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<document ID-DOI="10.5733/afin.056.0109" ID-ISSN="2305-2562" ID-Zenodo-Dep="7661863" approvalRequired="20" approvalRequired_for_textStreams="18" approvalRequired_for_treatments="2" checkinTime="1676998671595" checkinUser="felipe" docAuthor="Vannozzi, Angelo, Pizzini, Mauro &amp; Raines, Bret" docDate="2015" docId="D6171E5CFFEFFFBC27D5BFCE832A320B" docLanguage="en" docName="AfrInvertebr.56.1.99-136.pdf" docOrigin="African Invertebrates 56 (1)" docSource="http://www.bioone.org/doi/10.5733/afin.056.0109" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.2:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="2" docTitle="Caecum knysnaense Vannozzi &amp; Pizzini &amp; Raines 2015, sp. n." docType="treatment" docVersion="2" lastPageNumber="108" masterDocId="2A2E6624FFE7FFB5260DBC7A81623678" masterDocTitle="Revision of South African Caecidae (Mollusca: Gastropoda)" masterLastPageNumber="99" masterPageNumber="99" pageNumber="107" updateTime="1676999323622" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Revision of South African Caecidae (Mollusca: Gastropoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vannozzi, Angelo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pizzini, Mauro</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Raines, Bret</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
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<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-05-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>56</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>99</mods:start>
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<mods:url>http://www.bioone.org/doi/10.5733/afin.056.0109</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.056.0109</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7661863</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:D6171E5CFFEFFFBC27D5BFCE832A320B" httpUri="http://treatment.plazi.org/id/D6171E5CFFEFFFBC27D5BFCE832A320B" lastPageId="9" lastPageNumber="108" pageId="8" pageNumber="107">
<subSubSection box="[472,775,948,975]" pageId="8" pageNumber="107" type="nomenclature">
<paragraph blockId="8.[144,1104,948,1676]" box="[472,775,948,975]" pageId="8" pageNumber="107">
<heading box="[472,775,948,975]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="107" reason="2">
<taxonomicName ID-CoL="P8TP" authority="Vannozzi &amp; Pizzini &amp; Raines, 2015" authorityName="Vannozzi &amp; Pizzini &amp; Raines" authorityYear="2015" box="[472,708,948,975]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="107" phylum="Mollusca" rank="species" species="knysnaense" status="sp. n.">
<emphasis bold="true" box="[472,708,948,975]" italics="true" pageId="8" pageNumber="107">Caecum knysnaense</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[715,775,949,975]" pageId="8" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="108" pageId="8" pageNumber="107" type="description">
<paragraph blockId="8.[144,1104,948,1676]" box="[439,809,988,1015]" pageId="8" pageNumber="107">
<figureCitation box="[439,562,988,1015]" captionStart="Fig" captionStartId="9.[181,214,634,656]" captionTargetBox="[203,1098,220,601]" captionTargetId="figure-331@9.[189,1134,201,621]" captionTargetPageId="9" captionText="Fig.5.Caecum knysnaense sp. n., SouthAfrica:(AC) holotype, Knysna,Gericke Point,Sedgefield (NMSA S7625/T3362) — (A) right side, (B) ventral side, (C) microsculpture; (D, E) paratype, operculum, Walkers Point, W of Knysna (NMSA W9672/T3363) — (D) inner side, (E) outer side. Scale bar = 1 mm (A, B); 200 µm (D, E); 50 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.7661873" httpUri="https://zenodo.org/record/7661873/files/figure.png" pageId="8" pageNumber="107">Figs 5AE</figureCitation>
,
<figureCitation box="[574,590,989,1015]" captionStart="Fig" captionStartId="11.[181,214,758,780]" captionTargetBox="[221,1116,207,734]" captionTargetId="figure-383@11.[189,1134,203,771]" captionTargetPageId="11" captionText="Fig. 7. Distribution map of Caecum knysnaense sp. n. („), C. austrafricanum sp. n. (S) and C. intortum sp. n. (T). White symbol indicates the type locality." figureDoi="http://doi.org/10.5281/zenodo.7661877" httpUri="https://zenodo.org/record/7661877/files/figure.png" pageId="8" pageNumber="107">7</figureCitation>
