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<document id="246E8B4929F193F375C4A6BA0C1D50F0" ID-DOI="10.5281/zenodo.198822" ID-GBIF-Dataset="760a6ca0-02d5-493e-af11-be554ff45ff8" ID-ISSN="1175-5326" ID-Zenodo-Dep="198822" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460315517728" checkinUser="plazi" docAuthor="Morkel, Carsten" docDate="2010" docId="BB6D7E4FCD71383B44C15AF0FB337CAD" docLanguage="en" docName="zt02651p068.pdf" docOrigin="Zootaxa 2651" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Aradus serbicus Horvath 1888" docType="treatment" docVersion="7" lastPageNumber="68" masterDocId="47540637CD70383F44565F05FF9D7F6F" masterDocTitle="First records of Heterotoma merioptera (Scopoli, 1763) and Aradus serbicus (Horváth, 1888) (Heteroptera: Miridae et Aradidae) from Germany" masterLastPageNumber="68" masterPageNumber="64" pageNumber="65" updateTime="1698254891466" updateUser="plazi">
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<mods:title id="2A68635C17D4785E59EF2C85D437A163">First records of Heterotoma merioptera (Scopoli, 1763) and Aradus serbicus (Horváth, 1888) (Heteroptera: Miridae et Aradidae) from Germany</mods:title>
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<treatment id="BB6D7E4FCD71383B44C15AF0FB337CAD" ID-DOI="http://doi.org/10.5281/zenodo.6199969" ID-GBIF-Taxon="127802147" ID-Zenodo-Dep="6199969" LSID="urn:lsid:plazi:treatment:BB6D7E4FCD71383B44C15AF0FB337CAD" httpUri="http://treatment.plazi.org/id/BB6D7E4FCD71383B44C15AF0FB337CAD" lastPageId="4" lastPageNumber="68" pageId="1" pageNumber="65">
<subSubSection id="7BDE9CD2CD71383E44C15AF0FD2A7961" box="[151,695,1524,1551]" pageId="1" pageNumber="65" type="nomenclature">
<paragraph id="337BCF59CD71383E44C15AF0FD2A7961" blockId="1.[151,695,1524,1551]" box="[151,695,1524,1551]" pageId="1" pageNumber="65">
<heading id="68337835CD71383E44C15AF0FD2A7961" bold="true" box="[151,695,1524,1551]" fontSize="11" level="1" pageId="1" pageNumber="65" reason="1">
<taxonomicName id="F4C4B4DACD71383E44C15AF0FD2A7961" authority="Horvath, 1888" authorityName="Horvath" authorityYear="1888" box="[151,695,1524,1551]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD71383E44C15AF0FD2A7961" bold="true" box="[151,695,1524,1551]" pageId="1" pageNumber="65">
<emphasis id="01B0134BCD71383E44C15AF0FEC47960" bold="true" box="[151,345,1525,1551]" italics="true" pageId="1" pageNumber="65">Aradus serbicus</emphasis>
(Horváth, 1888) (Aradidae)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7BDE9CD2CD71383B44C15939FB337CAD" lastPageId="4" lastPageNumber="68" pageId="1" pageNumber="65" type="materials_examined">
<paragraph id="337BCF59CD71383E44C15939FCA17913" blockId="1.[151,1437,1596,2033]" pageId="1" pageNumber="65">
<emphasis id="01B0134BCD71383E44C15939FE8E7939" bold="true" box="[151,275,1596,1622]" pageId="1" pageNumber="65">Material:</emphasis>
<collectingCountry id="4BD38FC9CD71383E454A5938FE127938" box="[284,399,1597,1623]" name="Germany" pageId="1" pageNumber="65">Germany</collectingCountry>
, Hesse, Kellerwald-Edersee National Park,
<quantity id="F43C62BCCD71383E47FB5938FC677938" box="[941,1018,1597,1623]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="1" pageNumber="65" unit="m" value="425.0">425 m</quantity>
a.s.l., on
<taxonomicName id="F4C4B4DACD71383E40235938FAA57939" box="[1141,1336,1597,1622]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="65" phylum="Tracheophyta" rank="species" species="sylvatica">
<emphasis id="01B0134BCD71383E40235938FAA57939" box="[1141,1336,1597,1622]" italics="true" pageId="1" pageNumber="65">Fagus sylvatica</emphasis>
</taxonomicName>
, 27.v. 24.
