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<document id="21BE11C9F407614DB41581FA2E1146B5" ID-DOI="10.5281/zenodo.210126" ID-GBIF-Dataset="8b9b0a46-ad8c-4ebe-8249-233a9e41d11a" ID-ISSN="1175-5326" ID-Zenodo-Dep="210126" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460492767072" checkinUser="plazi" docAuthor="Lezama, Antônio Queiroz, Triques, Mauro L. &amp; Santos, Patrícia S." docDate="2012" docId="BB5E1948FF8CFFC0FF621DBAAF9885B9" docLanguage="en" docName="zt03352p068.pdf" docOrigin="Zootaxa 3352" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Trichomycterus argos Lezama &amp; Triques &amp; Santos 2012, new species" docType="treatment" docVersion="8" lastPageNumber="65" masterDocId="47676130FF8EFFC5FFF51D2DAF4D827A" masterDocTitle="Trichomycterus argos (Teleostei: Siluriformes: Trichomycteridae), a new species from the Doce River Basin, Eastern Brazil" masterLastPageNumber="68" masterPageNumber="60" pageNumber="62" updateTime="1698310383722" updateUser="plazi">
<mods:mods id="D12D0E4D3F17A61BA1F6EB87526A479C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1730AB50DB28CF67EFEA1511C7E08B22">
<mods:title id="33F70A49006DD6D23157FA8A76791CD5">Trichomycterus argos (Teleostei: Siluriformes: Trichomycteridae), a new species from the Doce River Basin, Eastern Brazil</mods:title>
</mods:titleInfo>
<mods:name id="EA69E4FC3BAFE0A319CC3C9D89391917" type="personal">
<mods:role id="A8B148D9D5D2874B2AC5A2A3E1E840AE">
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</mods:role>
<mods:namePart id="EE1617C6C138D48A9C44202BFF9326AF">Lezama, Antônio Queiroz</mods:namePart>
</mods:name>
<mods:name id="69B604A83BCD6CA56C070FA2351C2C91" type="personal">
<mods:role id="118D843C3FAD33965424BECC18AE5B6A">
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<mods:namePart id="849530213E467D3DB04B5FF9275A2D82">Triques, Mauro L.</mods:namePart>
</mods:name>
<mods:name id="528D24E82888D3049993B16E3F0AD5FA" type="personal">
<mods:role id="A28273356989B53D0F0FC55FB6F2E450">
<mods:roleTerm id="BDD29526A25365C201106ADF4CF12C73">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="639340F25C3D2DD66EA8A867A6FF3536">Santos, Patrícia S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="183E980C9777769479E5D3984EADFD3F">text</mods:typeOfResource>
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<mods:title id="586021AD47AF5649F34A539A787ADF43">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="14D67EF3BE8E54EEBF1AC661A9919B26">
<mods:date id="D76731FA1F3ED4841F2DE2C362B0EA07">2012</mods:date>
<mods:detail id="990B0A16096B7A9511B875F0AE80105B" type="volume">
<mods:number id="839804B3C5BE8DEC2E85579B47D52334">3352</mods:number>
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<mods:start id="44548D11A6CA9A2802B99FF0D5D3BC50">60</mods:start>
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<mods:classification id="603AD96C21CD80DC03D177865178702C">journal article</mods:classification>
<mods:identifier id="75D1CD115346AEF654AB33B2DCF523C5" type="DOI">10.5281/zenodo.210126</mods:identifier>
<mods:identifier id="19869679610F7FF771F616A6D8590AB6" type="GBIF-Dataset">8b9b0a46-ad8c-4ebe-8249-233a9e41d11a</mods:identifier>
<mods:identifier id="14A1C21D8103F56D385A0F71D70C0F71" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5FC51F4E3823365F4D80F5C22D70B762" type="Zenodo-Dep">210126</mods:identifier>
</mods:mods>
<treatment id="BB5E1948FF8CFFC0FF621DBAAF9885B9" ID-DOI="http://doi.org/10.5281/zenodo.5662444" ID-GBIF-Taxon="119520156" ID-Zenodo-Dep="5662444" LSID="urn:lsid:plazi:treatment:BB5E1948FF8CFFC0FF621DBAAF9885B9" httpUri="http://treatment.plazi.org/id/BB5E1948FF8CFFC0FF621DBAAF9885B9" lastPageId="5" lastPageNumber="65" pageId="2" pageNumber="62">
<subSubSection id="7BEDFBD5FF8CFFC7FF621DBAAFA682A8" pageId="2" pageNumber="62" type="nomenclature">
<paragraph id="3348A85EFF8CFFC7FF621DBAAD7082CB" blockId="2.[151,573,151,210]" box="[151,573,151,177]" pageId="2" pageNumber="62">
<heading id="68001F32FF8CFFC7FF621DBAAD7082CB" bold="true" box="[151,573,151,177]" fontSize="11" level="1" pageId="2" pageNumber="62" reason="1">
<emphasis id="0183744CFF8CFFC7FF621DBAAD7082CB" bold="true" box="[151,573,151,177]" pageId="2" pageNumber="62">
<taxonomicName id="F4F7D3DDFF8CFFC7FF621DBAAED282CB" ID-CoL="58FF8" authority="Lezama &amp; Triques &amp; Santos, 2012" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[151,415,151,177]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="argos" status="sp. nov.">
<emphasis id="0183744CFF8CFFC7FF621DBAAED282CB" bold="true" box="[151,415,151,177]" italics="true" pageId="2" pageNumber="62">Trichomycterus argos</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="1AB0C937FF8CFFC7FE581DBAAD7082CB" box="[429,573,151,177]" pageId="2" pageNumber="62" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3348A85EFF8CFFC7FF621D97AFA682A8" blockId="2.[151,573,151,210]" box="[151,235,186,210]" pageId="2" pageNumber="62">
(
<figureCitation id="ABCCB4DBFF8CFFC7FF6A1D97AFAE82A8" box="[159,227,186,210]" captionStart="FIGURE 2" captionStartId="4.[151,250,1424,1447]" captionTargetBox="[151,1435,193,1402]" captionTargetId="figure@4.[151,1436,193,1403]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of Trichomycterus argos, DZUFMG 103, 95.4 mm SL, in dorsal (1), lateral (2), and ventral (3) views." httpUri="https://zenodo.org/record/210128/files/figure.png" pageId="2" pageNumber="62">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7BEDFBD5FF8CFFC7FF621C2EABF883D0" pageId="2" pageNumber="62" type="materials_examined">
<paragraph id="3348A85EFF8CFFC7FF621C2EAADB8318" blockId="2.[151,1437,258,1974]" pageId="2" pageNumber="62">
<emphasis id="0183744CFF8CFFC7FF621C2EAE498361" bold="true" box="[151,260,259,283]" pageId="2" pageNumber="62">
<typeStatus id="EC4C16FCFF8CFFC7FF621C2EAE498361" box="[151,260,259,283]" pageId="2" pageNumber="62" type="holotype">Holotype</typeStatus>
</emphasis>
. DZUFMG 103, 95.4mm SL;
<collectingCountry id="4BE0E8CEFF8CFFC7FD9D1C2FADFC8360" box="[616,689,258,282]" name="Brazil" pageId="2" pageNumber="62">Brazil</collectingCountry>
: Minas Gerais State: Araponga municipality, Parque Estadual da Serra do Brigadeiro, Córrego Serra Nova, headwaters of the Rio Casca, right bank tributary of Rio Doce at the limits of Rio Doce and Rio Paraíba do Sul basins,
<geoCoordinate id="56C3CE99FF8CFFC7FD501C67AC6C8318" box="[677,801,330,354]" direction="south" orientation="latitude" pageId="2" pageNumber="62" precision="15" value="-20.721945">20°43'19''S</geoCoordinate>
/
<geoCoordinate id="56C3CE99FF8CFFC7FCC71C67ACF68318" box="[818,955,330,354]" direction="west" orientation="longitude" pageId="2" pageNumber="62" precision="15" value="-42.47861">42°28'43''W</geoCoordinate>
,
<quantity id="F40F05BBFF8CFFC7FC331C67AB568318" box="[966,1051,330,354]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="62" unit="m" value="1400.0">1400 m</quantity>
asl; P. S. Santos,
<date id="47498E9EFF8CFFC7FB151C66AADF8318" box="[1248,1426,330,354]" pageId="2" pageNumber="62" value="2001-10-07">7 October 2001</date>
.
