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<document id="307CA08E67C28DE08B0B117296D7536F" ID-CLB-Dataset="22814" ID-DOI="10.11646/zootaxa.4766.4.1" ID-GBIF-Dataset="1e2996e2-e31f-45e3-9149-60ebce934708" ID-ISSN="1175-5334" ID-Zenodo-Dep="3765652" ID-ZooBank="urn:lsid:zoobank.org:pub:2F8FEDC1-6891-46D1-B372-858CDCBE4051" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587754498722" checkinUser="carolina" docAuthor="Kompier, Tom, Dow, Rory A. &amp; Steinhoff, Philip O. M." docDate="2020" docId="AF6A87D5436E6174FF6ADB91493D3830" docLanguage="en" docName="zootaxa.4766.4.1.pdf.imf" docOrigin="Zootaxa 4766 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coeliccia caerulea Kompier &amp; Dow &amp; Steinhoff 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="505" masterDocId="5353FFAD436C6170FFFDDB7048083A53" masterDocTitle="Five new species of Coeliccia Kirby, 1890 from Vietnam (Odonata: Platycnemididae), and information on several other species of the genus" masterLastPageNumber="538" masterPageNumber="501" pageNumber="503" updateTime="1698835840532" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2AF2F33F1187A66817A38CB6B9C9EB0A">Five new species of Coeliccia Kirby, 1890 from Vietnam (Odonata: Platycnemididae), and information on several other species of the genus</mods:title>
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<mods:namePart id="1C371E3E7100EB9ADD764C17A3D29B39">Kompier, Tom</mods:namePart>
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<mods:namePart id="A9A0C4EABC6EEC611E3A4D5EFEB8C330">Dow, Rory A.</mods:namePart>
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<mods:namePart id="7BA987D2717A60E1E29C51630F58007B">Steinhoff, Philip O. M.</mods:namePart>
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<mods:title id="4E8C7246285BCF5E3A7350C8516E0002">Zootaxa</mods:title>
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<mods:classification id="F7802A3E57188BB5119ADC870075D4B3">journal article</mods:classification>
<mods:identifier id="579418B8235C666DA8C8AB8B8EDEE2D9" type="CLB-Dataset">22814</mods:identifier>
<mods:identifier id="E5F670E96E1172C49F5297D68657BD80" type="DOI">10.11646/zootaxa.4766.4.1</mods:identifier>
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<treatment id="AF6A87D5436E6174FF6ADB91493D3830" ID-DOI="http://doi.org/10.5281/zenodo.3803426" ID-GBIF-Taxon="163739037" ID-Zenodo-Dep="3803426" LSID="urn:lsid:plazi:treatment:AF6A87D5436E6174FF6ADB91493D3830" httpUri="http://treatment.plazi.org/id/AF6A87D5436E6174FF6ADB91493D3830" lastPageId="4" lastPageNumber="505" pageId="2" pageNumber="503">
<subSubSection id="6FD96548436E6172FF6ADB9149F33AAF" box="[151,507,225,252]" pageId="2" pageNumber="503" type="nomenclature">
<paragraph id="277C36C3436E6172FF6ADB9149F33AAF" blockId="2.[151,982,225,288]" box="[151,507,225,252]" pageId="2" pageNumber="503">
<heading id="7C3481AF436E6172FF6ADB9149F33AAF" bold="true" box="[151,507,225,252]" fontSize="10" level="2" pageId="2" pageNumber="503" reason="2">
<emphasis id="15B7EAD1436E6172FF6ADB9149F33AAF" bold="true" box="[151,507,225,252]" pageId="2" pageNumber="503">
1.
<taxonomicName id="E0C34D40436E6172FF48DB91499C3AAF" ID-CoL="9X2BH" authority="Kompier &amp; Dow &amp; Steinhoff, 2020" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[181,404,225,252]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="caerulea" status="sp. nov.">
<emphasis id="15B7EAD1436E6172FF48DB91499C3AAF" bold="true" box="[181,404,225,252]" italics="true" pageId="2" pageNumber="503">Coeliccia caerulea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0E8457AA436E6172FE61DB9249F33AAF" box="[412,507,226,252]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="6FD96548436E6172FF6ADA754BDE3B73" box="[151,982,261,288]" pageId="2" pageNumber="503" type="description">
<paragraph id="277C36C3436E6172FF6ADA754BDE3B73" blockId="2.[151,982,225,288]" box="[151,982,261,288]" pageId="2" pageNumber="503">
(
<figureCitation id="BFF82A46436E6172FF62DA7548FB3B4C" box="[159,243,261,287]" captionStart="FIGURE 1" captionStartId="3.[151,250,1981,2006]" captionTargetBox="[151,1436,993,1956]" captionTargetId="figure@3.[151,1436,993,1956]" captionTargetPageId="3" captionText="FIGURE 1. Head of male Coeliccia spp. in dorsolateral view. (a) C. caerulea sp. nov., holotype; (b) C. coronata sp. nov., holotype; (c) C. diomedea sp. nov., holotype; (d) C. curua sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3765654" httpUri="https://zenodo.org/record/3765654/files/figure.png" pageId="2" pageNumber="503">Figs 1a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FEFDDA7549153B4C" box="[256,285,261,287]" captionStart="FIGURE 2" captionStartId="4.[151,250,1615,1640]" captionTargetBox="[151,1436,633,1590]" captionTargetId="figure@4.[151,1436,633,1590]" captionTargetPageId="4" captionText="FIGURE 2. Prothorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765656" httpUri="https://zenodo.org/record/3765656/files/figure.png" pageId="2" pageNumber="503">2a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FED4DA75494E3B4C" box="[297,326,261,287]" captionStart="FIGURE 3" captionStartId="6.[151,250,1882,1907]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure@6.[151,1436,417,1857]" captionTargetPageId="6" captionText="FIGURE 3. Prothorax of Coeliccia spp. in dorsal view. [a-e] male, [f] female. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype; (e) C. pulchella sp. nov., holotype; (f) C. caerulea, paratype used for description." figureDoi="http://doi.org/10.5281/zenodo.3765658" httpUri="https://zenodo.org/record/3765658/files/figure.png" pageId="2" pageNumber="503">3a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FEACDA7549653B4C" box="[337,365,261,287]" captionStart="FIGURE 4" captionStartId="7.[151,250,1090,1115]" captionTargetBox="[151,1437,181,1063]" captionTargetId="figure@7.[151,1437,181,1064]" captionTargetPageId="7" captionText="FIGURE 4. Thorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765660" httpUri="https://zenodo.org/record/3765660/files/figure.png" pageId="2" pageNumber="503">4a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FE84DA75499E3B73" box="[377,406,261,288]" captionStart="FIGURE 5" captionStartId="8.[151,250,910,935]" captionTargetBox="[151,1436,181,885]" captionTargetId="figure@8.[151,1436,181,885]" captionTargetPageId="8" captionText="FIGURE 5. Thorax of male Coeliccia spp. in dorsal view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765662" httpUri="https://zenodo.org/record/3765662/files/figure.png" pageId="2" pageNumber="503">5a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FE5FDA7549B83B4C" box="[418,432,261,287]" captionStart="FIGURE 7" captionStartId="9.[151,250,1813,1838]" captionTargetBox="[151,1436,181,1786]" captionTargetId="figure@9.[151,1436,181,1787]" captionTargetPageId="9" captionText="FIGURE 7. Abdomen tip of male Coeliccia spp. [a,e,g,i] lateral view, [c] dorsolateral view, [b,d,f,h,j] dorsal view. (a,b) C. caerulea, holotype; (c,d) C. pulchella, holotype; (e,f) C. curua, holotype; (g,h) C. coronata, holotype; (i,j) C. diomedea, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765666" httpUri="https://zenodo.org/record/3765666/files/figure.png" pageId="2" pageNumber="503">7</figureCitation>
a–b, 8e–f, 11g–h, 15a–b, 21d, 22f, 24a–b, 26a–c)
</paragraph>
</subSubSection>
<subSubSection id="6FD96548436E6172FF6ADA3E491F3BFC" pageId="2" pageNumber="503" type="materials_examined">
<paragraph id="277C36C3436E6172FF6ADA3E4D2C3B34" blockId="2.[151,1437,333,2016]" box="[151,1316,333,360]" pageId="2" pageNumber="503">
<materialsCitation id="97AB3C9E436E6172FF6ADA3E4D2C3B34" ID-GBIF-Occurrence="2608670277" box="[151,1316,333,360]" collectingDate="2016-06-13" elevation="700" latitude="11.449" location="Bao Loc" longLatPrecision="77" longitude="107.712" pageId="2" pageNumber="503" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong Prov." typeStatus="holotype">
<emphasis id="15B7EAD1436E6172FF6ADA3E49033B3B" bold="true" box="[151,267,334,360]" pageId="2" pageNumber="503">
<typeStatus id="F8788861436E6172FF6ADA3E490E3B3B" box="[151,262,334,360]" pageId="2" pageNumber="503" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="31C5FD4A436E6172FEEFDA3E49213B3B" box="[274,297,334,360]" pageId="2" pageNumber="503" type="male">♂</specimenCount>
,
<location id="221C6018436E6172FEC8DA3E499F3B3B" LSID="urn:lsid:plazi:treatment:AF6A87D5436E6174FF6ADB91493D3830:221C6018436E6172FEC8DA3E499F3B3B" box="[309,407,334,360]" latitude="11.449" longLatPrecision="77" longitude="107.712" name="Bao Loc" pageId="2" pageNumber="503" stateProvince="Lam Dong Prov.">Bao Loc</location>
,
<collectingRegion id="E507F821436E6172FE5EDA3E4A553B3B" box="[419,605,334,360]" country="Vietnam" name="Lam Dong" pageId="2" pageNumber="503">Lam Dong Prov.</collectingRegion>
(appr.
