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<document id="CF17D220B3E80A900E41A74BA2402F40" ID-DOI="10.5281/zenodo.174552" ID-GBIF-Dataset="56c8eab9-101c-4396-8c7a-99497b0218d5" ID-ISSN="1175­5326" ID-Zenodo-Dep="174552" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459784744581" checkinUser="plazi" docAuthor="Holzinger, Werner E. &amp; Kunz, Gernot" docDate="2006" docId="E06E87BD6D49FFD7FED10B7DFA9DF899" docLanguage="en" docName="zt01353p061.pdf" docOrigin="Zootaxa 1353" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Noabennarella Holzinger &amp; Kunz, 2006, gen. nov." docType="treatment" docVersion="9" lastPageNumber="56" masterDocId="1C57FFC56D48FFD4FFD90971FFBFFFD0" masterDocTitle="A new genus and species of Bennarellini from Costa Rica (Hemiptera: Fulgoromorpha: Cixiidae)" masterLastPageNumber="61" masterPageNumber="53" pageNumber="54" updateTime="1698218363684" updateUser="plazi">
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<mods:title id="5C34869ABE79E0C503F643DB6874FB96">A new genus and species of Bennarellini from Costa Rica (Hemiptera: Fulgoromorpha: Cixiidae)</mods:title>
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<mods:namePart id="3549641EC59780CF9353CA15ED0F529E">Holzinger, Werner E.</mods:namePart>
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<mods:namePart id="9B8E92C0B6337EC3E2D7D643D175D48B">Kunz, Gernot</mods:namePart>
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<mods:date id="313B8C74F63F41656FD49FA97ECD6302">2006</mods:date>
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<treatment id="E06E87BD6D49FFD7FED10B7DFA9DF899" ID-DOI="http://doi.org/10.5281/zenodo.6258622" ID-GBIF-Taxon="119348806" ID-Zenodo-Dep="6258622" LSID="urn:lsid:plazi:treatment:E06E87BD6D49FFD7FED10B7DFA9DF899" httpUri="http://treatment.plazi.org/id/E06E87BD6D49FFD7FED10B7DFA9DF899" lastPageId="3" lastPageNumber="56" pageId="1" pageNumber="54">
<subSubSection id="20DD65206D49FFD5FED10B7DFD8DFDF5" box="[264,562,523,550]" pageId="1" pageNumber="54" type="nomenclature">
<paragraph id="687836AB6D49FFD5FED10B7DFD8DFDF5" blockId="1.[264,562,523,550]" box="[264,562,523,550]" pageId="1" pageNumber="54">
<heading id="333081C76D49FFD5FED10B7DFD8DFDF5" bold="true" box="[264,562,523,550]" fontSize="11" level="1" pageId="1" pageNumber="54" reason="1">
<emphasis id="5AB3EAB96D49FFD5FED10B7DFD8DFDF5" bold="true" box="[264,562,523,550]" pageId="1" pageNumber="54">
<taxonomicName id="AFC74D286D49FFD5FED10B7DFE03FDF6" ID-CoL="64JM" box="[264,444,524,550]" class="Insecta" family="Cixiidae" genus="Noabennarella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5AB3EAB96D49FFD5FED10B7DFE03FDF6" bold="true" box="[264,444,524,550]" italics="true" pageId="1" pageNumber="54">Noabennarella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="418057C26D49FFD5FE1D0B7AFD8DFDF5" box="[452,562,523,550]" pageId="1" pageNumber="54" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="20DD65206D49FFD7FED10B26FA9DF899" lastPageId="3" lastPageNumber="56" pageId="1" pageNumber="54" type="materials_examined">
<paragraph id="687836AB6D49FFD5FED10B26FCE0FDA1" blockId="1.[264,1324,599,1828]" box="[264,863,599,625]" pageId="1" pageNumber="54">
<typeStatus id="B77C88096D49FFD5FED10B26FEFDFDA1" box="[264,322,599,625]" pageId="1" pageNumber="54">Type</typeStatus>
species:
<taxonomicName id="AFC74D286D49FFD5FE770B26FD42FDA0" box="[430,765,599,624]" class="Insecta" family="Cixiidae" genus="Noabennarella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="costaricensis" status="sp. nov.">