,
<figureCitation box="[602,642,988,1014]" captionStart="Fig" captionStartId="29.[181,215,1162,1184]" captionTargetBox="[189,1134,204,1149]" captionTargetId="figure-53@29.[189,1134,204,1149]" captionTargetPageId="29" captionText="Fig. 22. Detail of the septum of South and western African Caecum species (right side): (AC) C. subquadratum Carpenter, 1859 — (A) Walkers Point, W of Knysna (NMSA S3729), (B) Walkers Point, W of Knysna (JPM), (C) juv, Langebaan Lagoon, Saldanha Bay (JPM); (D, E) C. morgan sp. n., Cape Morgan Lighthouse — (D) holotype (NMSA B7132/T3377), (E) paratype (NMSA W9676/ T3378); (FH) C. austrafricanum sp. n., Algoa Bay — (F) holotype (NMSA W9655/T3349), (G) paratype (JPM),(H) paratype (juv) (JPM); (I) C. knysnaense sp.n., holotype, Knysna, Gericke Point, Sedgefield (NMSA S7625/T3362); (J) C. intortum sp. n., holotype, Walkers Point, W of Knysna (NMSA W9668/T3358); (K) C. inhacaense Albano &amp; Pizzini, 2011, Aliwal Shoal, off Umkomaas area (JPM); (L, M) C. incisum sp. n. — (L) holotype, Walkers Point, W of Knysna (NMSA S3728/ T3355), (M) paratype, Knysna,Gericke Point, Sedgefield (NMSA W9661/T3357);(N, O) C.maraisi sp.n.—(N) holotype, Knysna, GerickePoint,Sedgefield (NMSA W9765/T3376),(O) paratype (juv), Fish R. mouth (between Port Alfred and East London) (JPM 2117); (P, Q) C. leilae sp. n. — (P) holotype, Mzamba (NMSA D3032/T3374), (Q) paratype, Umtamvuna R. mouth (NMSA V3999/ T3884); (R) C. cf. glabrum (Montagu, 1803), Knysna, Gericke Point, Sedgefield (NMSA W9658); (S, T) C. glabrum (Montagu, 1803) — (S) Porto Mindelo, Ilha de São Vicente, Cape Verde (MP), (T) St. Malo, France (MP). Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7661907" httpUri="https://zenodo.org/record/7661907/files/figure.png" pageId="8" pageNumber="107">22I</figureCitation>
,
<figureCitation box="[653,693,988,1014]" captionStart="Fig" captionStartId="30.[143,176,1164,1186]" captionTargetBox="[151,1096,204,1149]" captionTargetId="figure-63@30.[151,1096,204,1149]" captionTargetPageId="30" captionText="Fig. 23. Detail of the septum of South and western African Caecum species (ventral side): (AC) C. subquadratum Carpenter, 1859 — (A) Walkers Point, W of Knysna (NMSA S3729), (B) Walkers Point, W of Knysna (JPM), (C) juv, Langebaan Lagoon, Saldanha Bay (JPM); (D, E) C. morgan sp. n., Cape Morgan Lighthouse — (D) holotype (NMSA B7132/T3377), (E) paratype (NMSA W9676/ T3378); (FH) C. austrafricanum sp. n., Algoa Bay — (F) holotype (NMSA W9655/T3349), (G) paratype (JPM); (H)paratype (juv) (JPM); (I) C. knysnaense sp.n., holotype, Knysna, Gericke Point, Sedgefield (NMSA S7625/T3362); (J) C. intortum sp. n., holotype, Walkers Point, W of Knysna (NMSA W9668/T3358); (K) C. inhacaense Albano &amp; Pizzini, 2011, Aliwal Shoal, off Umkomaas area (JPM); (L, M) C. incisum sp. n. — (L) holotype, Walkers Point, W of Knysna (NMSA S3728/ T3355), (M) paratype, Knysna, Gericke Point, Sedgefield (NMSA W9661/T3357);(N, O) C. maraisi sp.n.— (N) holotype, Knysna, Gericke Point,Sedgefield (NMSA W9765/T3376), (O) paratype (juv), Fish R. mouth (between Port Alfred and East London) (JPM 2117); (P, Q) C. leilae sp. n. — (P) holotype, Mzamba (NMSA D3032/T3374), (Q) paratype, Umtamvuna R. mouth (NMSA V3999/ T3884); (R) C. cf. glabrum (Montagu, 1803), Knysna, Gericke Point, Sedgefield (NMSA W9658); (S, T) C. glabrum (Montagu, 1803) — (S) Porto Mindelo, Ilha de São Vicente, Cape Verde (MP), (T) St. Malo, France (MP). Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7661909" httpUri="https://zenodo.org/record/7661909/files/figure.png" pageId="8" pageNumber="107">23I</figureCitation>
,
