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<document id="1FF9D38FE37997E9ECEE52E77A092488" ID-CLB-Dataset="9140" ID-DOI="10.11646/zootaxa.4845.2.8" ID-GBIF-Dataset="6a994e4b-2dd0-455f-8380-f7bfdcbaae68" ID-ISSN="1175-5326" ID-Zenodo-Dep="4406616" ID-ZooBank="748422C9-347E-4830-99A8-4D4309107E23" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609423818215" checkinUser="plazi" docAuthor="Wagner, Rüdiger &amp; Rada, Tonći" docDate="2020" docId="BD200242A414FFCDFF4150D5FE78BD89" docLanguage="en" docName="zootaxa.4845.2.8.pdf" docOrigin="Zootaxa 4845 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psychoda glamocensis Wagner &amp; Rada 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="281" masterDocId="41197A3AA417FFCBFFD65042FFDDBC24" masterDocTitle="Moth flies (Diptera, Psychodidae) living in the dark of caves in the Dinaric Karst" masterLastPageNumber="282" masterPageNumber="275" pageNumber="278" updateTime="1698850486362" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="282052ED51F8496C49AC4AC36733E1E2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="CB86C4C37EBD6286EBA7461BB0DB99EF">Moth flies (Diptera, Psychodidae) living in the dark of caves in the Dinaric Karst</mods:title>
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<mods:name id="91B31DB647CA5380C34420B7C0B27F64" type="personal">
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<mods:namePart id="BC93B789EE6A913539F1765C217BDB59">Wagner, Rüdiger</mods:namePart>
<mods:affiliation id="1704940B5B296065B5819446A4A2155A">University of Kassel, FB 10, Biology-Zoology, Heinrich-Plett-Strasse 40, D- 34132 Kassel, Germany</mods:affiliation>
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<mods:namePart id="E40A29CCCC6A58F543E7CB0139341549">Rada, Tonći</mods:namePart>
<mods:nameIdentifier id="A3087D7FAA7D2F71292AD1222E2F2B52" type="ORCID">0000-0001-5287-6336</mods:nameIdentifier>
<mods:nameIdentifier id="98FFFE0651D467D589D418185943E1EE" type="email">tonci.radja1@gmail.com</mods:nameIdentifier>
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<mods:title id="92ED9906FBA61BE73CDA0934E4F58DE0">Zootaxa</mods:title>
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<mods:part id="1C35176540A6540C76DA98E1B722D9EA">
<mods:date id="1089C4FF684F7088FEAD7EB95209129C">2020</mods:date>
<mods:detail id="EDAB6B9B80343FBE049311E22E29BE41" type="pubDate">
<mods:number id="E10D8749E1EC9656F39744A4704145ED">2020-09-02</mods:number>
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<mods:number id="AD41A09A27B72691008EBC114E6F01C4">4845</mods:number>
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<mods:classification id="74606A78C960A2756B589397A2635A28">journal article</mods:classification>
<mods:identifier id="1F2718F56F8FC3D81BA3227CC8517497" type="CLB-Dataset">9140</mods:identifier>
<mods:identifier id="6FEA6A42751952D2B74BB18E785EDC81" type="DOI">10.11646/zootaxa.4845.2.8</mods:identifier>
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<treatment id="BD200242A414FFCDFF4150D5FE78BD89" ID-DOI="http://doi.org/10.5281/zenodo.4407361" ID-GBIF-Taxon="176349868" ID-Zenodo-Dep="4407361" LSID="urn:lsid:plazi:treatment:BD200242A414FFCDFF4150D5FE78BD89" httpUri="http://treatment.plazi.org/id/BD200242A414FFCDFF4150D5FE78BD89" lastPageId="6" lastPageNumber="281" pageId="3" pageNumber="278">
<subSubSection id="7D93E0DFA414FFC8FF4150D5FDD6BC95" box="[151,523,151,178]" pageId="3" pageNumber="278" type="nomenclature">
<paragraph id="3536B354A414FFC8FF4150D5FDD6BC95" blockId="3.[151,523,151,214]" box="[151,523,151,178]" pageId="3" pageNumber="278">
<heading id="6E7E0438A414FFC8FF4150D5FDD6BC95" bold="true" box="[151,523,151,178]" fontSize="11" level="1" pageId="3" pageNumber="278" reason="1">
<emphasis id="07FD6F46A414FFC8FF4150D5FDD6BC95" bold="true" box="[151,523,151,178]" pageId="3" pageNumber="278">
<taxonomicName id="F289C8D7A414FFC8FF4150D5FE7BBC96" ID-CoL="9HYSP" authority="Wagner &amp; Rada, 2020" authorityName="Wagner &amp; Rada" authorityYear="2020" box="[151,422,151,178]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="glamocensis" status="sp. nov.">
<emphasis id="07FD6F46A414FFC8FF4150D5FE7BBC96" bold="true" box="[151,422,151,178]" italics="true" pageId="3" pageNumber="278">Psychoda glamocensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1CCED23DA414FFC8FE7850D5FDD6BC95" box="[430,523,151,177]" pageId="3" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7D93E0DFA414FFC8FF4150F9FEFABCF2" box="[151,295,187,214]" pageId="3" pageNumber="278" type="description">
<paragraph id="3536B354A414FFC8FF4150F9FEFABCF2" blockId="3.[151,523,151,214]" box="[151,295,187,214]" pageId="3" pageNumber="278">
(