<emphasis id="5AB3EAB96D49FFD5FE770B26FD42FDA0" box="[430,765,599,624]" italics="true" pageId="1" pageNumber="54">Noabennarella costaricensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="418057C26D49FFD5FCDD0B26FCE0FDA1" box="[772,863,599,625]" pageId="1" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="687836AB6D49FFD5FEE10B0EFB64FD49" blockId="1.[264,1324,599,1828]" box="[312,1243,639,665]" pageId="1" pageNumber="54">Etymology: The name is an arbitrary combination of letters. Gender: Feminine.</paragraph>
<paragraph id="687836AB6D49FFD5FEE10BD6FE7FFD11" blockId="1.[264,1324,599,1828]" box="[312,448,679,705]" pageId="1" pageNumber="54">Description</paragraph>
<paragraph id="687836AB6D49FFD5FEE10BBEFE89FCE9" blockId="1.[264,1324,599,1828]" pageId="1" pageNumber="54">
Medium sized cixiids (body length incl. wings in males about
<quantity id="AF3F9B4E6D49FFD5FBCB0BBEFBE6FD39" box="[1042,1113,719,745]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="54" unit="mm" value="6.0">6 mm</quantity>
, females 8.5 mm) with body laterally compressed and wings in resting position steeply inclined (
<figureCitation id="F0FC2A2E6D49FFD5FB6C0B86FA9CFCC1" box="[1205,1315,759,785]" captionStart="FIGURES 1 2" captionStartId="2.[264,383,1816,1840]" captionTargetBox="[284,1309,326,1788]" captionTargetId="figure@2.[284,1310,326,1789]" captionTargetPageId="2" captionText="FIGURES 1 2. Habitus of Noabennarella costaricensis spec. nov., lateral and dorsal view, with abdominal appendages clearly visible." httpUri="https://zenodo.org/record/174553/files/figure.png" pageId="1" pageNumber="54">Figs 1, 2</figureCitation>
,
<figureCitation id="F0FC2A2E6D49FFD5FED10A6EFE96FCE9" box="[264,297,799,825]" captionStart="FIGURES 23 26. 23" captionStartId="7.[264,382,1775,1799]" captionTargetBox="[277,1310,337,1751]" captionTargetId="figure@7.[277,1310,326,1751]" captionTargetPageId="7" captionText="FIGURES 23 26. 23, Noabennarella costaricensis spec. nov., hind leg; 24, same, apex of tibia of hind leg; 25, Noabennarella spec., inner female genitalia; 26, Noabennarella spec., habitus of female." httpUri="https://zenodo.org/record/174558/files/figure.png" pageId="1" pageNumber="54">26</figureCitation>
).
</paragraph>
<paragraph id="687836AB6D49FFD5FEE10A36FB82FC09" blockId="1.[264,1324,599,1828]" pageId="1" pageNumber="54">
Frons long and slender (
<figureCitation id="F0FC2A2E6D49FFD5FD810A36FD0BFCB1" box="[600,692,839,865]" captionStart="FIGURES 7 10. 7" captionStartId="4.[264,382,1810,1834]" captionTargetBox="[321,1274,551,1795]" captionTargetId="figure@4.[321,1275,551,1795]" captionTargetPageId="4" captionText="FIGURES 7 10. 7, Amazobenna reticulata Penny, head of holotype, lateral aspect (photo: A. Henriques); 8, same, frontal aspect (photo: A. Henriques); 9, Noabennarella costaricensis spec. nov., head, lateral aspect; 10, same, frontal aspect." httpUri="https://zenodo.org/record/174555/files/figure.png" pageId="1" pageNumber="54">Figs 10</figureCitation>
,
<figureCitation id="F0FC2A2E6D49FFD5FD180A36FD5FFCB1" box="[705,736,839,865]" captionStart="FIGURES 11 15. 11" captionStartId="5.[264,382,1543,1567]" captionTargetBox="[316,1271,319,1524]" captionTargetId="figure@5.[316,1279,312,1540]" captionTargetPageId="5" captionText="FIGURES 11 15. 11, Noabennarella costaricensis spec. nov., hind wing; 12, same, fore wing; 13, same, head in frontal aspect; 14, same, head and thorax in dorsal aspect; 15, Bennarella bicoloripennis Muir, head and thorax in dorsal aspect. Scale bar: 1 mm." httpUri="https://zenodo.org/record/174556/files/figure.png" pageId="1" pageNumber="54">13</figureCitation>
), evenly rounded in lateral aspect, without carina on vertex.
