treatments-xml/data/E6/16/87/E61687C1FFA4FFDD7BB4BFACFCF91843.xml
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<document id="455997434A457D6B0A7CABBFB49F7CB2" ID-CLB-Dataset="2767" ID-DOI="10.11646/zootaxa.5087.3.2" ID-GBIF-Dataset="43c1b1e7-1797-4ed7-817d-606114b807ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="5826858" ID-ZooBank="6253FEED-A4FB-4832-AE51-91E4F51655C4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641539129469" checkinUser="plazi" docAuthor="Paz, Fernando, Ratcliffe, Brett C. &amp; Figueroa, Luis" docDate="2022" docId="E61687C1FFA4FFDD7BB4BFACFCF91843" docLanguage="en" docName="zootaxa.5087.3.2.pdf" docOrigin="Zootaxa 5087 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyclocephala hylaea Ratcliffe &amp; Paz. A 2022, new species" docType="treatment" docUuid="55F91F65-3C63-4AFD-9CA4-3F54DFE5E3A4" docUuidSource="ZooBank" docVersion="7" lastPageNumber="435" masterDocId="1A2FFFB9FFA1FFD57B23BF3BFFAE1A5A" masterDocTitle="Three new species of Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae) from Amazonian Peru and a checklist of Cyclocephala species in Peru" masterLastPageNumber="440" masterPageNumber="427" pageNumber="432" updateTime="1699296570230" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BC6FF05937818C87FC534D34C3C7009F">Three new species of Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae) from Amazonian Peru and a checklist of Cyclocephala species in Peru</mods:title>
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<mods:namePart id="7DE1C803C6DC8BE19A2620D8ACCA683E">Paz, Fernando</mods:namePart>
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<mods:affiliation id="4EDE7F9004031A7AC992D0CECBC25908">Departamento de Entomología, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Apartado Postal 14 - 0434, Lima, Peru. fernandopaz 683 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6331 - 4416</mods:affiliation>
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<mods:namePart id="8A28C8B22ED83A05FB7712C4F0C4898B">Ratcliffe, Brett C.</mods:namePart>
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<mods:affiliation id="580F7D89E941D7E972BCAB1584F43AAD">Systematics Research Collections, University of Nebraska State Museum, University of Nebraska, Lincoln, Nebraska, 68588 - 0338, United States of America. bratcliffe 1 @ unl. edu; http: // orcid. org / 0000 - 0002 - 0589 - 8791</mods:affiliation>
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<mods:namePart id="CC45C1EDFB0438FC23C6F03778B8D4C6">Figueroa, Luis</mods:namePart>
<mods:affiliation id="5C906189F9C7193CD9D928C0F72A1B20">Departamento de Entomología, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Lima, Peru.</mods:affiliation>
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<taxonomicName id="A9BF4D54FFA4FFD07BB4BFACFDF41AEB" authority="Ratcliffe &amp; Paz" authorityName="Ratcliffe &amp; Paz. A" authorityYear="2022" box="[151,602,151,178]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="432" phylum="Arthropoda" rank="species" species="hylaea" status="sp. nov.">
<emphasis id="5CCBEAC5FFA4FFD07BB4BFACFE3F1AE8" bold="true" box="[151,401,151,178]" italics="true" pageId="5" pageNumber="432">Cyclocephala hylaea</emphasis>
Ratcliffe &amp; Paz
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,
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(
<figureCitation id="F6842A52FFA4FFD07BBCBF80FE881A8F" box="[159,294,187,213]" captionStart-0="FIGURES 1319" captionStart-1="FIGURE 20" captionStartId-0="6.[151,264,1944,1968]" captionStartId-1="7.[151,250,1943,1967]" captionTargetBox-0="[224,1362,184,1917]" captionTargetBox-1="[198,1390,181,1919]" captionTargetId-0="figure-23@6.[214,1374,181,1920]" captionTargetId-1="figure-18@7.[198,1390,181,1920]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 1319. Cyclocephala hylaea. 13, Male holotype; 14, female allotype; 15, variation of pronotal and elytral markings, all specimens from same collecting event; 1617, parameres of holotype; 18, protarsal claw of holotype; 19, epipleuron (ventral view) of allotype. Scale bar = 5 mm." captionText-1="FIGURE 20. Distribution map of the new species of Peruvian Cyclocephala." figureDoi-0="http://doi.org/10.5281/zenodo.5826866" figureDoi-1="http://doi.org/10.5281/zenodo.5826870" httpUri-0="https://zenodo.org/record/5826866/files/figure.png" httpUri-1="https://zenodo.org/record/5826870/files/figure.png" pageId="5" pageNumber="432">Figs. 1320</figureCitation>
)
</paragraph>
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<emphasis id="5CCBEAC5FFA4FFD07BB4BE38FEE81B47" bold="true" box="[151,326,259,285]" pageId="5" pageNumber="432">Type material.</emphasis>
<materialsCitation id="DED73C8AFFA4FFD07A6DBE38FE0B1B3F" ID-GBIF-Occurrence="3427663319" collectingDate="1979-11-11" collectingDateMax="1979-11-15" collectingDateMin="1979-11-11" collectorName="J. B. Heppner" country="Peru" elevation="290" location="Rio Tambopata Res" pageId="5" pageNumber="432" specimenCount="290" stateProvince="Madre de Dios" typeStatus="holotype">
<typeStatus id="B1048875FFA4FFD07A6DBE38FE181B47" box="[334,438,259,285]" pageId="5" pageNumber="432" type="holotype">Holotype</typeStatus>
male and
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female labeled: “
<collectingCountry id="16A87647FFA4FFD0786EBE38FC351B47" box="[845,923,259,285]" name="Peru" pageId="5" pageNumber="432">PERU</collectingCountry>
:
<collectingRegion id="AC7BF835FFA4FFD07885BE38FBFF1B47" box="[934,1105,259,285]" country="Peru" name="Madre de Dios" pageId="5" pageNumber="432">Madre de Dios</collectingRegion>
; /
<location id="6B60600CFFA4FFD07F4DBE38FAE01B44" LSID="urn:lsid:plazi:treatment:E61687C1FFA4FFDD7BB4BFACFCF91843:6B60600CFFA4FFD07F4DBE38FAE01B44" box="[1134,1358,259,286]" country="Peru" name="Rio Tambopata Res" pageId="5" pageNumber="432" stateProvince="Madre de Dios">Rio Tambopata Res</location>
; 30 air / km. SW Pto. Maldonado,
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<quantity id="A9479B32FFA4FFD07AEABE1CFDA31B1B" box="[457,525,295,321]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="432" unit="m" value="290.0">
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</quantity>
.
</specimenCount>
/
<date id="1A011017FFA4FFD07905BE1CFD611B1B" box="[550,719,295,322]" pageId="5" pageNumber="432" value="1979-11-11" valueMax="1979-11-15" valueMin="1979-11-11">
<collectingDate id="0A45E9FFFFA4FFD07905BE1CFD611B1B" box="[550,719,295,322]" pageId="5" pageNumber="432" value="1979-11-11" valueMax="1979-11-15" valueMin="1979-11-11">1115 XI 1979</collectingDate>
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<collectorName id="C34A5301FFA4FFD079F5BE13FCC11B18" box="[726,879,296,322]" pageId="5" pageNumber="432">J. B. Heppner</collectorName>
/ subtropical moist forest” and with our red
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and
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labels
</materialsCitation>
.