</paragraph>
<paragraph id="3348A85EFF8CFFC7FF321C42ABF883D0" blockId="2.[151,1437,258,1974]" pageId="2" pageNumber="62">
<emphasis id="0183744CFF8CFFC7FF321C42AE7283FD" bold="true" box="[199,319,367,391]" pageId="2" pageNumber="62">
<typeStatus id="EC4C16FCFF8CFFC7FF321C42AE7283FD" box="[199,319,367,391]" pageId="2" pageNumber="62" type="paratype">Paratypes</typeStatus>
</emphasis>
. DZUFMG 0 58, 0 1 ex., 114.7 mm SL; DZUFMG 0 59, 14 ex., 56–116.8 mm SL; DZUFMG 0 67, 1 ex.,
<quantity id="F40F05BBFF8CFFC7FF331CBFAE6E83D0" box="[198,291,402,426]" metricMagnitude="-1" metricUnit="m" metricValue="1.01" pageId="2" pageNumber="62" unit="mm" value="101.0">101 mm</quantity>
SL, C&amp;S;
<collectionCode id="55E6309BFF8CFFC7FE551CBFAEB083D0" box="[416,509,402,426]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="62">MZUSP</collectionCode>
106274, 3 ex., 57.4–91.8 mm SL; all collected with
<typeStatus id="EC4C16FCFF8CFFC7FBBE1CBFABFC83D0" box="[1099,1201,402,426]" pageId="2" pageNumber="62" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="7BEDFBD5FF8CFFC7FF321C9BADD2847C" pageId="2" pageNumber="62" type="diagnosis">
<paragraph id="3348A85EFF8CFFC7FF321C9BADD2847C" blockId="2.[151,1437,258,1974]" pageId="2" pageNumber="62">
<emphasis id="0183744CFF8CFFC7FF321C9BAE0C83B5" bold="true" box="[199,321,438,463]" pageId="2" pageNumber="62">Diagnosis.</emphasis>
<taxonomicName id="F4F7D3DDFF8CFFC7FEB21C9BAD7683B5" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[327,571,438,463]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8CFFC7FEB21C9BAD7683B5" box="[327,571,438,463]" italics="true" pageId="2" pageNumber="62">Trichomycterus argos</emphasis>
</taxonomicName>
differs from its congeners, excluding
<taxonomicName id="F4F7D3DDFF8CFFC7FC171C95AA4783B4" authority="(Eigenmann)" authorityName="Eigenmann" baseAuthorityName="Eigenmann" baseAuthorityYear="1917" box="[994,1290,438,463]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="alternatus">
<emphasis id="0183744CFF8CFFC7FC171C95AB2383B4" box="[994,1134,438,463]" italics="true" pageId="2" pageNumber="62">T. alternatus</emphasis>
(Eigenmann)
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FAE11C95AE408388" authority="Lutken" authorityName="Lutken" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="0183744CFF8CFFC7FAE11C95AFF88389" italics="true" pageId="2" pageNumber="62">T. brasiliensis</emphasis>
Lütken
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FEEC1CF1AD058388" authority="Barbosa &amp; Costa" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[281,584,474,499]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="brunoi">
<emphasis id="0183744CFF8CFFC7FEEC1CF1AE328389" box="[281,383,475,499]" italics="true" pageId="2" pageNumber="62">T. brunoi</emphasis>
Barbosa &amp; Costa
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FDA61CF1ACE78388" authority="(Miranda Ribeiro)" authorityName="Miranda Ribeiro" baseAuthorityName="Miranda Ribeiro" baseAuthorityYear="1949" box="[595,938,474,499]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="candidus">
<emphasis id="0183744CFF8CFFC7FDA61CF1AD998389" box="[595,724,475,499]" italics="true" pageId="2" pageNumber="62">T. candidus</emphasis>
(Miranda Ribeiro)
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FC401CF1ABB78388" authority="Barbosa &amp; Costa" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[949,1274,474,499]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="claudiae">
<emphasis id="0183744CFF8CFFC7FC401CF1AB7F8389" box="[949,1074,475,499]" italics="true" pageId="2" pageNumber="62">T. claudiae</emphasis>
Barbosa &amp; Costa
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FAF01CF1AE10806C" authority="Barbosa &amp; Costa" authorityName="Barbosa &amp; Costa" authorityYear="2010" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="0183744CFF8CFFC7FAF01CF1AAD18389" box="[1285,1436,475,499]" italics="true" pageId="2" pageNumber="62">T. fuliginosus</emphasis>
Barbosa &amp; Costa
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FE9F1F2DAC68806C" authority="Barbosa &amp; Costa" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[362,805,510,535]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="macrotrichopterus">
<emphasis id="0183744CFF8CFFC7FE9F1F2DAD1B806C" box="[362,598,510,535]" italics="true" pageId="2" pageNumber="62">T. macrotrichopterus</emphasis>
Barbosa &amp; Costa
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FCC71F2DABF3806C" authority="Bockmann &amp; Sazima" authorityName="Bockmann &amp; Sazima" authorityYear="2004" box="[818,1214,510,535]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="maracaya">
<emphasis id="0183744CFF8CFFC7FCC71F2DACF0806D" box="[818,957,512,535]" italics="true" pageId="2" pageNumber="62">T. maracaya</emphasis>
Bockmann &amp; Sazima
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FB3E1F2DAE788040" authority="Barbosa &amp; Costa" authorityName="Barbosa &amp; Costa" authorityYear="2010" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="mariamole">
<emphasis id="0183744CFF8CFFC7FB3E1F2DAA2F806C" box="[1227,1378,510,535]" italics="true" pageId="2" pageNumber="62">T. mariamole</emphasis>
Barbosa &amp; Costa
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FEB71F09AD5F8040" authority="Costa" authorityName="Costa" authorityYear="1992" box="[322,530,546,571]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="mimonha">
<emphasis id="0183744CFF8CFFC7FEB71F09AE8A8041" box="[322,455,547,571]" italics="true" pageId="2" pageNumber="62">T. mimonha</emphasis>
Costa
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FDD51F09AC178040" authority="Valenciennes" authorityName="Valenciennes" box="[544,858,546,571]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="mirissumba">
<emphasis id="0183744CFF8CFFC7FDD51F09AD8D8041" box="[544,704,547,571]" italics="true" pageId="2" pageNumber="62">T. mirissumba</emphasis>
Valenciennes
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FC981F09ABA18040" authority="Barbosa &amp; Costa" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[877,1260,546,571]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="novalimensis">
<emphasis id="0183744CFF8CFFC7FC981F09AB518041" box="[877,1052,547,571]" italics="true" pageId="2" pageNumber="62">T. novalimensis</emphasis>
Barbosa &amp; Costa
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FB0C1F09AE158024" authority="Alencar &amp; Costa" authorityName="Alencar &amp; Costa" authorityYear="2004" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="pantherinus">
<emphasis id="0183744CFF8CFFC7FB0C1F09AAD18041" box="[1273,1436,547,571]" italics="true" pageId="2" pageNumber="62">T. pantherinus</emphasis>
Alencar &amp; Costa
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FE911F65ADD38024" authority="Barbosa &amp; Costa" authorityName="Barbosa &amp; Costa" authorityYear="2003" box="[356,670,582,607]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="potschi">
<emphasis id="0183744CFF8CFFC7FE911F65AE3C8025" box="[356,369,584,607]" italics="true" pageId="2" pageNumber="62">T</emphasis>
.
<emphasis id="0183744CFF8CFFC7FE741F6BAE9F8024" box="[385,466,582,606]" italics="true" pageId="2" pageNumber="62">potschi</emphasis>
Barbosa &amp; Costa
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FD5F1F65AB558024" authority="Barbosa &amp; Costa" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[682,1048,582,607]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="rubiginosus">
<emphasis id="0183744CFF8CFFC7FD5F1F65AC068024" box="[682,843,582,607]" italics="true" pageId="2" pageNumber="62">T. rubiginosus</emphasis>
Barbosa &amp; Costa
</taxonomicName>
, and
<taxonomicName id="F4F7D3DDFF8CFFC7FBA11F65AAD18024" authority="(Eigenmann)" authorityName="Eigenmann" baseAuthorityName="Eigenmann" baseAuthorityYear="1917" box="[1108,1436,582,607]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="vermiculatus">
<emphasis id="0183744CFF8CFFC7FBA11F65AA4F8024" box="[1108,1282,582,607]" italics="true" pageId="2" pageNumber="62">T. vermiculatus</emphasis>
(Eigenmann)
</taxonomicName>
by the presence of six branched rays in the pectoral fin, and by having the first pectoral-fin ray prolonged as a filament.