<collectingCountry id="10D0763B6D49FFD5FE560A1EFE75FC59" box="[399,458,879,905]" name="India" pageId="1" pageNumber="54">Hind</collectingCountry>
border straight. Median keel distinct, lateral keels strongly produced, semicircular in lateral view (
<figureCitation id="F0FC2A2E6D49FFD5FD830AE6FD6CFC61" box="[602,723,919,945]" captionStart="FIGURES 7 10. 7" captionStartId="4.[264,382,1810,1834]" captionTargetBox="[321,1274,551,1795]" captionTargetId="figure@4.[321,1275,551,1795]" captionTargetPageId="4" captionText="FIGURES 7 10. 7, Amazobenna reticulata Penny, head of holotype, lateral aspect (photo: A. Henriques); 8, same, frontal aspect (photo: A. Henriques); 9, Noabennarella costaricensis spec. nov., head, lateral aspect; 10, same, frontal aspect." httpUri="https://zenodo.org/record/174555/files/figure.png" pageId="1" pageNumber="54">Figs 9, 10</figureCitation>
,
<figureCitation id="F0FC2A2E6D49FFD5FD380AE6FC91FC61" box="[737,814,919,945]" captionStart="FIGURES 11 15. 11" captionStartId="5.[264,382,1543,1567]" captionTargetBox="[316,1271,319,1524]" captionTargetId="figure@5.[316,1279,312,1540]" captionTargetPageId="5" captionText="FIGURES 11 15. 11, Noabennarella costaricensis spec. nov., hind wing; 12, same, fore wing; 13, same, head in frontal aspect; 14, same, head and thorax in dorsal aspect; 15, Bennarella bicoloripennis Muir, head and thorax in dorsal aspect. Scale bar: 1 mm." httpUri="https://zenodo.org/record/174556/files/figure.png" pageId="1" pageNumber="54">13, 14</figureCitation>
). Median and lateral ocelli distinct. Scapus short, ringlike, pedicellus cylindrical, about 1.5 times as long as wide.
</paragraph>
<paragraph id="687836AB6D49FFD5FEE10A96FCAFFB19" blockId="1.[264,1324,599,1828]" pageId="1" pageNumber="54">
Pronotum short, lateral carinae distinct. Mesonotum with indistinct median and distinct lateral carinae. Fore wings long, slender, surpassing tip of abdomen by half their total length (
<figureCitation id="F0FC2A2E6D49FFD5FE460D46FE5AFB81" box="[415,485,1079,1105]" captionStart="FIGURES 1 2" captionStartId="2.[264,383,1816,1840]" captionTargetBox="[284,1309,326,1788]" captionTargetId="figure@2.[284,1310,326,1789]" captionTargetPageId="2" captionText="FIGURES 1 2. Habitus of Noabennarella costaricensis spec. nov., lateral and dorsal view, with abdominal appendages clearly visible." httpUri="https://zenodo.org/record/174553/files/figure.png" pageId="1" pageNumber="54">Fig. 1</figureCitation>
). R and MA three­forked, MP bifurcate. Cubital area of the fore wing reduced in favour of the medial area, thus fore wings slightly widened apically (