<date id="477AE999CD71383E44EC5967FEAF7913" box="[186,306,1634,1660]" pageId="1" pageNumber="65" value="2008-06-01">vi.2008, 1</date>
Ψ, U. Schaffrath leg., C. Morkel det. (
<figureCitation id="ABFFD3DCCD71383E46BB5967FCAF7913" box="[749,818,1634,1660]" captionStart="FIGURE 2" captionStartId="2.[151,260,1939,1963]" captionTargetBox="[227,1354,206,1902]" captionTargetId="figure@2.[217,1365,194,1917]" captionTargetPageId="2" captionText="FIGURE 2. Aradus serbicus. Compared to closely related species of the Aradus depressus - group, among other characters, females of Aradus serbicus show regularly rounded paratergites VIII similar to those of the males. Photo: G. Strauß." httpUri="https://zenodo.org/record/198824/files/figure.png" pageId="1" pageNumber="65">Fig. 2</figureCitation>
)
</paragraph>
<paragraph id="337BCF59CD71383E44905982FDDA7858" blockId="1.[151,1437,1596,2033]" pageId="1" pageNumber="65">
The distribution area of the extremely rare
<taxonomicName id="F4C4B4DACD71383E4687598DFCCA79CE" box="[721,855,1672,1697]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD71383E4687598DFCCA79CE" box="[721,855,1672,1697]" italics="true" pageId="1" pageNumber="65">A. serbicus</emphasis>
</taxonomicName>
known so far, ranges from southern
<collectingCountry id="4BD38FC9CD71383E41415982FAFA79CE" box="[1303,1383,1671,1697]" name="France" pageId="1" pageNumber="65">France</collectingCountry>
and
<collectingCountry id="4BD38FC9CD71383E44C159A8FF4D79A8" box="[151,208,1709,1735]" name="Italy" pageId="1" pageNumber="65">Italy</collectingCountry>
to
<collectingCountry id="4BD38FC9CD71383E44AE59A8FECF79A8" box="[248,338,1709,1735]" name="Greece" pageId="1" pageNumber="65">Greece</collectingCountry>
, including
<collectingCountry id="4BD38FC9CD71383E458D59A8FDA879A8" box="[475,565,1709,1735]" name="Austria" pageId="1" pageNumber="65">Austria</collectingCountry>
and the Balkan peninsula with the countries
<collectingCountry id="4BD38FC9CD71383E403559A8FB5D79A8" box="[1123,1216,1709,1735]" name="Croatia" pageId="1" pageNumber="65">Croatia</collectingCountry>
,
<collectingCountry id="4BD38FC9CD71383E409B59A8FA8379A8" box="[1229,1310,1709,1735]" name="Serbia" pageId="1" pageNumber="65">Serbia</collectingCountry>
,
<collectingCountry id="4BD38FC9CD71383E417A59A8FA0579A8" box="[1324,1432,1709,1735]" name="Hungary" pageId="1" pageNumber="65">Hungary</collectingCountry>
,
<collectingCountry id="4BD38FC9CD71383E44C159D7FE947983" box="[151,265,1746,1772]" name="Bulgaria" pageId="1" pageNumber="65">Bulgaria</collectingCountry>
, and
<collectingCountry id="4BD38FC9CD71383E451859D7FE467983" box="[334,475,1746,1772]" name="Macedonia" pageId="1" pageNumber="65">Macedonia</collectingCountry>
(
<bibRefCitation id="5755B2A8CD71383E45BB59D7FC8E7983" author="Heiss" box="[493,787,1746,1772]" pageId="1" pageNumber="65" refString="Heiss, E. &amp; Pericart (2007) Hemipteres Aradidae Piesmatidae et Dipsocoromorphes euro-mediterraneens. Faune de France 91, 509 pp. + 8 plates." type="book" year="2007">Heiss &amp; Péricart 2007</bibRefCitation>
). Among the associated trees, chiefly
<taxonomicName id="F4C4B4DACD71383E415759D7FACD7984" box="[1281,1360,1746,1771]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="65" phylum="Tracheophyta" rank="genus">
<emphasis id="01B0134BCD71383E415759D7FACD7984" box="[1281,1360,1746,1771]" italics="true" pageId="1" pageNumber="65">Fagus</emphasis>
</taxonomicName>
sp. is documented; however, also
<taxonomicName id="F4C4B4DACD71383E458859FDFDDC787E" box="[478,577,1784,1809]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="65" phylum="Tracheophyta" rank="genus">
<emphasis id="01B0134BCD71383E458859FDFDDC787E" box="[478,577,1784,1809]" italics="true" pageId="1" pageNumber="65">Quercus</emphasis>
</taxonomicName>
sp. in combination with