<geoCoordinate id="42F75004436E6172FD56DA3D4B063B34" box="[683,782,333,359]" degrees="11.449" direction="north" orientation="latitude" pageId="2" pageNumber="503" precision="55" value="11.449">11.449N</geoCoordinate>
,
<geoCoordinate id="42F75004436E6172FCE4DA3D4B813B34" box="[793,905,333,359]" degrees="107.712" direction="east" orientation="longitude" pageId="2" pageNumber="503" precision="55" value="107.712">107.712E</geoCoordinate>
; alt.
<elevation id="ACEED1F0436E6172FC43DA3D4C383B3B" box="[958,1072,333,360]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="503" unit="m" value="700.0">
<quantity id="E03B9B26436E6172FC43DA3D4C0D3B3B" box="[958,1029,333,360]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="503" unit="m" value="700.0">700 m</quantity>
asl
</elevation>
.),
<date id="537D1003436E6172FBB4DA3D4CCC3B34" box="[1097,1220,333,359]" pageId="2" pageNumber="503" value="2016-06-13">
<collectingDate id="4339E9EB436E6172FBB4DA3D4CCC3B34" box="[1097,1220,333,359]" pageId="2" pageNumber="503" value="2016-06-13">13 vi 2016</collectingDate>
</date>
, TK leg.
</materialsCitation>
</paragraph>
<paragraph id="277C36C3436E6172FF3ADA02491F3BFC" blockId="2.[151,1437,333,2016]" pageId="2" pageNumber="503">
<emphasis id="15B7EAD1436E6172FF3ADA02494E3BDF" bold="true" box="[199,326,370,396]" pageId="2" pageNumber="503">
<typeStatus id="F8788861436E6172FF3ADA0249493BDF" box="[199,321,370,396]" pageId="2" pageNumber="503" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="31C5FD4A436E6172FEB0DA0149863BDF" box="[333,398,369,396]" pageId="2" pageNumber="503" type="male">2 ♂♂</specimenCount>
, same date, location and collector as
<typeStatus id="F8788861436E6172FCCCDA014B9F3BD8" box="[817,919,369,395]" pageId="2" pageNumber="503" type="holotype">holotype</typeStatus>
;
<specimenCount id="31C5FD4A436E6172FC5FDA014BC43BDF" box="[930,972,369,396]" pageId="2" pageNumber="503" type="female">1 ♀</specimenCount>
, same location and collector as
<typeStatus id="F8788861436E6172FACEDA014D903BD8" box="[1331,1432,369,395]" pageId="2" pageNumber="503" type="holotype">holotype</typeStatus>
,
<date id="537D1003436E6172FF6ADAE5491B3BFC" box="[151,275,405,431]" pageId="2" pageNumber="503" value="2016-06-16">16 vi 2016</date>
.
</paragraph>
</subSubSection>
<subSubSection id="6FD96548436E6172FF3ADACA48F83BA4" pageId="2" pageNumber="503" type="etymology">
<paragraph id="277C36C3436E6172FF3ADACA48F83BA4" blockId="2.[151,1437,333,2016]" pageId="2" pageNumber="503">
<emphasis id="15B7EAD1436E6172FF3ADACA49403B87" bold="true" box="[199,328,442,468]" pageId="2" pageNumber="503">Etymology</emphasis>
. The adjective
<taxonomicName id="E0C34D40436E6172FE07DAC94A543B80" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[506,604,441,467]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis id="15B7EAD1436E6172FE07DAC94A543B80" box="[506,604,441,467]" italics="true" pageId="2" pageNumber="503">caerulea</emphasis>
</taxonomicName>
means blue in Latin and refers to the blue markings of the thorax of the species.
</paragraph>
</subSubSection>
<subSubSection id="6FD96548436E6173FF3AD9714B853848" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503" type="description">
<paragraph id="277C36C3436E6172FF3AD97149C138A0" blockId="2.[151,1437,333,2016]" pageId="2" pageNumber="503">
<emphasis id="15B7EAD1436E6172FF3AD97149E9384F" bold="true" box="[199,481,513,540]" pageId="2" pageNumber="503">
Description of
<typeStatus id="F8788861436E6172FE87D97249E9384F" box="[378,481,514,540]" pageId="2" pageNumber="503" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="BFF82A46436E6172FE0DD9714A4E3848" box="[496,582,513,539]" captionStart="FIGURE 1" captionStartId="3.[151,250,1981,2006]" captionTargetBox="[151,1436,993,1956]" captionTargetId="figure@3.[151,1436,993,1956]" captionTargetPageId="3" captionText="FIGURE 1. Head of male Coeliccia spp. in dorsolateral view. (a) C. caerulea sp. nov., holotype; (b) C. coronata sp. nov., holotype; (c) C. diomedea sp. nov., holotype; (d) C. curua sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3765654" httpUri="https://zenodo.org/record/3765654/files/figure.png" pageId="2" pageNumber="503">Figs 1a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FDA9D9714A793848" box="[596,625,513,539]" captionStart="FIGURE 2" captionStartId="4.[151,250,1615,1640]" captionTargetBox="[151,1436,633,1590]" captionTargetId="figure@4.[151,1436,633,1590]" captionTargetPageId="4" captionText="FIGURE 2. Prothorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765656" httpUri="https://zenodo.org/record/3765656/files/figure.png" pageId="2" pageNumber="503">2a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FD83D9714A933848" box="[638,667,513,539]" captionStart="FIGURE 3" captionStartId="6.[151,250,1882,1907]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure@6.[151,1436,417,1857]" captionTargetPageId="6" captionText="FIGURE 3. Prothorax of Coeliccia spp. in dorsal view. [a-e] male, [f] female. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype; (e) C. pulchella sp. nov., holotype; (f) C. caerulea, paratype used for description." figureDoi="http://doi.org/10.5281/zenodo.3765658" httpUri="https://zenodo.org/record/3765658/files/figure.png" pageId="2" pageNumber="503">3a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FD55D9714ACC3848" box="[680,708,513,539]" captionStart="FIGURE 4" captionStartId="7.[151,250,1090,1115]" captionTargetBox="[151,1437,181,1063]" captionTargetId="figure@7.[151,1437,181,1064]" captionTargetPageId="7" captionText="FIGURE 4. Thorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765660" httpUri="https://zenodo.org/record/3765660/files/figure.png" pageId="2" pageNumber="503">4a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FD2CD9714AE6384F" box="[721,750,513,540]" captionStart="FIGURE 5" captionStartId="8.[151,250,910,935]" captionTargetBox="[151,1436,181,885]" captionTargetId="figure@8.[151,1436,181,885]" captionTargetPageId="8" captionText="FIGURE 5. Thorax of male Coeliccia spp. in dorsal view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765662" httpUri="https://zenodo.org/record/3765662/files/figure.png" pageId="2" pageNumber="503">5a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FD06D9714B013848" box="[763,777,513,539]" captionStart="FIGURE 7" captionStartId="9.[151,250,1813,1838]" captionTargetBox="[151,1436,181,1786]" captionTargetId="figure@9.[151,1436,181,1787]" captionTargetPageId="9" captionText="FIGURE 7. Abdomen tip of male Coeliccia spp. [a,e,g,i] lateral view, [c] dorsolateral view, [b,d,f,h,j] dorsal view. (a,b) C. caerulea, holotype; (c,d) C. pulchella, holotype; (e,f) C. curua, holotype; (g,h) C. coronata, holotype; (i,j) C. diomedea, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765666" httpUri="https://zenodo.org/record/3765666/files/figure.png" pageId="2" pageNumber="503">7</figureCitation>
a–b, 8e–f).
<emphasis id="15B7EAD1436E6172FC76D9714BC03848" box="[907,968,513,539]" italics="true" pageId="2" pageNumber="503">Head</emphasis>
(
<figureCitation id="BFF82A46436E6172FC24D9714C223848" box="[985,1066,513,539]" captionStart="FIGURE 1" captionStartId="3.[151,250,1981,2006]" captionTargetBox="[151,1436,993,1956]" captionTargetId="figure@3.[151,1436,993,1956]" captionTargetPageId="3" captionText="FIGURE 1. Head of male Coeliccia spp. in dorsolateral view. (a) C. caerulea sp. nov., holotype; (b) C. coronata sp. nov., holotype; (c) C. diomedea sp. nov., holotype; (d) C. curua sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3765654" httpUri="https://zenodo.org/record/3765654/files/figure.png" pageId="2" pageNumber="503">Fig. 1a</figureCitation>
). Labrum and postclypeus shiny black. Anteclypeus cream bluish-white, with two vaguely darker areas left and right on lower half. Genae and base of mandibles whitish, the white narrowly extending inward over the antefrons above the postclypeus, but not connecting, remainder of frons matte black. Dorsal surface of head matte black, apart from two small white spots between base of antennae and lateral ocelli and two oblong white postocular spots. Antennae black, top of first segment and base of second white. Compound eyes three-colored in life, very dark chestnut above, green below, shading to blue posteriorly.
