[628,842,1593,1616]" italics="true" pageId="12" pageNumber="103">Psammogammarus</emphasis>
(
<bibRefCitation id="EFB59F85FF96FFC2FCB4F9BEFBB8F99B" author="Jaume" box="[862,1081,1591,1616]" pageId="12" pageNumber="103" refString="Jaume, D., Iliffe, T. M. &amp; van der Ham, J. L. (2013) A new Psammogammarus (Amphipoda: Eriopisidae) from anchialine pools on the Exuma Cays, Bahamas. Zootaxa, 3700 (1), 48 - 64. http: // dx. doi. org / 10.11646 / zootaxa. 3700.1.2" type="journal article" year="2013">
Jaume
<emphasis id="B9503E66FF96FFC2FC5EF9BFFC70F99B" box="[948,1009,1591,1616]" italics="true" pageId="12" pageNumber="103">et al.</emphasis>
2013
</bibRefCitation>
)
</taxonomicName>
, none of the species has this combination of the characters [see
<bibRefCitation id="EFB59F85FF96FFC2FDAEF9DAFCE2F9BF" box="[580,867,1628,1653]" pageId="12" pageNumber="103" refString="Tomikawa, K., Kakui, K. &amp; Yamasaki, H. (2010) A new species of Psammogammarus (Amphipoda: Melitidae) from Kuchinoerabu Island, Japan, with a note on its feeding habits. Zoological Science, 27, 615 - 626. http: // dx. doi. org / 10.2108 / zsj. 27.615" type="journal article">
Tomikawa
<emphasis id="B9503E66FF96FFC2FD22F9DBFC81F9BE" box="[712,768,1628,1653]" italics="true" pageId="12" pageNumber="103">et al</emphasis>
. (2010)
</bibRefCitation>
,
<bibRefCitation id="EFB59F85FF96FFC2FC9CF9DAFBD6F9BF" box="[886,1111,1628,1653]" pageId="12" pageNumber="103" refString="Vonk, R., Hoeksema, B. W. &amp; Jaume, D. (2011) A new marine interstitial Psammogammarus (Crustacea, Amphipoda, Melitidae) from Gura Ici Island, off western Halmahera (North Moluccas, Indonesia), and an overview of the genus. ZooKeys, 128, 53 - 73." type="journal article">
Vonk
<emphasis id="B9503E66FF96FFC2FC55F9DBFC7EF9BE" box="[959,1023,1628,1653]" italics="true" pageId="12" pageNumber="103">et al.</emphasis>
(2011)
</bibRefCitation>
and
<bibRefCitation id="EFB59F85FF96FFC2FB77F9DBFA13F9BF" box="[1181,1426,1628,1653]" pageId="12" pageNumber="103" refString="Jaume, D., Iliffe, T. M. &amp; van der Ham, J. L. (2013) A new Psammogammarus (Amphipoda: Eriopisidae) from anchialine pools on the Exuma Cays, Bahamas. Zootaxa, 3700 (1), 48 - 64. http: // dx. doi. org / 10.11646 / zootaxa. 3700.1.2" type="journal article">
Jaume
<emphasis id="B9503E66FF96FFC2FB19F9DBFAAAF9BE" box="[1267,1323,1628,1653]" italics="true" pageId="12" pageNumber="103">et al</emphasis>
. (2013)
</bibRefCitation>
].
<taxonomicName id="4C2499F7FF96FFC2FF7DF907FE69F953" box="[151,488,1665,1688]" class="Malacostraca" family="Melitidae" genus="Psammogammarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="103" phylum="Arthropoda" rank="species" species="mawatarii">
<emphasis id="B9503E66FF96FFC2FF7DF907FE69F953" box="[151,488,1665,1688]" italics="true" pageId="12" pageNumber="103">Psammogammarus mawatarii</emphasis>
</taxonomicName>
from
<collectingCountry id="F333A2E4FF96FFC2FDC6F907FDEDF953" box="[556,620,1665,1688]" name="Japan" pageId="12" pageNumber="103">Japan</collectingCountry>
can be distinguished from this new species by the shape of posterodistal lobes of bases in the pereopods 57.
</paragraph>
<paragraph id="8B9BE274FF96FFC2FF2DF941FB5DF92B" blockId="12.[151,1437,151,1760]" box="[199,1244,1735,1760]" pageId="12" pageNumber="103">
<emphasis id="B9503E66FF96FFC2FF2DF941FEABF92B" bold="true" box="[199,298,1735,1760]" pageId="12" pageNumber="103">Habitat.</emphasis>
Brackish with fluctuating salinity. Reverse surface of large stone on sandy tidal flat.
</paragraph>
</subSubSection>
</treatment>
</document>