<taxonomicName id="F4C4B4DACD71383E473259FDFC30787E" box="[868,941,1784,1809]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="65" phylum="Tracheophyta" rank="genus">
<emphasis id="01B0134BCD71383E473259FDFC30787E" box="[868,941,1784,1809]" italics="true" pageId="1" pageNumber="65">Fagus</emphasis>
</taxonomicName>
from records from
<collectingCountry id="4BD38FC9CD71383E40C559F2FB74787E" box="[1171,1257,1783,1809]" name="Greece" pageId="1" pageNumber="65">Greece</collectingCountry>
, are mentioned (
<bibRefCitation id="5755B2A8CD71383E44C95818FEBA7858" author="Heiss" box="[159,295,1821,1847]" pageId="1" pageNumber="65" refString="Heiss, E. (2006) New records of Aradidae from the Balkan countries (Heteroptera, Aradidae). Acta entomologica serbica, 11 (1 / 2), 1 - 10." type="journal article" year="2006">Heiss 2006</bibRefCitation>
,
<bibRefCitation id="5755B2A8CD71383E45645818FDA67858" author="Heiss" box="[306,571,1821,1847]" pageId="1" pageNumber="65" refString="Heiss, E. &amp; Pericart (2007) Hemipteres Aradidae Piesmatidae et Dipsocoromorphes euro-mediterraneens. Faune de France 91, 509 pp. + 8 plates." type="book" year="2007">Heiss &amp; Péricart 2007</bibRefCitation>
).
</paragraph>
<paragraph id="337BCF59CD71383B44935847FB737FB8" blockId="1.[151,1437,1596,2033]" lastBlockId="4.[151,1437,152,962]" lastPageId="4" lastPageNumber="68" pageId="1" pageNumber="65">
The current extralimital record of
<taxonomicName id="F4C4B4DACD71383E460B5847FD7F7834" box="[605,738,1858,1883]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD71383E460B5847FD7F7834" box="[605,738,1858,1883]" italics="true" pageId="1" pageNumber="65">A. serbicus</emphasis>
</taxonomicName>
was obtained from a pitfall trap exposed in a living beech tree (
<figureCitation id="ABFFD3DCCD71383E44815862FEBE78EE" box="[215,291,1895,1921]" captionStart="FIGURE 3" captionStartId="3.[151,256,1947,1971]" captionTargetBox="[204,1351,194,1915]" captionTargetId="figure@3.[204,1351,194,1915]" captionTargetPageId="3" captionText="FIGURE 3. Habitat of Aradus serbicus within Kellerwald-Edersee National Park: Old-stand Fagus sylvatica, infested by Oudemansiella mucida. Photo: C. Morkel." httpUri="https://zenodo.org/record/198825/files/figure.png" pageId="1" pageNumber="65">Fig. 3</figureCitation>
) split by lightning several decades ago, containing a large rot-hole as well as specific fungi infestation, including
<taxonomicName id="F4C4B4DACD71383E45C95888FCE178C8" authority="Schrad. ex Fr." authorityName="Schrad. ex Fr." box="[415,892,1933,1959]" class="Agaricomycetes" family="Physalacriaceae" genus="Oudemansiella" kingdom="Fungi" order="Agaricales" pageId="1" pageNumber="65" phylum="Basidiomycota" rank="species" species="mucida">
<emphasis id="01B0134BCD71383E45C95888FD2878C9" box="[415,693,1933,1958]" italics="true" pageId="1" pageNumber="65">Oudemansiella mucida</emphasis>
(Schrad. ex Fr.)
</taxonomicName>
, a species characteristic of old beech forests. The associated coleopterous fauna contained several species (Schaffrath, pers. comm. 2010) classified as primeval forest relict species according to
<bibRefCitation id="5755B2A8CD71383E46FB58D2FC05789E" box="[685,920,2007,2033]" pageId="1" pageNumber="65" refString="Muller, J., Bußler, H., Bense, U., Brustel, H., Flechtner, G., Fowles, A., Kahlen, M., Moller, G., Muhle, H., Schmidl, J. &amp; Zabransky, P. (2005) Urwald relict species - Saproxylic beetle indicating structural qualities and habitat tradition. waldoekologie online, 2, 106 - 113." type="journal article">
Müller
<emphasis id="01B0134BCD71383E475258DDFCA2789E" box="[772,831,2008,2033]" italics="true" pageId="1" pageNumber="65">et al.</emphasis>
(2005)
</bibRefCitation>
. The habitat itself, located on a steep slope, represents one of the small patches of virgin forest within Kellerwald-Edersee National Park showing a very long continuity concerning dead wood quality and quantity due to the impossibility of forestry.