<taxonomicName id="F4F7D3DDFF8CFFC7FF141FA3AE9780DD" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[225,474,654,679]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8CFFC7FF141FA3AE9780DD" box="[225,474,654,679]" italics="true" pageId="2" pageNumber="62">Trichomycterus argos</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="F4F7D3DDFF8CFFC7FD101FBDACCB80DC" box="[741,902,654,679]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="0183744CFF8CFFC7FD101FBDACCB80DC" box="[741,902,654,679]" italics="true" pageId="2" pageNumber="62">T. brasiliensis</emphasis>
</taxonomicName>
by the absence of series of spots, as well as absence of confluent spots forming elongated marks or vermiculations (vs. presence of series of large [three times eye-diameter or more] spots dorsolaterally, laterally and ventrolaterally on flank in
<taxonomicName id="F4F7D3DDFF8CFFC7FBC31FF5AB9F8094" box="[1078,1234,726,751]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="0183744CFF8CFFC7FBC31FF5AB9F8094" box="[1078,1234,726,751]" italics="true" pageId="2" pageNumber="62">T. brasiliensis</emphasis>
</taxonomicName>
, more clearly visible in small specimens), by a transverse and straight border between the parietosupraoccipital and frontal bones (
<figureCitation id="ABCCB4DBFF8CFFC7FF6A1E33AFAD814C" box="[159,224,798,822]" captionStart="FIGURE 3" captionStartId="5.[151,250,1151,1174]" captionTargetBox="[156,1426,220,1119]" captionTargetId="figure@5.[151,1436,193,1130]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal views of the cranium and Weberian complex of a) Trichomycterus argos (DZUFMG 0 67, 101 mm SL, paratype) and b) Trichomycterus brasiliensis (DZUFMG 0 64, 102, 1 mm SL). Abbreviations: AP, autopalatine; EP, epioccipital; FF, frontal fontanels; FR, frontal; IO, tendon-bone infraorbital; LE, lateral ethmoid; ME, mesethmoid; MX, maxilla; PM, premaxilla; PS, posttemporosupracleithrum; PT, pterotic; RF, foramen for the ramus lateralis accessorius facialis; SC, cephalic sensory canal; SO, tendon-bone supraorbital; SPP, sphenotic-prootic-pterosphenoid compound bone; SU, parietosupraoccipital; WC, Weberian capsule. Scale bar = 1 cm." httpUri="https://zenodo.org/record/210129/files/figure.png" pageId="2" pageNumber="62">Fig. 3</figureCitation>
a) (vs. oblique and irregular border, in
<taxonomicName id="F4F7D3DDFF8CFFC7FD641E32AC7D814C" box="[657,816,798,822]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="brasiliensis">T. brasiliensis</taxonomicName>
;
<figureCitation id="ABCCB4DBFF8CFFC7FCCE1E33AC36814C" box="[827,891,798,822]" captionStart="FIGURE 3" captionStartId="5.[151,250,1151,1174]" captionTargetBox="[156,1426,220,1119]" captionTargetId="figure@5.[151,1436,193,1130]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal views of the cranium and Weberian complex of a) Trichomycterus argos (DZUFMG 0 67, 101 mm SL, paratype) and b) Trichomycterus brasiliensis (DZUFMG 0 64, 102, 1 mm SL). Abbreviations: AP, autopalatine; EP, epioccipital; FF, frontal fontanels; FR, frontal; IO, tendon-bone infraorbital; LE, lateral ethmoid; ME, mesethmoid; MX, maxilla; PM, premaxilla; PS, posttemporosupracleithrum; PT, pterotic; RF, foramen for the ramus lateralis accessorius facialis; SC, cephalic sensory canal; SO, tendon-bone supraorbital; SPP, sphenotic-prootic-pterosphenoid compound bone; SU, parietosupraoccipital; WC, Weberian capsule. Scale bar = 1 cm." httpUri="https://zenodo.org/record/210129/files/figure.png" pageId="2" pageNumber="62">Fig. 3</figureCitation>
b) and by a large foramen for the
<emphasis id="0183744CFF8CFFC7FB071E0DAE398121" italics="true" pageId="2" pageNumber="62">ramus lateralis accessorius facialis</emphasis>
, visible in dorsal view, in the parietosupraoccipital bone (vs. foramen for the
<emphasis id="0183744CFF8CFFC7FB051E69AE3E8104" italics="true" pageId="2" pageNumber="62">ramus lateralis accessorius facialis</emphasis>
minute, at least four times smaller than in the new species, located on a ridge of the bone, and visible only in lateral view, in
<taxonomicName id="F4F7D3DDFF8CFFC7FE1A1EA1ADC081D9" box="[495,653,907,931]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="0183744CFF8CFFC7FE1A1EA1ADC081D9" box="[495,653,907,931]" italics="true" pageId="2" pageNumber="62">T. brasiliensis</emphasis>
</taxonomicName>
;
<figureCitation id="ABCCB4DBFF8CFFC7FD691EA7AD9381D8" box="[668,734,906,930]" captionStart="FIGURE 3" captionStartId="5.[151,250,1151,1174]" captionTargetBox="[156,1426,220,1119]" captionTargetId="figure@5.[151,1436,193,1130]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal views of the cranium and Weberian complex of a) Trichomycterus argos (DZUFMG 0 67, 101 mm SL, paratype) and b) Trichomycterus brasiliensis (DZUFMG 0 64, 102, 1 mm SL). Abbreviations: AP, autopalatine; EP, epioccipital; FF, frontal fontanels; FR, frontal; IO, tendon-bone infraorbital; LE, lateral ethmoid; ME, mesethmoid; MX, maxilla; PM, premaxilla; PS, posttemporosupracleithrum; PT, pterotic; RF, foramen for the ramus lateralis accessorius facialis; SC, cephalic sensory canal; SO, tendon-bone supraorbital; SPP, sphenotic-prootic-pterosphenoid compound bone; SU, parietosupraoccipital; WC, Weberian capsule. Scale bar = 1 cm." httpUri="https://zenodo.org/record/210129/files/figure.png" pageId="2" pageNumber="62">Fig. 3</figureCitation>
b).
<taxonomicName id="F4F7D3DDFF8CFFC7FCF11EA6ACB481D9" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[772,1017,907,931]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8CFFC7FCF11EA6ACB481D9" box="[772,1017,907,931]" italics="true" pageId="2" pageNumber="62">Trichomycterus argos</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="F4F7D3DDFF8CFFC7FB021EA1AA3581D9" baseAuthorityName="Miranda Ribeiro" baseAuthorityYear="1949" box="[1271,1400,907,931]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="candidus">
<emphasis id="0183744CFF8CFFC7FB021EA1AA3581D9" box="[1271,1400,907,931]" italics="true" pageId="2" pageNumber="62">T. candidus</emphasis>
</taxonomicName>
by the presence of pelvic fins (vs. absence of pelvic fins, in
<taxonomicName id="F4F7D3DDFF8CFFC7FCFE1E9DACC681BC" baseAuthorityName="Miranda Ribeiro" baseAuthorityYear="1949" box="[779,907,942,967]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="candidus">
<emphasis id="0183744CFF8CFFC7FCFE1E9DACC681BC" box="[779,907,942,967]" italics="true" pageId="2" pageNumber="62">T. candidus</emphasis>
</taxonomicName>
).
<taxonomicName id="F4F7D3DDFF8CFFC7FC571E83ABD881BD" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[930,1173,942,967]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8CFFC7FC571E83ABD881BD" box="[930,1173,942,967]" italics="true" pageId="2" pageNumber="62">Trichomycterus argos</emphasis>
</taxonomicName>
differs from
<taxonomicName id="F4F7D3DDFF8CFFC7FAD21E9DAFF58191" baseAuthorityName="Eigenmann" baseAuthorityYear="1917" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="alternatus">
<emphasis id="0183744CFF8CFFC7FAD21E9DAFF58191" italics="true" pageId="2" pageNumber="62">T. alternatus</emphasis>
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FF321EF9AE1F8191" authorityName="Bockmann &amp; Sazima" authorityYear="2004" box="[199,338,980,1003]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="maracaya">
<emphasis id="0183744CFF8CFFC7FF321EF9AE1F8191" box="[199,338,980,1003]" italics="true" pageId="2" pageNumber="62">T. maracaya</emphasis>
</taxonomicName>
and
<taxonomicName id="F4F7D3DDFF8CFFC7FE781EF9AD7D8191" authorityName="Alencar &amp; Costa" authorityYear="2004" box="[397,560,979,1003]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="pantherinus">
<emphasis id="0183744CFF8CFFC7FE781EF9AD7D8191" box="[397,560,979,1003]" italics="true" pageId="2" pageNumber="62">T. pantherinus</emphasis>
</taxonomicName>
by having superficial and deeper layers of small spots that seldom reach eye diameter in size (vs. consistently different color patterns in these species).
<taxonomicName id="F4F7D3DDFF8CFFC7FC001EDBABA18675" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[1013,1260,1014,1039]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8CFFC7FC001EDBABA18675" box="[1013,1260,1014,1039]" italics="true" pageId="2" pageNumber="62">Trichomycterus argos</emphasis>
</taxonomicName>
differs from
<taxonomicName id="F4F7D3DDFF8CFFC7FA721ED5AFB28649" authorityName="Costa" authorityYear="1992" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="mimonha">
<emphasis id="0183744CFF8CFFC7FA721ED5AFB28649" italics="true" pageId="2" pageNumber="62">T. mimonha</emphasis>
</taxonomicName>
by the shorter head width (79.1–87.6% HL vs. 93.8– 98.9 % HL, in
<taxonomicName id="F4F7D3DDFF8CFFC7FC0F1931AB308649" authorityName="Costa" authorityYear="1992" box="[1018,1149,1051,1075]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="mimonha">
<emphasis id="0183744CFF8CFFC7FC0F1931AB308649" box="[1018,1149,1051,1075]" italics="true" pageId="2" pageNumber="62">T. mimonha</emphasis>
</taxonomicName>
), and by having the tip of nasal barbels extending to posterior border of opercular plate of odontodes (vs. not reaching origin odontodes of interopercular plate, in
<taxonomicName id="F4F7D3DDFF8CFFC7FE691949AD6D8601" authorityName="Costa" authorityYear="1992" box="[412,544,1123,1147]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="mimonha">
<emphasis id="0183744CFF8CFFC7FE691949AD6D8601" box="[412,544,1123,1147]" italics="true" pageId="2" pageNumber="62">T. mimonha</emphasis>
</taxonomicName>
).
<taxonomicName id="F4F7D3DDFF8CFFC7FDC2194EAC618601" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[567,812,1123,1147]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8CFFC7FDC2194EAC618601" box="[567,812,1123,1147]" italics="true" pageId="2" pageNumber="62">Trichomycterus argos</emphasis>
</taxonomicName>
differs from
<taxonomicName id="F4F7D3DDFF8CFFC7FC351949AB138601" box="[960,1118,1123,1147]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="mirissumba">
<emphasis id="0183744CFF8CFFC7FC351949AB138601" box="[960,1118,1123,1147]" italics="true" pageId="2" pageNumber="62">T. mirissumba</emphasis>
</taxonomicName>
by the absence of stripes on the dorsum (vs. two dorsal stripes, from the nape towards dorsal-fin origin, in
<taxonomicName id="F4F7D3DDFF8CFFC7FBD119A5AB8B86E4" box="[1060,1222,1158,1183]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="mirissumba">
<emphasis id="0183744CFF8CFFC7FBD119A5AB8B86E4" box="[1060,1222,1158,1183]" italics="true" pageId="2" pageNumber="62">T. mirissumba</emphasis>
</taxonomicName>
) as well as by the absence of confluence of dark spots bellow horizontal midline.