</paragraph>
<paragraph id="277C36C3436E6172FF3AD98D4AE5399F" blockId="2.[151,1437,333,2016]" pageId="2" pageNumber="503">
<emphasis id="15B7EAD1436E6172FF3AD98D49103944" box="[199,280,765,791]" italics="true" pageId="2" pageNumber="503">Thorax</emphasis>
(
<figureCitation id="BFF82A46436E6172FEDAD98D49753944" box="[295,381,765,791]" captionStart="FIGURE 2" captionStartId="4.[151,250,1615,1640]" captionTargetBox="[151,1436,633,1590]" captionTargetId="figure@4.[151,1436,633,1590]" captionTargetPageId="4" captionText="FIGURE 2. Prothorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765656" httpUri="https://zenodo.org/record/3765656/files/figure.png" pageId="2" pageNumber="503">Figs 2a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FE74D98D49AE3944" box="[393,422,765,791]" captionStart="FIGURE 3" captionStartId="6.[151,250,1882,1907]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure@6.[151,1436,417,1857]" captionTargetPageId="6" captionText="FIGURE 3. Prothorax of Coeliccia spp. in dorsal view. [a-e] male, [f] female. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype; (e) C. pulchella sp. nov., holotype; (f) C. caerulea, paratype used for description." figureDoi="http://doi.org/10.5281/zenodo.3765658" httpUri="https://zenodo.org/record/3765658/files/figure.png" pageId="2" pageNumber="503">3a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FE4CD98D49C63944" box="[433,462,765,791]" captionStart="FIGURE 4" captionStartId="7.[151,250,1090,1115]" captionTargetBox="[151,1437,181,1063]" captionTargetId="figure@7.[151,1437,181,1064]" captionTargetPageId="7" captionText="FIGURE 4. Thorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765660" httpUri="https://zenodo.org/record/3765660/files/figure.png" pageId="2" pageNumber="503">4a</figureCitation>
,
<figureCitation id="BFF82A46436E6172FE27D98D49F0394B" box="[474,504,765,792]" captionStart="FIGURE 5" captionStartId="8.[151,250,910,935]" captionTargetBox="[151,1436,181,885]" captionTargetId="figure@8.[151,1436,181,885]" captionTargetPageId="8" captionText="FIGURE 5. Thorax of male Coeliccia spp. in dorsal view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765662" httpUri="https://zenodo.org/record/3765662/files/figure.png" pageId="2" pageNumber="503">5a</figureCitation>
). Prothorax: pronotum matte black, propleuron whitish. Synthorax: mesepisternum black with distinct, slightly curved, blue antehumeral stripe following mesopleural suture from mesinfraepisternum to two-thirds length (
<figureCitation id="BFF82A46436E6172FE78D83549D03933" box="[389,472,837,864]" captionStart="FIGURE 5" captionStartId="8.[151,250,910,935]" captionTargetBox="[151,1436,181,885]" captionTargetId="figure@8.[151,1436,181,885]" captionTargetPageId="8" captionText="FIGURE 5. Thorax of male Coeliccia spp. in dorsal view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765662" httpUri="https://zenodo.org/record/3765662/files/figure.png" pageId="2" pageNumber="503">Fig. 5a</figureCitation>
). Mesepimeron and mesinfraepisternum black. Metepimeron and metepisternum blue with black stripe along entire metapleural suture and black adjacent to wing bases (
<figureCitation id="BFF82A46436E6172FBA9D8194CA039D0" box="[1108,1192,873,899]" captionStart="FIGURE 4" captionStartId="7.[151,250,1090,1115]" captionTargetBox="[151,1437,181,1063]" captionTargetId="figure@7.[151,1437,181,1064]" captionTargetPageId="7" captionText="FIGURE 4. Thorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765660" httpUri="https://zenodo.org/record/3765660/files/figure.png" pageId="2" pageNumber="503">Fig. 4a</figureCitation>
). Metinfraepisternum white, with thin black line adjacent to metepisternum. Legs: coxae and trochanters white. Femur and tibia pale grayish with black lines. Spines and claws brownish grey.
</paragraph>
<paragraph id="277C36C3436E6172FF3AD8A64BFA39BC" blockId="2.[151,1437,333,2016]" box="[199,1010,981,1008]" pageId="2" pageNumber="503">
<emphasis id="15B7EAD1436E6172FF3AD8A6490339BC" box="[199,267,982,1007]" italics="true" pageId="2" pageNumber="503">Wings</emphasis>
. Hyaline, 17 Px in FW,
<quantity id="E03B9B26436E6172FDE7D8A54A5B39BC" box="[538,595,981,1007]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="2" pageNumber="503" unit="in" value="16.0">16 in</quantity>
HW. Pt brown, covering 1–1.5 cells.
</paragraph>
<paragraph id="277C36C3436E6172FF3AD8894A663E0F" blockId="2.[151,1437,333,2016]" pageId="2" pageNumber="503">
<emphasis id="15B7EAD1436E6172FF3AD88949393E40" box="[199,305,1017,1043]" italics="true" pageId="2" pageNumber="503">Abdomen</emphasis>
. S1–2 black dorsally, white ventrally; S3–7 black dorsally and white ventrally, with subapical white annulus; S8 black dorsally, narrowly white ventrally; S9 black; S10 black, but white on dorsum with small irregular black dorsal marking apically (
<figureCitation id="BFF82A46436E6172FE0EDF314A3D3E0F" box="[499,565,1089,1116]" captionStart="FIGURE 7" captionStartId="9.[151,250,1813,1838]" captionTargetBox="[151,1436,181,1786]" captionTargetId="figure@9.[151,1436,181,1787]" captionTargetPageId="9" captionText="FIGURE 7. Abdomen tip of male Coeliccia spp. [a,e,g,i] lateral view, [c] dorsolateral view, [b,d,f,h,j] dorsal view. (a,b) C. caerulea, holotype; (c,d) C. pulchella, holotype; (e,f) C. curua, holotype; (g,h) C. coronata, holotype; (i,j) C. diomedea, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765666" httpUri="https://zenodo.org/record/3765666/files/figure.png" pageId="2" pageNumber="503">Fig. 7</figureCitation>
a–b).
</paragraph>
<paragraph id="277C36C3436E6172FF3ADF1548E73F43" blockId="2.[151,1437,333,2016]" pageId="2" pageNumber="503">
<emphasis id="15B7EAD1436E6172FF3ADF1549823E2C" box="[199,394,1125,1151]" italics="true" pageId="2" pageNumber="503">Anal appendages</emphasis>
(
<figureCitation id="BFF82A46436E6172FE67DF1549D53ED3" box="[410,477,1125,1152]" captionStart="FIGURE 8" captionStartId="10.[151,250,1840,1865]" captionTargetBox="[166,1421,181,1813]" captionTargetId="figure@10.[166,1421,181,1814]" captionTargetPageId="10" captionText="FIGURE 8. S10 and caudal appendages of male Coeliccia spp. [a,c,e,] lateral view, [g] dorsolateral view, [b,d,f,h] dorsal view. (a,b) C. coronata, holotype; (c,d) C. diomedea, holotype; (e,f) C. caerulea, holotype; (g,h) C. pulchella, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765668" httpUri="https://zenodo.org/record/3765668/files/figure.png" pageId="2" pageNumber="503">Fig. 8</figureCitation>
e–f). White. In lateral view (
<figureCitation id="BFF82A46436E6172FCDCDF154B793E2C" box="[801,881,1125,1151]" captionStart="FIGURE 8" captionStartId="10.[151,250,1840,1865]" captionTargetBox="[166,1421,181,1813]" captionTargetId="figure@10.[166,1421,181,1814]" captionTargetPageId="10" captionText="FIGURE 8. S10 and caudal appendages of male Coeliccia spp. [a,c,e,] lateral view, [g] dorsolateral view, [b,d,f,h] dorsal view. (a,b) C. coronata, holotype; (c,d) C. diomedea, holotype; (e,f) C. caerulea, holotype; (g,h) C. pulchella, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765668" httpUri="https://zenodo.org/record/3765668/files/figure.png" pageId="2" pageNumber="503">Fig. 8e</figureCitation>
) the cerci twice as thick as and distinctly shorter than paraprocts (about four-fifths of their length). Cerci with apex smoothly rounded and a ventral expansion, tipped with a black tooth, at three quarters of their length, also expanded on inner margin in this position in dorsal view (
<figureCitation id="BFF82A46436E6172FF5DDFA148E63EBF" box="[160,238,1233,1260]" captionStart="FIGURE 8" captionStartId="10.[151,250,1840,1865]" captionTargetBox="[166,1421,181,1813]" captionTargetId="figure@10.[166,1421,181,1814]" captionTargetPageId="10" captionText="FIGURE 8. S10 and caudal appendages of male Coeliccia spp. [a,c,e,] lateral view, [g] dorsolateral view, [b,d,f,h] dorsal view. (a,b) C. coronata, holotype; (c,d) C. diomedea, holotype; (e,f) C. caerulea, holotype; (g,h) C. pulchella, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765668" httpUri="https://zenodo.org/record/3765668/files/figure.png" pageId="2" pageNumber="503">Fig. 8f</figureCitation>
). No basal interior tooth visible. The paraprocts of usual
<typeStatus id="F8788861436E6172FC94DFA24B943EBF" box="[873,924,1234,1260]" pageId="2" pageNumber="503">type</typeStatus>
, apically curved inwards and with black tooth at apex.
</paragraph>
<paragraph id="277C36C3436E6172FF3ADE694CBB3F28" blockId="2.[151,1437,333,2016]" pageId="2" pageNumber="503">
<emphasis id="15B7EAD1436E6172FF3ADE69496B3F60" box="[199,355,1305,1331]" italics="true" pageId="2" pageNumber="503">Genital ligula</emphasis>
(as in
<figureCitation id="BFF82A46436E6172FE4DDE6949F53F60" box="[432,509,1305,1331]" captionStart="FIGURE 11" captionStartId="16.[151,250,1577,1602]" captionTargetBox="[158,1416,166,1551]" captionTargetId="figure@16.[158,1416,166,1551]" captionTargetPageId="16" captionText="FIGURE 11. Genital ligula of Coeliccia spp. [a,c,e,g] ventral view, [b,d,f,h] lateral view. (a,b) C. diomedea, holotype; (c,d) C. pulchella, holotype; (e,f) C. coronata, paratype, Da Lat, Lam Dong Prov., 16 iv 2016; (g,h) C. caerulea, paratype, flagellum missing on one side." figureDoi="http://doi.org/10.5281/zenodo.3765674" httpUri="https://zenodo.org/record/3765674/files/figure.png" pageId="2" pageNumber="503">Fig. 11</figureCitation>
g–h, which shows a
<typeStatus id="F8788861436E6172FD1EDE6A4B413F67" box="[739,841,1306,1332]" pageId="2" pageNumber="503" type="paratype">paratype</typeStatus>
). The terminal segment of the genital ligula with two apico-lateral flagella, originating laterally and angled backwards for apical half, and with an apical flap, covering the apical third of the segment, and becoming progressively broader towards its distal margin.