<materialsCitation id="DED73C8AFFA4FFD07A93BE77FA861B3C" ID-GBIF-Occurrence="3427663312" box="[432,1320,331,358]" collectingDate="1979-11-02" collectingDateMax="1979-11-05" collectingDateMin="1979-11-02" collectorName="J. B. Heppner" country="Peru" elevation="290" location="Rio Tambopata Res" pageId="5" pageNumber="432" specimenCount="16" specimenCount-female="10" specimenCount-male="6" stateProvince="Madre de Dios" typeStatus="paratype">
<typeStatus id="B1048875FFA4FFD07A93BE77FDB31B3C" box="[432,541,332,358]" pageId="5" pageNumber="432" type="paratype">Paratypes</typeStatus>
with same data except dates of “
<date id="1A011017FFA4FFD078ADBE70FB8F1B3F" box="[910,1057,331,358]" pageId="5" pageNumber="432" value="1979-11-02" valueMax="1979-11-05" valueMin="1979-11-02">
<collectingDate id="0A45E9FFFFA4FFD078ADBE70FB8F1B3F" box="[910,1057,331,358]" pageId="5" pageNumber="432" value="1979-11-02" valueMax="1979-11-05" valueMin="1979-11-02">25 XI 1979</collectingDate>
</date>
” (
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,
<specimenCount id="78B9FD5EFFA4FFD07F82BE70FA8E1B3C" box="[1185,1312,331,358]" pageId="5" pageNumber="432" type="female">10 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD07E10BE70FE4D1BD0" ID-GBIF-Occurrence="3427663316" collectingDate="1979-11-06" collectingDateMax="1979-11-10" collectingDateMin="1979-11-06" collectorName="J. B. Heppner" country="Peru" elevation="290" isTailEnumeration="true" location="Rio Tambopata Res" pageId="5" pageNumber="432" specimenCount="49" specimenCount-female="33" specimenCount-male="16" stateProvince="Madre de Dios" typeStatus="paratype">
<date id="1A011017FFA4FFD07E1EBE70FF7F1BD3" pageId="5" pageNumber="432" value="1979-11-06" valueMax="1979-11-10" valueMin="1979-11-06">
<collectingDate id="0A45E9FFFFA4FFD07E1EBE70FF7F1BD3" pageId="5" pageNumber="432" value="1979-11-06" valueMax="1979-11-10" valueMin="1979-11-06">610 XI 1979</collectingDate>
</date>
” (
<specimenCount id="78B9FD5EFFA4FFD07BC8BE54FEFC1BD3" box="[235,338,367,393]" pageId="5" pageNumber="432" type="male">16 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07A7EBE54FE721BD0" box="[349,476,367,394]" pageId="5" pageNumber="432" type="female">33 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD07ACDBE54FC6A1BD0" ID-GBIF-Occurrence="3427663301" box="[494,964,367,394]" collectingDate="1979-11-11" collectingDateMax="1979-11-15" collectingDateMin="1979-11-11" country="Peru" pageId="5" pageNumber="432" specimenCount="198" specimenCount-female="101" specimenCount-male="97" stateProvince="Madre de Dios" typeStatus="paratype">
<date id="1A011017FFA4FFD07ADABE54FD0A1BD3" box="[505,676,367,394]" pageId="5" pageNumber="432" value="1979-11-11" valueMax="1979-11-15" valueMin="1979-11-11">
<collectingDate id="0A45E9FFFFA4FFD07ADABE54FD0A1BD3" box="[505,676,367,394]" pageId="5" pageNumber="432" value="1979-11-11" valueMax="1979-11-15" valueMin="1979-11-11">1115 XI 1979</collectingDate>
</date>
” (
<specimenCount id="78B9FD5EFFA4FFD0799EBE54FC8B1BD3" box="[701,805,367,393]" pageId="5" pageNumber="432" type="male">97 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07813BE54FC131BD0" box="[816,957,367,394]" pageId="5" pageNumber="432" type="female">101 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD078ECBE54FA361BD0" ID-GBIF-Occurrence="3427663323" box="[975,1432,367,394]" collectingDate="1979-11-16" collectingDateMax="1979-11-20" collectingDateMin="1979-11-16" country="Peru" pageId="5" pageNumber="432" specimenCount="75" specimenCount-female="51" specimenCount-male="24" stateProvince="Madre de Dios" typeStatus="paratype">
<date id="1A011017FFA4FFD078FABE54FB281BD3" box="[985,1158,367,394]" pageId="5" pageNumber="432" value="1979-11-16" valueMax="1979-11-20" valueMin="1979-11-16">
<collectingDate id="0A45E9FFFFA4FFD078FABE54FB281BD3" box="[985,1158,367,394]" pageId="5" pageNumber="432" value="1979-11-16" valueMax="1979-11-20" valueMin="1979-11-16">1620 XI 1979</collectingDate>
</date>
” (
<specimenCount id="78B9FD5EFFA4FFD07FBCBE54FAA91BD3" box="[1183,1287,367,393]" pageId="5" pageNumber="432" type="male">24 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07E31BE54FA3F1BD0" box="[1298,1425,367,394]" pageId="5" pageNumber="432" type="female">51 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD07BB4BEA8FDC31BF4" ID-GBIF-Occurrence="3427663313" box="[151,621,403,430]" collectingDate="1979-11-21" collectingDateMax="1979-11-25" collectingDateMin="1979-11-21" country="Peru" pageId="5" pageNumber="432" specimenCount="75" specimenCount-female="43" specimenCount-male="32" stateProvince="Madre de Dios" typeStatus="paratype">
<date id="1A011017FFA4FFD07B82BEA8FEFC1BF7" box="[161,338,403,430]" pageId="5" pageNumber="432" value="1979-11-21" valueMax="1979-11-25" valueMin="1979-11-21">
<collectingDate id="0A45E9FFFFA4FFD07B82BEA8FEFC1BF7" box="[161,338,403,430]" pageId="5" pageNumber="432" value="1979-11-21" valueMax="1979-11-25" valueMin="1979-11-21">2125 XI 1979</collectingDate>
</date>
” (
<specimenCount id="78B9FD5EFFA4FFD07A4EBEA8FE791BF7" box="[365,471,403,429]" pageId="5" pageNumber="432" type="male">32 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07AC7BEA8FDCB1BF4" box="[484,613,403,430]" pageId="5" pageNumber="432" type="female">43 females</specimenCount>
)
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, and “
<date id="1A011017FFA4FFD07997BEA8FCCB1BF7" box="[692,869,403,430]" pageId="5" pageNumber="432" value="1979-11-26" valueMax="1979-11-30" valueMin="1979-11-26">2630 XI 1979</date>
” (
<specimenCount id="78B9FD5EFFA4FFD078A2BEA8FC441BF7" box="[897,1002,403,429]" pageId="5" pageNumber="432" type="male">12 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD078D4BEA8FBD61BF4" box="[1015,1144,403,430]" pageId="5" pageNumber="432" type="female">16 females</specimenCount>
).