<taxonomicName id="F4F7D3DDFF8CFFC7FC911986AB1486B9" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[868,1113,1195,1219]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8CFFC7FC911986AB1486B9" box="[868,1113,1195,1219]" italics="true" pageId="2" pageNumber="62">Trichomycterus argos</emphasis>
</taxonomicName>
differs from
<taxonomicName id="F4F7D3DDFF8CFFC7FB1A1981AAD186B9" baseAuthorityName="Eigenmann" baseAuthorityYear="1917" box="[1263,1436,1195,1219]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="vermiculatus">
<emphasis id="0183744CFF8CFFC7FB1A1981AAD186B9" box="[1263,1436,1195,1219]" italics="true" pageId="2" pageNumber="62">T. vermiculatus</emphasis>
</taxonomicName>
by the absence of vermiculations, formed by fused spots on the body, and by having pelvic-fin origin placed one or two ocular diameters anterior to dorsal-fin origin (vs. pelvic-fin origin in a vertical through dorsal-fin origin, in
<taxonomicName id="F4F7D3DDFF8CFFC7FA7219D9AE658754" baseAuthorityName="Eigenmann" baseAuthorityYear="1917" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="vermiculatus">
<emphasis id="0183744CFF8CFFC7FA7219D9AE658754" italics="true" pageId="2" pageNumber="62">T. vermiculatus</emphasis>
</taxonomicName>
).
<taxonomicName id="F4F7D3DDFF8CFFC7FEB5183BAD788755" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[320,565,1302,1327]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8CFFC7FEB5183BAD788755" box="[320,565,1302,1327]" italics="true" pageId="2" pageNumber="62">Trichomycterus argos</emphasis>
</taxonomicName>
differs from
<taxonomicName id="F4F7D3DDFF8CFFC7FD391835AC758754" authorityName="Barbosa &amp; Costa" authorityYear="2003" box="[716,824,1302,1327]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="potschi">
<emphasis id="0183744CFF8CFFC7FD391835AC758754" box="[716,824,1302,1327]" italics="true" pageId="2" pageNumber="62">T. potschi</emphasis>
</taxonomicName>
by the absence of spots fused as stripes on the lateral region of the body (vs. one or more stripes, in
<taxonomicName id="F4F7D3DDFF8CFFC7FD581811AC548729" authorityName="Barbosa &amp; Costa" authorityYear="2003" box="[685,793,1339,1363]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="potschi">
<emphasis id="0183744CFF8CFFC7FD581811AC548729" box="[685,793,1339,1363]" italics="true" pageId="2" pageNumber="62">T. potschi</emphasis>
</taxonomicName>
).
<taxonomicName id="F4F7D3DDFF8CFFC7FCC11816AB678729" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[820,1066,1339,1363]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8CFFC7FCC11816AB678729" box="[820,1066,1339,1363]" italics="true" pageId="2" pageNumber="62">Trichomycterus argos</emphasis>
</taxonomicName>
differes from
<taxonomicName id="F4F7D3DDFF8CFFC7FB3B1811AA7B8729" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[1230,1334,1339,1363]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="brunoi">
<emphasis id="0183744CFF8CFFC7FB3B1811AA7B8729" box="[1230,1334,1339,1363]" italics="true" pageId="2" pageNumber="62">T. brunoi</emphasis>
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FAB01811AF8A870C" authorityName="Barbosa &amp; Costa" authorityYear="2010" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="claudiae">
<emphasis id="0183744CFF8CFFC7FAB01811AF8A870C" italics="true" pageId="2" pageNumber="62">T. claudiae</emphasis>
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FF23184DAE23870C" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[214,366,1374,1399]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="0183744CFF8CFFC7FF23184DAE23870C" box="[214,366,1374,1399]" italics="true" pageId="2" pageNumber="62">T. fuliginosus</emphasis>
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FE89184DAD5F870C" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[380,530,1374,1399]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="mariamole">
<emphasis id="0183744CFF8CFFC7FE89184DAD5F870C" box="[380,530,1374,1399]" italics="true" pageId="2" pageNumber="62">T. mariamole</emphasis>
</taxonomicName>
,
<taxonomicName id="F4F7D3DDFF8CFFC7FDD4184DAD9D870C" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[545,720,1374,1399]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="novalimensis">
<emphasis id="0183744CFF8CFFC7FDD4184DAD9D870C" box="[545,720,1374,1399]" italics="true" pageId="2" pageNumber="62">T. novalimensis</emphasis>
</taxonomicName>
and
<taxonomicName id="F4F7D3DDFF8CFFC7FCFD184DACB9870C" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[776,1012,1374,1399]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="macrotrichopterus">
<emphasis id="0183744CFF8CFFC7FCFD184DACB9870C" box="[776,1012,1374,1399]" italics="true" pageId="2" pageNumber="62">T. macrotrichopterus</emphasis>
</taxonomicName>
by the absence of stripes, vermiculations or reticulations, present in these species (
<bibRefCitation id="5766D5AFFF8CFFC7FD3018AFAC9687E0" author="Barbosa" box="[709,987,1410,1434]" pageId="2" pageNumber="62" refString="Barbosa, M. A. &amp; Costa, W. J. E. M. (2010) Seven new species of the catfish genus Trichomycterus (Teleostei: Siluriformes: Trichomycteridae) from Southeastern Brazil and redescription of T. brasiliensis. Ichthyological Exploration of Freshwaters, 21, 97 - 122." type="journal article" year="2010">Barbosa &amp; Costa, 2010</bibRefCitation>
).
<taxonomicName id="F4F7D3DDFF8CFFC7FC0418AEABA587E1" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[1009,1256,1411,1435]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8CFFC7FC0418AEABA587E1" box="[1009,1256,1411,1435]" italics="true" pageId="2" pageNumber="62">Trichomycterus argos</emphasis>
</taxonomicName>
differs from
<taxonomicName id="F4F7D3DDFF8CFFC7FA7218A9AE5687C4" authorityName="Barbosa &amp; Costa" authorityYear="2010" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="rubiginosus">
<emphasis id="0183744CFF8CFFC7FA7218A9AE5687C4" italics="true" pageId="2" pageNumber="62">T. rubiginosus</emphasis>
</taxonomicName>
by a wider head (head width 79.1–87.6% of head length, vs. 72.2–78.9% in
<taxonomicName id="F4F7D3DDFF8CFFC7FB831885AA5B87C4" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[1142,1302,1446,1471]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="rubiginosus">
<emphasis id="0183744CFF8CFFC7FB831885AA5B87C4" box="[1142,1302,1446,1471]" italics="true" pageId="2" pageNumber="62">T. rubiginosus</emphasis>
</taxonomicName>
), larger eye (eye diameter 10.5–17.5% of head length, vs. 7.3–9.2 in
<taxonomicName id="F4F7D3DDFF8CFFC7FCD618E1AC898799" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[803,964,1483,1507]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="rubiginosus">
<emphasis id="0183744CFF8CFFC7FCD618E1AC898799" box="[803,964,1483,1507]" italics="true" pageId="2" pageNumber="62">T. rubiginosus</emphasis>
</taxonomicName>
) and smaller anal-fin base (7.2–8.8 % of standard length, vs. 9.0–9.9 in
<taxonomicName id="F4F7D3DDFF8CFFC7FE1A18DDADC2847C" authorityName="Barbosa &amp; Costa" authorityYear="2010" box="[495,655,1518,1543]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="62" phylum="Chordata" rank="species" species="rubiginosus">
<emphasis id="0183744CFF8CFFC7FE1A18DDADC2847C" box="[495,655,1518,1543]" italics="true" pageId="2" pageNumber="62">T. rubiginosus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="7BEDFBD5FF8CFFC1FF321B3FAE5785A6" lastPageId="4" lastPageNumber="64" pageId="2" pageNumber="62" type="description">
<paragraph id="3348A85EFF8CFFC7FF321B3FAA608434" blockId="2.[151,1437,258,1974]" pageId="2" pageNumber="62">
<emphasis id="0183744CFF8CFFC7FF321B3FAE148451" bold="true" box="[199,345,1554,1579]" pageId="2" pageNumber="62">Description.</emphasis>
Morphometric data of
<typeStatus id="EC4C16FCFF8CFFC7FD901B3FAD8A8450" box="[613,711,1554,1578]" pageId="2" pageNumber="62" type="holotype">holotype</typeStatus>
and
<typeStatus id="EC4C16FCFF8CFFC7FCF71B3EAC208451" box="[770,877,1555,1579]" pageId="2" pageNumber="62" type="paratype">paratypes</typeStatus>
are given in
<tableCitation id="7E759DE5FF8CFFC7FBFC1B3FAB128450" box="[1033,1119,1554,1578]" captionStart="TABLE 1" captionStartId="3.[151,235,152,173]" captionTargetBox="[159,1225,200,1169]" captionText="TABLE 1. Morphometric data for holotype and paratypes (n=19) of Trichomycterus argos, new species." httpUri="http://table.plazi.org/id/6788F8D6FF8DFFC6FF621DB5ABF482D4" pageId="2" pageNumber="62" tableUuid="6788F8D6FF8DFFC6FF621DB5ABF482D4">Table 1</tableCitation>
. Body elongate, cylindrical near head and gradually becoming more compressed towards tail. Caudal peduncle strongly compressed.