<figureCitation id="ADB2AFD1A414FFC8FF4950F9FEC2BCF2" box="[159,287,187,214]" captionStart-0="FIGURES 10–21" captionStart-1="FIGURES 22–26" captionStartId-0="4.[151,264,1500,1524]" captionStartId-1="5.[151,264,1748,1772]" captionTargetBox-0="[159,1426,187,1468]" captionTargetBox-1="[165,1430,393,1713]" captionTargetId-0="figure-23@4.[151,1436,181,1477]" captionTargetId-1="figure-126@5.[151,1436,381,1725]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 10–21. Psychoda glamocensis. 10. Eyebridge of male holotype. 11–12. Eyebridges of male paratypes. 13. Same of female paratype. 14. Flagellomeres 10–13. 15. 2nd flagellomere with ascoid. 16. Male mouth parts.17–18. Female mouth parts. 19. Wing. 20. Female subgenital plate. 21. Internal genital sclerites, female. Scale: 1.0 mm (19), 0.1 mm (10–15, 20–21), 0.05 mm (16–18)." captionText-1="FIGURES 22–26. Psychoda glamocensis. 22. Hypandrium, gonostyli and aedeagus, holotype. 23. Epandrium, subepandrial plate and bases of epandrial processes. 24. Right epandrial process with retinaculum. 25. Left gonocoxite, parameral bridges and position of distiphallus sclerite, holotype. 26. Basiphallus and distiphallus sclerites and periaedeagal membrane. Scale: 0.1 mm (22–26)." figureDoi-0="http://doi.org/10.5281/zenodo.4406622" figureDoi-1="http://doi.org/10.5281/zenodo.4406626" httpUri-0="https://zenodo.org/record/4406622/files/figure.png" httpUri-1="https://zenodo.org/record/4406626/files/figure.png" pageId="3" pageNumber="278">Figs 10–26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7D93E0DFA414FFC8FF415141FEF3BD65" pageId="3" pageNumber="278" type="etymology">
<paragraph id="3536B354A414FFC8FF415141FEF3BD65" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="278">
<emphasis id="07FD6F46A414FFC8FF415141FEC3BD39" bold="true" box="[151,286,259,285]" pageId="3" pageNumber="278">Etymology.</emphasis>
Named after Glamoč, a town and municipality located in Canton 10 of the Federation of
<collectingCountry id="4D9EF3C4A414FFC8FAC85141FEF7BD65" name="Bosnia and Herzegovina" pageId="3" pageNumber="278">Bosnia and Herzegovina</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="7D93E0DFA414FFC8FF115109FAFFBDAD" pageId="3" pageNumber="278" type="diagnosis">
<paragraph id="3536B354A414FFC8FF115109FAFFBDAD" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="278">
<emphasis id="07FD6F46A414FFC8FF115109FE9FBD41" bold="true" box="[199,322,331,357]" pageId="3" pageNumber="278">Diagnosis.</emphasis>
<taxonomicName id="F289C8D7A414FFC8FE9E5109FD98BD41" authority="Wagner &amp; Rada, 2020" authorityName="Wagner &amp; Rada" authorityYear="2020" box="[328,581,331,357]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="glamocensis" status="sp. nov.">
<emphasis id="07FD6F46A414FFC8FE9E5109FD98BD41" box="[328,581,331,357]" italics="true" pageId="3" pageNumber="278">Psychoda glamocensis</emphasis>
</taxonomicName>
<emphasis id="07FD6F46A414FFC8FD9A5109FD78BD41" bold="true" box="[588,677,331,357]" pageId="3" pageNumber="278">
<taxonomicNameLabel id="1CCED23DA414FFC8FD9A5109FD78BD41" box="[588,677,331,357]" pageId="3" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from its congeners by the reduced eyebridge, gonostyli with a very long seta in the basal third and the subtriangular aedeagus with the axe-shaped sclerite.
</paragraph>
</subSubSection>
<subSubSection id="7D93E0DFA414FFC8FF1151D1FB53BF31" pageId="3" pageNumber="278" type="materials_examined">
<paragraph id="3536B354A414FFC8FF1151D1FBD7BE3D" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="278">
<emphasis id="07FD6F46A414FFC8FF1151D1FEA8BD89" bold="true" box="[199,373,403,429]" pageId="3" pageNumber="278">Type material.</emphasis>
<materialsCitation id="85E1B909A414FFC8FEAA51D1FBD7BE3D" ID-GBIF-Occurrence="3007911315" collectingDate="2018-07" collectingDateMax="2019-02-15" collectingDateMin="2018-07" country="Croatia" county="Kraljevkom Pit" elevation="1" location="Radosic" municipality="Golubinka" pageId="3" pageNumber="278" specimenCount="101" specimenCount-female="35" specimenCount-male="66" typeStatus="holotype">
<typeStatus id="EA320DF6A414FFC8FEAA51D1FE39BD89" box="[380,484,403,429]" pageId="3" pageNumber="278" type="holotype">Holotype</typeStatus>
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, Bosnia and Hercegovina, Golubnjača kod Skucana, Skucani, Glamoč (43˚58’31.9’’N/016˚54’43.6’’E,
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<quantity id="F2711EB1A414FFC8FE4551F5FE2DBDF6" box="[403,496,439,466]" metricMagnitude="0" metricUnit="m" metricValue="1.005" pageId="3" pageNumber="278" unit="m" value="1.005">1.005 m</quantity>
a.s.l.
</elevation>
).
<date id="41379594A414FFC8FD9251F5FD6BBDF5" box="[580,694,439,465]" pageId="3" pageNumber="278" value="2018-07">
<collectingDate id="51736C7CA414FFC8FD9251F5FD6BBDF5" box="[580,694,439,465]" pageId="3" pageNumber="278" value="2018-07">July 2018</collectingDate>
</date>
;
<typeStatus id="EA320DF6A414FFC8FD1451FAFCEDBDF6" box="[706,816,440,466]" pageId="3" pageNumber="278" type="paratype">paratypes</typeStatus>
,
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,
<specimenCount id="238F78DDA414FFC8FCB851F5FC4CBDF5" box="[878,913,439,465]" pageId="3" pageNumber="278" type="female">1♀</specimenCount>
same as
<typeStatus id="EA320DF6A414FFC8FC2C51F5FBBDBDF5" box="[1018,1120,439,465]" pageId="3" pageNumber="278" type="holotype">holotype</typeStatus>
;
<specimenCount id="238F78DDA414FFC8FBBA51F5FB4CBDF6" box="[1132,1169,439,466]" pageId="3" pageNumber="278" type="male">8♂</specimenCount>
,
<specimenCount id="238F78DDA414FFC8FB4851F5FB1FBDF5" box="[1182,1218,439,465]" pageId="3" pageNumber="278" type="female">2♀</specimenCount>
, (
<typeStatus id="EA320DF6A414FFC8FB0151F5FA95BDF5" box="[1239,1352,439,465]" pageId="3" pageNumber="278" type="paratype">paratypes</typeStatus>
), Croatia, Golubinka pod Kraljevcom, Radošić (43˚35’45.2’’ N/ 016˚20’32’’E,
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<quantity id="F2711EB1A414FFC8FC1E5199FBCDBDD2" box="[968,1040,475,502]" metricMagnitude="2" metricUnit="m" metricValue="2.33" pageId="3" pageNumber="278" unit="m" value="233.0">233 m</quantity>
asl
</elevation>
.)