</paragraph>
<paragraph id="277C36C3436E6172FF3ADEF64B853FCC" blockId="2.[151,1437,333,2016]" box="[199,909,1413,1440]" pageId="2" pageNumber="503">Measurements (in mm). HW 26; abdomen incl. appendages 41.</paragraph>
<paragraph id="277C36C3436E6172FF3ADED949553C5F" blockId="2.[151,1437,333,2016]" pageId="2" pageNumber="503">
<emphasis id="15B7EAD1436E6172FF3ADED94A123F97" bold="true" box="[199,538,1449,1476]" pageId="2" pageNumber="503">
Variation in
<typeStatus id="F8788861436E6172FE9FDEDA49C53F97" box="[354,461,1450,1476]" pageId="2" pageNumber="503" type="paratype">paratype</typeStatus>
males
</emphasis>
. Middle pronotal lobe varies from completely black to adorned with a pair of round white spots. Dorsum of S10 almost entirely black in one. 17–18 Px in FW,
<quantity id="E03B9B26436E6172FBD5DEBD4C833FB4" box="[1064,1163,1485,1511]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.318" metricValueMin="4.064" pageId="2" pageNumber="503" unit="in" value="16.5" valueMax="17.0" valueMin="16.0">16–17 in</quantity>
HW. Abdomen incl. appendages
<quantity id="E03B9B26436E6172FEFBDE8149503C5F" box="[262,344,1521,1548]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="503" unit="mm" value="38.0">38 mm</quantity>
.
</paragraph>
<paragraph id="277C36C3436E6172FF3ADD654CD23D54" blockId="2.[151,1437,333,2016]" pageId="2" pageNumber="503">
<emphasis id="15B7EAD1436E6172FF3ADD654A323C63" bold="true" box="[199,570,1557,1584]" pageId="2" pageNumber="503">
Description of female
<typeStatus id="F8788861436E6172FE32DD664A323C63" box="[463,570,1558,1584]" pageId="2" pageNumber="503" type="paratype">paratype</typeStatus>
</emphasis>
(
<figureCitation id="BFF82A46436E6172FDB7DD654A953C7C" box="[586,669,1557,1583]" captionStart="FIGURE 3" captionStartId="6.[151,250,1882,1907]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure@6.[151,1436,417,1857]" captionTargetPageId="6" captionText="FIGURE 3. Prothorax of Coeliccia spp. in dorsal view. [a-e] male, [f] female. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype; (e) C. pulchella sp. nov., holotype; (f) C. caerulea, paratype used for description." figureDoi="http://doi.org/10.5281/zenodo.3765658" httpUri="https://zenodo.org/record/3765658/files/figure.png" pageId="2" pageNumber="503">Figs 3f</figureCitation>
,
<figureCitation id="BFF82A46436E6172FD54DD654AC03C63" box="[681,712,1557,1584]" captionStart="FIGURE 5" captionStartId="8.[151,250,910,935]" captionTargetBox="[151,1436,181,885]" captionTargetId="figure@8.[151,1436,181,885]" captionTargetPageId="8" captionText="FIGURE 5. Thorax of male Coeliccia spp. in dorsal view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765662" httpUri="https://zenodo.org/record/3765662/files/figure.png" pageId="2" pageNumber="503">5b</figureCitation>
,
<figureCitation id="BFF82A46436E6172FD29DD654AFA3C63" box="[724,754,1557,1584]" captionStart="FIGURE 15" captionStartId="20.[151,250,680,705]" captionTargetBox="[151,1436,273,655]" captionTargetId="figure@20.[151,1436,273,655]" captionTargetPageId="20" captionText="FIGURE 15. Female Coeliccia caerulea used for description. (a) head in frontal view; (b) prothorax in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3765682" httpUri="https://zenodo.org/record/3765682/files/figure.png" pageId="2" pageNumber="503">15</figureCitation>
,
<figureCitation id="BFF82A46436E6172FD02DD654B243C7C" box="[767,812,1557,1583]" captionStart="FIGURE 16" captionStartId="21.[151,250,1653,1678]" captionTargetBox="[151,1435,181,1627]" captionTargetId="figure@21.[151,1435,181,1627]" captionTargetPageId="21" captionText="FIGURE 16. Prothorax of female Coeliccia spp. used for description in lateral view. (a) C. coronata; (b) C. diomedea; (c) C. hayashii; (d) C. galbina; (e) C. pulchella; (f) C. suoitia." figureDoi="http://doi.org/10.5281/zenodo.3765684" httpUri="https://zenodo.org/record/3765684/files/figure.png" pageId="2" pageNumber="503">16d</figureCitation>
,
<figureCitation id="BFF82A46436E6172FCC5DD654B573C7C" box="[824,863,1557,1583]" captionStart="FIGURE 17" captionStartId="22.[151,250,1779,1804]" captionTargetBox="[151,1436,309,1754]" captionTargetId="figure@22.[151,1436,309,1754]" captionTargetPageId="22" captionText="FIGURE 17. Prothorax of female Coeliccia spp. used for description in dorsal view. (a) C. coronata; (b) C. diomedea; (c) C. hayashii; (d) C. galbina; (e) C. pulchella; (f) C. suoitia." figureDoi="http://doi.org/10.5281/zenodo.3765686" httpUri="https://zenodo.org/record/3765686/files/figure.png" pageId="2" pageNumber="503">17f</figureCitation>
,
<figureCitation id="BFF82A46436E6172FC91DD654B9F3C7C" box="[876,919,1557,1583]" captionStart="FIGURE 21" captionStartId="26.[151,250,1018,1043]" captionTargetBox="[151,1436,181,993]" captionTargetId="figure@26.[151,1436,181,993]" captionTargetPageId="26" captionText="FIGURE 21. Posterior pronotal lobe of Coeliccia spp. used for description in oblique dorsal view. (a) C. suoitia; (b) C. pulchella; (c) C. galbina; (d) C. caerulea." figureDoi="http://doi.org/10.5281/zenodo.3765694" httpUri="https://zenodo.org/record/3765694/files/figure.png" pageId="2" pageNumber="503">21d</figureCitation>
,
<figureCitation id="BFF82A46436E6172FC59DD654BC43C7C" box="[932,972,1557,1583]" captionStart="FIGURE 22" captionStartId="27.[151,250,1506,1531]" captionTargetBox="[157,1419,345,1481]" captionTargetId="figure@27.[157,1419,345,1481]" captionTargetPageId="27" captionText="FIGURE 22. Abdomen tip of female Coeliccia spp. used for description in lateral view. (a) C. coronata; (b) C. diomedea; (c) C. hayashii; (d) C. suoitia, covered in dirt; (e) C. galbina; (f) C. caerulea, alive in hand." figureDoi="http://doi.org/10.5281/zenodo.3765696" httpUri="https://zenodo.org/record/3765696/files/figure.png" pageId="2" pageNumber="503">22f</figureCitation>
,
<figureCitation id="BFF82A46436E6172FC25DD654C0D3C7C" box="[984,1029,1557,1583]" captionStart="FIGURE 24" captionStartId="29.[151,250,1493,1518]" captionTargetBox="[151,1436,181,1468]" captionTargetId="figure@29.[151,1436,181,1468]" captionTargetPageId="29" captionText="FIGURE 24. Thorax and head alive in hand of Coeliccia spp. [a,b] C. caerulea, [c,d] C. coronata, [e,f] C. diomedea, [a,c,e] male, [b,d,f] female. (a) holotype in lateral view; (b) female used for description in dorsolateral view; (c) paratype in dorsolateral view, Da Lat, Lam Dong Prov., 16 iv 2016; (d) female used for description in lateral view; (e) dorsolateral view, Quang Nam Prov., along QL14, 26 ix 2015; (f) female used for description in dorsolateral view." figureDoi="http://doi.org/10.5281/zenodo.3765700" httpUri="https://zenodo.org/record/3765700/files/figure.png" pageId="2" pageNumber="503">24b</figureCitation>
,
<figureCitation id="BFF82A46436E6172FBECDD654C353C63" box="[1041,1085,1557,1584]" captionStart="FIGURE 26" captionStartId="31.[151,250,1942,1967]" captionTargetBox="[151,1436,633,1917]" captionTargetId="figure@31.[151,1436,633,1917]" captionTargetPageId="31" captionText="FIGURE 26. Habitus of Coeliccia spp. from Bao Loc, Lam Dong Prov. [a-c] C. caerulea, [d-f] C. coronata (a) male, 13 vi 2016; (b) male, 18 v 2016; (c) female, 16 vi 2016; (d) female, 13 vi 2016; (e) male, 16 v 2016; (f) male, 16 v 2016." figureDoi="http://doi.org/10.5281/zenodo.3765704" httpUri="https://zenodo.org/record/3765704/files/figure.png" pageId="2" pageNumber="503">26c</figureCitation>
).
<emphasis id="15B7EAD1436E6172FBACDD654C863C7C" box="[1105,1166,1557,1583]" italics="true" pageId="2" pageNumber="503">Head</emphasis>
(
<figureCitation id="BFF82A46436E6172FB63DD654CF63C63" box="[1182,1278,1557,1584]" captionStart="FIGURE 15" captionStartId="20.[151,250,680,705]" captionTargetBox="[151,1436,273,655]" captionTargetId="figure@20.[151,1436,273,655]" captionTargetPageId="20" captionText="FIGURE 15. Female Coeliccia caerulea used for description. (a) head in frontal view; (b) prothorax in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3765682" httpUri="https://zenodo.org/record/3765682/files/figure.png" pageId="2" pageNumber="503">Fig. 15a</figureCitation>
). Labrum and postclypeus shiny black.Anteclypeus pale yellow, with two dark brown spots left and right on lower half. Genae and base of mandibles greenish yellow, this color continuing over the antefrons above the postclypeus about one-fourth inward, remainder of frons matte black. Dorsal surface of head matte black, apart from two irregular yellow lines from the edge of the compound eyes to the front of the lateral ocelli and continuing to the anterior ocellus, and two oblong greenish yellow postocular spots. Antennae black with top of first segment and base of second segment pale grey. Compound eyes in life blackish brown dorsally, green ventrally, shading to blue posteriorly.