</paragraph>
<paragraph id="3348A85EFF8CFFC7FF321B77AD6285CC" blockId="2.[151,1437,258,1974]" pageId="2" pageNumber="62">Head wide and depressed. In dorsal view, small specimens with head much wider posteriorly than anteriorly, nearly triangle-shaped; large specimens with rounded head anteriorly. Smaller specimens with straight dorsal head profile, inclined upward from anterior nostril to supraoccipital posterior border; larger specimens with dorsal head profile slightly convex anteriorly, almost horizontal, from eye to supraorbital posterior border; dorsal body profile horizontal or slightly convex from supraoccipital posterior border to dorsal-fin origin, which is somewhat elevated. Dorsal-fin base profile straight and inclined downward, along base of first three rays, continuing straight and horizontal or slightly inclined downward toward end of base, then nearly straight and inclined upward to insertion of caudal fin. Ventral profile of head horizontal or slightly convex; slightly convex or straight towards origin of anal fin; anal-fin base straight and inclined upward (horizontal in small specimens), then nearly straight and inclined downward to insertion of caudal fin.</paragraph>
<caption id="6788F8D6FF8DFFC6FF621DB5ABF482D4" ID-Table-UUID="6788F8D6FF8DFFC6FF621DB5ABF482D4" box="[151,1209,152,174]" httpUri="http://table.plazi.org/id/6788F8D6FF8DFFC6FF621DB5ABF482D4" pageId="3" pageNumber="63" startId="3.[151,235,152,173]" targetBox="[159,1225,200,1169]" targetIsTable="true" targetPageId="3">
<paragraph id="3348A85EFF8DFFC6FF621DB5ABF482D4" blockId="3.[151,1209,152,174]" box="[151,1209,152,174]" pageId="3" pageNumber="63">
<emphasis id="0183744CFF8DFFC6FF621DB5AE4982D7" bold="true" box="[151,260,152,173]" pageId="3" pageNumber="63">TABLE 1.</emphasis>
Morphometric data for holotype and paratypes (n=19) of
<taxonomicName id="F4F7D3DDFF8DFFC6FCBA1DB5AB2F82D4" authority="Lezama &amp; Triques &amp; Santos, 2012" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[847,1122,152,174]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="63" phylum="Chordata" rank="species" species="argos" status="new">
<emphasis id="0183744CFF8DFFC6FCBA1DB5AB7F82D4" box="[847,1074,152,174]" italics="true" pageId="3" pageNumber="63">Trichomycterus argos,</emphasis>
new
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph id="3348A85EFF8DFFC6FD931DE5ABF086EB" pageId="3" pageNumber="63">
<table id="41F75AFEFF8D003AFF6A1DE5AB8486EB" box="[159,1225,200,1169]" gridcols="4" gridrows="25" pageId="3" pageNumber="63">
<tr id="8DC7AA1CFF8D003AFF6A1DE5AB8482A4" box="[159,1225,200,222]" gridrow="0" pageId="3" pageNumber="63" rowspan-0="1">
<th id="CE16C360FF8D003AFD931DE5AD8982A4" box="[614,708,200,222]" gridcol="1" gridrow="0" pageId="3" pageNumber="63">Holotype</th>
<th id="CE16C360FF8D003AFC8E1DE5ACA082A4" box="[891,1005,200,222]" gridcol="2" gridrow="0" pageId="3" pageNumber="63">Range</th>
<th id="CE16C360FF8D003AFB651DE5AB8482A4" box="[1168,1225,200,222]" gridcol="3" gridrow="0" pageId="3" pageNumber="63">Mean</th>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1DDDAB84837D" box="[159,1225,240,263]" gridrow="1" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1DDDAE99837D" box="[159,468,240,263]" gridcol="0" gridrow="1" pageId="3" pageNumber="63">Standard length (mm)</th>
<td id="CE16C360FF8D003AFD931DDDAD89837D" box="[614,708,240,263]" gridcol="1" gridrow="1" pageId="3" pageNumber="63">95.4</td>
<td id="CE16C360FF8D003AFC8E1DDDACA0837D" box="[891,1005,240,263]" gridcol="2" gridrow="1" pageId="3" pageNumber="63">56.0–116.8</td>
<td id="CE16C360FF8D003AFB651DDDAB84837D" box="[1168,1225,240,263]" gridcol="3" gridrow="1" pageId="3" pageNumber="63">83.9</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1C3AAB848354" box="[159,1225,279,302]" gridrow="2" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1C3AAE998354" box="[159,468,279,302]" gridcol="0" gridrow="2" pageId="3" pageNumber="63">Total length (mm)</th>
<td id="CE16C360FF8D003AFD931C3AAD898354" box="[614,708,279,302]" gridcol="1" gridrow="2" pageId="3" pageNumber="63">109.5</td>
<td id="CE16C360FF8D003AFC8E1C3AACA08354" box="[891,1005,279,302]" gridcol="2" gridrow="2" pageId="3" pageNumber="63">65.1–134.9</td>
<td id="CE16C360FF8D003AFB651C3AAB848354" box="[1168,1225,279,302]" gridcol="3" gridrow="2" pageId="3" pageNumber="63">97.1</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1C12AB84832F" box="[159,1225,319,341]" gridrow="3" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1C12AE99832F" box="[159,468,319,341]" gridcol="0" gridrow="3" pageId="3" pageNumber="63">Head length (mm)</th>
<td id="CE16C360FF8D003AFD931C12AD89832F" box="[614,708,319,341]" gridcol="1" gridrow="3" pageId="3" pageNumber="63">20,3</td>
<td id="CE16C360FF8D003AFC8E1C12ACA0832F" box="[891,1005,319,341]" gridcol="2" gridrow="3" pageId="3" pageNumber="63">12.6–26.3</td>
<td id="CE16C360FF8D003AFB651C12AB84832F" box="[1168,1225,319,341]" gridcol="3" gridrow="3" pageId="3" pageNumber="63">18.6</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1C4BAB848306" box="[159,1225,358,380]" gridrow="4" pageId="3" pageNumber="63" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="CE16C360FF8D003AFF6A1C4BAE998306" box="[159,468,358,380]" gridcol="0" gridrow="4" pageId="3" pageNumber="63">Percentages of standard length</th>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1CA3AB8483DE" box="[159,1225,398,420]" gridrow="5" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1CA3AE9983DE" box="[159,468,398,420]" gridcol="0" gridrow="5" pageId="3" pageNumber="63">Total length</th>
<td id="CE16C360FF8D003AFD931CA3AD8983DE" box="[614,708,398,420]" gridcol="1" gridrow="5" pageId="3" pageNumber="63">114.8</td>
<td id="CE16C360FF8D003AFC8E1CA3ACA083DE" box="[891,1005,398,420]" gridcol="2" gridrow="5" pageId="3" pageNumber="63">114.8–118</td>
<td id="CE16C360FF8D003AFB651CA3AB8483DE" box="[1168,1225,398,420]" gridcol="3" gridrow="5" pageId="3" pageNumber="63">116.0</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1C98AB8483B1" box="[159,1225,437,459]" gridrow="6" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1C98AE9983B1" box="[159,468,437,459]" gridcol="0" gridrow="6" pageId="3" pageNumber="63">Body depth</th>
<td id="CE16C360FF8D003AFD931C98AD8983B1" box="[614,708,437,459]" gridcol="1" gridrow="6" pageId="3" pageNumber="63">13.8</td>
<td id="CE16C360FF8D003AFC8E1C98ACA083B1" box="[891,1005,437,459]" gridcol="2" gridrow="6" pageId="3" pageNumber="63">12.8–17.5</td>
<td id="CE16C360FF8D003AFB651C98AB8483B1" box="[1168,1225,437,459]" gridcol="3" gridrow="6" pageId="3" pageNumber="63">15.9</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1CF0AB848389" box="[159,1225,477,499]" gridrow="7" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1CF0AE998389" box="[159,468,477,499]" gridcol="0" gridrow="7" pageId="3" pageNumber="63">Caudal peduncle length</th>
<td id="CE16C360FF8D003AFD931CF0AD898389" box="[614,708,477,499]" gridcol="1" gridrow="7" pageId="3" pageNumber="63">20.4</td>
<td id="CE16C360FF8D003AFC8E1CF0ACA08389" box="[891,1005,477,499]" gridcol="2" gridrow="7" pageId="3" pageNumber="63">19.2–22.7</td>
<td id="CE16C360FF8D003AFB651CF0AB848389" box="[1168,1225,477,499]" gridcol="3" gridrow="7" pageId="3" pageNumber="63">21.0</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1F29AB848060" box="[159,1225,516,538]" gridrow="8" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1F29AE998060" box="[159,468,516,538]" gridcol="0" gridrow="8" pageId="3" pageNumber="63">Caudal peduncle depth</th>
<td id="CE16C360FF8D003AFD931F29AD898060" box="[614,708,516,538]" gridcol="1" gridrow="8" pageId="3" pageNumber="63">15.8</td>
<td id="CE16C360FF8D003AFC8E1F29ACA08060" box="[891,1005,516,538]" gridcol="2" gridrow="8" pageId="3" pageNumber="63">14.6–17.5</td>
<td id="CE16C360FF8D003AFB651F29AB848060" box="[1168,1225,516,538]" gridcol="3" gridrow="8" pageId="3" pageNumber="63">16.3</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1F06AB84803B" box="[159,1225,555,577]" gridrow="9" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1F06AE99803B" box="[159,468,555,577]" gridcol="0" gridrow="9" pageId="3" pageNumber="63">Predorsal length</th>
<td id="CE16C360FF8D003AFD931F06AD89803B" box="[614,708,555,577]" gridcol="1" gridrow="9" pageId="3" pageNumber="63">63.1</td>
<td id="CE16C360FF8D003AFC8E1F06ACA0803B" box="[891,1005,555,577]" gridcol="2" gridrow="9" pageId="3" pageNumber="63">60.9–65.9</td>
<td id="CE16C360FF8D003AFB651F06AB84803B" box="[1168,1225,555,577]" gridcol="3" gridrow="9" pageId="3" pageNumber="63">64.0</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1F7EAB848013" box="[159,1225,595,617]" gridrow="10" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1F7EAE998013" box="[159,468,595,617]" gridcol="0" gridrow="10" pageId="3" pageNumber="63">Preanal length</th>
<td id="CE16C360FF8D003AFD931F7EAD898013" box="[614,708,595,617]" gridcol="1" gridrow="10" pageId="3" pageNumber="63">72.0</td>
<td id="CE16C360FF8D003AFC8E1F7EACA08013" box="[891,1005,595,617]" gridcol="2" gridrow="10" pageId="3" pageNumber="63">70.1–73.7</td>