<date id="41379594A414FFC8FB985199FAC4BDD1" box="[1102,1305,475,502]" pageId="3" pageNumber="278" value="2019-02-15">
<collectingDate id="51736C7CA414FFC8FB985199FAC4BDD1" box="[1102,1305,475,502]" pageId="3" pageNumber="278" value="2019-02-15">15 February 2019</collectingDate>
</date>
;
<specimenCount id="238F78DDA414FFC8FAF35199FA85BDD2" box="[1317,1368,475,502]" pageId="3" pageNumber="278" type="male">49♂</specimenCount>
,
<specimenCount id="238F78DDA414FFC8FAB25199FA4ABDD1" box="[1380,1431,475,501]" pageId="3" pageNumber="278" type="female">32♀</specimenCount>
, (
<typeStatus id="EA320DF6A414FFC8FF4951BDFECDBE3D" box="[159,272,511,537]" pageId="3" pageNumber="278" type="paratype">paratypes</typeStatus>
),
<collectingCountry id="4D9EF3C4A414FFC8FEF551BDFEA5BE3D" box="[291,376,511,537]" name="Croatia" pageId="3" pageNumber="278">Croatia</collectingCountry>
,
<location id="3056E58FA414FFC8FE5251BDFE3CBE3D" LSID="urn:lsid:plazi:treatment:BD200242A414FFCDFF4150D5FE78BD89:3056E58FA414FFC8FE5251BDFE3CBE3D" box="[388,481,511,537]" country="Croatia" county="Kraljevkom Pit" municipality="Golubinka" name="Radosic" pageId="3" pageNumber="278">Radošić</location>
,
<collectingMunicipality id="D552292EA414FFC8FE3A51BDFDBFBE3D" box="[492,610,511,537]" pageId="3" pageNumber="278">Golubinka</collectingMunicipality>
pod
<collectingCounty id="DC57CBD8A414FFC8FD4C51BDFC94BE3D" box="[666,841,511,537]" pageId="3" pageNumber="278">Kraljevkom Pit</collectingCounty>
,
<date id="41379594A414FFC8FC8251BDFBDBBE3D" box="[852,1030,511,537]" pageId="3" pageNumber="278" value="2018-10-08">
<collectingDate id="51736C7CA414FFC8FC8251BDFBDBBE3D" box="[852,1030,511,537]" pageId="3" pageNumber="278" value="2018-10-08">8 October 2018</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="3536B354A414FFC8FF115261FB54BEEA" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="278">
<emphasis id="07FD6F46A414FFC8FF115261FE4FBE19" bold="true" box="[199,402,547,573]" pageId="3" pageNumber="278">Further material</emphasis>
.
<materialsCitation id="85E1B909A414FFC8FE765261FBFCBE19" ID-GBIF-Occurrence="3007911301" box="[416,1057,547,574]" collectingDate="2019-04-06" country="Croatia" location="Rusica" municipality="Ercegovici" pageId="3" pageNumber="278" specimenCount="41" specimenCount-female="5" specimenCount-male="36">
<specimenCount id="238F78DDA414FFC8FE765261FE09BE19" box="[416,468,547,573]" pageId="3" pageNumber="278" type="male">36♂</specimenCount>
,
<specimenCount id="238F78DDA414FFC8FE095261FDDEBE1A" box="[479,515,547,574]" pageId="3" pageNumber="278" type="female">5♀</specimenCount>
,
<collectingCountry id="4D9EF3C4A414FFC8FDC65261FDB8BE19" box="[528,613,547,573]" name="Croatia" pageId="3" pageNumber="278">Croatia</collectingCountry>
,
<collectingMunicipality id="D552292EA414FFC8FDA75261FD30BE19" box="[625,749,547,573]" pageId="3" pageNumber="278">Ercegovici</collectingMunicipality>
,
<location id="3056E58FA414FFC8FD2E5261FC99BE19" LSID="urn:lsid:plazi:treatment:BD200242A414FFCDFF4150D5FE78BD89:3056E58FA414FFC8FD2E5261FC99BE19" box="[760,836,547,573]" country="Croatia" municipality="Ercegovici" name="Rusica" pageId="3" pageNumber="278">Rušića</location>
jama,
<date id="41379594A414FFC8FC465261FBFCBE19" box="[912,1057,547,573]" pageId="3" pageNumber="278" value="2019-04-06">
<collectingDate id="51736C7CA414FFC8FC465261FBFCBE19" box="[912,1057,547,573]" pageId="3" pageNumber="278" value="2019-04-06">6 April 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="85E1B909A414FFC8FBFA5261FED6BE45" ID-GBIF-Occurrence="3007911314" collectingDate="2019-06-05" country="Croatia" location="Bakusa" municipality="Trilj" pageId="3" pageNumber="278" specimenCount="57" specimenCount-female="8" specimenCount-male="49">
<specimenCount id="238F78DDA414FFC8FBFA5261FBBDBE19" box="[1068,1120,547,573]" pageId="3" pageNumber="278" type="male">49♂</specimenCount>
,
<specimenCount id="238F78DDA414FFC8FBBA5261FB4DBE19" box="[1132,1168,547,573]" pageId="3" pageNumber="278" type="female">8♀</specimenCount>
,
<collectingCountry id="4D9EF3C4A414FFC8FB4B5261FB32BE19" box="[1181,1263,547,573]" name="Croatia" pageId="3" pageNumber="278">Croatia</collectingCountry>
<collectingMunicipality id="D552292EA414FFC8FB205261FAF7BE19" box="[1270,1322,547,573]" pageId="3" pageNumber="278">Trilj</collectingMunicipality>
,
<location id="3056E58FA414FFC8FAE35261FA5ABE19" LSID="urn:lsid:plazi:treatment:BD200242A414FFCDFF4150D5FE78BD89:3056E58FA414FFC8FAE35261FA5ABE19" box="[1333,1415,547,573]" country="Croatia" municipality="Trilj" name="Bakusa" pageId="3" pageNumber="278">Bakuša</location>
<date id="41379594A414FFC8FA585261FED6BE45" pageId="3" pageNumber="278" value="2019-06-05">
<collectingDate id="51736C7CA414FFC8FA585261FED6BE45" pageId="3" pageNumber="278" value="2019-06-05">5 June 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="85E1B909A414FFC8FEC05205FC7FBE45" ID-GBIF-Occurrence="3007911302" box="[278,930,583,610]" collectingDate="2019-02-24" country="Croatia" county="Vostane" location="Ljutu" municipality="Golubinka" pageId="3" pageNumber="278" specimenCount="1" specimenCount-male="1">
<specimenCount id="238F78DDA414FFC8FEC05205FEE4BE45" box="[278,313,583,609]" pageId="3" pageNumber="278" type="male">1♂</specimenCount>
<collectingCountry id="4D9EF3C4A414FFC8FEE85205FE4EBE45" box="[318,403,583,609]" name="Croatia" pageId="3" pageNumber="278">Croatia</collectingCountry>
,