<materialsCitation id="DED73C8AFFA4FFD07FA9BEA8FF421BAC" ID-GBIF-Occurrence="3427663305" collectingDate="1979-12-08" collectingDateMax="1979-12-10" collectingDateMin="1979-12-08" collectorName="J. B. Heppner" country="Peru" location="Pilcopata" pageId="5" pageNumber="432" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cusco" typeStatus="paratype">
Additional
<typeStatus id="B1048875FFA4FFD07E28BEAFFAD81BF4" box="[1291,1398,404,430]" pageId="5" pageNumber="432" type="paratype">paratypes</typeStatus>
labeled “
<collectingCountry id="16A87647FFA4FFD07BC7BE8CFE9C1B8B" box="[228,306,439,465]" name="Peru" pageId="5" pageNumber="432">PERU</collectingCountry>
:
<collectingRegion id="AC7BF835FFA4FFD07A1EBE8CFE291B8B" box="[317,391,439,465]" country="Peru" name="Cusco" pageId="5" pageNumber="432">Cusco</collectingRegion>
; /
<location id="6B60600CFFA4FFD07ABCBE8CFDA41B8B" LSID="urn:lsid:plazi:treatment:E61687C1FFA4FFDD7BB4BFACFCF91843:6B60600CFFA4FFD07ABCBE8CFDA41B8B" box="[415,522,439,465]" country="Peru" name="Pilcopata" pageId="5" pageNumber="432" stateProvince="Cusco">Pilcopata</location>
, 600meters /
<date id="1A011017FFA4FFD079BDBE8CFCEC1B8B" box="[670,834,439,466]" pageId="5" pageNumber="432" value="1979-12-08" valueMax="1979-12-10" valueMin="1979-12-08">
<collectingDate id="0A45E9FFFFA4FFD079BDBE8CFCEC1B8B" box="[670,834,439,466]" pageId="5" pageNumber="432" value="1979-12-08" valueMax="1979-12-10" valueMin="1979-12-08">810 XII 1979</collectingDate>
</date>
/
<collectorName id="C34A5301FFA4FFD07874BE83FC5F1B88" box="[855,1009,440,466]" pageId="5" pageNumber="432">J. B. Heppner</collectorName>
, premontane / moist forest” (
<specimenCount id="78B9FD5EFFA4FFD07E1BBE8CFA2D1B8B" box="[1336,1411,439,465]" pageId="5" pageNumber="432" type="male">1 male</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07EADBE8CFF4B1BAC" pageId="5" pageNumber="432" type="female">1 female</specimenCount>
)
</materialsCitation>
<materialsCitation id="DED73C8AFFA4FFD07BD0BEE0FC0A1BAC" ID-GBIF-Occurrence="3427663311" box="[243,932,475,502]" collectingDate="1979-12-11" collectingDateMax="1979-12-14" collectingDateMin="1979-12-11" collectorName="J. B. Heppner" country="Peru" location="Pilcopata" pageId="5" pageNumber="432" specimenCount="29" specimenCount-female="21" specimenCount-male="8" stateProvince="Cusco" typeStatus="paratype">
and same data except “
<date id="1A011017FFA4FFD07AD3BEE0FD0B1BAF" box="[496,677,475,502]" pageId="5" pageNumber="432" value="1979-12-11" valueMax="1979-12-14" valueMin="1979-12-11">
<collectingDate id="0A45E9FFFFA4FFD07AD3BEE0FD0B1BAF" box="[496,677,475,502]" pageId="5" pageNumber="432" value="1979-12-11" valueMax="1979-12-14" valueMin="1979-12-11">1114 XII 1979</collectingDate>
</date>
” (
<specimenCount id="78B9FD5EFFA4FFD0799DBEE0FCB91BAF" box="[702,791,475,501]" pageId="5" pageNumber="432" type="male">8 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07802BEE0FC331BAC" box="[801,925,475,502]" pageId="5" pageNumber="432" type="female">21 females</specimenCount>
)
</materialsCitation>
<materialsCitation id="DED73C8AFFA4FFD07889BEE0FC271840" ID-GBIF-Occurrence="3427663303" collectingDate="1979-11-02" collectingDateMax="1979-11-05" collectingDateMin="1979-11-02" country="Peru" county="rio Tambopata" elevation="290" location="Pto. Maldonado" municipality="Km" pageId="5" pageNumber="432" specimenCount="11" specimenCount-female="8" specimenCount-male="3" stateProvince="Madre de Dios" typeStatus="paratype">
and “
<collectingRegion id="AC7BF835FFA4FFD078C1BEE0FB131BAF" box="[994,1213,475,502]" country="Peru" name="Madre de Dios" pageId="5" pageNumber="432">MADRE DE DIOS</collectingRegion>
/
<collectingCounty id="87614E5BFFA4FFD07FF1BEE0FAD81BAF" box="[1234,1398,475,501]" pageId="5" pageNumber="432">río Tambopata</collectingCounty>
, 30
<collectingMunicipality id="8E64ACADFFA4FFD07BB4BD3BFF6F1840" box="[151,193,512,538]" pageId="5" pageNumber="432">Km</collectingMunicipality>
SO
<location id="6B60600CFFA4FFD07BD0BD3BFE081843" LSID="urn:lsid:plazi:treatment:E61687C1FFA4FFDD7BB4BFACFCF91843:6B60600CFFA4FFD07BD0BD3BFE081843" box="[243,422,511,538]" country="Peru" county="rio Tambopata" municipality="Km" name="Pto. Maldonado" pageId="5" pageNumber="432" stateProvince="Madre de Dios">Pto. Maldonado</location>
/
<quantity id="A9479B32FFA4FFD07A99BEC4FDAD1840" box="[442,515,511,538]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="432" unit="m" value="290.0">
<elevation id="E592D1E4FFA4FFD07A99BEC4FDAD1840" box="[442,515,511,538]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="432" unit="m" value="290.0">290 m</elevation>
</quantity>
,
<date id="1A011017FFA4FFD0792CBEC4FD3F1840" box="[527,657,511,538]" pageId="5" pageNumber="432" value="1979-11-02" valueMax="1979-11-05" valueMin="1979-11-02">
<collectingDate id="0A45E9FFFFA4FFD0792CBEC4FD3F1840" box="[527,657,511,538]" pageId="5" pageNumber="432" value="1979-11-02" valueMax="1979-11-05" valueMin="1979-11-02">2-5.xi.1979</collectingDate>
</date>
” (
<specimenCount id="78B9FD5EFFA4FFD0798FBEC4FCAB1843" box="[684,773,511,537]" pageId="5" pageNumber="432" type="male">3 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07833BEC4FC2F1840" box="[784,897,511,538]" pageId="5" pageNumber="432" type="female">8 females</specimenCount>
)
</materialsCitation>
.
<typeStatus id="B1048875FFA4FFD078B0BD3BFBAE1840" box="[915,1024,512,538]" pageId="5" pageNumber="432" type="paratype">Paratypes</typeStatus>
(483) each with our yellow
<typeStatus id="B1048875FFA4FFD07E1FBD3BFA321840" box="[1340,1436,512,538]" pageId="5" pageNumber="432" type="paratype">paratype</typeStatus>
label.