<figureCitation box="[705,754,988,1014]" captionStart="Fig" captionStartId="32.[144,177,1039,1061]" captionTargetBox="[151,1096,204,1021]" captionTargetId="figure-159@32.[151,1096,204,1021]" captionTargetPageId="32" captionText="Fig. 24. Ventral side of South and western African Caecum species: (A) C. austrafricanum sp. n., paratype, Walkers Point, W of Knysna (NMSA W9656/T3351); (B) C. knysnaense sp. n., paratype, Knysna, Gericke Point,Sedgefield (NMSA W9673/T3364);(C, D) C. intortum sp.n.— (C)paratype,Walkers Point, W of Knysna (NMSA W9669/T3360), (D) the same specimen slightly rotated, detail of the macula; (E, F) C. leilae sp. n. — (E) paratype, Mzamba, near Natal border (JPM 2119), (F) the same specimen, detail of the macula; (G) C. inhacaense Albano &amp; Pizzini, 2011, Mzamba, near Natal border (JPM 2119); (H) C. macrum van der Linden &amp; Moolenbeek, 2000, Charbonnier, Senegal (MP); (I) C. crassum de Folin, 1870, Banc dArguin, Mauritania (MP). Scale bar = 1 mm (AC, E, GI); 300 µm (D, F)." figureDoi="http://doi.org/10.5281/zenodo.7661911" httpUri="https://zenodo.org/record/7661911/files/figure.png" pageId="8" pageNumber="107">24B</figureCitation>
,
<figureCitation box="[765,809,988,1015]" captionStart="Fig" captionStartId="33.[181,214,1342,1364]" captionTargetBox="[226,1097,204,1321]" captionTargetId="figure-13@33.[226,1097,204,1321]" captionTargetPageId="33" captionText="Fig. 25. South and western African Caecum and Parastrophia species: (A) P. avaricosa sp. n., holotype, Mzamba (NMSA E1665/T3354); (B, C) P.ornata sp. n., Mzamba,near Natal border — (B) holotype (NMSA W9677/T3379), (C) paratype (JPM 2119); (D) C. sepimentum de Folin, 1868, Kosi Bay (NMSA S2010); (E) C. cf. oahuense Pilsbry, 1921, Aliwal Shoal, off Umkomaas area (JPM); (F) Caecum cf. gulosum Hedley, 1899, Kosi Bay (NMSA S2869); (G, H) C. succineum de Folin, 1880 — (G) between Bhanga Nek andKosi Bay (NMSAS1586), (H)Aliwal Shoal, off Scottburgh (NMSA W9684); (I) C. cf. glabrum (Montagu, 1803), Knysna, Gericke Point, Sedgefield (NMSA W9658); (J) C. glabrum (Montagu, 1803), Rabat, Morocco (MP); (K) C.lindae sp. n., holotype, Cape Morgan Lighthouse (NMSA W9682/T3924); (L, M) C. cf. musorstomi Pizzini, Raines &amp; Vannozzi, 2013, Landers reef (JPM); (N, O) C.austrafricanum sp.n., paratypes,Walkers Point,W of Knysna (NMSA W9656/T3351);(P) C.knysnaense sp.n., paratype,Knysna,Gericke Point,Sedgefield (NMSAW9673/ T3364); (Q) C. intortum sp. n., paratype, Walkers Point, W of Knysna (NMSA W9669/T3360); (R) C. macrum van der Linden &amp; Moolenbeek, 2000, Charbonnier, Senegal (MP); (S, T) C. crassum de Folin, 1870, Gabon (MP): (S) adult; (T) juvenile. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7661913" httpUri="https://zenodo.org/record/7661913/files/figure.png" pageId="8" pageNumber="107">25P</figureCitation>
</paragraph>
<paragraph blockId="8.[144,1104,948,1676]" box="[144,628,1028,1054]" pageId="8" pageNumber="107">
Etymology: Derived from the
<typeStatus box="[483,531,1028,1054]" pageId="8" pageNumber="107">type</typeStatus>
locality.
</paragraph>
<paragraph blockId="8.[144,1104,948,1676]" pageId="8" pageNumber="107">Description: Tube subcylindrical, with a slight dextral torsion, evenly arched. Tube smooth, glossy, colourless, semitransparent. Aperture with a barely visible swelling, strongly oblique toward the ventral side, preceded by a white band. Surface with only fine oblique growth lines. Septum smooth, dome­shaped, protruding, with an ear­like mucro directed halfway between the dorsal and the right side. Macula large, with a frosted appearance, formed by two oblong bands fused in the middle. Juveniles rather similar to the adults, showing a narrower and more arched shape of the tube. Periostracum light brown. Operculum flat with a raised internal nucleus and internally thickened edge. Outer side showing a spiral of about three whorls. Larval stage unknown.</paragraph>
<paragraph blockId="8.[144,1104,948,1676]" box="[144,382,1362,1388]" pageId="8" pageNumber="107">
Length:
<quantity box="[238,377,1362,1388]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.9" metricValueMin="1.2" pageId="8" pageNumber="107" unit="mm" value="1.5499999999999998" valueMax="1.9" valueMin="1.2">1.21.9 mm</quantity>
.
</paragraph>
<paragraph blockId="8.[144,1104,948,1676]" box="[144,1086,1402,1428]" pageId="8" pageNumber="107">
Type locality:
<collectingCountry box="[305,511,1402,1428]" name="South Africa" pageId="8" pageNumber="107">SOUTH AFRICA</collectingCountry>
:
<emphasis box="[521,676,1402,1428]" italics="true" pageId="8" pageNumber="107">
<collectingRegion box="[521,676,1402,1428]" country="South Africa" name="Western Cape" pageId="8" pageNumber="107">Western Cape</collectingRegion>
</emphasis>
: Knysna, Gericke Point, Sedgefield.