<collectingCounty id="DC57CBD8A414FFC8FE4B5205FE25BE45" box="[413,504,583,609]" pageId="3" pageNumber="278">Voštane</collectingCounty>
,
<collectingMunicipality id="D552292EA414FFC8FDD45205FDA5BE45" box="[514,632,583,609]" pageId="3" pageNumber="278">Golubinka</collectingMunicipality>
u
<location id="3056E58FA414FFC8FD475205FD0CBE45" LSID="urn:lsid:plazi:treatment:BD200242A414FFCDFF4150D5FE78BD89:3056E58FA414FFC8FD475205FD0CBE45" box="[657,721,583,609]" country="Croatia" county="Vostane" municipality="Golubinka" name="Ljutu" pageId="3" pageNumber="278">Ljutu</location>
,
<date id="41379594A414FFC8FD0C5205FC7FBE45" box="[730,930,583,609]" pageId="3" pageNumber="278" value="2019-02-24">
<collectingDate id="51736C7CA414FFC8FD0C5205FC7FBE45" box="[730,930,583,609]" pageId="3" pageNumber="278" value="2019-02-24">24 February 2019</collectingDate>
</date>
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;
<materialsCitation id="85E1B909A414FFC8FC7A5205FE36BEA1" ID-GBIF-Occurrence="3007911310" collectingDate="2019-03-13" country="Croatia" location="Velika Zecica Donji" municipality="Labin Dalmatinski" pageId="3" pageNumber="278" specimenCount="44" specimenCount-female="2" specimenCount-male="42">
<specimenCount id="238F78DDA414FFC8FC7A5205FC3DBE45" box="[940,992,583,609]" pageId="3" pageNumber="278" type="male">42♂</specimenCount>
,
<specimenCount id="238F78DDA414FFC8FC3C5205FBD3BE45" box="[1002,1038,583,609]" pageId="3" pageNumber="278" type="female">2♀</specimenCount>
,
<collectingCountry id="4D9EF3C4A414FFC8FBCC5205FBB2BE45" box="[1050,1135,583,609]" name="Croatia" pageId="3" pageNumber="278">Croatia</collectingCountry>
,
<collectingMunicipality id="D552292EA414FFC8FBAF5205FA91BE45" box="[1145,1356,583,609]" pageId="3" pageNumber="278">Labin Dalmatinski</collectingMunicipality>
,
<location id="3056E58FA414FFC8FA835205FEF0BEA1" LSID="urn:lsid:plazi:treatment:BD200242A414FFCDFF4150D5FE78BD89:3056E58FA414FFC8FA835205FEF0BEA1" country="Croatia" municipality="Labin Dalmatinski" name="Velika Zecica Donji" pageId="3" pageNumber="278">Velika Zečica Donji</location>
,
<date id="41379594A414FFC8FEEC5229FE36BEA1" box="[314,491,619,645]" pageId="3" pageNumber="278" value="2019-03-13">
<collectingDate id="51736C7CA414FFC8FEEC5229FE36BEA1" box="[314,491,619,645]" pageId="3" pageNumber="278" value="2019-03-13">13 March 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="85E1B909A414FFC8FE2F5229FBB9BEA1" ID-GBIF-Occurrence="3007911305" box="[505,1124,619,646]" collectingDate="2019-05-19" country="Croatia" location="Elezova" municipality="Zelovo" pageId="3" pageNumber="278" specimenCount="7" specimenCount-female="2" specimenCount-male="5">
<specimenCount id="238F78DDA414FFC8FE2F5229FDC3BEA2" box="[505,542,619,646]" pageId="3" pageNumber="278" type="male">5♂</specimenCount>
,
<specimenCount id="238F78DDA414FFC8FDFA5229FD8DBEA1" box="[556,592,619,645]" pageId="3" pageNumber="278" type="female">2♀</specimenCount>
,
<collectingCountry id="4D9EF3C4A414FFC8FD885229FD6EBEA1" box="[606,691,619,645]" name="Croatia" pageId="3" pageNumber="278">Croatia</collectingCountry>
,
<collectingMunicipality id="D552292EA414FFC8FD175229FCCEBEA1" box="[705,787,619,645]" pageId="3" pageNumber="278">Zelovo</collectingMunicipality>
,
<location id="3056E58FA414FFC8FCF65229FCA7BEA1" LSID="urn:lsid:plazi:treatment:BD200242A414FFCDFF4150D5FE78BD89:3056E58FA414FFC8FCF65229FCA7BEA1" box="[800,890,619,645]" country="Croatia" municipality="Zelovo" name="Elezova" pageId="3" pageNumber="278">Elezova</location>
jama,
<date id="41379594A414FFC8FC1F5229FBB9BEA1" box="[969,1124,619,646]" pageId="3" pageNumber="278" value="2019-05-19">
<collectingDate id="51736C7CA414FFC8FC1F5229FBB9BEA1" box="[969,1124,619,646]" pageId="3" pageNumber="278" value="2019-05-19">19 May 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="85E1B909A414FFC8FBA45229FD93BE8D" ID-GBIF-Occurrence="3007911313" collectingDate="2019-03-10" country="Croatia" location="Dzelalijina Golubinka" municipality="Unesic" pageId="3" pageNumber="278" specimenCount="17" specimenCount-female="5" specimenCount-male="12">
<specimenCount id="238F78DDA414FFC8FBA45229FB78BEA1" box="[1138,1189,619,645]" pageId="3" pageNumber="278" type="male">12♂</specimenCount>
,
<specimenCount id="238F78DDA414FFC8FB645229FB0ABEA2" box="[1202,1239,619,646]" pageId="3" pageNumber="278" type="female">5♀</specimenCount>
,
<collectingCountry id="4D9EF3C4A414FFC8FB335229FAE7BEA1" box="[1253,1338,619,645]" name="Croatia" pageId="3" pageNumber="278">Croatia</collectingCountry>
,
<collectingMunicipality id="D552292EA414FFC8FA915229FA45BEA1" box="[1351,1432,619,645]" pageId="3" pageNumber="278">Unešić</collectingMunicipality>
,
<location id="3056E58FA414FFC8FF4152CDFE4EBE8D" LSID="urn:lsid:plazi:treatment:BD200242A414FFCDFF4150D5FE78BD89:3056E58FA414FFC8FF4152CDFE4EBE8D" box="[151,403,655,681]" country="Croatia" municipality="Unesic" name="Dzelalijina Golubinka" pageId="3" pageNumber="278">Dželalijina Golubinka</location>
,
<date id="41379594A414FFC8FE4852CDFD93BE8D" box="[414,590,655,681]" pageId="3" pageNumber="278" value="2019-03-10">
<collectingDate id="51736C7CA414FFC8FE4852CDFD93BE8D" box="[414,590,655,681]" pageId="3" pageNumber="278" value="2019-03-10">10 March 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="85E1B909A414FFC8FD8C52CDFCD5BE8D" ID-GBIF-Occurrence="3007911307" box="[602,776,655,682]" collectionCode="T" collectorName="Rada" country="Croatia" pageId="3" pageNumber="278" specimenCount="1">
all leg.