<typeStatus id="B1048875FFA4FFD07BF9BD18FEEB1867" box="[218,325,547,573]" pageId="5" pageNumber="432" type="holotype">Holotype</typeStatus>
,
<typeStatus id="B1048875FFA4FFD07A6CBD18FE021867" box="[335,428,547,573]" pageId="5" pageNumber="432" type="allotype">allotype</typeStatus>
, and
<specimenCount id="78B9FD5EFFA4FFD07AC7BD18FDD11864" box="[484,639,547,574]" pageId="5" pageNumber="432" type="generic" typeStatus="paratypes">392 paratypes</specimenCount>
(
<specimenCount id="78B9FD5EFFA4FFD079AEBD18FCAC1867" box="[653,770,547,573]" pageId="5" pageNumber="432" type="male">170 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD0782FBD18FC3A1864" box="[780,916,547,574]" pageId="5" pageNumber="432" type="female">222 females</specimenCount>
) deposited in
<materialsCitation id="DED73C8AFFA4FFD07F0DBD18FEAA183B" ID-GBIF-Occurrence="3427663317" collectionCode="USNM" country="Peru" pageId="5" pageNumber="432" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
<collectionCode id="08AEAE12FFA4FFD07F0DBD18FB2D1867" box="[1070,1155,547,573]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="432" type="Museum">USNM</collectionCode>
.Additional
<typeStatus id="B1048875FFA4FFD07E24BD1FFADC1864" box="[1287,1394,548,574]" pageId="5" pageNumber="432" type="paratype">paratypes</typeStatus>
deposited in
</materialsCitation>
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<collectionCode id="08AEAE12FFA4FFD07A29BD7CFEFB183B" box="[266,341,583,609]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="5" pageNumber="432">BCRC</collectionCode>
(
<specimenCount id="78B9FD5EFFA4FFD07A40BD7CFE12183B" box="[355,444,583,609]" pageId="5" pageNumber="432" type="male">7 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07AE5BD7CFD981838" box="[454,566,583,610]" pageId="5" pageNumber="432" type="female">6 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD0796BBD73FC2B1838" ID-GBIF-Occurrence="3427663327" box="[584,901,583,610]" collectionCode="BMNH" country="Peru" pageId="5" pageNumber="432" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Madre de Dios" typeStatus="holotype">
<collectionCode id="08AEAE12FFA4FFD0796BBD73FD321838" box="[584,668,584,610]" pageId="5" pageNumber="432">BMNH</collectionCode>
(
<specimenCount id="78B9FD5EFFA4FFD07988BD7CFCAD183B" box="[683,771,583,609]" pageId="5" pageNumber="432" type="male">3 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD0782DBD7CFCD01838" box="[782,894,583,610]" pageId="5" pageNumber="432" type="female">3 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD078ACBD7CFB651838" ID-GBIF-Occurrence="3427663328" box="[911,1227,583,610]" collectionCode="CMNC" country="Peru" pageId="5" pageNumber="432" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Madre de Dios" typeStatus="holotype">
<collectionCode id="08AEAE12FFA4FFD078ACBD7CFC4F183B" box="[911,993,583,609]" pageId="5" pageNumber="432">CMNC</collectionCode>
(
<specimenCount id="78B9FD5EFFA4FFD078D2BD7CFBE7183B" box="[1009,1097,583,609]" pageId="5" pageNumber="432" type="male">3 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07F70BD7CFB6D1838" box="[1107,1219,583,610]" pageId="5" pageNumber="432" type="female">3 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD07FF6BD7CFF5518DC" ID-GBIF-Occurrence="3427663306" collectionCode="EGPC" country="Peru" pageId="5" pageNumber="432" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Madre de Dios" typeStatus="holotype">
<collectionCode id="08AEAE12FFA4FFD07FF6BD7CFAB3183B" box="[1237,1309,583,609]" pageId="5" pageNumber="432">EGPC</collectionCode>
(
<specimenCount id="78B9FD5EFFA4FFD07E0FBD7CFA2A183B" box="[1324,1412,583,609]" pageId="5" pageNumber="432" type="male">3 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07EADBD7CFF5D18DC" pageId="5" pageNumber="432" type="female">3 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD07A26BD50FD9B18DC" ID-GBIF-Occurrence="3427663310" box="[261,565,619,646]" collectionCode="JNNC" country="Peru" pageId="5" pageNumber="432" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Madre de Dios" typeStatus="holotype">
<collectionCode id="08AEAE12FFA4FFD07A26BD50FEE518DF" box="[261,331,619,645]" pageId="5" pageNumber="432">JNNC</collectionCode>
(
<specimenCount id="78B9FD5EFFA4FFD07A79BD50FE1C18DF" box="[346,434,619,645]" pageId="5" pageNumber="432" type="male">3 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07A9EBD50FD8318DC" box="[445,557,619,646]" pageId="5" pageNumber="432" type="female">3 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD0791CBD50FCCB18DC" ID-GBIF-Occurrence="3427663314" box="[575,869,619,646]" collectionCode="JPSC" country="Peru" pageId="5" pageNumber="432" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Madre de Dios" typeStatus="holotype">
<collectionCode id="08AEAE12FFA4FFD0791CBD50FDD218DF" box="[575,636,619,645]" pageId="5" pageNumber="432">JPSC</collectionCode>
(
<specimenCount id="78B9FD5EFFA4FFD079A8BD50FD4D18DF" box="[651,739,619,645]" pageId="5" pageNumber="432" type="male">3 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD079CEBD50FCF018DC" box="[749,862,619,646]" pageId="5" pageNumber="432" type="female">3 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD0784CBD57FB1318DC" ID-GBIF-Occurrence="3427663308" box="[879,1213,619,646]" collectionCode="MEKRB" country="Peru" pageId="5" pageNumber="432" specimenCount="12" specimenCount-female="7" specimenCount-male="5" stateProvince="Madre de Dios" typeStatus="holotype">
<collectionCode id="08AEAE12FFA4FFD0784CBD57FC7D18DC" box="[879,979,620,646]" pageId="5" pageNumber="432">MEKRB</collectionCode>
(
<specimenCount id="78B9FD5EFFA4FFD078C1BD50FB9418DF" box="[994,1082,619,646]" pageId="5" pageNumber="432" type="male">5 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07F66BD57FB1B18DC" box="[1093,1205,619,646]" pageId="5" pageNumber="432" type="female">7 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD07FE5BD57FF5518F0" ID-GBIF-Occurrence="3427663324" collectionCode="MNHN" country="Peru" pageId="5" pageNumber="432" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Madre de Dios" typeStatus="holotype">
<collectionCode id="08AEAE12FFA4FFD07FE5BD57FAB218DC" box="[1222,1308,620,646]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="432">MNHN</collectionCode>
(
<specimenCount id="78B9FD5EFFA4FFD07E08BD50FA2A18DF" box="[1323,1412,619,645]" pageId="5" pageNumber="432" type="male">3 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07EADBD50FF5D18F0" pageId="5" pageNumber="432" type="female">3 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD07A27BDB4FD9618F0" ID-GBIF-Occurrence="3427663315" box="[260,568,655,682]" collectionCode="MSPC" country="Peru" pageId="5" pageNumber="432" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Madre de Dios" typeStatus="holotype">
<collectionCode id="08AEAE12FFA4FFD07A27BDB4FEE118F3" box="[260,335,655,681]" country="Italy" httpUri="http://grbio.org/cool/t9g4-v7fn" name="Museo di Storia Naturale &amp;quot;Pietro Calderini&amp;quot;" pageId="5" pageNumber="432">MSPC</collectionCode>
(
<specimenCount id="78B9FD5EFFA4FFD07A7DBDB4FE1818F3" box="[350,438,655,681]" pageId="5" pageNumber="432" type="male">2 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07AE3BDB4FD9E18F0" box="[448,560,655,682]" pageId="5" pageNumber="432" type="female">4 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD07962BDB4FC2018F0" ID-GBIF-Occurrence="3427663326" box="[577,910,655,682]" collectionCode="MUSM" country="Peru" pageId="5" pageNumber="432" specimenCount="20" specimenCount-female="12" specimenCount-male="8" stateProvince="Madre de Dios" typeStatus="holotype">
<collectionCode id="08AEAE12FFA4FFD07962BDB4FD3918F3" box="[577,663,655,681]" pageId="5" pageNumber="432">MUSM</collectionCode>
(
<specimenCount id="78B9FD5EFFA4FFD07985BDB4FD5018F3" box="[678,766,655,681]" pageId="5" pageNumber="432" type="male">8 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD0782BBDB4FC2818F0" box="[776,902,655,682]" pageId="5" pageNumber="432" type="female">12 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD078B4BDB4FAEC18F0" ID-GBIF-Occurrence="3427663318" box="[919,1346,655,682]" collectionCode="RDCC" country="Peru" pageId="5" pageNumber="432" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Madre de Dios" typeStatus="holotype">
<collectionCode id="08AEAE12FFA4FFD078B4BDB4FC4D18F3" box="[919,995,655,681]" pageId="5" pageNumber="432">RDCC</collectionCode>
(6), SLTC (
<specimenCount id="78B9FD5EFFA4FFD07F4BBDB4FB6E18F3" box="[1128,1216,655,681]" pageId="5" pageNumber="432" type="male">3 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07FE9BDB4FA9418F0" box="[1226,1338,655,682]" pageId="5" pageNumber="432" type="female">3 females</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA4FFD07E68BDB4FED31894" ID-GBIF-Occurrence="3427663304" collectionCode="UNSM" country="Peru" pageId="5" pageNumber="432" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Madre de Dios" typeStatus="holotype">
<collectionCode id="08AEAE12FFA4FFD07E68BDB4FA3218F3" box="[1355,1436,655,681]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="5" pageNumber="432">UNSM</collectionCode>
(
<specimenCount id="78B9FD5EFFA4FFD07B83BD88FF541897" box="[160,250,691,717]" pageId="5" pageNumber="432" type="male">3 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD07A26BD88FED81894" box="[261,374,691,718]" pageId="5" pageNumber="432" type="female">3 females</specimenCount>
)
</materialsCitation>
, and
<materialsCitation id="DED73C8AFFA4FFD07A9BBD8FFD561894" ID-GBIF-Occurrence="3427663307" box="[440,760,691,718]" collectionCode="ZMHU" country="Peru" pageId="5" pageNumber="432" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Madre de Dios" typeStatus="holotype">
<collectionCode id="08AEAE12FFA4FFD07A9BBD8FFDA41894" box="[440,522,692,718]" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="432">ZMHU</collectionCode>
(
<specimenCount id="78B9FD5EFFA4FFD07939BD88FDDD1897" box="[538,627,691,717]" pageId="5" pageNumber="432" type="male">3 males</specimenCount>
,
<specimenCount id="78B9FD5EFFA4FFD0795CBD88FD5E1894" box="[639,752,691,718]" pageId="5" pageNumber="432" type="female">3 females</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="26A5655CFFA4FFD07BE4BDECFD711CC3" pageId="5" pageNumber="432" type="description">
<paragraph id="6E0036D7FFA4FFD07BE4BDECFDEA1E9F" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="432">
<emphasis id="5CCBEAC5FFA4FFD07BE4BDECFE7818AB" bold="true" box="[199,470,727,753]" pageId="5" pageNumber="432">
Description.