<figureCitation id="F0FC2A2E6D49FFD5FB1A0D2EFAA2FBA9" box="[1219,1309,1119,1145]" captionStart="FIGURES 11 15. 11" captionStartId="5.[264,382,1543,1567]" captionTargetBox="[316,1271,319,1524]" captionTargetId="figure@5.[316,1279,312,1540]" captionTargetPageId="5" captionText="FIGURES 11 15. 11, Noabennarella costaricensis spec. nov., hind wing; 12, same, fore wing; 13, same, head in frontal aspect; 14, same, head and thorax in dorsal aspect; 15, Bennarella bicoloripennis Muir, head and thorax in dorsal aspect. Scale bar: 1 mm." httpUri="https://zenodo.org/record/174556/files/figure.png" pageId="1" pageNumber="54">Fig. 12</figureCitation>
). Legs very long, metatibiae without macrosetae, apically with 6 spines (
<figureCitation id="F0FC2A2E6D49FFD5FB980DF6FB77FB71" box="[1089,1224,1159,1185]" captionStart="FIGURES 23 26. 23" captionStartId="7.[264,382,1775,1799]" captionTargetBox="[277,1310,337,1751]" captionTargetId="figure@7.[277,1310,326,1751]" captionTargetPageId="7" captionText="FIGURES 23 26. 23, Noabennarella costaricensis spec. nov., hind leg; 24, same, apex of tibia of hind leg; 25, Noabennarella spec., inner female genitalia; 26, Noabennarella spec., habitus of female." httpUri="https://zenodo.org/record/174558/files/figure.png" pageId="1" pageNumber="54">Figs 23, 24</figureCitation>
). Tarsus with 6+5 apical spines, platellae not present.
</paragraph>
<paragraph id="687836AB6D49FFD5FEE10DA6FBBAFACC" blockId="1.[264,1324,599,1828]" pageId="1" pageNumber="54">
Segments four and five with large abdominal processes, bearing 3+2 sensory pits (
<figureCitation id="F0FC2A2E6D49FFD5FB2C0DA6FE89FACC" captionStart="FIGURES 1 2" captionStartId="2.[264,383,1816,1840]" captionTargetBox="[284,1309,326,1788]" captionTargetId="figure@2.[284,1310,326,1789]" captionTargetPageId="2" captionText="FIGURES 1 2. Habitus of Noabennarella costaricensis spec. nov., lateral and dorsal view, with abdominal appendages clearly visible." httpUri="https://zenodo.org/record/174553/files/figure.png" pageId="1" pageNumber="54">Figs 1, 2</figureCitation>
,
<figureCitation id="F0FC2A2E6D49FFD5FE9A0C73FECEFACC" box="[323,369,1282,1308]" captionStart="FIGURES 3 6. 3" captionStartId="3.[264,382,1499,1523]" captionTargetBox="[264,1323,326,1475]" captionTargetId="figure@3.[264,1323,326,1475]" captionTargetPageId="3" captionText="FIGURES 3 6. 3, Noabennarella sp., female abdomen, lateral view; 4, same, ventral aspect; 5, abdominal appendages of Noabennarella sp.; 6, abdominal appendages of Bebaiotes sp. (Achilixiidae)." httpUri="https://zenodo.org/record/174554/files/figure.png" pageId="1" pageNumber="54">3, 5</figureCitation>
). Two additional sensory pits on 6th abdominal segment.