<collectionCode id="53982B91A414FFC8FD7852D2FD1DBE8E" box="[686,704,656,682]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="278">T</collectionCode>
.
<collectorName id="987CD682A414FFC8FD1D52CDFCD5BE8D" box="[715,776,655,681]" pageId="3" pageNumber="278">Rada</collectorName>
</materialsCitation>
;
<materialsCitation id="85E1B909A414FFC8FCC252CDFD7ABEE9" ID-GBIF-Occurrence="3007911308" collectingDate="1972-10-12" collectorName="Malicky" country="Austria" location="Schreierbach" municipality="Lunz am See" pageId="3" pageNumber="278" specimenCount="1" specimenCount-male="1" stateProvince="Niederoesterreich">
<specimenCount id="238F78DDA414FFC8FCC252CDFCEABE8D" box="[788,823,655,681]" pageId="3" pageNumber="278" type="male">1♂</specimenCount>
<collectingCountry id="4D9EF3C4A414FFC8FCEB52CDFC4FBE8D" box="[829,914,655,681]" name="Austria" pageId="3" pageNumber="278">Austria</collectingCountry>
,
<collectingRegion id="F74D7DB6A414FFC8FC4852D2FB9FBE8D" box="[926,1090,655,682]" country="Austria" name="Niederoesterreich" pageId="3" pageNumber="278">Lower Austria</collectingRegion>
,
<collectingMunicipality id="D552292EA414FFC8FB9852D2FB39BE8D" box="[1102,1252,655,682]" pageId="3" pageNumber="278">Lunz am See</collectingMunicipality>
,
<collectingMethod id="ECC8CB43A414FFC8FB2652D2FA40BE8E" box="[1264,1437,656,682]" pageId="3" pageNumber="278">emergence trap</collectingMethod>
at
<location id="3056E58FA414FFC8FF6452F1FE9ABEE9" LSID="urn:lsid:plazi:treatment:BD200242A414FFCDFF4150D5FE78BD89:3056E58FA414FFC8FF6452F1FE9ABEE9" box="[178,327,691,717]" country="Austria" municipality="Lunz am See" name="Schreierbach" pageId="3" pageNumber="278" stateProvince="Niederoesterreich">Schreierbach</location>
,
<date id="41379594A414FFC8FE8552F1FDD2BEE9" box="[339,527,691,717]" pageId="3" pageNumber="278" value="1972-10-12">
<collectingDate id="51736C7CA414FFC8FE8552F1FDD2BEE9" box="[339,527,691,717]" pageId="3" pageNumber="278" value="1972-10-12">12 October 1972</collectingDate>
</date>
leg.
<collectorName id="987CD682A414FFC8FD9052F1FD7ABEE9" box="[582,679,691,717]" pageId="3" pageNumber="278">Malicky</collectorName>
</materialsCitation>
; 1
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report from
<collectingCountry id="4D9EF3C4A414FFC8FC8E52F1FC77BEE9" box="[856,938,691,717]" name="Austria" pageId="3" pageNumber="278">Austria</collectingCountry>
(in collection RW).
</paragraph>
<paragraph id="3536B354A414FFC8FF115295FB53BF31" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="278">
<materialsCitation id="85E1B909A414FFC8FF115295FB56BF31" ID-GBIF-Occurrence="3007911309" collectorName="Natural History Museum &amp; Zoo" country="Croatia" location="Croatia" pageId="3" pageNumber="278" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="holotype">
<typeStatus id="EA320DF6A414FFC8FF115295FEF2BED5" box="[199,303,727,753]" pageId="3" pageNumber="278" type="holotype">Holotype</typeStatus>
and
<typeStatus id="EA320DF6A414FFC8FEBD529AFE0BBED6" box="[363,470,728,754]" pageId="3" pageNumber="278" type="paratype">paratypes</typeStatus>
and the mentioned further material are deposited in the
<collectorName id="987CD682A414FFC8FBB65295FAA0BED6" box="[1120,1405,727,754]" pageId="3" pageNumber="278">Natural History Museum</collectorName>
&amp;
<collectorName id="987CD682A414FFC8FF4152BEFF1ABF32" box="[151,199,764,790]" pageId="3" pageNumber="278">Zoo</collectorName>
, Split,
<collectingCountry id="4D9EF3C4A414FFC8FEC352B9FEB6BF31" box="[277,363,763,789]" name="Croatia" pageId="3" pageNumber="278">Croatia</collectingCountry>
. Three
<typeStatus id="EA320DF6A414FFC8FE6B52BEFDF5BF32" box="[445,552,764,790]" pageId="3" pageNumber="278" type="paratype">paratypes</typeStatus>
(
<specimenCount id="238F78DDA414FFC8FDEF52B9FD83BF31" box="[569,606,763,789]" pageId="3" pageNumber="278" type="male">2♂</specimenCount>
,
<specimenCount id="238F78DDA414FFC8FDBC52B9FD56BF31" box="[618,651,763,789]" pageId="3" pageNumber="278" type="female">1♀</specimenCount>
) remain in the collection of the senior author
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7D93E0DFA414FFCFFF11535DFB2BBB31" lastPageId="4" lastPageNumber="279" pageId="3" pageNumber="278" type="description">
<paragraph id="3536B354A414FFC8FF11535DFEEAB87E" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="278">
<emphasis id="07FD6F46A414FFC8FF11535DFE7ABF1D" bold="true" box="[199,423,799,825]" pageId="3" pageNumber="278">Description. Male:</emphasis>
Eyes reniform, eyebridge with 2 or 3 rows of irregularly arranged facets, interocular suture wide U-shaped, distance between the eyes 2–2.5 facet diameter (figs 10–12). Antenna (fig. 14–15) with scape, pedicel and 13 flagellomeres; scape short, wider than long, pedicel spherical, flagellomeres bottle-shaped, with remarkably long necks, flagellomeres 1–10 with a pair of Y-shaped ascoids about as long as a flagellomere, apical flagellomeres 11–13 reduced in size, 12 and 13 with a subapical lobe and a short stiff bristle. Relative length of scape, pedicel and flagellomeres: 49-28-88-83-82-81-77-76-72-66-69-57-25-15-12; absolute length: 0.13-0.07- 0.23-0.22-0.22-0.21-0.20-0.20-0.19-0.17-0.18-0.15-0.07-0.04-
<quantity id="F2711EB1A414FFC8FC8453B5FC60B836" box="[850,957,1015,1042]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="278" unit="mm" value="0.03">0.03 mm</quantity>
. Four palpus segments, terminal segment flexible; relative length: 49-49-46-55; absolute length: 0.13-0.13-
<quantity id="F2711EB1A414FFC8FCA85459FBFEB812" box="[894,1059,1051,1078]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="3" pageNumber="278" unit="mm" value="0.13" valueMax="0.14" valueMin="0.12">0.12-0.14 mm</quantity>
. Mouth parts with 4 acute apical teeth (fig. 16).