</paragraph>
<paragraph blockId="8.[144,1104,948,1676]" pageId="8" pageNumber="107">
<materialsCitation collectionCode="NMSA" country="South Africa" county="Knysna" location="Sedgefield" municipality="Gericke Point" pageId="8" pageNumber="107" specimenCode="S7625" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<typeStatus box="[144,228,1438,1460]" pageId="8" pageNumber="107">Holotype</typeStatus>
(
<figureCitation box="[241,343,1438,1460]" captionStart="Fig" captionStartId="9.[181,214,634,656]" captionTargetBox="[203,1098,220,601]" captionTargetId="figure-331@9.[189,1134,201,621]" captionTargetPageId="9" captionText="Fig.5.Caecum knysnaense sp. n., SouthAfrica:(AC) holotype, Knysna,Gericke Point,Sedgefield (NMSA S7625/T3362) — (A) right side, (B) ventral side, (C) microsculpture; (D, E) paratype, operculum, Walkers Point, W of Knysna (NMSA W9672/T3363) — (D) inner side, (E) outer side. Scale bar = 1 mm (A, B); 200 µm (D, E); 50 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.7661873" httpUri="https://zenodo.org/record/7661873/files/figure.png" pageId="8" pageNumber="107">Figs 5AC</figureCitation>
,
<figureCitation box="[353,386,1438,1459]" captionStart="Fig" captionStartId="29.[181,215,1162,1184]" captionTargetBox="[189,1134,204,1149]" captionTargetId="figure-53@29.[189,1134,204,1149]" captionTargetPageId="29" captionText="Fig. 22. Detail of the septum of South and western African Caecum species (right side): (AC) C. subquadratum Carpenter, 1859 — (A) Walkers Point, W of Knysna (NMSA S3729), (B) Walkers Point, W of Knysna (JPM), (C) juv, Langebaan Lagoon, Saldanha Bay (JPM); (D, E) C. morgan sp. n., Cape Morgan Lighthouse — (D) holotype (NMSA B7132/T3377), (E) paratype (NMSA W9676/ T3378); (FH) C. austrafricanum sp. n., Algoa Bay — (F) holotype (NMSA W9655/T3349), (G) paratype (JPM),(H) paratype (juv) (JPM); (I) C. knysnaense sp.n., holotype, Knysna, Gericke Point, Sedgefield (NMSA S7625/T3362); (J) C. intortum sp. n., holotype, Walkers Point, W of Knysna (NMSA W9668/T3358); (K) C. inhacaense Albano &amp; Pizzini, 2011, Aliwal Shoal, off Umkomaas area (JPM); (L, M) C. incisum sp. n. — (L) holotype, Walkers Point, W of Knysna (NMSA S3728/ T3355), (M) paratype, Knysna,Gericke Point, Sedgefield (NMSA W9661/T3357);(N, O) C.maraisi sp.n.—(N) holotype, Knysna, GerickePoint,Sedgefield (NMSA W9765/T3376),(O) paratype (juv), Fish R. mouth (between Port Alfred and East London) (JPM 2117); (P, Q) C. leilae sp. n. — (P) holotype, Mzamba (NMSA D3032/T3374), (Q) paratype, Umtamvuna R. mouth (NMSA V3999/ T3884); (R) C. cf. glabrum (Montagu, 1803), Knysna, Gericke Point, Sedgefield (NMSA W9658); (S, T) C. glabrum (Montagu, 1803) — (S) Porto Mindelo, Ilha de São Vicente, Cape Verde (MP), (T) St. Malo, France (MP). Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7661907" httpUri="https://zenodo.org/record/7661907/files/figure.png" pageId="8" pageNumber="107">22I</figureCitation>
,
<figureCitation box="[395,429,1438,1460]" captionStart="Fig" captionStartId="30.[143,176,1164,1186]" captionTargetBox="[151,1096,204,1149]" captionTargetId="figure-63@30.[151,1096,204,1149]" captionTargetPageId="30" captionText="Fig. 23. Detail of the septum of South and western African Caecum species (ventral side): (AC) C. subquadratum Carpenter, 1859 — (A) Walkers Point, W of Knysna (NMSA S3729), (B) Walkers Point, W of Knysna (JPM), (C) juv, Langebaan Lagoon, Saldanha Bay (JPM); (D, E) C. morgan sp. n., Cape Morgan Lighthouse — (D) holotype (NMSA B7132/T3377), (E) paratype (NMSA W9676/ T3378); (FH) C. austrafricanum sp. n., Algoa Bay — (F) holotype (NMSA W9655/T3349), (G) paratype (JPM); (H)paratype (juv) (JPM); (I) C. knysnaense sp.n., holotype, Knysna, Gericke Point, Sedgefield (NMSA S7625/T3362); (J) C. intortum sp. n., holotype, Walkers Point, W of Knysna (NMSA W9668/T3358); (K) C. inhacaense Albano &amp; Pizzini, 2011, Aliwal Shoal, off Umkomaas area (JPM); (L, M) C. incisum sp. n. — (L) holotype, Walkers Point, W of Knysna (NMSA S3728/ T3355), (M) paratype, Knysna, Gericke Point, Sedgefield (NMSA W9661/T3357);(N, O) C. maraisi sp.n.— (N) holotype, Knysna, Gericke Point,Sedgefield (NMSA W9765/T3376), (O) paratype (juv), Fish R. mouth (between Port Alfred and East London) (JPM 2117); (P, Q) C. leilae sp. n. — (P) holotype, Mzamba (NMSA D3032/T3374), (Q) paratype, Umtamvuna R. mouth (NMSA V3999/ T3884); (R) C. cf. glabrum (Montagu, 1803), Knysna, Gericke Point, Sedgefield (NMSA W9658); (S, T) C. glabrum (Montagu, 1803) — (S) Porto Mindelo, Ilha de São Vicente, Cape Verde (MP), (T) St. Malo, France (MP). Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7661909" httpUri="https://zenodo.org/record/7661909/files/figure.png" pageId="8" pageNumber="107">23I</figureCitation>