</paragraph>
<caption id="67BB9FD1CD72383D44C15896FF7E7885" httpUri="https://zenodo.org/record/198824/files/figure.png" pageId="2" pageNumber="66" targetBox="[227,1354,206,1902]" targetPageId="2">
<paragraph id="337BCF59CD72383D44C15896FF7E7885" blockId="2.[151,1437,1939,2026]" pageId="2" pageNumber="66">
<emphasis id="01B0134BCD72383D44C15896FEBF78C4" bold="true" box="[151,290,1939,1963]" pageId="2" pageNumber="66">FIGURE 2.</emphasis>
<taxonomicName id="F4C4B4DACD72383D457C5896FE7978C5" box="[298,484,1939,1962]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD72383D457C5896FE7978C5" box="[298,484,1939,1962]" italics="true" pageId="2" pageNumber="66">Aradus serbicus</emphasis>
</taxonomicName>
. Compared to closely related species of the
<taxonomicName id="F4C4B4DACD72383D47B15896FB2C78C5" box="[999,1201,1939,1962]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="depressus">
<emphasis id="01B0134BCD72383D47B15896FB2C78C5" box="[999,1201,1939,1962]" italics="true" pageId="2" pageNumber="66">Aradus depressus</emphasis>
</taxonomicName>
-group, among other characters, females of
<taxonomicName id="F4C4B4DACD72383D45DE58B6FDA878A5" box="[392,565,1971,1994]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD72383D45DE58B6FDA878A5" box="[392,565,1971,1994]" italics="true" pageId="2" pageNumber="66">Aradus serbicus</emphasis>
</taxonomicName>
show regularly rounded paratergites VIII similar to those of the males. Photo: G. Strauß.
</paragraph>
</caption>
<caption id="67BB9FD1CD73383C44C1589EFDE078BD" httpUri="https://zenodo.org/record/198825/files/figure.png" pageId="3" pageNumber="67" targetBox="[204,1351,194,1915]" targetPageId="3">
<paragraph id="337BCF59CD73383C44C1589EFDE078BD" blockId="3.[151,1436,1947,2002]" pageId="3" pageNumber="67">
<emphasis id="01B0134BCD73383C44C1589EFE8778DC" bold="true" box="[151,282,1947,1971]" pageId="3" pageNumber="67">FIGURE 3.</emphasis>
Habitat of
<taxonomicName id="F4C4B4DACD73383C45C1589EFDDB78DD" box="[407,582,1947,1970]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD73383C45C1589EFDDB78DD" box="[407,582,1947,1970]" italics="true" pageId="3" pageNumber="67">Aradus serbicus</emphasis>
</taxonomicName>
within Kellerwald-Edersee National Park: Old-stand
<taxonomicName id="F4C4B4DACD73383C40D8589EFAA578DD" box="[1166,1336,1947,1970]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="67" phylum="Tracheophyta" rank="species" species="sylvatica">
<emphasis id="01B0134BCD73383C40D8589EFAA578DD" box="[1166,1336,1947,1970]" italics="true" pageId="3" pageNumber="67">Fagus sylvatica</emphasis>
</taxonomicName>
, infested by
<taxonomicName id="F4C4B4DACD73383C44EE58BEFE3278BD" box="[184,431,1979,2002]" class="Agaricomycetes" family="Physalacriaceae" genus="Oudemansiella" kingdom="Fungi" order="Agaricales" pageId="3" pageNumber="67" phylum="Basidiomycota" rank="species" species="mucida">
<emphasis id="01B0134BCD73383C44EE58BEFE3278BD" box="[184,431,1979,2002]" italics="true" pageId="3" pageNumber="67">Oudemansiella mucida</emphasis>
</taxonomicName>
. Photo: C. Morkel.