</paragraph>
<paragraph id="3536B354A414FFC8FF115421FD7DB929" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="278">
Wing length
<quantity id="F2711EB1A414FFC8FE8E5421FE6CB85A" box="[344,433,1123,1150]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="278" unit="mm" value="3.1">3.1 mm</quantity>
, width
<quantity id="F2711EB1A414FFC8FDD55421FD81B85A" box="[515,604,1123,1150]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="278" unit="mm" value="1.1">1.1 mm</quantity>
. C basally with 2 swellings; Sc short, a setose area, terminates in wing. R
<subScript id="A90DB111A414FFC8FA425434FA41B8A0" attach="left" box="[1428,1436,1142,1156]" fontSize="6" pageId="3" pageNumber="278">1</subScript>
terminates in the distal half of the wing; division of Rs with fork R
<subScript id="A90DB111A414FFC8FCA754D8FCA4B88C" attach="left" box="[881,889,1178,1192]" fontSize="6" pageId="3" pageNumber="278">2</subScript>
/ R
<subScript id="A90DB111A414FFC8FC4C54D8FC6EB88C" attach="left" box="[922,947,1178,1192]" fontSize="6" pageId="3" pageNumber="278">3+4</subScript>
at level of Sc tip. R
<subScript id="A90DB111A414FFC8FB5C54DBFB4FB883" attach="left" box="[1162,1170,1177,1191]" fontSize="6" pageId="3" pageNumber="278">5</subScript>
terminates short behind wing tip; radial and medial fork complete; medial fork basal of radial fork. CuA basally broader; in the basal part of the wing all veins are broken, here is fold that enables specimens to hold the wing in roof shape at rest. Cross veins not detected. Thorax without specific features.
</paragraph>
<paragraph id="3536B354A414FFC8FF115555FB5ABA75" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="278">Abdomen with 8 segments, genitalia (figs 22–26) inverted; hypandrium broad, slightly broader in the midst. Gonocoxites short tubular about 1.5 times longer than wide, probably without setae. Dorsal parameral bridge (fig. 25) of gonocoxites thin in the middle, laterally widened; ventral bridge of parameres are blade shaped sclerites with sharp bent tips that do not meet in the middle. Gonostyli elongate triangular. almost straight; behind the basal swelling with a remarkably long and strong seta, about as long as the gonostylus. Epandrium (fig. 23) with 1 median foramen; edges roundish. basal edge deeply curved, apical edge almost straight. Subepandial plate increasingly wide apically with a broad median connection to the proctiger and to lateral plates connecting with the bases of the basally swollen epandrial appendages. Proctiger not clearly discernable; the epandrial processes clearly longer than the epandrium, strongly bent, apically with a sharp tip beside the basis of the retinaculum.</paragraph>
<paragraph id="3536B354A414FFC8FF115619FC90BABD" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="278">Aedeagus consists of an asymmetric bilobed basal sclerite, lobes apically curved inwards; the distal sclerite is shorter than the basal sclerite, asymmetric, tomahawk shaped.</paragraph>
<paragraph id="3536B354A414FFC8FF1156E1FCD1BAC5" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="278">
<emphasis id="07FD6F46A414FFC8FF1156E1FEF5BA99" bold="true" box="[199,296,1699,1725]" pageId="3" pageNumber="278">Female:</emphasis>
in most respects similar to male but poorly preserved. Differences are minor in the antenna and wing venation. Eye bridge fig. 13, mouth parts fig. 17 and 18.
</paragraph>
<paragraph id="3536B354A414FFC8FF1156A9FADABB69" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="278">
Relative length of antennal articles: 34-24-57-51-53-48-52-48-42-; absolute length: 0,09-0,06-0,15-0,13-0,14- 0,13-0,14-0,13-0,11- mm, apical segments missing. Four palpus segments, terminal segment flexible; relative length: 30-28-28-34; absolute length: 0.08-0.07-
<quantity id="F2711EB1A414FFC8FD655771FC8BBB6A" box="[691,854,1843,1870]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7" pageId="3" pageNumber="278" unit="mm" value="0.08" valueMax="0.09" valueMin="0.07">0.07-0.09 mm</quantity>
. Mouth parts with 4 acute apical teeth.
</paragraph>
<paragraph id="3536B354A414FFC8FF115715FDB1BB56" blockId="3.[151,1437,259,2013]" box="[199,620,1879,1906]" pageId="3" pageNumber="278">
Wing length approximately
<quantity id="F2711EB1A414FFC8FDD75715FDB5BB56" box="[513,616,1879,1906]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="278" unit="mm" value="2.2">2.20 mm</quantity>
.
</paragraph>
<paragraph id="3536B354A414FFC8FF115739FD3CBBF9" blockId="3.[151,1437,259,2013]" pageId="3" pageNumber="278">Genitalia (figs. 20, 21) with a bilobed subgenital plate with an elongate tubular structure, laterally accompanied by a pair of sclerites that appear to be superficially similar to the parameral complex of the male. Basally is a complicatedly folded sclerite of probably specific shape.</paragraph>
<caption id="61F6E3DCA413FFCFFF41559EFEC1BA45" ID-DOI="http://doi.org/10.5281/zenodo.4406622" ID-Zenodo-Dep="4406622" httpUri="https://zenodo.org/record/4406622/files/figure.png" pageId="4" pageNumber="279" startId="4.[151,264,1500,1524]" targetBox="[159,1426,187,1468]" targetPageId="4">
<paragraph id="3536B354A413FFCFFF41559EFEC1BA45" blockId="4.[151,1437,1500,1633]" pageId="4" pageNumber="279">
<emphasis id="07FD6F46A413FFCFFF41559EFD96B9D1" bold="true" box="[151,587,1500,1525]" pageId="4" pageNumber="279">
FIGURES 10–21.