</paragraph>
<paragraph id="277C36C3436E6173FF3ADC6149003A84" blockId="2.[151,1437,333,2016]" lastBlockId="3.[151,1437,153,971]" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503">
<emphasis id="15B7EAD1436E6172FF3ADC6149103D78" box="[199,280,1809,1835]" italics="true" pageId="2" pageNumber="503">Thorax</emphasis>
(
<figureCitation id="BFF82A46436E6172FED4DC6149743D78" box="[297,380,1809,1835]" captionStart="FIGURE 3" captionStartId="6.[151,250,1882,1907]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure@6.[151,1436,417,1857]" captionTargetPageId="6" captionText="FIGURE 3. Prothorax of Coeliccia spp. in dorsal view. [a-e] male, [f] female. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype; (e) C. pulchella sp. nov., holotype; (f) C. caerulea, paratype used for description." figureDoi="http://doi.org/10.5281/zenodo.3765658" httpUri="https://zenodo.org/record/3765658/files/figure.png" pageId="2" pageNumber="503">Figs 3f</figureCitation>
,
<figureCitation id="BFF82A46436E6172FE77DC6149BE3D7F" box="[394,438,1809,1836]" captionStart="FIGURE 15" captionStartId="20.[151,250,680,705]" captionTargetBox="[151,1436,273,655]" captionTargetId="figure@20.[151,1436,273,655]" captionTargetPageId="20" captionText="FIGURE 15. Female Coeliccia caerulea used for description. (a) head in frontal view; (b) prothorax in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3765682" httpUri="https://zenodo.org/record/3765682/files/figure.png" pageId="2" pageNumber="503">15b</figureCitation>
,
<figureCitation id="BFF82A46436E6172FE39DC6149E73D78" box="[452,495,1809,1835]" captionStart="FIGURE 21" captionStartId="26.[151,250,1018,1043]" captionTargetBox="[151,1436,181,993]" captionTargetId="figure@26.[151,1436,181,993]" captionTargetPageId="26" captionText="FIGURE 21. Posterior pronotal lobe of Coeliccia spp. used for description in oblique dorsal view. (a) C. suoitia; (b) C. pulchella; (c) C. galbina; (d) C. caerulea." figureDoi="http://doi.org/10.5281/zenodo.3765694" httpUri="https://zenodo.org/record/3765694/files/figure.png" pageId="2" pageNumber="503">21d</figureCitation>
,
<figureCitation id="BFF82A46436E6172FE00DC614A233D7F" box="[509,555,1809,1836]" captionStart="FIGURE 24" captionStartId="29.[151,250,1493,1518]" captionTargetBox="[151,1436,181,1468]" captionTargetId="figure@29.[151,1436,181,1468]" captionTargetPageId="29" captionText="FIGURE 24. Thorax and head alive in hand of Coeliccia spp. [a,b] C. caerulea, [c,d] C. coronata, [e,f] C. diomedea, [a,c,e] male, [b,d,f] female. (a) holotype in lateral view; (b) female used for description in dorsolateral view; (c) paratype in dorsolateral view, Da Lat, Lam Dong Prov., 16 iv 2016; (d) female used for description in lateral view; (e) dorsolateral view, Quang Nam Prov., along QL14, 26 ix 2015; (f) female used for description in dorsolateral view." figureDoi="http://doi.org/10.5281/zenodo.3765700" httpUri="https://zenodo.org/record/3765700/files/figure.png" pageId="2" pageNumber="503">24b</figureCitation>
). Prothorax: black, middle pronotal lobe with a rounded yellow spot laterally and propleuron yellow. Central part of posterior pronotal lobe semi-circular, not raised, broader at base than it is long, and two distinct lapels (
<figureCitation id="BFF82A46436E6172FE17DC294A363D20" box="[490,574,1881,1907]" captionStart="FIGURE 3" captionStartId="6.[151,250,1882,1907]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure@6.[151,1436,417,1857]" captionTargetPageId="6" captionText="FIGURE 3. Prothorax of Coeliccia spp. in dorsal view. [a-e] male, [f] female. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype; (e) C. pulchella sp. nov., holotype; (f) C. caerulea, paratype used for description." figureDoi="http://doi.org/10.5281/zenodo.3765658" httpUri="https://zenodo.org/record/3765658/files/figure.png" pageId="2" pageNumber="503">Figs 3f</figureCitation>
,
<figureCitation id="BFF82A46436E6172FDB1DC294A703D27" box="[588,632,1881,1908]" captionStart="FIGURE 15" captionStartId="20.[151,250,680,705]" captionTargetBox="[151,1436,273,655]" captionTargetId="figure@20.[151,1436,273,655]" captionTargetPageId="20" captionText="FIGURE 15. Female Coeliccia caerulea used for description. (a) head in frontal view; (b) prothorax in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3765682" httpUri="https://zenodo.org/record/3765682/files/figure.png" pageId="2" pageNumber="503">15b</figureCitation>
,
<figureCitation id="BFF82A46436E6172FD7BDC294ABB3D27" box="[646,691,1881,1908]" captionStart="FIGURE 21" captionStartId="26.[151,250,1018,1043]" captionTargetBox="[151,1436,181,993]" captionTargetId="figure@26.[151,1436,181,993]" captionTargetPageId="26" captionText="FIGURE 21. Posterior pronotal lobe of Coeliccia spp. used for description in oblique dorsal view. (a) C. suoitia; (b) C. pulchella; (c) C. galbina; (d) C. caerulea." figureDoi="http://doi.org/10.5281/zenodo.3765694" httpUri="https://zenodo.org/record/3765694/files/figure.png" pageId="2" pageNumber="503">21d</figureCitation>
). Synthorax: mesepisternum black, with a distinct yellowish blue antehumeral stripe along almost the complete length (
<figureCitation id="BFF82A46436E6172FD05DC0D4B543DCB" box="[760,860,1917,1944]" captionStart="FIGURE 23" captionStartId="28.[151,250,1893,1918]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure@28.[151,1436,181,1868]" captionTargetPageId="28" captionText="FIGURE 23. Holotype of Coeliccia montana Fraser, 1932. Photos courtesy of Dr. Ben Price, Natural History Museum, London. (a) thorax and head in lateral view; (b) abdomen tip and caudal appendages in lateral view; (c) thorax in dorsal view; (d) type on card with labels." figureDoi="http://doi.org/10.5281/zenodo.3765698" httpUri="https://zenodo.org/record/3765698/files/figure.png" pageId="2" pageNumber="503">Fig. 23b</figureCitation>
). Mesepimeron black, lower one-fourth bluish yellow and mesinfraepisternum black with some yellow bordering coxa. Metepisternum bluish yellow, narrowly black along wing base, and with distinct black line over metapleural suture. Metepimeron and metinfraepisternum green- ish yellow. Legs: Coxae and trochanters whitish yellow. Femur and tibia grey with black lines. Claws and spines dark grey.
</paragraph>
<paragraph id="277C36C3436F6173FF3ADB924BDE3AA8" blockId="3.[151,1437,153,971]" box="[199,982,224,251]" pageId="3" pageNumber="504">
<emphasis id="15B7EAD1436F6173FF3ADB9249033AA8" box="[199,267,226,251]" italics="true" pageId="3" pageNumber="504">Wings</emphasis>
. Hyaline, 19 Px in FW,
<quantity id="E03B9B26436F6173FDE7DB914A5B3AA8" box="[538,595,225,251]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="3" pageNumber="504" unit="in" value="18.0">18 in</quantity>
HW. Pt brown, covering 1.5 cells.
</paragraph>
<paragraph id="277C36C3436F6173FF3ADA754CA53BFC" blockId="3.[151,1437,153,971]" pageId="3" pageNumber="504">
<emphasis id="15B7EAD1436F6173FF3ADA7549393B4C" box="[199,305,261,287]" italics="true" pageId="3" pageNumber="504">Abdomen</emphasis>
(
<figureCitation id="BFF82A46436F6173FEBCDA7549AA3B4C" box="[321,418,261,287]" captionStart="FIGURE 22" captionStartId="27.[151,250,1506,1531]" captionTargetBox="[157,1419,345,1481]" captionTargetId="figure@27.[157,1419,345,1481]" captionTargetPageId="27" captionText="FIGURE 22. Abdomen tip of female Coeliccia spp. used for description in lateral view. (a) C. coronata; (b) C. diomedea; (c) C. hayashii; (d) C. suoitia, covered in dirt; (e) C. galbina; (f) C. caerulea, alive in hand." figureDoi="http://doi.org/10.5281/zenodo.3765696" httpUri="https://zenodo.org/record/3765696/files/figure.png" pageId="3" pageNumber="504">Figs 22f</figureCitation>
,
<figureCitation id="BFF82A46436F6173FE53DA7549D03B73" box="[430,472,261,288]" captionStart="FIGURE 26" captionStartId="31.[151,250,1942,1967]" captionTargetBox="[151,1436,633,1917]" captionTargetId="figure@31.[151,1436,633,1917]" captionTargetPageId="31" captionText="FIGURE 26. Habitus of Coeliccia spp. from Bao Loc, Lam Dong Prov. [a-c] C. caerulea, [d-f] C. coronata (a) male, 13 vi 2016; (b) male, 18 v 2016; (c) female, 16 vi 2016; (d) female, 13 vi 2016; (e) male, 16 v 2016; (f) male, 16 v 2016." figureDoi="http://doi.org/10.5281/zenodo.3765704" httpUri="https://zenodo.org/record/3765704/files/figure.png" pageId="3" pageNumber="504">26c</figureCitation>
). S1 yellow with posterior margin and dorsum narrowly black; S2 black dorsally and yellow ventrally, with apical black ring; S3 blackish brown dorsally and white ventrally, with a subbasal annulus; S4–7 blackish brown dorsally and whitish ventrally, with increasingly prominent subapical whitish annulus interrupted dorsally; S8 blackish brown on anterior two-thirds of dorsum, remainder yellowish white; S9–10 blackish brown, S9 with narrow pale dorsal line from anterior margin to about two-thirds of its length.