</paragraph>
<paragraph id="687836AB6D49FFD5FEE10C5BFC2CFA44" blockId="1.[264,1324,599,1828]" pageId="1" pageNumber="54">
Male genital and anal segment more or less symmetrical (
<figureCitation id="F0FC2A2E6D49FFD5FC300C5BFB1EFA94" box="[1001,1185,1322,1348]" captionStart="FIGURES 16 22. 16" captionStartId="6.[264,382,1735,1759]" captionTargetBox="[308,1282,513,1714]" captionTargetId="figure@6.[302,1295,498,1729]" captionTargetPageId="6" captionText="FIGURES 16 22. 16, Noabennarella costaricensis spec. nov., aedeagus, left lateral view; 17, same, right lateral view; 18, genital style, inner maximum view; 19, male genital and anal segment (aedeagus and styli omitted), left lateral view; 20, ventromedian process of male genital segment, ventral view; 21, male anal segment, caudal aspect; 22, same, dorsal aspect. Scale bar: 0.25 mm for Figs 16 18, 0.5 mm for Figs 19 22." httpUri="https://zenodo.org/record/174557/files/figure.png" pageId="1" pageNumber="54">Figs 19, 21, 22</figureCitation>
). Aedeagus with shaft and flagellum well developed, shaft with set of movable spines, flagellum long and slender, in repose bent to the left side (
<figureCitation id="F0FC2A2E6D49FFD5FD240C0BFC3AFA44" box="[765,901,1402,1428]" captionStart="FIGURES 16 22. 16" captionStartId="6.[264,382,1735,1759]" captionTargetBox="[308,1282,513,1714]" captionTargetId="figure@6.[302,1295,498,1729]" captionTargetPageId="6" captionText="FIGURES 16 22. 16, Noabennarella costaricensis spec. nov., aedeagus, left lateral view; 17, same, right lateral view; 18, genital style, inner maximum view; 19, male genital and anal segment (aedeagus and styli omitted), left lateral view; 20, ventromedian process of male genital segment, ventral view; 21, male anal segment, caudal aspect; 22, same, dorsal aspect. Scale bar: 0.25 mm for Figs 16 18, 0.5 mm for Figs 19 22." httpUri="https://zenodo.org/record/174557/files/figure.png" pageId="1" pageNumber="54">Figs 16, 17</figureCitation>
).
</paragraph>
<paragraph id="687836AB6D49FFD5FEE10CD3FE56F9DC" blockId="1.[264,1324,599,1828]" pageId="1" pageNumber="54">
Apex of female abdomen truncate, without wax plate. Ovipositor evenly curved, at rest adjacenting to abdomen (
<figureCitation id="F0FC2A2E6D49FFD5FDBA0CBBFD71FA34" box="[611,718,1482,1508]" captionStart="FIGURES 3 6. 3" captionStartId="3.[264,382,1499,1523]" captionTargetBox="[264,1323,326,1475]" captionTargetId="figure@3.[264,1323,326,1475]" captionTargetPageId="3" captionText="FIGURES 3 6. 3, Noabennarella sp., female abdomen, lateral view; 4, same, ventral aspect; 5, abdominal appendages of Noabennarella sp.; 6, abdominal appendages of Bebaiotes sp. (Achilixiidae)." httpUri="https://zenodo.org/record/174554/files/figure.png" pageId="1" pageNumber="54">Figs 3, 4</figureCitation>
). Ductus ejaculatorius wound helix­like, with three windings (
<figureCitation id="F0FC2A2E6D49FFD5FE5A0C83FE64F9DC" box="[387,475,1522,1548]" captionStart="FIGURES 23 26. 23" captionStartId="7.[264,382,1775,1799]" captionTargetBox="[277,1310,337,1751]" captionTargetId="figure@7.[277,1310,326,1751]" captionTargetPageId="7" captionText="FIGURES 23 26. 23, Noabennarella costaricensis spec. nov., hind leg; 24, same, apex of tibia of hind leg; 25, Noabennarella spec., inner female genitalia; 26, Noabennarella spec., habitus of female." httpUri="https://zenodo.org/record/174558/files/figure.png" pageId="1" pageNumber="54">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="687836AB6D49FFD5FEE10F6BFD68F8F4" blockId="1.[264,1324,599,1828]" pageId="1" pageNumber="54">
Note: We studied one female
<taxonomicName id="AFC74D286D49FFD5FD1D0F6BFCCCF9E3" box="[708,883,1562,1587]" class="Insecta" family="Cixiidae" genus="Noabennarella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="5AB3EAB96D49FFD5FD1D0F6BFCCCF9E3" box="[708,883,1562,1587]" italics="true" pageId="1" pageNumber="54">Noabennarella</emphasis>
</taxonomicName>
specimen with a wing coloration distinctly different from that of