<taxonomicName id="F289C8D7A413FFCFFE8C559EFD99B9D0" authorityName="Wagner &amp; Rada" authorityYear="2020" box="[346,580,1500,1524]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="glamocensis">
<emphasis id="07FD6F46A413FFCFFE8C559EFD99B9D0" bold="true" box="[346,580,1500,1524]" italics="true" pageId="4" pageNumber="279">Psychoda glamocensis</emphasis>
</taxonomicName>
.
</emphasis>
10. Eyebridge of male holotype. 11–12. Eyebridges of male paratypes. 13. Same of female paratype. 14. Flagellomeres 10–13. 15. 2
<superScript id="C2FC1E1CA413FFCFFD555643FD4FBA2B" attach="left" box="[643,658,1537,1551]" fontSize="6" pageId="4" pageNumber="279">nd</superScript>
flagellomere with ascoid. 16. Male mouth parts.17–18. Female mouth parts. 19. Wing. 20. Female subgenital plate. 21. Internal genital sclerites, female. Scale: 1.0 mm (19), 0.1 mm (10–15, 20–21), 0.05 mm (16–18).
</paragraph>
</caption>
<paragraph id="3536B354A413FFCFFF1156CDFB2BBB31" blockId="4.[151,1437,1679,2030]" pageId="4" pageNumber="279">
<emphasis id="07FD6F46A413FFCFFF1156CDFE52BA8D" bold="true" box="[199,399,1679,1705]" pageId="4" pageNumber="279">DNA Barcoding.</emphasis>
The availability of a DNA barcode permits comparison with other
<taxonomicName id="F289C8D7A413FFCFFB4C56CDFADABA8D" box="[1178,1287,1679,1705]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="07FD6F46A413FFCFFB4C56CDFADABA8D" box="[1178,1287,1679,1705]" italics="true" pageId="4" pageNumber="279">Psychoda</emphasis>
</taxonomicName>
species. The two sequences (ZFMK-TIS-2617210 consensus sequence (reversed), ZFMK-TIS-2617212 consensus sequence (reversed)) indicate the closest relation with specimens of
<taxonomicName id="F289C8D7A413FFCFFCE15695FB7ABAD5" authority="Vaillant 1973" authorityName="Vaillant" authorityYear="1973" box="[823,1191,1751,1777]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="07FD6F46A413FFCFFCE15695FBDEBAD5" box="[823,1027,1751,1777]" italics="true" pageId="4" pageNumber="279">Psychoda alticola</emphasis>
Vaillant 1973
</taxonomicName>
, from the Norwegian
<collectingRegion id="F74D7DB6A413FFCFFF4156B9FED9BB31" box="[151,260,1787,1813]" country="Norway" name="Finnmark" pageId="4" pageNumber="279">Finnmark</collectingRegion>
(
<bibRefCitation id="5118CEA5A413FFCFFEC556B9FDF0BB31" author="Kvifte, G. M. &amp; Andersen, T." box="[275,557,1787,1813]" pageId="4" pageNumber="279" pagination="108 - 119" refId="ref4262" refString="Kvifte, G. M. &amp; Andersen, T. (2012) Moth flies (Diptera, Psychodidae) from Finnmark, northern Norway. Norwegian Journal of Entomology, 59, 108 - 119." type="journal article" year="2012">Kvifte &amp; Andersen 2012</bibRefCitation>
). Sequences from Alpine specimens are presently not available.
</paragraph>
</subSubSection>
<subSubSection id="7D93E0DFA413FFCDFF11575DFE78BD89" lastPageId="6" lastPageNumber="281" pageId="4" pageNumber="279" type="discussion">
<paragraph id="3536B354A413FFCEFF11575DFB23BCF1" blockId="4.[151,1437,1679,2030]" lastBlockId="5.[151,1436,151,357]" lastPageId="5" lastPageNumber="280" pageId="4" pageNumber="279">
<emphasis id="07FD6F46A413FFCFFF11575DFEE6BB1D" bold="true" box="[199,315,1823,1849]" pageId="4" pageNumber="279">Remarks.</emphasis>
The most striking feature of both new species is the reduction of the eyes probably due to life in caves The number of facets and facet rows varies extremely and differs even between specimens. In
<emphasis id="07FD6F46A413FFCFFAC35701FF61BBA5" italics="true" pageId="4" pageNumber="279">
<taxonomicName id="F289C8D7A413FFCFFAC35701FA5FBB79" box="[1301,1410,1859,1885]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus" sensu="lato">Psychoda</taxonomicName>
<taxonomicNameLabel id="1CCED23DA413FFCFFA5C5706FF61BBA5" pageId="4" pageNumber="279" sensu="lato">s. lat.</taxonomicNameLabel>
</emphasis>
species the eye bridge usually consists of 4 parallel regular rows but in
<taxonomicName id="F289C8D7A413FFCFFC35572AFB4ABBA5" authority="Wagner &amp; Rada, 2020" authorityName="Wagner &amp; Rada" authorityYear="2020" box="[995,1175,1895,1921]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="glamocensis" status="sp. nov.">
<emphasis id="07FD6F46A413FFCFFC35572AFB4ABBA5" box="[995,1175,1895,1921]" italics="true" pageId="4" pageNumber="279">Ps. glamocensis</emphasis>
</taxonomicName>
<emphasis id="07FD6F46A413FFCFFB485725FB25BBA5" bold="true" box="[1182,1272,1895,1921]" pageId="4" pageNumber="279">
<taxonomicNameLabel id="1CCED23DA413FFCFFB485725FB25BBA5" box="[1182,1272,1895,1921]" pageId="4" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the number of facet rows is reduced to mostly 2, with irregular arranged facets. Eyes are widely separate in both sexes so that the shape of the eyebridge varies between individuals. All specimens were collected deep in the caves, at Golubnjača kod Skucana at a depth of
<quantity id="F2711EB1A413FFCFFE135791FE22BBC9" box="[453,511,2003,2029]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="279" unit="m" value="30.0">30 m</quantity>
at permanent darkness, and at Golubinka pod Kraljevcom
<quantity id="F2711EB1A413FFCFFB4C5791FB11BBCA" box="[1178,1228,2003,2030]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="279" unit="m" value="25.0">25m</quantity>
deep at semidarkness. The malformation of palpus segments in several specimens is probably due to in-breeding in small populations in different caves. Number of teeth on mouthparts is as well variable and differs between specimens.