</paragraph>
<paragraph id="277C36C3436F6173FF3ADAC94A4F3BA4" blockId="3.[151,1437,153,971]" pageId="3" pageNumber="504">
<emphasis id="15B7EAD1436F6173FF3ADAC949823B80" box="[199,394,441,467]" italics="true" pageId="3" pageNumber="504">Anal appendages</emphasis>
(
<figureCitation id="BFF82A46436F6173FE66DAC949F03B87" box="[411,504,441,468]" captionStart="FIGURE 22" captionStartId="27.[151,250,1506,1531]" captionTargetBox="[157,1419,345,1481]" captionTargetId="figure@27.[157,1419,345,1481]" captionTargetPageId="27" captionText="FIGURE 22. Abdomen tip of female Coeliccia spp. used for description in lateral view. (a) C. coronata; (b) C. diomedea; (c) C. hayashii; (d) C. suoitia, covered in dirt; (e) C. galbina; (f) C. caerulea, alive in hand." figureDoi="http://doi.org/10.5281/zenodo.3765696" httpUri="https://zenodo.org/record/3765696/files/figure.png" pageId="3" pageNumber="504">Fig. 22f</figureCitation>
). Cerci blackish brown, half as long as S10, ovipositor blackish brown, extending almost the length of S10 beyond cerci.
</paragraph>
<paragraph id="277C36C3436F6173FF3AD9714B853848" blockId="3.[151,1437,153,971]" box="[199,909,513,539]" pageId="3" pageNumber="504">Measurements (in mm). HW 26; abdomen incl. appendages 37.</paragraph>
</subSubSection>
<subSubSection id="6FD96548436F6174FF3AD9544D053BA4" lastPageId="4" lastPageNumber="505" pageId="3" pageNumber="504" type="diagnosis">
<paragraph id="277C36C3436F6173FF3AD9544B9A3998" blockId="3.[151,1437,153,971]" pageId="3" pageNumber="504">
<emphasis id="15B7EAD1436F6173FF3AD95449C6386C" bold="true" box="[199,462,548,575]" pageId="3" pageNumber="504">Differential diagnosis.</emphasis>
The structure of the genital ligula, with apico-lateral flagella and apical flap (
<figureCitation id="BFF82A46436F6173FACCD9554D76386C" box="[1329,1406,549,575]" captionStart="FIGURE 11" captionStartId="16.[151,250,1577,1602]" captionTargetBox="[158,1416,166,1551]" captionTargetId="figure@16.[158,1416,166,1551]" captionTargetPageId="16" captionText="FIGURE 11. Genital ligula of Coeliccia spp. [a,c,e,g] ventral view, [b,d,f,h] lateral view. (a,b) C. diomedea, holotype; (c,d) C. pulchella, holotype; (e,f) C. coronata, paratype, Da Lat, Lam Dong Prov., 16 iv 2016; (g,h) C. caerulea, paratype, flagellum missing on one side." figureDoi="http://doi.org/10.5281/zenodo.3765674" httpUri="https://zenodo.org/record/3765674/files/figure.png" pageId="3" pageNumber="504">Fig. 11</figureCitation>
gh), places
<taxonomicName id="E0C34D40436F6173FEF4D93949D43830" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[265,476,585,611]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis id="15B7EAD1436F6173FEF4D93949D43830" box="[265,476,585,611]" italics="true" pageId="3" pageNumber="504">Coeliccia caerulea</emphasis>
</taxonomicName>
in the
<taxonomicName id="E0C34D40436F6173FDD5D9394A953830" authorityName="Laidlaw" authorityYear="1932" box="[552,669,585,611]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="15B7EAD1436F6173FDD5D9394A953830" box="[552,669,585,611]" italics="true" pageId="3" pageNumber="504">pyriformis</emphasis>
</taxonomicName>
-group. It differs clearly in coloration from
<emphasis id="15B7EAD1436F6173FB7DD9394D453830" box="[1152,1357,585,611]" italics="true" pageId="3" pageNumber="504">
<taxonomicName id="E0C34D40436F6173FB7DD9394CEC3830" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[1152,1252,585,611]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="curua">C. curua</taxonomicName>
,
<taxonomicName id="E0C34D40436F6173FB12D9394D453830" authorityName="Asahina" authorityYear="1997" box="[1263,1357,585,611]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="uenoi">C. uenoi</taxonomicName>
</emphasis>
and
<taxonomicName id="E0C34D40436F6173FA7ED93948F138D4" authorityName="Phan &amp; Tran" authorityYear="2018" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="phamiha">
<emphasis id="15B7EAD1436F6173FA7ED93948F138D4" italics="true" pageId="3" pageNumber="504">C. phamiha</emphasis>
</taxonomicName>
, which have large blue antehumeral shields, and in the case of
<taxonomicName id="E0C34D40436F6173FC38D91D4C2F38D4" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[965,1063,621,647]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="curua">
<emphasis id="15B7EAD1436F6173FC38D91D4C2F38D4" box="[965,1063,621,647]" italics="true" pageId="3" pageNumber="504">C. curua</emphasis>
</taxonomicName>
and
<taxonomicName id="E0C34D40436F6173FB9DD91D4CB738D4" authorityName="Asahina" authorityYear="1997" box="[1120,1215,621,647]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="15B7EAD1436F6173FB9DD91D4CB738D4" box="[1120,1215,621,647]" italics="true" pageId="3" pageNumber="504">C. uenoi</emphasis>
</taxonomicName>
yellow S9–10. The coloration of its thorax (
<figureCitation id="BFF82A46436F6173FE5AD9E149F438F8" box="[423,508,657,683]" captionStart="FIGURE 4" captionStartId="7.[151,250,1090,1115]" captionTargetBox="[151,1437,181,1063]" captionTargetId="figure@7.[151,1437,181,1064]" captionTargetPageId="7" captionText="FIGURE 4. Thorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765660" httpUri="https://zenodo.org/record/3765660/files/figure.png" pageId="3" pageNumber="504">Figs 4a</figureCitation>
,
<figureCitation id="BFF82A46436F6173FDF4D9E04A2D38F8" box="[521,549,656,683]" captionStart="FIGURE 5" captionStartId="8.[151,250,910,935]" captionTargetBox="[151,1436,181,885]" captionTargetId="figure@8.[151,1436,181,885]" captionTargetPageId="8" captionText="FIGURE 5. Thorax of male Coeliccia spp. in dorsal view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765662" httpUri="https://zenodo.org/record/3765662/files/figure.png" pageId="3" pageNumber="504">5a</figureCitation>
,
<figureCitation id="BFF82A46436F6173FDCCD9E14A4538FF" box="[561,589,657,684]" captionStart="FIGURE 26" captionStartId="31.[151,250,1942,1967]" captionTargetBox="[151,1436,633,1917]" captionTargetId="figure@31.[151,1436,633,1917]" captionTargetPageId="31" captionText="FIGURE 26. Habitus of Coeliccia spp. from Bao Loc, Lam Dong Prov. [a-c] C. caerulea, [d-f] C. coronata (a) male, 13 vi 2016; (b) male, 18 v 2016; (c) female, 16 vi 2016; (d) female, 13 vi 2016; (e) male, 16 v 2016; (f) male, 16 v 2016." figureDoi="http://doi.org/10.5281/zenodo.3765704" httpUri="https://zenodo.org/record/3765704/files/figure.png" pageId="3" pageNumber="504">26</figureCitation>
a–b) is more similar to
<taxonomicName id="E0C34D40436F6173FCAED9E14BEF38F8" authorityName="Kompier &amp; Phan" authorityYear="2017" box="[851,999,657,683]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="mientrung">
<emphasis id="15B7EAD1436F6173FCAED9E14BEF38F8" box="[851,999,657,683]" italics="true" pageId="3" pageNumber="504">C. mientrung</emphasis>
</taxonomicName>
and
<taxonomicName id="E0C34D40436F6173FBE3D9E14CBC38F8" authorityName="Laidlaw" authorityYear="1932" box="[1054,1204,657,683]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="15B7EAD1436F6173FBE3D9E14CBC38F8" box="[1054,1204,657,683]" italics="true" pageId="3" pageNumber="504">C. pyriformis</emphasis>
</taxonomicName>
. The slightly curved blue antehumeral stripes are similar to those of the generally larger
<emphasis id="15B7EAD1436F6173FC75D9C54C2C389C" box="[904,1060,693,719]" italics="true" pageId="3" pageNumber="504">
<taxonomicName id="E0C34D40436F6173FC75D9C54C28389C" authorityName="Laidlaw" authorityYear="1932" box="[904,1056,693,719]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="pyriformis">C. pyriformis</taxonomicName>
,
</emphasis>
whereas in
<taxonomicName id="E0C34D40436F6173FB54D9C54D34389C" authorityName="Kompier &amp; Phan" authorityYear="2017" box="[1193,1340,693,719]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="mientrung">
<emphasis id="15B7EAD1436F6173FB54D9C54D34389C" box="[1193,1340,693,719]" italics="true" pageId="3" pageNumber="504">C. mientrung</emphasis>
</taxonomicName>
they are clearly shorter (
<bibRefCitation id="43524B32436F6173FEB8D9A94A4038A0" author="Kompier, T. &amp; Phan, Q. T." box="[325,584,729,755]" pageId="3" pageNumber="504" pagination="131 - 140" refId="ref18689" refString="Kompier, T. &amp; Phan, Q. T. (2017) Coeliccia mientrung spec. nov. from Central Vietnam (Odonata: Platycnemididae). Zootaxa, 4247 (2), 131 - 140. https: // doi. org / 10.11646 / zootaxa. 4247.2.4" type="journal article" year="2017">Kompier &amp; Phan 2017</bibRefCitation>
). Both these species have yellow S9–10 and
<taxonomicName id="E0C34D40436F6173FBC5D9A94CC538A0" authorityName="Laidlaw" authorityYear="1932" box="[1080,1229,729,755]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="15B7EAD1436F6173FBC5D9A94CC538A0" box="[1080,1229,729,755]" italics="true" pageId="3" pageNumber="504">C. pyriformis</emphasis>
</taxonomicName>
also has bicolored compound eyes in life, lacking green. Its yellow cerci have the ventral expansion placed slightly further distally than in
<taxonomicName id="E0C34D40436F6173FF49D851493F3968" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[180,311,801,827]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis id="15B7EAD1436F6173FF49D851493F3968" box="[180,311,801,827]" italics="true" pageId="3" pageNumber="504">C. caerulea</emphasis>
</taxonomicName>
, where it is placed one-fourth of cerci length back from the apex. The genital ligula of
<taxonomicName id="E0C34D40436F6173FAFAD8514D953968" authorityName="Laidlaw" authorityYear="1932" box="[1287,1437,801,827]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="15B7EAD1436F6173FAFAD8514D953968" box="[1287,1437,801,827]" italics="true" pageId="3" pageNumber="504">C. pyriformis</emphasis>
</taxonomicName>
has flagella with hammerhead shaped apices. The male of
<taxonomicName id="E0C34D40436F6173FCE1D8354BA7390C" authorityName="Kompier &amp; Phan" authorityYear="2017" box="[796,943,837,863]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="mientrung">
<emphasis id="15B7EAD1436F6173FCE1D8354BA7390C" box="[796,943,837,863]" italics="true" pageId="3" pageNumber="504">C. mientrung</emphasis>
</taxonomicName>
has similar genital ligula as
<taxonomicName id="E0C34D40436F6173FB17D8354D63390C" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[1258,1387,837,863]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis id="15B7EAD1436F6173FB17D8354D63390C" box="[1258,1387,837,863]" italics="true" pageId="3" pageNumber="504">C. caerulea</emphasis>
</taxonomicName>
, but the ventral expansion of its cerci is placed even further distally than in
<taxonomicName id="E0C34D40436F6173FC4FD8194C4039D0" authorityName="Laidlaw" authorityYear="1932" box="[946,1096,873,899]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="15B7EAD1436F6173FC4FD8194C4039D0" box="[946,1096,873,899]" italics="true" pageId="3" pageNumber="504">C. pyriformis</emphasis>
</taxonomicName>
.