<typeStatus id="B1048875FFA4FFD07A41BDECFE7F18AB" box="[354,465,727,753]" pageId="5" pageNumber="432" type="holotype">Holotype</typeStatus>
,
</emphasis>
male. (
<figureCitation id="F6842A52FFA4FFD0790CBDECFD2A18AB" box="[559,644,727,753]" captionStart="FIGURES 1319" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[224,1362,184,1917]" captionTargetId="figure-23@6.[214,1374,181,1920]" captionTargetPageId="6" captionText="FIGURES 1319. Cyclocephala hylaea. 13, Male holotype; 14, female allotype; 15, variation of pronotal and elytral markings, all specimens from same collecting event; 1617, parameres of holotype; 18, protarsal claw of holotype; 19, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826866" httpUri="https://zenodo.org/record/5826866/files/figure.png" pageId="5" pageNumber="432">Fig. 13</figureCitation>
). Length
<quantity id="A9479B32FFA4FFD079D7BDECFCF018A8" box="[756,862,727,754]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="5" pageNumber="432" unit="mm" value="12.7">12.7 mm</quantity>
; width
<quantity id="A9479B32FFA4FFD07896BDECFBBE18A8" box="[949,1040,727,754]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="432" unit="mm" value="4.9">4.9 mm</quantity>
. Color testaceous except for black frons; 4 small, indistinct black marks on each side of pronotum; and 3 small black maculae on each elytron; pygidium, abdominal sternites, and tarsomeres reddish brown.
<emphasis id="5CCBEAC5FFA4FFD07834BC24FCCE1963" bold="true" box="[791,864,799,825]" pageId="5" pageNumber="432">Head:</emphasis>
Frons and clypeus with punctures moderate in size and density, setigerous; setae short, tawny. Clypeal apex subtruncate, slightly reflexed. Interocular width equals 3.5 transverse eye diameters. Antenna with 10 antennomeres, club subequal in length to antennomeres 27.
<emphasis id="5CCBEAC5FFA4FFD07E39BC5CFA3219DB" bold="true" box="[1306,1436,871,897]" pageId="5" pageNumber="432">Pronotum:</emphasis>
Surface with punctures similar to those of frons, setigerous in anterior angles and along lateral margins; setae short, tawny. Base with marginal bead.
<emphasis id="5CCBEAC5FFA4FFD07928BC94FDCE1993" bold="true" box="[523,608,943,969]" pageId="5" pageNumber="432">Elytra:</emphasis>
Surface weakly punctate-striate, punctures slightly larger and denser than those of pronotum, setigerous; setae minute, tawny, in rows.
<emphasis id="5CCBEAC5FFA4FFD07875BCE8FC7D19B7" bold="true" box="[854,979,979,1005]" pageId="5" pageNumber="432">Pygidium:</emphasis>
Surface opaque with small, moderately dense, setigerous punctures; setae long, dense, tawny. In lateral view, surface convex.
<emphasis id="5CCBEAC5FFA4FFD07FA3BCCCFB6F1E4B" bold="true" box="[1152,1217,1015,1041]" pageId="5" pageNumber="432">Legs:</emphasis>
Protibia bidentate. Protarsomere 4 with large, ventral flange directed forward. Protarsomere 5 enlarged, slightly bent, venter concave; medial claw large, strongly bent, apex entire (
<figureCitation id="F6842A52FFA4FFD079B8BB04FD411E03" box="[667,751,1087,1113]" captionStart="FIGURES 1319" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[224,1362,184,1917]" captionTargetId="figure-23@6.[214,1374,181,1920]" captionTargetPageId="6" captionText="FIGURES 1319. Cyclocephala hylaea. 13, Male holotype; 14, female allotype; 15, variation of pronotal and elytral markings, all specimens from same collecting event; 1617, parameres of holotype; 18, protarsal claw of holotype; 19, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826866" httpUri="https://zenodo.org/record/5826866/files/figure.png" pageId="5" pageNumber="432">Fig. 18</figureCitation>
). Metatarsus much longer than metatibia.
<emphasis id="5CCBEAC5FFA4FFD07FE9BB04FA8D1E03" bold="true" box="[1226,1315,1087,1113]" pageId="5" pageNumber="432">Venter:</emphasis>
Prosternal process small, conical, apex narrowly rounded.
<emphasis id="5CCBEAC5FFA4FFD07992BB58FC951E27" bold="true" box="[689,827,1123,1149]" pageId="5" pageNumber="432">Parameres:</emphasis>
In caudal view, form almost horseshoe-shaped, tapering, inner margin of each paramere with long, dense, tawny setae (
<figureCitation id="F6842A52FFA4FFD078B6BBBCFC441EF8" box="[917,1002,1159,1186]" captionStart="FIGURES 1319" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[224,1362,184,1917]" captionTargetId="figure-23@6.[214,1374,181,1920]" captionTargetPageId="6" captionText="FIGURES 1319. Cyclocephala hylaea. 13, Male holotype; 14, female allotype; 15, variation of pronotal and elytral markings, all specimens from same collecting event; 1617, parameres of holotype; 18, protarsal claw of holotype; 19, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826866" httpUri="https://zenodo.org/record/5826866/files/figure.png" pageId="5" pageNumber="432">Fig. 16</figureCitation>
). In lateral view, basal piece almost 3 times longer than parameres (
<figureCitation id="F6842A52FFA4FFD07AC0BB90FD981E9F" box="[483,566,1195,1221]" captionStart="FIGURES 1319" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[224,1362,184,1917]" captionTargetId="figure-23@6.[214,1374,181,1920]" captionTargetPageId="6" captionText="FIGURES 1319. Cyclocephala hylaea. 13, Male holotype; 14, female allotype; 15, variation of pronotal and elytral markings, all specimens from same collecting event; 1617, parameres of holotype; 18, protarsal claw of holotype; 19, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826866" httpUri="https://zenodo.org/record/5826866/files/figure.png" pageId="5" pageNumber="432">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="6E0036D7FFA4FFD07BE4BBF4FE461F9B" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="432">
<emphasis id="5CCBEAC5FFA4FFD07BE4BBF4FE9D1EB3" bold="true" box="[199,307,1231,1257]" pageId="5" pageNumber="432">
<typeStatus id="B1048875FFA4FFD07BE4BBF4FE801EB3" box="[199,302,1231,1257]" pageId="5" pageNumber="432" type="allotype">Allotype</typeStatus>
,
</emphasis>
female (
<figureCitation id="F6842A52FFA4FFD07ABBBBF4FE421EB3" box="[408,492,1231,1257]" captionStart="FIGURES 1319" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[224,1362,184,1917]" captionTargetId="figure-23@6.[214,1374,181,1920]" captionTargetPageId="6" captionText="FIGURES 1319. Cyclocephala hylaea. 13, Male holotype; 14, female allotype; 15, variation of pronotal and elytral markings, all specimens from same collecting event; 1617, parameres of holotype; 18, protarsal claw of holotype; 19, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826866" httpUri="https://zenodo.org/record/5826866/files/figure.png" pageId="5" pageNumber="432">Fig. 14</figureCitation>
). Length 12.0 mm; width across humeri
<quantity id="A9479B32FFA4FFD07898BBF4FBBB1EB0" box="[955,1045,1231,1258]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="432" unit="mm" value="5.2">5.2 mm</quantity>
. As
<typeStatus id="B1048875FFA4FFD07F65BBF4FB061EB3" box="[1094,1192,1231,1257]" pageId="5" pageNumber="432" type="holotype">holotype</typeStatus>
except in the following respects: Pronotum with 2 small spots and 1 short line on each side; elytral maculae slightly larger; pygidium testaceous.