</paragraph>
<paragraph id="3536B354A412FFCEFF11509DFABDBD41" blockId="5.[151,1436,151,357]" pageId="5" pageNumber="280">This indicates that populations live for a long time in the caves. Long term low temperatures and enduring darkness are probable preconditions to develop morphological changes of the eyes and palpus segments. The antenna appears longer compared with the head diameter and the long seta on the gonostyli are further indications that antennae and setae may play major roles as tactile instruments to detect and locate conspecifics or avoid predators.</paragraph>
<caption id="61F6E3DCA412FFCEFF415696FEC1BB7C" ID-DOI="http://doi.org/10.5281/zenodo.4406626" ID-Zenodo-Dep="4406626" httpUri="https://zenodo.org/record/4406626/files/figure.png" pageId="5" pageNumber="280" startId="5.[151,264,1748,1772]" targetBox="[165,1430,393,1713]" targetPageId="5">
<paragraph id="3536B354A412FFCEFF415696FEC1BB7C" blockId="5.[151,1437,1748,1880]" pageId="5" pageNumber="280">
<emphasis id="07FD6F46A412FFCEFF415696FD8DBAC9" bold="true" box="[151,592,1748,1773]" pageId="5" pageNumber="280">
FIGURES 22–26.
<taxonomicName id="F289C8D7A412FFCEFE885696FD97BAC8" authorityName="Wagner &amp; Rada" authorityYear="2020" box="[350,586,1748,1772]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="280" phylum="Arthropoda" rank="species" species="glamocensis">
<emphasis id="07FD6F46A412FFCEFE885696FD97BAC8" bold="true" box="[350,586,1748,1772]" italics="true" pageId="5" pageNumber="280">Psychoda glamocensis</emphasis>
</taxonomicName>
.
</emphasis>
22. Hypandrium, gonostyli and aedeagus, holotype. 23. Epandrium, subepandrial plate and bases of epandrial processes. 24. Right epandrial process with retinaculum. 25. Left gonocoxite, parameral bridges and position of distiphallus sclerite, holotype. 26. Basiphallus and distiphallus sclerites and periaedeagal membrane. Scale: 0.1 mm (22–26).
</paragraph>
</caption>
<paragraph id="3536B354A412FFCDFF1157C5FB1FBCF1" blockId="5.[151,1437,1927,2025]" lastBlockId="6.[151,1436,151,429]" lastPageId="6" lastPageNumber="281" pageId="5" pageNumber="280">
The sex ratio in collections from all localities is unusual for
<taxonomicName id="F289C8D7A412FFCEFC4657C5FC20BB85" box="[912,1021,1927,1953]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="07FD6F46A412FFCEFC4657C5FC20BB85" box="[912,1021,1927,1953]" italics="true" pageId="5" pageNumber="280">Psychoda</emphasis>
</taxonomicName>
species; light trap collections often contain exclusively females. However, in Golubnjača kod Skucana cave 90%, in the Golubinka pod Kraljevcom cave 80%, and in Golubinka pod Kraljevkom cave 60% males were collected. The first two caves give shelter to wild pigeon (
<taxonomicName id="F289C8D7A411FFCDFEFA50D5FDF6BC96" authority="Gmelin" authorityName="Gmelin" box="[300,555,151,178]" class="Aves" family="Columbidae" genus="Columba" kingdom="Animalia" order="Columbiformes" pageId="6" pageNumber="281" phylum="Chordata" rank="species" species="livia">
<emphasis id="07FD6F46A411FFCDFEFA50D5FE11BC95" box="[300,460,151,177]" italics="true" pageId="6" pageNumber="281">Columba livia</emphasis>
Gmelin
</taxonomicName>
); probably the larvae of
<taxonomicName id="F289C8D7A411FFCDFCED50D5FBE5BC95" authority="Wagner &amp; Rada, 2020" authorityName="Wagner &amp; Rada" authorityYear="2020" box="[827,1080,151,177]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="glamocensis" status="sp. nov.">
<emphasis id="07FD6F46A411FFCDFCED50D5FBE5BC95" box="[827,1080,151,177]" italics="true" pageId="6" pageNumber="281">Psychoda glamocensis</emphasis>
</taxonomicName>
<emphasis id="07FD6F46A411FFCDFB9650D5FB44BC95" bold="true" box="[1088,1177,151,177]" pageId="6" pageNumber="281">
<taxonomicNameLabel id="1CCED23DA411FFCDFB9650D5FB44BC95" box="[1088,1177,151,177]" pageId="6" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
feed on their feces and remains; bats that are usually further inhabitants of caves may provide further organic material.
</paragraph>
<paragraph id="3536B354A411FFCDFF11509DFE78BD89" blockId="6.[151,1436,151,429]" pageId="6" pageNumber="281">
The additional male from
<collectingCountry id="4D9EF3C4A411FFCDFE24509DFD99BCDD" box="[498,580,223,249]" name="Austria" pageId="6" pageNumber="281">Austria</collectingCountry>
was found as undetermined but close to
<taxonomicName id="F289C8D7A411FFCDFBCB509DFB4DBCDD" authorityName="Vaillant" authorityYear="1973" box="[1053,1168,223,249]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="07FD6F46A411FFCDFBCB509DFB4DBCDD" box="[1053,1168,223,249]" italics="true" pageId="6" pageNumber="281">P. alticola</emphasis>
</taxonomicName>
in the collection of the senior author; identification was finally easy based on the reduced eye bridge and male genitalia. The locality Schreierbach is a permanent cold stream (probably a very large rheocrene) with constant water temperature of about 8°C all year round. Here
<taxonomicName id="F289C8D7A411FFCDFE625109FD86BD41" authorityName="Wagner &amp; Rada" authorityYear="2020" box="[436,603,331,357]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="glamocensis">
<emphasis id="07FD6F46A411FFCDFE625109FD86BD41" box="[436,603,331,357]" italics="true" pageId="6" pageNumber="281">P. glamocensis</emphasis>
</taxonomicName>
was collected together with specimens that are doubtless
<taxonomicName id="F289C8D7A411FFCDFB3F5109FA86BD41" authorityName="Vaillant" authorityYear="1973" box="[1257,1371,331,357]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="07FD6F46A411FFCDFB3F5109FA86BD41" box="[1257,1371,331,357]" italics="true" pageId="6" pageNumber="281">P. alticola</emphasis>
</taxonomicName>
; both species seem to prefer cold conditions that occur in the Alps at higher elevation, but as well in cold springs, and caves at lower altitudes.
</paragraph>
</subSubSection>
</treatment>
</document>
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