<td id="CE16C360FF8D003AFB651F7EAB848013" box="[1168,1225,595,617]" gridcol="3" gridrow="10" pageId="3" pageNumber="63">71.5</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1F57AB8480EA" box="[159,1225,634,656]" gridrow="11" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1F57AE9980EA" box="[159,468,634,656]" gridcol="0" gridrow="11" pageId="3" pageNumber="63">Prepelvic length</th>
<td id="CE16C360FF8D003AFD931F57AD8980EA" box="[614,708,634,656]" gridcol="1" gridrow="11" pageId="3" pageNumber="63">58.7</td>
<td id="CE16C360FF8D003AFC8E1F57ACA080EA" box="[891,1005,634,656]" gridcol="2" gridrow="11" pageId="3" pageNumber="63">55.4–76.6</td>
<td id="CE16C360FF8D003AFB651F57AB8480EA" box="[1168,1225,634,656]" gridcol="3" gridrow="11" pageId="3" pageNumber="63">60.3</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1F8FAB8480C2" box="[159,1225,674,696]" gridrow="12" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1F8FAE9980C2" box="[159,468,674,696]" gridcol="0" gridrow="12" pageId="3" pageNumber="63">Dorsal fin base length</th>
<td id="CE16C360FF8D003AFD931F8FAD8980C2" box="[614,708,674,696]" gridcol="1" gridrow="12" pageId="3" pageNumber="63">10.9</td>
<td id="CE16C360FF8D003AFC8E1F8FACA080C2" box="[891,1005,674,696]" gridcol="2" gridrow="12" pageId="3" pageNumber="63">8.7–11.2</td>
<td id="CE16C360FF8D003AFB651F8FAB8480C2" box="[1168,1225,674,696]" gridcol="3" gridrow="12" pageId="3" pageNumber="63">10.0</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1FE7AB84809A" box="[159,1225,714,736]" gridrow="13" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1FE7AE99809A" box="[159,468,714,736]" gridcol="0" gridrow="13" pageId="3" pageNumber="63">Anal fin base length</th>
<td id="CE16C360FF8D003AFD931FE7AD89809A" box="[614,708,714,736]" gridcol="1" gridrow="13" pageId="3" pageNumber="63">8.4</td>
<td id="CE16C360FF8D003AFC8E1FE7ACA0809A" box="[891,1005,714,736]" gridcol="2" gridrow="13" pageId="3" pageNumber="63">7.2–8.8</td>
<td id="CE16C360FF8D003AFB651FE7AB84809A" box="[1168,1225,714,736]" gridcol="3" gridrow="13" pageId="3" pageNumber="63">8.1</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1FDCAB84817D" box="[159,1225,753,775]" gridrow="14" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1FDCAE99817D" box="[159,468,753,775]" gridcol="0" gridrow="14" pageId="3" pageNumber="63">Dorsal-caudal length</th>
<td id="CE16C360FF8D003AFD931FDCAD89817D" box="[614,708,753,775]" gridcol="1" gridrow="14" pageId="3" pageNumber="63">37.3</td>
<td id="CE16C360FF8D003AFC8E1FDCACA0817D" box="[891,1005,753,775]" gridcol="2" gridrow="14" pageId="3" pageNumber="63">34.5–39.2</td>
<td id="CE16C360FF8D003AFB651FDCAB84817D" box="[1168,1225,753,775]" gridcol="3" gridrow="14" pageId="3" pageNumber="63">36.6</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1E35AB848154" box="[159,1225,792,814]" gridrow="15" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1E35AE998154" box="[159,468,792,814]" gridcol="0" gridrow="15" pageId="3" pageNumber="63">Pelvic-caudal length</th>
<td id="CE16C360FF8D003AFD931E35AD898154" box="[614,708,792,814]" gridcol="1" gridrow="15" pageId="3" pageNumber="63">42.7</td>
<td id="CE16C360FF8D003AFC8E1E35ACA08154" box="[891,1005,792,814]" gridcol="2" gridrow="15" pageId="3" pageNumber="63">40.1–43.6</td>
<td id="CE16C360FF8D003AFB651E35AB848154" box="[1168,1225,792,814]" gridcol="3" gridrow="15" pageId="3" pageNumber="63">41.8</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1E6DAB84812C" box="[159,1225,832,854]" gridrow="16" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1E6DAE99812C" box="[159,468,832,854]" gridcol="0" gridrow="16" pageId="3" pageNumber="63">Head length</th>
<td id="CE16C360FF8D003AFD931E6DAD89812C" box="[614,708,832,854]" gridcol="1" gridrow="16" pageId="3" pageNumber="63">21.3</td>
<td id="CE16C360FF8D003AFC8E1E6DACA0812C" box="[891,1005,832,854]" gridcol="2" gridrow="16" pageId="3" pageNumber="63">20.8–23.3</td>
<td id="CE16C360FF8D003AFB651E6DAB84812C" box="[1168,1225,832,854]" gridcol="3" gridrow="16" pageId="3" pageNumber="63">22.3</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1E4AAB848107" box="[159,1225,871,893]" gridrow="17" pageId="3" pageNumber="63" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="CE16C360FF8D003AFF6A1E4AAE998107" box="[159,468,871,893]" gridcol="0" gridrow="17" pageId="3" pageNumber="63">Percentages of head length</th>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1EA2AB8481DF" box="[159,1225,911,933]" gridrow="18" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1EA2AE9981DF" box="[159,468,911,933]" gridcol="0" gridrow="18" pageId="3" pageNumber="63">Head width</th>
<td id="CE16C360FF8D003AFD931EA2AD8981DF" box="[614,708,911,933]" gridcol="1" gridrow="18" pageId="3" pageNumber="63">85.2</td>
<td id="CE16C360FF8D003AFC8E1EA2ACA081DF" box="[891,1005,911,933]" gridcol="2" gridrow="18" pageId="3" pageNumber="63">79.1–87.6</td>
<td id="CE16C360FF8D003AFB651EA2AB8481DF" box="[1168,1225,911,933]" gridcol="3" gridrow="18" pageId="3" pageNumber="63">84.3</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1E9BAB8481B6" box="[159,1225,950,972]" gridrow="19" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1E9BAE9981B6" box="[159,468,950,972]" gridcol="0" gridrow="19" pageId="3" pageNumber="63">Head depth</th>
<td id="CE16C360FF8D003AFD931E9BAD8981B6" box="[614,708,950,972]" gridcol="1" gridrow="19" pageId="3" pageNumber="63">55.2</td>
<td id="CE16C360FF8D003AFC8E1E9BACA081B6" box="[891,1005,950,972]" gridcol="2" gridrow="19" pageId="3" pageNumber="63">47.2–60.2</td>
<td id="CE16C360FF8D003AFB651E9BAB8481B6" box="[1168,1225,950,972]" gridcol="3" gridrow="19" pageId="3" pageNumber="63">53.0</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1EF3AB84818E" box="[159,1225,990,1012]" gridrow="20" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1EF3AE99818E" box="[159,468,990,1012]" gridcol="0" gridrow="20" pageId="3" pageNumber="63">Interorbital width</th>
<td id="CE16C360FF8D003AFD931EF3AD89818E" box="[614,708,990,1012]" gridcol="1" gridrow="20" pageId="3" pageNumber="63">26.6</td>
<td id="CE16C360FF8D003AFC8E1EF3ACA0818E" box="[891,1005,990,1012]" gridcol="2" gridrow="20" pageId="3" pageNumber="63">24.6–29.4</td>
<td id="CE16C360FF8D003AFB651EF3AB84818E" box="[1168,1225,990,1012]" gridcol="3" gridrow="20" pageId="3" pageNumber="63">26.9</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1928AB848661" box="[159,1225,1029,1051]" gridrow="21" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1928AE998661" box="[159,468,1029,1051]" gridcol="0" gridrow="21" pageId="3" pageNumber="63">Internasal width</th>
<td id="CE16C360FF8D003AFD931928AD898661" box="[614,708,1029,1051]" gridcol="1" gridrow="21" pageId="3" pageNumber="63">19.2</td>
<td id="CE16C360FF8D003AFC8E1928ACA08661" box="[891,1005,1029,1051]" gridcol="2" gridrow="21" pageId="3" pageNumber="63">16.4–20.6</td>
<td id="CE16C360FF8D003AFB651928AB848661" box="[1168,1225,1029,1051]" gridcol="3" gridrow="21" pageId="3" pageNumber="63">18.9</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1900AB848639" box="[159,1225,1069,1091]" gridrow="22" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1900AE998639" box="[159,468,1069,1091]" gridcol="0" gridrow="22" pageId="3" pageNumber="63">Snout length</th>
<td id="CE16C360FF8D003AFD931900AD898639" box="[614,708,1069,1091]" gridcol="1" gridrow="22" pageId="3" pageNumber="63">40.9</td>
<td id="CE16C360FF8D003AFC8E1900ACA08639" box="[891,1005,1069,1091]" gridcol="2" gridrow="22" pageId="3" pageNumber="63">37.4–46.9</td>
<td id="CE16C360FF8D003AFB651900AB848639" box="[1168,1225,1069,1091]" gridcol="3" gridrow="22" pageId="3" pageNumber="63">42.0</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1979AB848610" box="[159,1225,1108,1130]" gridrow="23" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1979AE998610" box="[159,468,1108,1130]" gridcol="0" gridrow="23" pageId="3" pageNumber="63">Eye diameter</th>
<td id="CE16C360FF8D003AFD931979AD898610" box="[614,708,1108,1130]" gridcol="1" gridrow="23" pageId="3" pageNumber="63">13.3</td>
<td id="CE16C360FF8D003AFC8E1979ACA08610" box="[891,1005,1108,1130]" gridcol="2" gridrow="23" pageId="3" pageNumber="63">10.5–17.5</td>
<td id="CE16C360FF8D003AFB651979AB848610" box="[1168,1225,1108,1130]" gridcol="3" gridrow="23" pageId="3" pageNumber="63">9.7</td>
</tr>
<tr id="8DC7AA1CFF8D003AFF6A1956AB8486EB" box="[159,1225,1147,1169]" gridrow="24" pageId="3" pageNumber="63">
<th id="CE16C360FF8D003AFF6A1956AE9986EB" box="[159,468,1147,1169]" gridcol="0" gridrow="24" pageId="3" pageNumber="63">Mouth width</th>
<td id="CE16C360FF8D003AFD931956AD8986EB" box="[614,708,1147,1169]" gridcol="1" gridrow="24" pageId="3" pageNumber="63">44.3</td>
<td id="CE16C360FF8D003AFC8E1956ACA086EB" box="[891,1005,1147,1169]" gridcol="2" gridrow="24" pageId="3" pageNumber="63">32.8–47.4</td>
<td id="CE16C360FF8D003AFB651956AB8486EB" box="[1168,1225,1147,1169]" gridcol="3" gridrow="24" pageId="3" pageNumber="63">39.2</td>
</tr>
</table>
</paragraph>
<paragraph id="3348A85EFF8DFFC6FF3219FBAE5087BF" blockId="3.[151,1436,1237,2017]" pageId="3" pageNumber="63">Eyes small and elliptical, uniformly covered by skin; eyes dorsally on head, at middle or slightly at head anterior region, the former condition more common in large and medium-sized specimens, and the latter in small specimens. Mouth sub-terminal, with two fleshy lobes posterolaterally to inferior lip and medially to origin of rictal barbels. Branchial openings wide, with thick branchial membranes, anteriorly united to isthmus at a vertical through posterior edge of eyes. Opercle small and round, with 20–22 conical and thick odontodes. Interopercle oblique, ranging from two to three times longer than opercle, with 38–41 conical and thick odontodes, arranged in three series.</paragraph>