<emphasis id="5CCBEAC5FFA4FFD07A3BBA2CFECF1F6B" bold="true" box="[280,353,1303,1329]" pageId="5" pageNumber="432">Head:</emphasis>
Surface glabrous.
<emphasis id="5CCBEAC5FFA4FFD07911BA2CFD261F6B" bold="true" box="[562,648,1303,1329]" pageId="5" pageNumber="432">Elytra:</emphasis>
Epipleuron (ventral view) expanded at level of metacoxa and abruptly constricted with a subacute tooth at level of abdominal sternite 3 (
<figureCitation id="F6842A52FFA4FFD078ABBA00FC731F0F" box="[904,989,1339,1365]" captionStart="FIGURES 1319" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[224,1362,184,1917]" captionTargetId="figure-23@6.[214,1374,181,1920]" captionTargetPageId="6" captionText="FIGURES 1319. Cyclocephala hylaea. 13, Male holotype; 14, female allotype; 15, variation of pronotal and elytral markings, all specimens from same collecting event; 1617, parameres of holotype; 18, protarsal claw of holotype; 19, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826866" httpUri="https://zenodo.org/record/5826866/files/figure.png" pageId="5" pageNumber="432">Fig. 19</figureCitation>
). In dorsal view, lateral margin slightly expanded above sternite 3.
<emphasis id="5CCBEAC5FFA4FFD07AF3BA64FDED1F23" bold="true" box="[464,579,1375,1401]" pageId="5" pageNumber="432">Pygidium</emphasis>
: Surface finely reticulated, glabrous except for short, sparse setae on basal half. In lateral view, surface nearly flat.
<emphasis id="5CCBEAC5FFA4FFD0797CBAB8FD0E1FC7" bold="true" box="[607,672,1411,1437]" pageId="5" pageNumber="432">Legs:</emphasis>
Protibia tridentate. Protarsus simple, not enlarged. Metatarsus only slightly longer than metatibia.
</paragraph>
<paragraph id="6E0036D7FFA4FFD07BE4BAF0FD711CC3" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="432">
<emphasis id="5CCBEAC5FFA4FFD07BE4BAF0FDF01FBF" bold="true" box="[199,606,1483,1509]" pageId="5" pageNumber="432">
Variation. Males (
<specimenCount id="78B9FD5EFFA4FFD07A81BAF0FDFC1FBF" box="[418,594,1483,1509]" pageId="5" pageNumber="432" type="generic" typeStatus="paratypes">199 paratypes</specimenCount>
).
</emphasis>
Length 10.513.0 mm; width across humeri 5.0
<quantity id="A9479B32FFA4FFD07FBDBAF0FB541FBC" box="[1182,1274,1483,1510]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="5" pageNumber="432" unit="mm" value="6.2">6.2 mm</quantity>
.
<emphasis id="5CCBEAC5FFA4FFD07E2BBAF0FEB01C53" bold="true" pageId="5" pageNumber="432">
Female (
<specimenCount id="78B9FD5EFFA4FFD07E52BAF0FEBF1C53" pageId="5" pageNumber="432" type="generic" typeStatus="paratypes">284 paratypes</specimenCount>
).
</emphasis>
Length
<quantity id="A9479B32FFA4FFD07A5CBAD4FD891C50" box="[383,551,1519,1546]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" metricValueMax="1.28" metricValueMin="1.06" pageId="5" pageNumber="432" unit="mm" value="11.7" valueMax="12.8" valueMin="10.6">10.612.8 mm</quantity>
; width across humeri
<quantity id="A9479B32FFA4FFD07807BAD4FC011C50" box="[804,943,1519,1546]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="5.8" metricValueMin="4.8" pageId="5" pageNumber="432" unit="mm" value="5.3" valueMax="5.8" valueMin="4.8">4.85.8 mm</quantity>
. The
<typeStatus id="B1048875FFA4FFD078D3BACBFBF51C50" box="[1008,1115,1520,1546]" pageId="5" pageNumber="432" type="paratype">paratypes</typeStatus>
of both sexes display slight variation in the size of the black marks on the pronotum and elytra (
<figureCitation id="F6842A52FFA4FFD078B3B928FC4F1C74" box="[912,993,1555,1582]" captionStart="FIGURES 1319" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[224,1362,184,1917]" captionTargetId="figure-23@6.[214,1374,181,1920]" captionTargetPageId="6" captionText="FIGURES 1319. Cyclocephala hylaea. 13, Male holotype; 14, female allotype; 15, variation of pronotal and elytral markings, all specimens from same collecting event; 1617, parameres of holotype; 18, protarsal claw of holotype; 19, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826866" httpUri="https://zenodo.org/record/5826866/files/figure.png" pageId="5" pageNumber="432">Fig. 15</figureCitation>
) as follows:
<emphasis id="5CCBEAC5FFA4FFD07F51B928FB5A1C77" bold="true" box="[1138,1268,1555,1581]" pageId="5" pageNumber="432">Pronotum:</emphasis>
Spots variably reduced or rarely coalesced into 2 longitudinal, black bands or more rarely coalesced into an almost
<taxonomicName id="A9BF4D54FFA4FFD07E3FB903FEF41C2F" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="432" phylum="Arthropoda" rank="species" species="lunulata">
<emphasis id="5CCBEAC5FFA4FFD07E3FB903FA811C08" box="[1308,1327,1592,1618]" italics="true" pageId="5" pageNumber="432">C</emphasis>
.
<emphasis id="5CCBEAC5FFA4FFD07E1CB90CFA321C0B" box="[1343,1436,1591,1617]" italics="true" pageId="5" pageNumber="432">lunulata</emphasis>
Burmeister, 1847
</taxonomicName>
-like pattern.
<emphasis id="5CCBEAC5FFA4FFD07AD0B960FDE71C2F" bold="true" box="[499,585,1627,1653]" pageId="5" pageNumber="432">Elytra:</emphasis>
Spots variably reduced or rarely expanded in size where the post-scutellar and post-humeral spots connected by a zigzag band.
