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<document id="4A58AB3FFC1ECDDA7F1825B702111870" ID-DOI="10.11646/zootaxa.4037.1.1" ID-GBIF-Dataset="0dd60c86-99b2-43f8-a06e-18d012818103" ID-ISSN="1175-5326" ID-Zenodo-Dep="233519" ID-ZooBank="670819B1-3840-4C0A-ABFF-4D5AE3A263C0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461308665603" checkinUser="plazi" docAuthor="Kaczmarek, Łukasz, Bartels, Paul J., Roszkowska, Milena &amp; Nelson, Diane R." docDate="2015" docId="A5435063FF08FF06FF18F881FC37FD01" docLanguage="en" docName="zt04037p189.pdf" docOrigin="Zootaxa 4037 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Echiniscoides sigismundi subsp. hispaniensis Kristensen &amp; Hallas 1980" docType="treatment" docVersion="9" lastPageNumber="152" masterDocId="597A281BFF9EFF91FF8FFFBCFFFFFFCD" masterDocTitle="The Zoogeography of Marine Tardigrada" masterLastPageNumber="189" masterPageNumber="1" pageNumber="151" updateTime="1698618093469" updateUser="plazi">
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<mods:title id="569CB33EA39EC1CEFFB45A301440C896">The Zoogeography of Marine Tardigrada</mods:title>
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<mods:namePart id="2C0CFD819E34DA9BC454652143A507D0">Kaczmarek, Łukasz</mods:namePart>
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<mods:namePart id="ED1FEAA2BDC5673EC5C4D482A489CCED">Roszkowska, Milena</mods:namePart>
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<mods:namePart id="04E88B861C7F87AFF6D8CEA630F156C3">Nelson, Diane R.</mods:namePart>
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<mods:date id="A95A25D960FC30B20651A44DE40F1EF3">2015</mods:date>
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<subSubSection id="65F0B2FEFF08FF07FF18F881FC35F89B" box="[151,970,1853,1878]" pageId="150" pageNumber="151" type="nomenclature">
<paragraph id="2D55E175FF08FF07FF18F881FC35F89B" blockId="150.[151,1436,1853,2017]" box="[151,970,1853,1878]" pageId="150" pageNumber="151">
<heading id="761D5619FF08FF07FF18F881FC35F89B" bold="true" box="[151,970,1853,1878]" fontSize="10" level="2" pageId="150" pageNumber="151" reason="2">
<emphasis id="1F9E3D67FF08FF07FF18F881FC35F89B" bold="true" box="[151,970,1853,1878]" pageId="150" pageNumber="151">
187.
<taxonomicName id="EAEA9AF6FF08FF07FF40F881FC35F89B" ID-CoL="5H278" authority="Kristensen &amp; Hallas, 1980" authorityName="Kristensen &amp; Hallas" authorityYear="1980" box="[207,970,1853,1878]" class="Heterotardigrada" family="Echiniscoididae" genus="Echiniscoides" kingdom="Animalia" order="Echiniscoidea" pageId="150" pageNumber="151" phylum="Tardigrada" rank="subSpecies" species="sigismundi" subSpecies="hispaniensis">
<emphasis id="1F9E3D67FF08FF07FF40F881FD75F89B" bold="true" box="[207,650,1853,1878]" italics="true" pageId="150" pageNumber="151">Echiniscoides sigismundi hispaniensis</emphasis>
<bibRefCitation id="497B9C84FF08FF07FD1EF881FC35F89B" author="Kristensen" box="[657,970,1853,1878]" pageId="150" pageNumber="151" refString="Kristensen, R. M. &amp; Hallas, T. E. (1980) The tidal genus Echiniscoides and its variability, with erection of Echiniscoididae fam. n. (Tardigrada). Zoologica Scripta, 9, 113 - 127. http: // dx. doi. org / 10.1111 / j. 1463 - 6409.1980. tb 00657. x" type="journal article" year="1980">Kristensen &amp; Hallas, 1980</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