<taxonomicName id="E0C34D40436F6173FBABD8194D3339D0" authorityName="Kompier &amp; Phan" authorityYear="2017" box="[1110,1339,873,899]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="mientrung">
<emphasis id="15B7EAD1436F6173FBABD8194D3339D0" box="[1110,1339,873,899]" italics="true" pageId="3" pageNumber="504">Coeliccia mientrung</emphasis>
</taxonomicName>
also has larger white spots laterally on the middle pronotal lobe of the prothorax.
<taxonomicName id="E0C34D40436F6173FC38D8FD4CB439F4" authorityName="Ris" authorityYear="1912" box="[965,1212,909,935]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="cyanomelas">
<emphasis id="15B7EAD1436F6173FC38D8FD4CB439F4" box="[965,1212,909,935]" italics="true" pageId="3" pageNumber="504">Coeliccia cyanomelas</emphasis>
</taxonomicName>
has blue abdominal tip, different thoracic pattern and shape of the cerci (
<bibRefCitation id="43524B32436F6173FD1FD8C14B8E3998" author="Yu, X. &amp; Zhang, M. &amp; Ning, X." box="[738,902,945,971]" pageId="3" pageNumber="504" pagination="155 - 165" refId="ref19492" refString="Yu, X., Zhang, M. &amp; Ning, X. (2019) A study of Coeliccia cyanomelas Ris, 1912 (Odonata: Platycnemididiae). International Journal of Odonatology, 22 (3 - 4), 155 - 165. https: // doi. org / 10.1080 / 13887890.2019.1641853" type="journal article" year="2019">
Yu
<emphasis id="15B7EAD1436F6173FCF7D8C24B4A3998" box="[778,834,945,971]" italics="true" pageId="3" pageNumber="504">et al.</emphasis>
2019
</bibRefCitation>
).
</paragraph>
<caption id="73BC664B436F6173FF6ADCCD4B993DA9" ID-DOI="http://doi.org/10.5281/zenodo.3765654" ID-Zenodo-Dep="3765654" httpUri="https://zenodo.org/record/3765654/files/figure.png" pageId="3" pageNumber="504" startId="3.[151,250,1981,2006]" targetBox="[151,1436,993,1956]" targetPageId="3">
<paragraph id="277C36C3436F6173FF6ADCCD4B993DA9" blockId="3.[151,1436,1981,2042]" pageId="3" pageNumber="504">
<emphasis id="15B7EAD1436F6173FF6ADCCD491B3D85" bold="true" box="[151,275,1981,2006]" pageId="3" pageNumber="504">FIGURE 1.</emphasis>
Head of male
<taxonomicName id="E0C34D40436F6173FE54DCCD4A023D86" authorityName="Kirby" authorityYear="1890" box="[425,522,1981,2005]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="15B7EAD1436F6173FE54DCCD4A023D86" box="[425,522,1981,2005]" italics="true" pageId="3" pageNumber="504">Coeliccia</emphasis>
</taxonomicName>
spp. in dorsolateral view. (a)
<taxonomicName id="E0C34D40436F6173FCC5DCCE4BA73D86" authority="Kompier &amp; Dow &amp; Steinhoff, 2020" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[824,943,1981,2005]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="caerulea" status="sp. nov.">
<emphasis id="15B7EAD1436F6173FCC5DCCE4BA73D86" box="[824,943,1981,2005]" italics="true" pageId="3" pageNumber="504">C. caerulea</emphasis>
</taxonomicName>
<emphasis id="15B7EAD1436F6173FC48DCCE4C0E3D85" bold="true" box="[949,1030,1982,2006]" pageId="3" pageNumber="504">
<taxonomicNameLabel id="0E8457AA436F6173FC48DCCE4C0E3D85" box="[949,1030,1982,2006]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype; (b)
<taxonomicName id="E0C34D40436F6173FB63DCCE4D1E3D86" authority="Kompier &amp; Dow &amp; Steinhoff, 2020" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[1182,1302,1982,2005]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="coronata" status="sp. nov.">
<emphasis id="15B7EAD1436F6173FB63DCCE4D1E3D86" box="[1182,1302,1982,2005]" italics="true" pageId="3" pageNumber="504">C. coronata</emphasis>
</taxonomicName>
<emphasis id="15B7EAD1436F6173FAE1DCCE4D653D85" bold="true" box="[1308,1389,1982,2006]" pageId="3" pageNumber="504">
<taxonomicNameLabel id="0E8457AA436F6173FAE1DCCE4D653D85" box="[1308,1389,1982,2006]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype; (c)
<taxonomicName id="E0C34D40436F6173FEFBDC92498E3DAA" authority="Kompier &amp; Dow &amp; Steinhoff, 2020" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[262,390,2017,2041]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="diomedea" status="sp. nov.">
<emphasis id="15B7EAD1436F6173FEFBDC92498E3DAA" box="[262,390,2017,2041]" italics="true" pageId="3" pageNumber="504">C. diomedea</emphasis>
</taxonomicName>
<emphasis id="15B7EAD1436F6173FE70DC9249D53DA9" bold="true" box="[397,477,2018,2042]" pageId="3" pageNumber="504">
<taxonomicNameLabel id="0E8457AA436F6173FE70DC9249D53DA9" box="[397,477,2018,2042]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype; (d)
<taxonomicName id="E0C34D40436F6173FD89DC924AC53DAA" authority="Kompier &amp; Dow &amp; Steinhoff, 2020" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[628,717,2018,2041]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="curua" status="sp. nov.">
<emphasis id="15B7EAD1436F6173FD89DC924AC53DAA" box="[628,717,2018,2041]" italics="true" pageId="3" pageNumber="504">C. curua</emphasis>
</taxonomicName>
<emphasis id="15B7EAD1436F6173FD2EDC924B2C3DA9" bold="true" box="[723,804,2018,2042]" pageId="3" pageNumber="504">
<taxonomicNameLabel id="0E8457AA436F6173FD2EDC924B2C3DA9" box="[723,804,2018,2042]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype.
</paragraph>
</caption>
<paragraph id="277C36C343686174FF3ADBE94D053BA4" blockId="4.[151,1437,153,611]" pageId="4" pageNumber="505">
The female of
<taxonomicName id="E0C34D4043686174FE92DBE949FB3AE0" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[367,499,153,179]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis id="15B7EAD143686174FE92DBE949FB3AE0" box="[367,499,153,179]" italics="true" pageId="4" pageNumber="505">C. caerulea</emphasis>
</taxonomicName>
shares the general structure of the prothorax with the females of
<emphasis id="15B7EAD143686174FB20DBE948F13A84" italics="true" pageId="4" pageNumber="505">
<taxonomicName id="E0C34D4043686174FB20DBE94D7F3AE0" authorityName="Laidlaw" authorityYear="1932" box="[1245,1399,153,179]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="pyriformis">C. pyriformis</taxonomicName>
,
<taxonomicName id="E0C34D4043686174FA7EDBE948F13A84" authorityName="Phan &amp; Tran" authorityYear="2018" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="phamiha">C. phamiha</taxonomicName>
</emphasis>
and
<emphasis id="15B7EAD143686174FEC9DBCD49D93A84" box="[308,465,189,215]" italics="true" pageId="4" pageNumber="505">
<taxonomicName id="E0C34D4043686174FEC9DBCD49C63A84" authorityName="Kompier &amp; Phan" authorityYear="2017" box="[308,462,189,215]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="mientrung">C. mientrung</taxonomicName>
.