):
<collectingCountry box="[446,613,1438,1459]" name="South Africa" pageId="8" pageNumber="107">SOUTH AFRICA</collectingCountry>
:
<emphasis box="[622,748,1438,1459]" italics="true" pageId="8" pageNumber="107">
<collectingRegion box="[622,748,1438,1459]" country="South Africa" name="Western Cape" pageId="8" pageNumber="107">Western Cape</collectingRegion>
</emphasis>
: lv,
<collectingCounty box="[789,861,1438,1459]" pageId="8" pageNumber="107">Knysna</collectingCounty>
,
<collectingMunicipality box="[870,998,1438,1460]" pageId="8" pageNumber="107">Gericke Point</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:D6171E5CFFEFFFBC27D5BFCE832A320B:5B61F991FFEFFFBD25FDB9E4853233CC" box="[1008,1104,1438,1460]" country="South Africa" county="Knysna" municipality="Gericke Point" name="Sedgefield" pageId="8" pageNumber="107" stateProvince="Western Cape">Sedgefield</location>
(
<collectionCode box="[150,217,1462,1483]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="8" pageNumber="107" type="Museum">NMSA</collectionCode>
<specimenCode box="[220,277,1462,1484]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="8" pageNumber="107" type="Museum">S7625</specimenCode>
/T3362), length
<quantity box="[423,494,1462,1484]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="107" unit="mm" value="1.6">1.6 mm</quantity>
, min. diam. (posterior end)
<quantity box="[741,823,1462,1483]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="107" unit="mm" value="0.31">0.31 mm</quantity>
, diam. in the middle of the tube
<quantity box="[144,227,1486,1508]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="8" pageNumber="107" unit="mm" value="0.36">0.36 mm</quantity>
, max. diam. (apertural end)
<quantity box="[482,566,1486,1507]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="8" pageNumber="107" unit="mm" value="0.37">0.37 mm</quantity>
.
</materialsCitation>
</paragraph>
<paragraph blockId="8.[144,1104,948,1676]" pageId="8" pageNumber="107">
<materialsCitation box="[144,1101,1518,1540]" collectionCode="NMSA" country="South Africa" county="Knysna" location="Sedgefield" municipality="Gericke Point" pageId="8" pageNumber="107" specimenCode="W9673" specimenCount="1" specimenCount-larva="1" stateProvince="Western Cape" typeStatus="paratype">
<typeStatus box="[144,236,1518,1539]" pageId="8" pageNumber="107">Paratypes</typeStatus>
:
<collectingCountry box="[243,413,1518,1539]" name="South Africa" pageId="8" pageNumber="107">SOUTH AFRICA</collectingCountry>
:
<emphasis box="[424,549,1518,1540]" italics="true" pageId="8" pageNumber="107">
<collectingRegion box="[424,549,1518,1540]" country="South Africa" name="Western Cape" pageId="8" pageNumber="107">Western Cape</collectingRegion>
</emphasis>
:
<specimenCount box="[560,593,1518,1540]" pageId="8" pageNumber="107" type="larva">1 lv</specimenCount>
and 6 sh, same data as holotype (
<collectionCode box="[888,954,1518,1539]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="8" pageNumber="107" type="Museum">NMSA</collectionCode>
<specimenCode box="[958,1023,1518,1540]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="8" pageNumber="107" type="Museum">W9673</specimenCode>
/T3364)
</materialsCitation>
;
<materialsCitation box="[144,1053,1542,1564]" collectingDate="1991-03" collectionCode="NMSA" collectorName="Walker's Point &amp; J. P. Marais" country="South Africa" location="Knysna" pageId="8" pageNumber="107" specimenCode="W9672" specimenCount="2" specimenCount-larva="2" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount box="[144,179,1542,1564]" pageId="8" pageNumber="107" type="larva">2 lv</specimenCount>
and 6 sh (1 juv),
<collectorName box="[337,474,1542,1564]" pageId="8" pageNumber="107">Walkers Point</collectorName>
, W of
<location LSID="urn:lsid:plazi:treatment:D6171E5CFFEFFFBC27D5BFCE832A320B:5B61F991FFEFFFBD241BBA7C833C3063" box="[534,606,1542,1563]" country="South Africa" name="Knysna" pageId="8" pageNumber="107" stateProvince="Western Cape">Knysna</location>
, leg.
<collectorName box="[653,752,1542,1564]" pageId="8" pageNumber="107">J.P. Marais</collectorName>
<date box="[758,827,1542,1564]" pageId="8" pageNumber="107" value="1991-03">
<collectingDate box="[758,827,1542,1564]" pageId="8" pageNumber="107" value="1991-03">iii.1991</collectingDate>
</date>
(
<collectionCode box="[839,905,1542,1563]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="8" pageNumber="107" type="Museum">NMSA</collectionCode>
<specimenCode box="[910,977,1542,1564]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="8" pageNumber="107" type="Museum">W9672</specimenCode>
/T3363)
</materialsCitation>
.