<taxonomicName id="AFC74D286D49FFD5FD5B0F33FC81F98B" box="[642,830,1602,1627]" class="Insecta" family="Cixiidae" genus="Noabennarella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="5AB3EAB96D49FFD5FD5B0F33FC81F98B" box="[642,830,1602,1627]" italics="true" pageId="1" pageNumber="54">N. costaricensis</emphasis>
</taxonomicName>
. This specimen (see
<figureCitation id="F0FC2A2E6D49FFD5FBE30F33FB13F98C" box="[1082,1196,1602,1628]" captionStart="FIGURES 3 6. 3" captionStartId="3.[264,382,1499,1523]" captionTargetBox="[264,1323,326,1475]" captionTargetId="figure@3.[264,1323,326,1475]" captionTargetPageId="3" captionText="FIGURES 3 6. 3, Noabennarella sp., female abdomen, lateral view; 4, same, ventral aspect; 5, abdominal appendages of Noabennarella sp.; 6, abdominal appendages of Bebaiotes sp. (Achilixiidae)." httpUri="https://zenodo.org/record/174554/files/figure.png" pageId="1" pageNumber="54">Figs. 3, 4</figureCitation>
,
<figureCitation id="F0FC2A2E6D49FFD5FB630F33FAB9F98C" box="[1210,1286,1602,1628]" captionStart="FIGURES 23 26. 23" captionStartId="7.[264,382,1775,1799]" captionTargetBox="[277,1310,337,1751]" captionTargetId="figure@7.[277,1310,326,1751]" captionTargetPageId="7" captionText="FIGURES 23 26. 23, Noabennarella costaricensis spec. nov., hind leg; 24, same, apex of tibia of hind leg; 25, Noabennarella spec., inner female genitalia; 26, Noabennarella spec., habitus of female." httpUri="https://zenodo.org/record/174558/files/figure.png" pageId="1" pageNumber="54">25, 26</figureCitation>
) is labelled “
<collectingCountry id="10D0763B6D49FFD5FEA40F1BFE06F954" box="[381,441,1642,1668]" name="Peru" pageId="1" pageNumber="54">Peru</collectingCountry>
: Monon Valley, Tingo Maria,
<date id="1C79106B6D49FFD5FCE50F1BFC79F954" box="[828,966,1642,1668]" pageId="1" pageNumber="54" value="1954-11-21">XI­21­1954</date>
// E.I.Schlinger &amp; E.S.Ross collectors //
<taxonomicName id="AFC74D286D49FFD5FE710FE3FD7DF97C" box="[424,706,1682,1708]" class="Insecta" family="Cixiidae" genus="Amazobenna" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="reticulata">Amazobenna reticulata</taxonomicName>
” and stored in the collection of the Californian Academy of Sciences. We do not know if different wing coloration patterns or sexual dimorphism exist within
<taxonomicName id="AFC74D286D49FFD5FDED0F93FD5CF92B" box="[564,739,1762,1787]" class="Insecta" family="Cixiidae" genus="Noabennarella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="5AB3EAB96D49FFD5FDED0F93FD5CF92B" box="[564,739,1762,1787]" italics="true" pageId="1" pageNumber="54">Noabennarella</emphasis>
</taxonomicName>
and if this female belongs to
<taxonomicName id="AFC74D286D49FFD5FB950F93FAB6F92B" box="[1100,1289,1762,1787]" class="Insecta" family="Cixiidae" genus="Noabennarella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="5AB3EAB96D49FFD5FB950F93FAB6F92B" box="[1100,1289,1762,1787]" italics="true" pageId="1" pageNumber="54">N. costaricensis</emphasis>
</taxonomicName>
or represents another undescribed species.
</paragraph>
<caption id="3CB866236D4AFFD6FED10E69FD1FF882" httpUri="https://zenodo.org/record/174553/files/figure.png" pageId="2" pageNumber="55" targetBox="[284,1309,326,1788]" targetPageId="2">
<paragraph id="687836AB6D4AFFD6FED10E69FD1FF882" blockId="2.[264,1322,1816,1874]" pageId="2" pageNumber="55">
<emphasis id="5AB3EAB96D4AFFD6FED10E69FE09F8E0" bold="true" box="[264,438,1816,1840]" pageId="2" pageNumber="55">FIGURES 12.</emphasis>
Habitus of
<taxonomicName id="AFC74D286D4AFFD6FDE30E68FCD2F8E0" box="[570,877,1817,1840]" class="Insecta" family="Cixiidae" genus="Noabennarella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="costaricensis" status="sp. nov.">
<emphasis id="5AB3EAB96D4AFFD6FDE30E68FCD2F8E0" box="[570,877,1817,1840]" italics="true" pageId="2" pageNumber="55">Noabennarella costaricensis</emphasis>
</taxonomicName>
<emphasis id="5AB3EAB96D4AFFD6FCAF0E69FC5AF8E0" bold="true" box="[886,997,1816,1840]" pageId="2" pageNumber="55">
<taxonomicNameLabel id="418057C26D4AFFD6FCAF0E69FC5AF8E0" box="[886,997,1816,1840]" pageId="2" pageNumber="55" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, lateral and dorsal view, with abdominal appendages clearly visible.