</paragraph>
</subSubSection>
<subSubSection id="26A5655CFFA4FFD07BE4B998FC9B1CBB" pageId="5" pageNumber="432" type="etymology">
<paragraph id="6E0036D7FFA4FFD07BE4B998FC9B1CBB" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="432">
<emphasis id="5CCBEAC5FFA4FFD07BE4B998FEE01CE7" bold="true" box="[199,334,1699,1725]" pageId="5" pageNumber="432">Etymology.</emphasis>
The species name
<taxonomicName id="A9BF4D54FFA4FFD0793EB998FDC61CE7" authorityName="Ratcliffe &amp; Paz. A" authorityYear="2022" box="[541,616,1699,1725]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="432" phylum="Arthropoda" rank="species" species="hylaea">
<emphasis id="5CCBEAC5FFA4FFD0793EB998FDC61CE7" box="[541,616,1699,1725]" italics="true" pageId="5" pageNumber="432">hylaea</emphasis>
</taxonomicName>
is from the Latin meaning “of the forest”, indicating its habitat. This name should be treated as an adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="26A5655CFFA4FFD07BE4B9D0FCFB1D17" pageId="5" pageNumber="432" type="distribution">
<paragraph id="6E0036D7FFA4FFD07BE4B9D0FDCD1D73" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="432">
<emphasis id="5CCBEAC5FFA4FFD07BE4B9D0FEF71D5F" bold="true" box="[199,345,1771,1797]" pageId="5" pageNumber="432">Distribution</emphasis>
.
<specimenCount id="78B9FD5EFFA4FFD07A48B9D0FDBA1D5F" box="[363,532,1771,1798]" pageId="5" pageNumber="432" type="generic">485 specimens</specimenCount>
examined (
<figureCitation id="F6842A52FFA4FFD079BEB9D0FD5B1D5F" box="[669,757,1771,1797]" captionStart="FIGURE 20" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[198,1390,181,1919]" captionTargetId="figure-18@7.[198,1390,181,1920]" captionTargetPageId="7" captionText="FIGURE 20. Distribution map of the new species of Peruvian Cyclocephala." figureDoi="http://doi.org/10.5281/zenodo.5826870" httpUri="https://zenodo.org/record/5826870/files/figure.png" pageId="5" pageNumber="432">Fig. 20</figureCitation>
).
<collectingCountry id="16A87647FFA4FFD07828B9D7FCEC1D59" box="[779,834,1772,1798]" name="Peru" pageId="5" pageNumber="432">PERU</collectingCountry>
(485):
<collectingRegion id="AC7BF835FFA4FFD078B8B9D0FC561D5F" box="[923,1016,1771,1797]" country="Peru" name="Cusco" pageId="5" pageNumber="432">CUSCO</collectingRegion>
(31): Pilcopata.
<collectingRegion id="AC7BF835FFA4FFD07F9FB9D0FA321D5F" box="[1212,1436,1771,1798]" country="Peru" name="Madre de Dios" pageId="5" pageNumber="432">MADRE DE DIOS</collectingRegion>
(454): Reserva Nacional Rio Tambopata.
</paragraph>
<paragraph id="6E0036D7FFA4FFD07BE4B808FCFB1D17" blockId="5.[151,1437,259,2014]" box="[199,853,1843,1870]" pageId="5" pageNumber="432">
<emphasis id="5CCBEAC5FFA4FFD07BE4B808FE781D17" bold="true" box="[199,470,1843,1869]" pageId="5" pageNumber="432">Temporal distribution.</emphasis>
November (454), December (31).
</paragraph>
</subSubSection>
<subSubSection id="26A5655CFFA4FFDD7BE4B86CFCF91843" lastPageId="8" lastPageNumber="435" pageId="5" pageNumber="432" type="diagnosis">
<paragraph id="6E0036D7FFA4FFDD7BE4B86CFD561B47" blockId="5.[151,1437,259,2014]" lastBlockId="8.[151,1437,151,537]" lastPageId="8" lastPageNumber="435" pageId="5" pageNumber="432">
<emphasis id="5CCBEAC5FFA4FFD07BE4B86CFEEC1D2B" bold="true" box="[199,322,1879,1905]" pageId="5" pageNumber="432">Diagnosis.</emphasis>
The most heavily patterned specimens display an almost “
<taxonomicName id="A9BF4D54FFA4FFD078C2B863FBCE1D2B" authorityName="Burmeister" authorityYear="1847" box="[993,1120,1879,1906]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="432" phylum="Arthropoda" rank="species" species="lunulata">
<emphasis id="5CCBEAC5FFA4FFD078C2B863FC5A1D28" box="[993,1012,1880,1906]" italics="true" pageId="5" pageNumber="432">C</emphasis>
.
<emphasis id="5CCBEAC5FFA4FFD07F20B86CFBCE1D2B" box="[1027,1120,1879,1905]" italics="true" pageId="5" pageNumber="432">lunulata</emphasis>
</taxonomicName>
”-
<typeStatus id="B1048875FFA4FFD07F50B86CFB091D2B" box="[1139,1191,1879,1905]" pageId="5" pageNumber="432">type</typeStatus>
pattern (
<figureCitation id="F6842A52FFA4FFD07E2DB86CFACC1D28" box="[1294,1378,1879,1906]" captionStart="FIGURES 1319" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[224,1362,184,1917]" captionTargetId="figure-23@6.[214,1374,181,1920]" captionTargetPageId="6" captionText="FIGURES 1319. Cyclocephala hylaea. 13, Male holotype; 14, female allotype; 15, variation of pronotal and elytral markings, all specimens from same collecting event; 1617, parameres of holotype; 18, protarsal claw of holotype; 19, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826866" httpUri="https://zenodo.org/record/5826866/files/figure.png" pageId="5" pageNumber="432">Fig. 15</figureCitation>
), but these specimens are not common.