</paragraph>
<paragraph blockId="8.[144,1104,948,1676]" box="[144,1085,1574,1596]" pageId="8" pageNumber="107">
<materialsCitation box="[144,1082,1574,1596]" collectingDate="1991-03" collectionCode="JPM" country="South Africa" location="Walker's Bay" municipality="Doubtful" pageId="8" pageNumber="107" specimenCount="1" stateProvince="Western Cape">
<collectingMunicipality box="[144,229,1574,1596]" pageId="8" pageNumber="107">Doubtful</collectingMunicipality>
:
<collectingCountry box="[237,404,1574,1595]" name="South Africa" pageId="8" pageNumber="107">SOUTH AFRICA</collectingCountry>
:
<emphasis box="[412,538,1574,1596]" italics="true" pageId="8" pageNumber="107">
<collectingRegion box="[412,538,1574,1596]" country="South Africa" name="Western Cape" pageId="8" pageNumber="107">Western Cape</collectingRegion>
</emphasis>
: 1 sh (broken),
<location LSID="urn:lsid:plazi:treatment:D6171E5CFFEFFFBC27D5BFCE832A320B:5B61F991FFEFFFBD24A7BA5C82473043" box="[682,805,1574,1596]" country="South Africa" municipality="Doubtful" name="Walker's Bay" pageId="8" pageNumber="107" stateProvince="Western Cape">Walkers Bay</location>
,
<locationDeviation box="[814,938,1574,1596]" location="Knysna" pageId="8" pageNumber="107">W of Knysna</locationDeviation>
,
<date box="[946,1016,1574,1596]" pageId="8" pageNumber="107" value="1991-03">
<collectingDate box="[946,1016,1574,1596]" pageId="8" pageNumber="107" value="1991-03">iii.1991</collectingDate>
</date>
(
<collectionCode box="[1028,1074,1574,1596]" pageId="8" pageNumber="107">JPM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="8.[144,1104,948,1676]" pageId="8" pageNumber="107">
<materialsCitation collectionCode="MP" collectorName="J. Pelorce" country="Senegal" elevation="34" location="Comparative" municipality="Charbonnier" pageId="8" pageNumber="107" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:D6171E5CFFEFFFBC27D5BFCE832A320B:5B61F991FFEFFFBD269DBA3C80643024" box="[144,262,1606,1628]" country="Senegal" municipality="Charbonnier" name="Comparative" pageId="8" pageNumber="107">Comparative</location>
material examined:
<taxonomicName authority="van der Linden &amp; Moolenbeek, 2000" authorityName="van der Linden &amp; Moolenbeek" authorityYear="2000" box="[458,926,1606,1628]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="107" phylum="Mollusca" rank="species" species="macrum">
<emphasis box="[458,562,1606,1628]" italics="true" pageId="8" pageNumber="107">C. macrum</emphasis>
<bibRefCitation author="VAN DER LINDEN, J. &amp; MOOLENBEEK, R. G." box="[571,926,1606,1628]" pageId="8" pageNumber="107" pagination="81 - 92" refId="ref24720" refString="VAN DER LINDEN, J. &amp; MOOLENBEEK, R. G. 2000. Caecidae (Mollusca: Gastropoda) from Mauritania. Vita Marina 47 (3): 81 - 92." type="journal article" year="2000">van der Linden &amp; Moolenbeek, 2000</bibRefCitation>
</taxonomicName>
,
<collectingCountry box="[938,1046,1606,1627]" name="Senegal" pageId="8" pageNumber="107">SENEGAL</collectingCountry>
: 2 sh,
<collectingMunicipality box="[144,257,1630,1652]" pageId="8" pageNumber="107">Charbonnier</collectingMunicipality>
, petite épave, -
<quantity box="[386,434,1630,1652]" metricMagnitude="1" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="107" unit="m" value="34.0">
<elevation box="[386,434,1630,1652]" metricMagnitude="1" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="107" unit="m" value="34.0">34 m</elevation>
</quantity>
, fond sable coquiller légerement vaseux, leg.
<collectorName box="[832,917,1630,1652]" pageId="8" pageNumber="107">J. Pelorce</collectorName>
(
<collectionCode box="[928,965,1630,1652]" country="0" name="Mohonk Preserve, Inc." pageId="8" pageNumber="107">MP</collectionCode>
); 1 sh, Somone, -
<quantity box="[151,188,1655,1676]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="107" unit="m" value="7.0">
<elevation box="[151,188,1655,1676]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="107" unit="m" value="7.0">7 m</elevation>
</quantity>
, sable sur rochers plats avec curvettes sable, leg.