<paragraph id="3348A85EFF8DFFC6FF3218FCAE558403" blockId="3.[151,1436,1237,2017]" pageId="3" pageNumber="63">Anterior nostrils surrounded by a fleshy skin fold medially and one nasal barbel laterally. Posterior nostrils antero-posteriorly elongated, surrounded anteriorly by a fleshy skin fold. Tip of nasal barbels reaching posterior border of opercular plate of odontodes. Tip of maxillary barbells extending to end of pectoral-fin base. Tip of rictal barbels reaching origin of pectoral fin. Barbels laminated and wide basally, gradually narrowing towards tip, thread-like.</paragraph>
<paragraph id="3348A85EFF8DFFC6FF321BA8AE82859B" blockId="3.[151,1436,1237,2017]" pageId="3" pageNumber="63">Anal and urogenital openings below anterior half of dorsal-fin base. Integument thick at base of all fins. Dorsal-fin rays i+7 (13*), ii+7 (6) or ii+6 (1); two unsegmented rays in C&amp;S specimen; distal margin convex; origin nearly at two thirds of the standard length. Pectoral-fin rays i+6; distal margin convex; origin nearly at vertical by end of opercular plate of odontodes; first ray nearly an ocular diameter longer than others. Pelvic-fin rays i+4 (pelvic splint visible in C&amp;S specimen); distal margin convex; origin from one to two ocular diameters in front of dorsal-fin origin and equidistant to tip of snout and end of caudal fin, or nearer to last one; distal margin surpassing anus and reaching one or two ocular diameters in front of anal-fin origin, being generally more anterior in large specimens than in small ones. Anal-fin rays i+5 (7*) or ii+5 (13); three unsegmented rays in C&amp;S specimen; distal margin convex; origin at same vertical or slightly anterior to end of dorsal-fin base. Caudal-fin rays i+5; i+6; distal margin rounded or truncate.</paragraph>
<caption id="6788F8D6FF8AFFC1FF6218BDAACB87DC" box="[151,1414,1424,1447]" httpUri="https://zenodo.org/record/210128/files/figure.png" pageId="4" pageNumber="64" targetBox="[151,1435,193,1402]" targetPageId="4">
<paragraph id="3348A85EFF8AFFC1FF6218BDAACB87DC" blockId="4.[151,1414,1424,1447]" box="[151,1414,1424,1447]" pageId="4" pageNumber="64">
<emphasis id="0183744CFF8AFFC1FF6218BDAE5E87DF" bold="true" box="[151,275,1424,1447]" pageId="4" pageNumber="64">FIGURE 2.</emphasis>
Holotype of
<taxonomicName id="F4F7D3DDFF8AFFC1FE6C18BDAD3887DC" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[409,629,1424,1446]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="64" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8AFFC1FE6C18BDAD3887DC" box="[409,629,1424,1446]" italics="true" pageId="4" pageNumber="64">Trichomycterus argos</emphasis>
</taxonomicName>
, DZUFMG 103, 95.4 mm SL, in dorsal (1), lateral (2), and ventral (3) views.
</paragraph>
</caption>
<paragraph id="3348A85EFF8AFFC1FF3218E1AC73857E" blockId="4.[151,1436,1484,2012]" pageId="4" pageNumber="64">
<emphasis id="0183744CFF8AFFC1FF3218E1AE02879F" bold="true" box="[199,335,1484,1509]" pageId="4" pageNumber="64">Coloration.</emphasis>
Body and head background color cream, in small specimens, and dark brown dorsally to cream ventrally on flanks, in adults. Head and body with spots of superficial chromatophores that seldom reach eye diameter in size, coarse dorsally, progressively more sparse ventrally, to the level of pectoral and pelvic-fin insertions, not forming vermiculations. Smaller spots of deeper chromatophores present, from dorsum to level of pectoral and pelvic-fin insertions, over head and body, with nearly same concentration over its range of distribution, not forming vermiculations. Below level of pectoral and pelvic-fin insertions, spots very few and sparse, or absent over the greater part of this region; presence of a mid-ventral line of chromatophores, from anterior part of the isthmus towards level of the pelvic-fin origin, forming, in some cases, a narrow continuous stripe, rarely absent. Opercular membrane border with line of concentrated chromatophores.
</paragraph>
<paragraph id="3348A85EFF8AFFC1FF321A3DAE5785A6" blockId="4.[151,1436,1484,2012]" pageId="4" pageNumber="64">Chromatophores present on rays and ray-borders all through length of fins. Bases of dorsal, caudal and dorsal side of pectoral fin with superficial and deep spots. Fin membranes hyaline distally, exception in large specimens with dorsal, anal and caudal fins presenting clear brown background color, with dark roundish marks proximally, and hyaline membrane distally (only dorsal fin pigmented distally on membrane). Medial area of fins variably pigmented or hyaline. Barbels pigmented on each edge, becoming lighter towards tip; rictal barbels slightly lighter than others.</paragraph>
</subSubSection>
<caption id="6788F8D6FF8BFFC0FF621952ADF08746" httpUri="https://zenodo.org/record/210129/files/figure.png" pageId="5" pageNumber="65" targetBox="[156,1426,220,1119]" targetPageId="5">
<paragraph id="3348A85EFF8BFFC0FF621952ADF08746" blockId="5.[151,1436,1151,1341]" pageId="5" pageNumber="65">
<emphasis id="0183744CFF8BFFC0FF621952AE5886EF" bold="true" box="[151,277,1151,1174]" pageId="5" pageNumber="65">FIGURE 3.</emphasis>
Dorsal views of the cranium and Weberian complex of
<emphasis id="0183744CFF8BFFC0FC961952AC3586EC" bold="true" box="[867,888,1151,1174]" pageId="5" pageNumber="65">a)</emphasis>
<taxonomicName id="F4F7D3DDFF8BFFC0FC741952AB2D86EC" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[897,1120,1151,1174]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="65" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8BFFC0FC741952AB2D86EC" box="[897,1120,1151,1174]" italics="true" pageId="5" pageNumber="65">Trichomycterus argos</emphasis>
</taxonomicName>
(DZUFMG 0 67, 101 mm SL, paratype) and
<emphasis id="0183744CFF8BFFC0FEDC198DAE0D86CD" bold="true" box="[297,320,1184,1207]" pageId="5" pageNumber="65">b)</emphasis>
<taxonomicName id="F4F7D3DDFF8BFFC0FEB2198CAD2D86CD" box="[327,608,1185,1207]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="65" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="0183744CFF8BFFC0FEB2198CAD2D86CD" box="[327,608,1185,1207]" italics="true" pageId="5" pageNumber="65">Trichomycterus brasiliensis</emphasis>
</taxonomicName>
(DZUFMG 0 64, 102,1 mm SL). Abbreviations:
<emphasis id="0183744CFF8BFFC0FBA1198CAB3686CC" bold="true" box="[1108,1147,1185,1206]" pageId="5" pageNumber="65">AP,</emphasis>
autopalatine;
<emphasis id="0183744CFF8BFFC0FAFE198CAA7D86CC" bold="true" box="[1291,1328,1185,1206]" pageId="5" pageNumber="65">EP,</emphasis>
epioccipital;
<emphasis id="0183744CFF8BFFC0FF4819EFAFAC86AD" bold="true" box="[189,225,1218,1239]" pageId="5" pageNumber="65">FF,</emphasis>
frontal fontanels;
<emphasis id="0183744CFF8BFFC0FE6919EFAE8986AD" bold="true" box="[412,452,1218,1239]" pageId="5" pageNumber="65">FR,</emphasis>
frontal;
<emphasis id="0183744CFF8BFFC0FDEE19ECAD7286A2" bold="true" box="[539,575,1217,1240]" pageId="5" pageNumber="65">IO,</emphasis>
tendon-bone infraorbital;
<emphasis id="0183744CFF8BFFC0FCBF19EFAC3F86AD" bold="true" box="[842,882,1218,1239]" pageId="5" pageNumber="65">LE,</emphasis>
lateral ethmoid;
<emphasis id="0183744CFF8BFFC0FBEA19EFAB0386AD" bold="true" box="[1055,1102,1218,1239]" pageId="5" pageNumber="65">ME,</emphasis>
mesethmoid;
<emphasis id="0183744CFF8BFFC0FB2B19EFAA4386AD" bold="true" box="[1246,1294,1218,1239]" pageId="5" pageNumber="65">MX,</emphasis>
maxilla;
<emphasis id="0183744CFF8BFFC0FA9B19EFAAD186AD" bold="true" box="[1390,1436,1218,1239]" pageId="5" pageNumber="65">PM,</emphasis>
premaxilla;
<emphasis id="0183744CFF8BFFC0FEE019CEAE748680" bold="true" box="[277,313,1251,1274]" pageId="5" pageNumber="65">PS,</emphasis>
posttemporosupracleithrum;
<emphasis id="0183744CFF8BFFC0FD9819C9ADDF8683" bold="true" box="[621,658,1252,1273]" pageId="5" pageNumber="65">PT,</emphasis>
pterotic;
<emphasis id="0183744CFF8BFFC0FCFC19C9AC628683" bold="true" box="[777,815,1252,1273]" pageId="5" pageNumber="65">RF,</emphasis>
foramen for the
<emphasis id="0183744CFF8BFFC0FC1919C9AA2E8683" box="[1004,1379,1251,1273]" italics="true" pageId="5" pageNumber="65">ramus lateralis accessorius facialis</emphasis>
;
<emphasis id="0183744CFF8BFFC0FA8019CEAAD18680" bold="true" box="[1397,1436,1251,1274]" pageId="5" pageNumber="65">SC,</emphasis>
cephalic sensory canal;
<emphasis id="0183744CFF8BFFC0FE7E1829AEF78760" bold="true" box="[395,442,1284,1307]" pageId="5" pageNumber="65">SO,</emphasis>
tendon-bone supraorbital;
<emphasis id="0183744CFF8BFFC0FD3B1829AC4D8761" bold="true" box="[718,768,1284,1307]" pageId="5" pageNumber="65">SPP,</emphasis>
sphenotic-prootic-pterosphenoid compound bone;
<emphasis id="0183744CFF8BFFC0FAFC1829AA628761" bold="true" box="[1289,1327,1284,1307]" pageId="5" pageNumber="65">SU,</emphasis>
parietosupraoccipital;
<emphasis id="0183744CFF8BFFC0FEEE180BAE008747" bold="true" box="[283,333,1318,1341]" pageId="5" pageNumber="65">WC,</emphasis>
Weberian capsule. Scale bar = 1cm.