</paragraph>
</caption>
<paragraph id="337BCF59CD74383B44935FE7FB8C7EFD" blockId="4.[151,1437,152,962]" pageId="4" pageNumber="68">
The vast majority of palaearctic
<taxonomicName id="F4C4B4DACD74383B46175FE7FD317F93" box="[577,684,226,252]" class="Insecta" family="Aradidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="family">Aradidae</taxonomicName>
are known to be dendrobiont and mycetophagous, relying more or less specifically on wood- and tree-infesting fungi (
<bibRefCitation id="5755B2A8CD74383B47705E0DFBA57E4D" author="Heiss" box="[806,1080,264,290]" pageId="4" pageNumber="68" refString="Heiss, E. &amp; Pericart (2007) Hemipteres Aradidae Piesmatidae et Dipsocoromorphes euro-mediterraneens. Faune de France 91, 509 pp. + 8 plates." type="book" year="2007">Heiss &amp; Pericart 2007</bibRefCitation>
). For some of the most rarely found species within Central Europe, a temporal continuity of several centuries in the occurrence of dead wood structures of various decomposition stages (habitat tradition) necessary for the colonisation by particular fungi species (Goßner
<emphasis id="01B0134BCD74383B464E5E7DFDCE7EFE" box="[536,595,376,401]" italics="true" pageId="4" pageNumber="68">et al.</emphasis>
2007) is hypothesized to be essential.
</paragraph>
<paragraph id="337BCF59CD74383B44935E98FD2E7D48" blockId="4.[151,1437,152,962]" pageId="4" pageNumber="68">
Judging from the circumstances of finding of the recent record as well as published data (
<bibRefCitation id="5755B2A8CD74383B40B25E98FAF37ED8" author="Heiss" box="[1252,1390,413,439]" pageId="4" pageNumber="68" refString="Heiss, E. (2006) New records of Aradidae from the Balkan countries (Heteroptera, Aradidae). Acta entomologica serbica, 11 (1 / 2), 1 - 10." type="journal article" year="2006">Heiss 2006</bibRefCitation>
),
<taxonomicName id="F4C4B4DACD74383B41D55E98FF647EB4" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD74383B41D55E98FF647EB4" italics="true" pageId="4" pageNumber="68">A. serbicus</emphasis>
</taxonomicName>
is considered to feed on a specific range of wood-dwelling fungi limited to broad-leafed trees or even exclusively to
<taxonomicName id="F4C4B4DACD74383B45155EEDFE617D6E" box="[323,508,488,513]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="68" phylum="Tracheophyta" rank="species" species="sylvatica">
<emphasis id="01B0134BCD74383B45155EEDFE617D6E" box="[323,508,488,513]" italics="true" pageId="4" pageNumber="68">Fagus sylvatica</emphasis>
</taxonomicName>
. Moreover, the recent record strongly indicates a long habitat tradition to have been critical for the occurrence of
<taxonomicName id="F4C4B4DACD74383B467C5D08FD317D49" box="[554,684,525,550]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD74383B467C5D08FD317D49" box="[554,684,525,550]" italics="true" pageId="4" pageNumber="68">A. serbicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="337BCF59CD74383B44905D37FC747DD3" blockId="4.[151,1437,152,962]" pageId="4" pageNumber="68">
Thus, considering the extralimital documentation of
<taxonomicName id="F4C4B4DACD74383B473F5D37FC6B7D24" box="[873,1014,562,587]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD74383B473F5D37FC6B7D24" box="[873,1014,562,587]" italics="true" pageId="4" pageNumber="68">A. serbicus</emphasis>
</taxonomicName>
in combination with the habitat conditions described, and the only 17 localities recorded so far (with the correct determination of
<taxonomicName id="F4C4B4DACD74383B414F5D5DFA067D1E" box="[1305,1435,600,625]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD74383B414F5D5DFA067D1E" box="[1305,1435,600,625]" italics="true" pageId="4" pageNumber="68">A. serbicus</emphasis>
</taxonomicName>
for ten of them confirmed by
<bibRefCitation id="5755B2A8CD74383B45A05D78FC9C7DF8" author="Heiss" box="[502,769,637,663]" pageId="4" pageNumber="68" refString="Heiss, E. &amp; Pericart (2007) Hemipteres Aradidae Piesmatidae et Dipsocoromorphes euro-mediterraneens. Faune de France 91, 509 pp. + 8 plates." type="book" year="2007">Heiss &amp; Péricart 2007</bibRefCitation>
), the species is considered to represent a primeval forest relict in the closest sense (see
<bibRefCitation id="5755B2A8CD74383B45AE5DA7FD507DD3" author="Muller" box="[504,717,674,700]" pageId="4" pageNumber="68" refString="Muller, J., Bußler, H., Bense, U., Brustel, H., Flechtner, G., Fowles, A., Kahlen, M., Moller, G., Muhle, H., Schmidl, J. &amp; Zabransky, P. (2005) Urwald relict species - Saproxylic beetle indicating structural qualities and habitat tradition. waldoekologie online, 2, 106 - 113." type="journal article" year="2005">
Müller
<emphasis id="01B0134BCD74383B46195DA7FD177DD4" box="[591,650,674,699]" italics="true" pageId="4" pageNumber="68">et al.</emphasis>
2005
</bibRefCitation>
) within Central Europe.