<collectorName box="[635,723,1654,1676]" pageId="8" pageNumber="107">J. Pelorce</collectorName>
(
<collectionCode box="[736,773,1654,1676]" country="0" name="Mohonk Preserve, Inc." pageId="8" pageNumber="107">MP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7661873" ID-Zenodo-Dep="7661873" httpUri="https://zenodo.org/record/7661873/files/figure.png" pageId="9" pageNumber="108" startId="9.[181,214,634,656]" targetBox="[203,1098,220,601]" targetPageId="9">
<paragraph blockId="9.[181,1142,634,728]" pageId="9" pageNumber="108">
Fig. 5.
<taxonomicName authority="Vannozzi &amp; Pizzini &amp; Raines, 2015" authorityName="Vannozzi &amp; Pizzini &amp; Raines" authorityYear="2015" box="[242,424,634,655]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="108" phylum="Mollusca" rank="species" species="knysnaense" status="sp. n.">
<emphasis box="[242,424,634,655]" italics="true" pageId="9" pageNumber="108">Caecum knysnaense</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[428,476,634,655]" pageId="9" pageNumber="108" rank="species">sp. n.</taxonomicNameLabel>
, South Africa: (AC) holotype, Knysna, Gericke Point, Sedgefield (NMSA S7625/T3362) — (A) right side, (B) ventral side, (C) microsculpture; (D, E) paratype, operculum, Walkers Point, W of Knysna (NMSA W9672/T3363) — (D) inner side, (E) outer side. Scale bar = 1 mm (A, B); 200 µm (D, E); 50 µm (C).
</paragraph>
</caption>
<paragraph blockId="9.[181,1142,784,1139]" box="[181,884,784,810]" pageId="9" pageNumber="108">
Distribution: Known with certainty only from Knysna (
<figureCitation box="[802,870,784,810]" captionStart="Fig" captionStartId="11.[181,214,758,780]" captionTargetBox="[221,1116,207,734]" captionTargetId="figure-383@11.[189,1134,203,771]" captionTargetPageId="11" captionText="Fig. 7. Distribution map of Caecum knysnaense sp. n. („), C. austrafricanum sp. n. (S) and C. intortum sp. n. (T). White symbol indicates the type locality." figureDoi="http://doi.org/10.5281/zenodo.7661877" httpUri="https://zenodo.org/record/7661877/files/figure.png" pageId="9" pageNumber="108">Fig. 7</figureCitation>
).
</paragraph>
<paragraph blockId="9.[181,1142,784,1139]" pageId="9" pageNumber="108">
Remarks:This species clearly resembles
<taxonomicName authority="Vannozzi &amp; Pizzini &amp; Raines, 2015" authorityName="Vannozzi &amp; Pizzini &amp; Raines" authorityYear="2015" box="[633,903,825,851]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="108" phylum="Mollusca" rank="species" species="austrafricanum" status="sp. n.">
<emphasis box="[633,903,825,851]" italics="true" pageId="9" pageNumber="108">Caecum austrafricanum</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[908,966,825,851]" pageId="9" pageNumber="108" rank="species">sp. n.</taxonomicNameLabel>
However, there are some consistent differences that set specimens of this species apart from the hundreds of
<taxonomicName authorityName="Vannozzi &amp; Pizzini &amp; Raines" authorityYear="2015" box="[211,415,888,914]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="108" phylum="Mollusca" rank="species" species="austrafricanum">
<emphasis box="[211,415,888,914]" italics="true" pageId="9" pageNumber="108">C. austrafricanum</emphasis>
</taxonomicName>
specimens found in the same sites. In fact,
<taxonomicName authority="Vannozzi &amp; Pizzini &amp; Raines, 2015" authorityName="Vannozzi &amp; Pizzini &amp; Raines" authorityYear="2015" box="[893,1053,888,914]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="108" phylum="Mollusca" rank="species" species="knysnaense" status="sp. n.">
<emphasis box="[893,1053,888,914]" italics="true" pageId="9" pageNumber="108">C. knysnaense</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1058,1117,888,914]" pageId="9" pageNumber="108" rank="species">sp. n.</taxonomicNameLabel>
is smaller and shows an aperture which is more oblique by ca. 10° and lacks a clear swelling. Additionally, the septum of
<taxonomicName authorityName="Vannozzi &amp; Pizzini &amp; Raines" authorityYear="2015" box="[499,661,951,978]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="108" phylum="Mollusca" rank="species" species="knysnaense">
<emphasis box="[499,661,951,978]" italics="true" pageId="9" pageNumber="108">C. knysnaense</emphasis>
</taxonomicName>
is convex while in
<taxonomicName authorityName="Vannozzi &amp; Pizzini &amp; Raines" authorityYear="2015" box="[885,1091,951,978]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="108" phylum="Mollusca" rank="species" species="austrafricanum">
<emphasis box="[885,1091,951,978]" italics="true" pageId="9" pageNumber="108">C. austrafricanum</emphasis>
</taxonomicName>
it is typically flat or S­shaped.Also the mucro and the general aspect of the tube are different.