</paragraph>
</caption>
<subSubSection id="7BEDFBD5FF8BFFC0FF321841AF9885B9" pageId="5" pageNumber="65" type="etymology">
<paragraph id="3348A85EFF8BFFC0FF321841AC8787DD" blockId="5.[151,1436,1387,1987]" pageId="5" pageNumber="65">
<emphasis id="0183744CFF8BFFC0FF321841AE0387FE" bold="true" box="[199,334,1388,1412]" pageId="5" pageNumber="65">Etymology.</emphasis>
The epithet “argos” (Gr.), meaning “a hundred-eyed monster” (
<bibRefCitation id="5766D5AFFF8BFFC0FBDC1846ABB087F9" author="Brown" box="[1065,1277,1387,1411]" pageId="5" pageNumber="65" refString="Brown, R. W. (1956) Composition of Scientific Words. A manual of methods and lexicon of materials for the practice of logotechnics. Smithsonian Institution Press, Washington D. C., 882 pp." type="book" year="1956">Brown, 1956: 312</bibRefCitation>
), refers to the eye-spotted color pattern of the new species, not forming vermiculations.
</paragraph>
<paragraph id="3348A85EFF8BFFC0FF32189EAD9884FA" blockId="5.[151,1436,1387,1987]" pageId="5" pageNumber="65">
<emphasis id="0183744CFF8BFFC0FF32189EAECD87B6" bold="true" box="[199,384,1459,1484]" pageId="5" pageNumber="65">Biological data.</emphasis>
Sampling for study of habitat use was conducted between
<date id="47498E9EFF8BFFC0FBE0189EABE387B1" box="[1045,1198,1459,1483]" pageId="5" pageNumber="65" value="2001-10">October 2001</date>
and
<date id="47498E9EFF8BFFC0FB10189EAAD187B1" box="[1253,1436,1459,1483]" pageId="5" pageNumber="65" value="2002-09">September 2002</date>
at Serra Nova stream and two of its tributaries, in the same area where the
<typeStatus id="EC4C16FCFF8BFFC0FC2718FAAB798795" box="[978,1076,1495,1519]" pageId="5" pageNumber="65" type="holotype">holotype</typeStatus>
was collected, in a region of the Parque Estadual da Serra do Brigadeiro called Fazenda da Neblina. The local climate presents two well defined seasons, one predominantly mesothermic (Köppen’s “Cwb” climate classification) with annual mean temperature of 15°C, with possible occurrence of minimum temperatures below 0°C. Mean annual precipitation is
<quantity id="F40F05BBFF8BFFC0FAD71B6EAADA8421" box="[1314,1431,1603,1627]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="65" unit="mm" value="1.5">1,500 mm</quantity>
, with the dry season extending from July to August.
</paragraph>
<paragraph id="3348A85EFF8BFFC0FF321BA1AFBA8501" blockId="5.[151,1436,1387,1987]" pageId="5" pageNumber="65">
Twenty funnel traps with double entrances were installed monthly in each stream. The height of the water column was measured, and each trap was used for 72 hours/month. The
<specimenCount id="25F163D7FF8BFFC0FC4E1B82AB1884BD" box="[955,1109,1711,1735]" pageId="5" pageNumber="65" type="generic">49 specimens</specimenCount>
of
<taxonomicName id="F4F7D3DDFF8BFFC0FB8A1B82AA6384BD" box="[1151,1326,1711,1735]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="65" phylum="Chordata" rank="genus">
<emphasis id="0183744CFF8BFFC0FB8A1B82AA6384BD" box="[1151,1326,1711,1735]" italics="true" pageId="5" pageNumber="65">Trichomycterus</emphasis>
</taxonomicName>
collected throughout the study period (SL between 4 and
<quantity id="F40F05BBFF8BFFC0FD5F1BFEADB48491" box="[682,761,1747,1771]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="65" unit="mm" value="17.0">17 mm</quantity>
) occupied habitats with depth and temperature varying from
<quantity id="F40F05BBFF8BFFC0FF621BDAAE568575" box="[151,283,1783,1807]" metricMagnitude="-1" metricUnit="m" metricValue="2.85" metricValueMax="4.4" metricValueMin="1.3" pageId="5" pageNumber="65" unit="cm" value="28.5" valueMax="44.0" valueMin="13.0">13 to 44 cm</quantity>
and from 14 to 19°C. A larger number of specimens (15) was collected in
<date id="47498E9EFF8BFFC0FB951BDAAB9D8575" box="[1120,1232,1783,1807]" pageId="5" pageNumber="65" value="2002-07">July 2002</date>
, possibly a reflection of the lower water flow which did not promote the downstream flow of specimens. Twenty nine of the
<specimenCount id="25F163D7FF8BFFC0FACA1A36AF97852D" pageId="5" pageNumber="65" type="generic">49 specimens</specimenCount>
belong to
<taxonomicName id="F4F7D3DDFF8BFFC0FEA41A6DAE92852D" baseAuthorityName="Eigenmann" baseAuthorityYear="1917" box="[337,479,1855,1879]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="65" phylum="Chordata" rank="species" species="alternatus">
<emphasis id="0183744CFF8BFFC0FEA41A6DAE13852D" box="[337,350,1856,1879]" italics="true" pageId="5" pageNumber="65">T</emphasis>
.
<emphasis id="0183744CFF8BFFC0FE981A12AE92852D" box="[365,479,1855,1879]" italics="true" pageId="5" pageNumber="65">alternatus</emphasis>
</taxonomicName>
, a clearly different species than
<taxonomicName id="F4F7D3DDFF8BFFC0FCBE1A6DACE8852D" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[843,933,1856,1879]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="65" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8BFFC0FCBE1A6DACE8852D" box="[843,933,1856,1879]" italics="true" pageId="5" pageNumber="65">T. argos</emphasis>
</taxonomicName>
, on the basis of body color pattern and other features.
</paragraph>
<paragraph id="3348A85EFF8BFFC0FF321AAAAF9885B9" blockId="5.[151,1436,1387,1987]" pageId="5" pageNumber="65">
Predation of tadpoles of
<taxonomicName id="F4F7D3DDFF8BFFC0FE291AA5AD6A85E5" box="[476,551,1928,1951]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="65" phylum="Chordata" rank="genus">
<emphasis id="0183744CFF8BFFC0FE291AA5AD6A85E5" box="[476,551,1928,1951]" italics="true" pageId="5" pageNumber="65">Scinax</emphasis>
</taxonomicName>
gr.
<emphasis id="0183744CFF8BFFC0FDA71AAAAD8685E5" box="[594,715,1927,1951]" italics="true" pageId="5" pageNumber="65">catharinae</emphasis>
by specimens of
<taxonomicName id="F4F7D3DDFF8BFFC0FC641AA5ACA185E5" authorityName="Lezama &amp; Triques &amp; Santos" authorityYear="2012" box="[913,1004,1928,1951]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="65" phylum="Chordata" rank="species" species="argos">
<emphasis id="0183744CFF8BFFC0FC641AA5ACA185E5" box="[913,1004,1928,1951]" italics="true" pageId="5" pageNumber="65">T. argos</emphasis>
</taxonomicName>
was observed under laboratory conditions.
</paragraph>
</subSubSection>
</treatment>
</document>
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