<taxonomicName id="A9BF4D54FFA4FFD0793DB840FCA21DCF" authorityName="Ratcliffe &amp; Paz. A" authorityYear="2022" box="[542,780,1915,1941]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="432" phylum="Arthropoda" rank="species" species="hylaea">
<emphasis id="5CCBEAC5FFA4FFD0793DB840FCA21DCF" box="[542,780,1915,1941]" italics="true" pageId="5" pageNumber="432">Cyclocephala hylaea</emphasis>
</taxonomicName>
is most similar to the sympatric
<taxonomicName id="A9BF4D54FFA4FFD07FA7B847FA321DCF" authority="Herbst, 1790" authorityName="Herbst" authorityYear="1790" box="[1156,1436,1915,1942]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="432" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="5CCBEAC5FFA4FFD07FA7B847FAAC1DCF" box="[1156,1282,1915,1942]" italics="true" pageId="5" pageNumber="432">C. discolor</emphasis>
Herbst,
<quantity id="A9479B32FFA4FFD07E47B840FA321DCF" box="[1380,1436,1915,1941]" metricMagnitude="1" metricUnit="m" metricValue="4.5466" pageId="5" pageNumber="432" unit="in" value="1790.0">1790</quantity>
</taxonomicName>
in
<emphasis id="5CCBEAC5FFA4FFD07B95B8A4FEAF1DE3" box="[182,257,1951,1977]" italics="true" pageId="5" pageNumber="432">gestalt</emphasis>
, but males can be differentiated by the form of the clypeus (apex subtruncate and marginal bead slender to obsolete versus clypeal apex rounded, parabolic, and with a raised, thick marginal bead in
<taxonomicName id="A9BF4D54FFA4FFD07E28B8FFFA251D87" authorityName="Herbst" authorityYear="1790" box="[1291,1419,1987,2014]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="432" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="5CCBEAC5FFA4FFD07E28B8FFFA251D87" box="[1291,1419,1987,2014]" italics="true" pageId="5" pageNumber="432">C. discolor</emphasis>
</taxonomicName>
); large protarsal claw entire versus finely split in
<taxonomicName id="A9BF4D54FFA9FFDD798DBFACFC851AEB" authorityName="Herbst" authorityYear="1790" box="[686,811,151,177]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="5CCBEAC5FFA9FFDD798DBFACFC851AEB" box="[686,811,151,177]" italics="true" pageId="8" pageNumber="435">C. discolor</emphasis>
</taxonomicName>
; and form of the parameres (inner margin of each paramere densely setose versus glabrous in
<taxonomicName id="A9BF4D54FFA9FFDD7973BF80FD621A8F" authorityName="Herbst" authorityYear="1790" box="[592,716,187,213]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="5CCBEAC5FFA9FFDD7973BF80FD621A8F" box="[592,716,187,213]" italics="true" pageId="8" pageNumber="435">C. discolor</emphasis>
</taxonomicName>
). Females are most easily distinguished by the form of the prosternal process: small and conical in
<taxonomicName id="A9BF4D54FFA9FFDD790EBFE4FD371AA3" authorityName="Ratcliffe &amp; Paz. A" authorityYear="2022" box="[557,665,223,249]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="hylaea">
<emphasis id="5CCBEAC5FFA9FFDD790EBFE4FD371AA3" box="[557,665,223,249]" italics="true" pageId="8" pageNumber="435">C. hylaea</emphasis>
</taxonomicName>
, but long, columnar, and with apex obliquely flattened with anterior half to 3/4 elevated into round “button” in
<taxonomicName id="A9BF4D54FFA9FFDD7956BE38FD5F1B47" authorityName="Herbst" authorityYear="1790" box="[629,753,259,285]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="5CCBEAC5FFA9FFDD7956BE38FD5F1B47" box="[629,753,259,285]" italics="true" pageId="8" pageNumber="435">C. discolor</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="3AC0665FFFA7FFD37BB4B8A3FDA51DAD" ID-DOI="http://doi.org/10.5281/zenodo.5826866" ID-Zenodo-Dep="5826866" httpUri="https://zenodo.org/record/5826866/files/figure.png" pageId="6" pageNumber="433" startId="6.[151,264,1944,1968]" targetBox="[224,1362,184,1917]" targetPageId="6">
<paragraph id="6E0036D7FFA7FFD37BB4B8A3FDA51DAD" blockId="6.[151,1437,1944,2040]" pageId="6" pageNumber="433">
<emphasis id="5CCBEAC5FFA7FFD37BB4B8A3FEFD1DEA" bold="true" box="[151,339,1944,1968]" pageId="6" pageNumber="433">FIGURES 1319.</emphasis>
<taxonomicName id="A9BF4D54FFA7FFD37A7AB8A3FD801DEA" authorityName="Ratcliffe &amp; Paz. A" authorityYear="2022" box="[345,558,1944,1968]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="hylaea">
<emphasis id="5CCBEAC5FFA7FFD37A7AB8A3FD801DEA" box="[345,558,1944,1968]" italics="true" pageId="6" pageNumber="433">Cyclocephala hylaea</emphasis>
</taxonomicName>
. 13, Male holotype; 14, female allotype; 15, variation of pronotal and elytral markings, all specimens from same collecting event; 1617, parameres of holotype; 18, protarsal claw of holotype; 19, epipleuron (ventral view) of allotype. Scale bar = 5 mm.
</paragraph>
</caption>
<caption id="3AC0665FFFA6FFD27BB4B8ACFC091DF5" ID-DOI="http://doi.org/10.5281/zenodo.5826870" ID-Zenodo-Dep="5826870" box="[151,935,1943,1968]" httpUri="https://zenodo.org/record/5826870/files/figure.png" pageId="7" pageNumber="434" startId="7.[151,250,1943,1967]" targetBox="[198,1390,181,1919]" targetPageId="7">
<paragraph id="6E0036D7FFA6FFD27BB4B8ACFC091DF5" blockId="7.[151,935,1943,1968]" box="[151,935,1943,1968]" pageId="7" pageNumber="434">
<emphasis id="5CCBEAC5FFA6FFD27BB4B8ACFE8E1DF5" bold="true" box="[151,288,1943,1967]" pageId="7" pageNumber="434">FIGURE 20.</emphasis>
Distribution map of the new species of Peruvian
<taxonomicName id="A9BF4D54FFA6FFD27836B8A3FC0E1DEA" authorityName="Dejean" authorityYear="1821" box="[789,928,1944,1968]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="5CCBEAC5FFA6FFD27836B8A3FC0E1DEA" box="[789,928,1944,1968]" italics="true" pageId="7" pageNumber="434">Cyclocephala</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="6E0036D7FFA9FFDD7BE4BE1CFE111B8B" blockId="8.[151,1437,151,537]" pageId="8" pageNumber="435">
The parameres resemble those of
<taxonomicName id="A9BF4D54FFA9FFDD791CBE1CFC241B18" authority="(Linnaeus, 1767)" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[575,906,295,322]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="5CCBEAC5FFA9FFDD791CBE1CFD6A1B1A" box="[575,708,295,321]" italics="true" pageId="8" pageNumber="435">C. amazona</emphasis>
(Linnaeus, 1767)
</taxonomicName>
(but are broader and not as narrowly elongate), and the male protarsomere 5 is simple, whereas in
<taxonomicName id="A9BF4D54FFA9FFDD79C2BE70FCC61B3E" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[737,872,331,357]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="5CCBEAC5FFA9FFDD79C2BE70FCC61B3E" box="[737,872,331,357]" italics="true" pageId="8" pageNumber="435">C. amazona</emphasis>
</taxonomicName>
there is a distinct tooth near the base on the medio-ventral margin of the tarsomere. Females are very different because the elytral lateral margin is only slightly expanded above sternite
<quantity id="A9479B32FFA9FFDD7A89BEA8FE7A1BF7" box="[426,468,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="435" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="A9BF4D54FFA9FFDD7AFABEA8FDEA1BF7" authorityName="Ratcliffe &amp; Paz. A" authorityYear="2022" box="[473,580,403,429]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="hylaea">
<emphasis id="5CCBEAC5FFA9FFDD7AFABEA8FDEA1BF7" box="[473,580,403,429]" italics="true" pageId="8" pageNumber="435">C. hylaea</emphasis>
</taxonomicName>
, but
<taxonomicName id="A9BF4D54FFA9FFDD7959BEA8FD511BF6" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[634,767,403,429]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="5CCBEAC5FFA9FFDD7959BEA8FD511BF6" box="[634,767,403,429]" italics="true" pageId="8" pageNumber="435">C. amazona</emphasis>
</taxonomicName>
has a large, deeply emarginate swelling on the elytral lateral margin behind the middle.
</paragraph>
<paragraph id="6E0036D7FFA9FFDD7BE4BEE0FCF91843" blockId="8.[151,1437,151,537]" pageId="8" pageNumber="435">
<emphasis id="5CCBEAC5FFA9FFDD7BE4BEE0FE2B1BAE" bold="true" box="[199,389,474,501]" pageId="8" pageNumber="435">Natural history.</emphasis>
<taxonomicName id="A9BF4D54FFA9FFDD7AAEBEE0FDD71BAF" authorityName="Ratcliffe &amp; Paz. A" authorityYear="2022" box="[397,633,475,501]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="hylaea">
<emphasis id="5CCBEAC5FFA9FFDD7AAEBEE0FDD71BAF" box="[397,633,475,501]" italics="true" pageId="8" pageNumber="435">Cyclocephala hylaea</emphasis>
</taxonomicName>
was abundantly collected at lights in lowland rainforest at Reserva Nacional Tambopata and at Pilcopata at elevations of
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290
<specimenCount id="78B9FD5EFFA9FFDD782ABEC4FCFD1843" box="[777,851,511,537]" pageId="8" pageNumber="435" type="generic">600 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>