</paragraph>
</caption>
<caption id="3CB866236D4BFFD7FED10CAAFE1AF9E8" httpUri="https://zenodo.org/record/174554/files/figure.png" pageId="3" pageNumber="56" targetBox="[264,1323,326,1475]" targetPageId="3">
<paragraph id="687836AB6D4BFFD7FED10CAAFE1AF9E8" blockId="3.[264,1323,1499,1592]" pageId="3" pageNumber="56">
<emphasis id="5AB3EAB96D4BFFD7FED10CAAFE09FA23" bold="true" box="[264,438,1499,1523]" pageId="3" pageNumber="56">FIGURES 36.</emphasis>
3,
<taxonomicName id="AFC74D286D4BFFD7FE070CADFDC2FA23" box="[478,637,1500,1523]" class="Insecta" family="Cixiidae" genus="Noabennarella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="5AB3EAB96D4BFFD7FE070CADFDC2FA23" box="[478,637,1500,1523]" italics="true" pageId="3" pageNumber="56">Noabennarella</emphasis>
</taxonomicName>
sp., female abdomen, lateral view; 4, same, ventral aspect; 5, abdominal appendages of
<taxonomicName id="AFC74D286D4BFFD7FD9D0C8EFD5CF9C6" box="[580,739,1535,1558]" class="Insecta" family="Cixiidae" genus="Noabennarella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="5AB3EAB96D4BFFD7FD9D0C8EFD5CF9C6" box="[580,739,1535,1558]" italics="true" pageId="3" pageNumber="56">Noabennarella</emphasis>
</taxonomicName>
sp.; 6, abdominal appendages of
<taxonomicName id="AFC74D286D4BFFD7FB490C8EFB46F9C6" box="[1168,1273,1535,1558]" class="Insecta" family="Achilixiidae" genus="Bebaiotes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="5AB3EAB96D4BFFD7FB490C8EFB46F9C6" box="[1168,1273,1535,1558]" italics="true" pageId="3" pageNumber="56">Bebaiotes</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="AFC74D286D4BFFD7FED60F50FE26F9E8" box="[271,409,1569,1592]" class="Insecta" family="Achilixiidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="family">Achilixiidae</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="687836AB6D4BFFD7FEE10F16FD89F951" blockId="3.[264,1323,1639,1865]" box="[312,566,1639,1665]" pageId="3" pageNumber="56">Differential diagnosis</paragraph>
<paragraph id="687836AB6D4BFFD7FEE10FFEFA9DF899" blockId="3.[264,1323,1639,1865]" pageId="3" pageNumber="56">
<taxonomicName id="AFC74D286D4BFFD7FEE10FFEFE59F978" box="[312,486,1679,1704]" class="Insecta" family="Cixiidae" genus="Noabennarella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="5AB3EAB96D4BFFD7FEE10FFEFE59F978" box="[312,486,1679,1704]" italics="true" pageId="3" pageNumber="56">Noabennarella</emphasis>
</taxonomicName>
can be easily distinguished from other Bennarellini by the shape of the head (especially frons) and its keels and by the venation of the fore wings. In
<taxonomicName id="AFC74D286D4BFFD7FB4A0FC6FA94F900" box="[1171,1323,1719,1744]" class="Insecta" family="Cixiidae" genus="Amazobenna" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="5AB3EAB96D4BFFD7FB4A0FC6FA94F900" box="[1171,1323,1719,1744]" italics="true" pageId="3" pageNumber="56">Amazobenna</emphasis>
</taxonomicName>
and