</paragraph>
<paragraph id="337BCF59CD74383B44935DCDFE9B7C18" blockId="4.[151,1437,152,962]" pageId="4" pageNumber="68">
Records of the closely related
<taxonomicName id="F4C4B4DACD74383B466A5DCDFC2E7D8D" authority="Fieber, 1860" authorityName="Fieber" authorityYear="1860" box="[572,947,712,738]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="01B0134BCD74383B466A5DCDFC8C7D8E" box="[572,785,712,737]" italics="true" pageId="4" pageNumber="68">Aradus truncatus</emphasis>
Fieber, 1860
</taxonomicName>
of European distribution are confirmed mainly from southern Europe by
<bibRefCitation id="5755B2A8CD74383B46685DE8FCEC7C68" author="Heiss" box="[574,881,749,775]" pageId="4" pageNumber="68" refString="Heiss, E. &amp; Pericart (2007) Hemipteres Aradidae Piesmatidae et Dipsocoromorphes euro-mediterraneens. Faune de France 91, 509 pp. + 8 plates." type="book" year="2007">Heiss &amp; Péricart (2007)</bibRefCitation>
; those from remaining countries, especially
<collectingCountry id="4BD38FC9CD74383B44C15C17FE947C43" box="[151,265,786,812]" name="Germany" pageId="4" pageNumber="68">Germany</collectingCountry>
,
<collectingCountry id="4BD38FC9CD74383B45405C17FEF37C43" box="[278,366,786,812]" name="Poland" pageId="4" pageNumber="68">Poland</collectingCountry>
,
<collectingCountry id="4BD38FC9CD74383B452D5C17FE727C43" box="[379,495,786,812]" name="Denmark" pageId="4" pageNumber="68">Denmark</collectingCountry>
,
<collectingCountry id="4BD38FC9CD74383B45AB5C17FDFC7C43" box="[509,609,786,812]" name="Sweden" pageId="4" pageNumber="68">Sweden</collectingCountry>
, and
<collectingCountry id="4BD38FC9CD74383B46F75C17FC9F7C43" box="[673,770,786,812]" name="Finland" pageId="4" pageNumber="68">Finland</collectingCountry>
, should be verified in order to clarify if some of them represent
<taxonomicName id="F4C4B4DACD74383B455B5C3DFE097C3E" box="[269,404,824,849]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD74383B455B5C3DFE097C3E" box="[269,404,824,849]" italics="true" pageId="4" pageNumber="68">A. serbicus</emphasis>
</taxonomicName>
, and if the occurrence of
<taxonomicName id="F4C4B4DACD74383B46985C3DFCC97C3E" box="[718,852,824,849]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD74383B46985C3DFCC97C3E" box="[718,852,824,849]" italics="true" pageId="4" pageNumber="68">A. serbicus</emphasis>
</taxonomicName>
corresponds with the distribution area of
<taxonomicName id="F4C4B4DACD74383B41075C3DFF627C19" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="68" phylum="Tracheophyta" rank="species" species="sylvatica">
<emphasis id="01B0134BCD74383B41075C3DFF627C19" italics="true" pageId="4" pageNumber="68">Fagus sylvatica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="337BCF59CD74383B44935C87FB337CAD" blockId="4.[151,1437,152,962]" pageId="4" pageNumber="68">
Further investigations concerning the biology of
<taxonomicName id="F4C4B4DACD74383B47515C87FC177CF4" box="[775,906,898,923]" class="Insecta" family="Aradidae" genus="Aradus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="01B0134BCD74383B47515C87FC177CF4" box="[775,906,898,923]" italics="true" pageId="4" pageNumber="68">A. serbicus</emphasis>
</taxonomicName>
should be conducted, in order to support the hypothesis regarding the species´habitat preferences and to obtain data on its host fungi..
</paragraph>
</subSubSection>
</treatment>
</document>