</emphasis>
The posterior pronotal lobe has a rounded central part (
<figureCitation id="BFF82A4643686174FBABDBCD4CB13A8B" box="[1110,1209,189,216]" captionStart="FIGURE 21" captionStartId="26.[151,250,1018,1043]" captionTargetBox="[151,1436,181,993]" captionTargetId="figure@26.[151,1436,181,993]" captionTargetPageId="26" captionText="FIGURE 21. Posterior pronotal lobe of Coeliccia spp. used for description in oblique dorsal view. (a) C. suoitia; (b) C. pulchella; (c) C. galbina; (d) C. caerulea." figureDoi="http://doi.org/10.5281/zenodo.3765694" httpUri="https://zenodo.org/record/3765694/files/figure.png" pageId="4" pageNumber="505">Fig. 21d</figureCitation>
), which is similarly rounded or more triangular in
<taxonomicName id="E0C34D4043686174FE14DB914A753AA8" authorityName="Kompier &amp; Phan" authorityYear="2017" box="[489,637,225,251]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="mientrung">
<emphasis id="15B7EAD143686174FE14DB914A753AA8" box="[489,637,225,251]" italics="true" pageId="4" pageNumber="505">C. mientrung</emphasis>
</taxonomicName>
and
<emphasis id="15B7EAD143686174FD49DB914B353AA8" box="[692,829,225,251]" italics="true" pageId="4" pageNumber="505">
<taxonomicName id="E0C34D4043686174FD49DB914B313AA8" authorityName="Phan &amp; Tran" authorityYear="2018" box="[692,825,225,251]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="phamiha">C. phamiha</taxonomicName>
,
</emphasis>
but squarish in
<taxonomicName id="E0C34D4043686174FC0DDB914C8E3AA8" authorityName="Laidlaw" authorityYear="1932" box="[1008,1158,225,251]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="15B7EAD143686174FC0DDB914C8E3AA8" box="[1008,1158,225,251]" italics="true" pageId="4" pageNumber="505">C. pyriformis</emphasis>
</taxonomicName>
. Unlike in these species, S9 is completely dark, without a prominent yellow dorsal spot (
<figureCitation id="BFF82A4643686174FCA1DA754BB23B73" box="[860,954,261,288]" captionStart="FIGURE 22" captionStartId="27.[151,250,1506,1531]" captionTargetBox="[157,1419,345,1481]" captionTargetId="figure@27.[157,1419,345,1481]" captionTargetPageId="27" captionText="FIGURE 22. Abdomen tip of female Coeliccia spp. used for description in lateral view. (a) C. coronata; (b) C. diomedea; (c) C. hayashii; (d) C. suoitia, covered in dirt; (e) C. galbina; (f) C. caerulea, alive in hand." figureDoi="http://doi.org/10.5281/zenodo.3765696" httpUri="https://zenodo.org/record/3765696/files/figure.png" pageId="4" pageNumber="505">Fig. 22f</figureCitation>
). In
<taxonomicName id="E0C34D4043686174FC17DA754C763B4C" authorityName="Kompier &amp; Phan" authorityYear="2017" box="[1002,1150,261,287]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="mientrung">
<emphasis id="15B7EAD143686174FC17DA754C763B4C" box="[1002,1150,261,287]" italics="true" pageId="4" pageNumber="505">C. mientrung</emphasis>
</taxonomicName>
the labrum is largely yellow, whereas in
<taxonomicName id="E0C34D4043686174FEB6DA5949E83B10" authorityName="Laidlaw" authorityYear="1932" box="[331,480,297,323]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="15B7EAD143686174FEB6DA5949E83B10" box="[331,480,297,323]" italics="true" pageId="4" pageNumber="505">C. pyriformis</emphasis>
</taxonomicName>
the yellow from the genae generally does not continue as far over the antefrons above the postclypeus. The yellow spot laterally on the middle pronotal lobe of the prothorax is much reduced compared to both
<taxonomicName id="E0C34D4043686174FF13DA01498A3BD8" authorityName="Kompier &amp; Phan" authorityYear="2017" box="[238,386,369,395]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="mientrung">
<emphasis id="15B7EAD143686174FF13DA01498A3BD8" box="[238,386,369,395]" italics="true" pageId="4" pageNumber="505">C. mientrung</emphasis>
</taxonomicName>
and
<taxonomicName id="E0C34D4043686174FE44DA014A473BD8" authorityName="Laidlaw" authorityYear="1932" box="[441,591,369,395]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="15B7EAD143686174FE44DA014A473BD8" box="[441,591,369,395]" italics="true" pageId="4" pageNumber="505">C. pyriformis</emphasis>
</taxonomicName>
, which are largely yellow laterally, whereas in
<taxonomicName id="E0C34D4043686174FB99DA014CEF3BD8" authorityName="Phan &amp; Tran" authorityYear="2018" box="[1124,1255,369,395]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="phamiha">
<emphasis id="15B7EAD143686174FB99DA014CEF3BD8" box="[1124,1255,369,395]" italics="true" pageId="4" pageNumber="505">C. phamiha</emphasis>
</taxonomicName>
the middle pronotal lobe appears to be completely yellow.
<taxonomicName id="E0C34D4043686174FD7CDAE54B393BFC" authorityName="Asahina" authorityYear="1997" box="[641,817,405,431]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="15B7EAD143686174FD7CDAE54B393BFC" box="[641,817,405,431]" italics="true" pageId="4" pageNumber="505">Coeliccia uenoi</emphasis>
</taxonomicName>
differs for instance by the shape of the extension of the posterior pronotal lobe, with has raised lateral corners (
<bibRefCitation id="43524B3243686174FD06DAC94B9F3B80" author="Asahina, S." box="[763,919,441,467]" pageId="4" pageNumber="505" pagination="17 - 34" refId="ref18407" refString="Asahina, S. (1997) Records of the Northern Vietnamese Odonata Taken by the Expedition Members from the Science Museum, Tokyo. 5. Coenagrionidae, Protoneuridae and Platycnemididae. Bulletin of the National Science Museum, Tokyo, Series A, 23 (1), 17 - 34." type="journal article" year="1997">Asahina 1997</bibRefCitation>
) and the largely yellow middle pronotal lobes.
<taxonomicName id="E0C34D4043686174FF6ADAAD49863BA4" authorityName="Ris" authorityYear="1912" box="[151,398,477,503]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="cyanomelas">
<emphasis id="15B7EAD143686174FF6ADAAD49863BA4" box="[151,398,477,503]" italics="true" pageId="4" pageNumber="505">Coeliccia cyanomelas</emphasis>
</taxonomicName>
has no central extension of note on the posterior pronotal lobe of the prothorax.
</paragraph>
</subSubSection>
<subSubSection id="6FD9654843686174FF3AD970493D3830" pageId="4" pageNumber="505" type="biology_ecology">
<paragraph id="277C36C343686174FF3AD970493D3830" blockId="4.[151,1437,153,611]" pageId="4" pageNumber="505">
<emphasis id="15B7EAD143686174FF3AD97049B43848" bold="true" box="[199,444,512,539]" pageId="4" pageNumber="505">Habitat and ecology.</emphasis>
<taxonomicName id="E0C34D4043686174FE3FD9714A9E3848" authority="Kompier &amp; Dow &amp; Steinhoff, 2020" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[450,662,513,539]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="caerulea" status="sp. nov.">
<emphasis id="15B7EAD143686174FE3FD9714A9E3848" box="[450,662,513,539]" italics="true" pageId="4" pageNumber="505">Coeliccia caerulea</emphasis>
</taxonomicName>
<emphasis id="15B7EAD143686174FD60D9714AFE3848" bold="true" box="[669,758,513,539]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="0E8457AA43686174FD60D9714AFE3848" box="[669,758,513,539]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found in degraded primary forest along a rocky and fast flowing mountain stream, along which
<taxonomicName id="E0C34D4043686174FDACD9554ADE386C" authority="Kompier &amp; Dow &amp; Steinhoff, 2020" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[593,726,549,575]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="coronata" status="sp. nov.">
<emphasis id="15B7EAD143686174FDACD9554ADE386C" box="[593,726,549,575]" italics="true" pageId="4" pageNumber="505">C. coronata</emphasis>
</taxonomicName>
<emphasis id="15B7EAD143686174FD20D9554B3F386C" bold="true" box="[733,823,549,575]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="0E8457AA43686174FD20D9554B3F386C" box="[733,823,549,575]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="E0C34D4043686174FCB8D9554CB3386C" authority="Kosterin, 2016" authorityName="Kosterin" authorityYear="2016" box="[837,1211,549,575]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="subSpecies" species="poungyi" subSpecies="dasha">
<emphasis id="15B7EAD143686174FCB8D9554C03386C" box="[837,1035,549,575]" italics="true" pageId="4" pageNumber="505">C. poungyi dasha</emphasis>
<bibRefCitation id="43524B3243686174FBEFD9554CB3386C" author="Kosterin, O. E." box="[1042,1211,549,575]" pageId="4" pageNumber="505" pagination="1 - 16" refId="ref18777" refString="Kosterin, O. E. (2016) Coeliccia poungyi dasha subsp. nov. (Odonata, Platycnemididae, Calicnemiinae) from eastern Cambodia. International Dragonfly Fund-Report, 97, 1 - 16." type="journal article" year="2016">Kosterin, 2016</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="E0C34D4043686174FB0AD9554D94386C" authorityName="Kosterin &amp; Kompier" authorityYear="2017" box="[1271,1436,549,575]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="rolandorum">
<emphasis id="15B7EAD143686174FB0AD9554D94386C" box="[1271,1436,549,575]" italics="true" pageId="4" pageNumber="505">C. rolandorum</emphasis>
</taxonomicName>
also occurred.
</paragraph>
</subSubSection>
</treatment>
</document>
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