<taxonomicName authorityName="Vannozzi &amp; Pizzini &amp; Raines" authorityYear="2015" box="[208,426,1017,1043]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="108" phylum="Mollusca" rank="species" species="knysnaense">
<emphasis box="[208,426,1017,1043]" italics="true" pageId="9" pageNumber="108">Caecum knysnaense</emphasis>
</taxonomicName>
also strongly resembles
<taxonomicName authority="van der Linden &amp; Moolenbeek, 2000" authorityName="van der Linden &amp; Moolenbeek" authorityYear="2000" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="108" phylum="Mollusca" rank="species" species="macrum">
<emphasis box="[685,804,1017,1043]" italics="true" pageId="9" pageNumber="108">C. macrum</emphasis>
<bibRefCitation author="VAN DER LINDEN, J. &amp; MOOLENBEEK, R. G." pageId="9" pageNumber="108" pagination="81 - 92" refId="ref24720" refString="VAN DER LINDEN, J. &amp; MOOLENBEEK, R. G. 2000. Caecidae (Mollusca: Gastropoda) from Mauritania. Vita Marina 47 (3): 81 - 92." type="journal article" year="2000">van der Linden &amp; Moolenbeek, 2000</bibRefCitation>
</taxonomicName>
(
<figureCitation box="[251,353,1048,1075]" captionStart="Fig" captionStartId="33.[181,214,1342,1364]" captionTargetBox="[226,1097,204,1321]" captionTargetId="figure-13@33.[226,1097,204,1321]" captionTargetPageId="33" captionText="Fig. 25. South and western African Caecum and Parastrophia species: (A) P. avaricosa sp. n., holotype, Mzamba (NMSA E1665/T3354); (B, C) P.ornata sp. n., Mzamba,near Natal border — (B) holotype (NMSA W9677/T3379), (C) paratype (JPM 2119); (D) C. sepimentum de Folin, 1868, Kosi Bay (NMSA S2010); (E) C. cf. oahuense Pilsbry, 1921, Aliwal Shoal, off Umkomaas area (JPM); (F) Caecum cf. gulosum Hedley, 1899, Kosi Bay (NMSA S2869); (G, H) C. succineum de Folin, 1880 — (G) between Bhanga Nek andKosi Bay (NMSAS1586), (H)Aliwal Shoal, off Scottburgh (NMSA W9684); (I) C. cf. glabrum (Montagu, 1803), Knysna, Gericke Point, Sedgefield (NMSA W9658); (J) C. glabrum (Montagu, 1803), Rabat, Morocco (MP); (K) C.lindae sp. n., holotype, Cape Morgan Lighthouse (NMSA W9682/T3924); (L, M) C. cf. musorstomi Pizzini, Raines &amp; Vannozzi, 2013, Landers reef (JPM); (N, O) C.austrafricanum sp.n., paratypes,Walkers Point,W of Knysna (NMSA W9656/T3351);(P) C.knysnaense sp.n., paratype,Knysna,Gericke Point,Sedgefield (NMSAW9673/ T3364); (Q) C. intortum sp. n., paratype, Walkers Point, W of Knysna (NMSA W9669/T3360); (R) C. macrum van der Linden &amp; Moolenbeek, 2000, Charbonnier, Senegal (MP); (S, T) C. crassum de Folin, 1870, Gabon (MP): (S) adult; (T) juvenile. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7661913" httpUri="https://zenodo.org/record/7661913/files/figure.png" pageId="9" pageNumber="108">Fig. 25R</figureCitation>
), from which it can be distinguished by the different shape of the mucro and by the more oblique aperture. Also the macula is different, being shorter and heartshaped in
<taxonomicName authorityName="van der Linden &amp; Moolenbeek" authorityYear="2000" box="[295,420,1113,1139]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="108" phylum="Mollusca" rank="species" species="macrum">
<emphasis box="[295,420,1113,1139]" italics="true" pageId="9" pageNumber="108">C. macrum</emphasis>
</taxonomicName>
(
<figureCitation box="[435,572,1112,1139]" captionStart="Fig" captionStartId="32.[144,177,1039,1061]" captionTargetBox="[151,1096,204,1021]" captionTargetId="figure-159@32.[151,1096,204,1021]" captionTargetPageId="32" captionText="Fig. 24. Ventral side of South and western African Caecum species: (A) C. austrafricanum sp. n., paratype, Walkers Point, W of Knysna (NMSA W9656/T3351); (B) C. knysnaense sp. n., paratype, Knysna, Gericke Point,Sedgefield (NMSA W9673/T3364);(C, D) C. intortum sp.n.— (C)paratype,Walkers Point, W of Knysna (NMSA W9669/T3360), (D) the same specimen slightly rotated, detail of the macula; (E, F) C. leilae sp. n. — (E) paratype, Mzamba, near Natal border (JPM 2119), (F) the same specimen, detail of the macula; (G) C. inhacaense Albano &amp; Pizzini, 2011, Mzamba, near Natal border (JPM 2119); (H) C. macrum van der Linden &amp; Moolenbeek, 2000, Charbonnier, Senegal (MP); (I) C. crassum de Folin, 1870, Banc dArguin, Mauritania (MP). Scale bar = 1 mm (AC, E, GI); 300 µm (D, F)." figureDoi="http://doi.org/10.5281/zenodo.7661911" httpUri="https://zenodo.org/record/7661911/files/figure.png" pageId="9" pageNumber="108">Fig. 24B, H</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>