<taxonomicName id="AFC74D286D4BFFD7FEE50FAEFE02F928" box="[316,445,1759,1784]" class="Insecta" family="Cixiidae" genus="Bennarella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="5AB3EAB96D4BFFD7FEE50FAEFE02F928" box="[316,445,1759,1784]" italics="true" pageId="3" pageNumber="56">Bennarella</emphasis>
</taxonomicName>
the frons is much broader, and the lateral keels are directed laterally (see
<figureCitation id="F0FC2A2E6D4BFFD7FED10E76FE08F8F1" box="[264,439,1799,1825]" captionStart="FIGURES 7 10. 7" captionStartId="4.[264,382,1810,1834]" captionTargetBox="[321,1274,551,1795]" captionTargetId="figure@4.[321,1275,551,1795]" captionTargetPageId="4" captionText="FIGURES 7 10. 7, Amazobenna reticulata Penny, head of holotype, lateral aspect (photo: A. Henriques); 8, same, frontal aspect (photo: A. Henriques); 9, Noabennarella costaricensis spec. nov., head, lateral aspect; 10, same, frontal aspect." httpUri="https://zenodo.org/record/174555/files/figure.png" pageId="3" pageNumber="56">Figs 7, 8, 9, 10</figureCitation>
,
<figureCitation id="F0FC2A2E6D4BFFD7FE1A0E76FD85F8F1" box="[451,570,1799,1825]" captionStart="FIGURES 11 15. 11" captionStartId="5.[264,382,1543,1567]" captionTargetBox="[316,1271,319,1524]" captionTargetId="figure@5.[316,1279,312,1540]" captionTargetPageId="5" captionText="FIGURES 11 15. 11, Noabennarella costaricensis spec. nov., hind wing; 12, same, fore wing; 13, same, head in frontal aspect; 14, same, head and thorax in dorsal aspect; 15, Bennarella bicoloripennis Muir, head and thorax in dorsal aspect. Scale bar: 1 mm." httpUri="https://zenodo.org/record/174556/files/figure.png" pageId="3" pageNumber="56">13, 14, 15</figureCitation>
). CuA and MP veins are strongly arched near the apex (
<figureCitation id="F0FC2A2E6D4BFFD7FB1F0E76FAA3F8F1" box="[1222,1308,1799,1825]" captionStart="FIGURES 11 15. 11" captionStartId="5.[264,382,1543,1567]" captionTargetBox="[316,1271,319,1524]" captionTargetId="figure@5.[316,1279,312,1540]" captionTargetPageId="5" captionText="FIGURES 11 15. 11, Noabennarella costaricensis spec. nov., hind wing; 12, same, fore wing; 13, same, head in frontal aspect; 14, same, head and thorax in dorsal aspect; 15, Bennarella bicoloripennis Muir, head and thorax in dorsal aspect. Scale bar: 1 mm." httpUri="https://zenodo.org/record/174556/files/figure.png" pageId="3" pageNumber="56">Fig. 12</figureCitation>
), whereas they are straight in
<taxonomicName id="AFC74D286D4BFFD7FD8B0E5EFD6CF898" box="[594,723,1839,1864]" class="Insecta" family="Cixiidae" genus="Bennarella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="5AB3EAB96D4BFFD7FD8B0E5EFD6CF898" box="[594,723,1839,1864]" italics="true" pageId="3" pageNumber="56">Bennarella</emphasis>
</taxonomicName>
, and in
<taxonomicName id="AFC74D286D4BFFD7FCE80E5EFC77F898" box="[817,968,1839,1864]" class="Insecta" family="Cixiidae" genus="Amazobenna" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="5AB3EAB96D4BFFD7FCE80E5EFC77F898" box="[817,968,1839,1864]" italics="true" pageId="3" pageNumber="56">Amazobenna</emphasis>
</taxonomicName>
only CuA is strongly arched.
</paragraph>
</subSubSection>
</treatment>
</document>