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<paragraph id="2D55E175FF08FF07FF40F8DDFD1AF853" blockId="150.[151,1436,1853,2017]" pageId="150" pageNumber="151">
<taxonomicName id="EAEA9AF6FF08FF07FF40F8DDFD82F8B7" box="[207,637,1889,1914]" class="Heterotardigrada" family="Echiniscoididae" genus="Echiniscoides" kingdom="Animalia" order="Echiniscoidea" pageId="150" pageNumber="151" phylum="Tardigrada" rank="subSpecies" species="sigismundi" status="subsp. nov." subSpecies="hispaniensis">
<emphasis id="1F9E3D67FF08FF07FF40F8DDFD82F8B7" box="[207,637,1889,1914]" italics="true" pageId="150" pageNumber="151">Echiniscoides sigismundi hispaniensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AD801CFF08FF07FD0BF8DDFD19F8B4" box="[644,742,1889,1914]" pageId="150" pageNumber="151" rank="subSpecies">subsp. n.</taxonomicNameLabel>
(
<bibRefCitation id="497B9C84FF08FF07FD7AF8DDFBE5F8B7" author="Kristensen" box="[757,1050,1889,1914]" pageId="150" pageNumber="151" refString="Kristensen, R. M. &amp; Hallas, T. E. (1980) The tidal genus Echiniscoides and its variability, with erection of Echiniscoididae fam. n. (Tardigrada). Zoologica Scripta, 9, 113 - 127. http: // dx. doi. org / 10.1111 / j. 1463 - 6409.1980. tb 00657. x" type="journal article" year="1980">Kristensen &amp; Hallas 1980</bibRefCitation>
)
<emphasis id="1F9E3D67FF08FF07FF40F83AFD1AF853" bold="true" box="[207,741,1925,1950]" pageId="150" pageNumber="151">
<taxonomicName id="EAEA9AF6FF08FF07FF40F83AFEA3F853" box="[207,348,1925,1950]" class="Insecta" family="Lycaenidae" genus="Terra" kingdom="Animalia" order="Lepidoptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="1F9E3D67FF08FF07FF40F83AFEA3F853" bold="true" box="[207,348,1925,1950]" italics="true" pageId="150" pageNumber="151">Terra typica</emphasis>
</taxonomicName>
: Atlantic Ocean (
<collectingCountry id="55FDA1E5FF08FF07FDA0F839FD88F853" box="[559,631,1925,1950]" name="Spain" pageId="150" pageNumber="151">Spain</collectingCountry>
, Europe)
</emphasis>
</paragraph>
<paragraph id="2D55E175FF08FF07FF18F816FECBF80E" blockId="150.[151,1436,1853,2017]" box="[151,308,1962,1987]" pageId="150" pageNumber="151">
<emphasis id="1F9E3D67FF08FF07FF18F816FECBF80E" bold="true" box="[151,308,1962,1987]" pageId="150" pageNumber="151">Adriatic Sea:</emphasis>
</paragraph>
<paragraph id="2D55E175FF08FF06FF18F875FDE5FF35" blockId="150.[151,1436,1853,2017]" lastBlockId="151.[151,1436,151,717]" lastPageId="151" lastPageNumber="152" pageId="150" pageNumber="151">
•
<geoCoordinate id="48DE87B2FF08FF07FF48F874FEDAF82C" box="[199,293,1992,2017]" direction="north" orientation="latitude" pageId="150" pageNumber="151" precision="925" value="42.1">
<emphasis id="1F9E3D67FF08FF07FF48F874FEF5F82C" bold="true" box="[199,266,1992,2017]" pageId="150" pageNumber="151">42°06</emphasis>
′
<emphasis id="1F9E3D67FF08FF07FE9FF874FEDAF82C" bold="true" box="[272,293,1992,2017]" pageId="150" pageNumber="151">N</emphasis>
</geoCoordinate>
,
<geoCoordinate id="48DE87B2FF08FF07FEBBF874FE6BF82C" box="[308,404,1992,2017]" direction="east" orientation="longitude" pageId="150" pageNumber="151" precision="925" value="15.483334">15°29′E</geoCoordinate>
;
<quantity id="EA124C90FF08FF07FE2CF874FE21F82C" box="[419,478,1992,2017]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="150" pageNumber="151" unit="m" value="10.0">10 m</quantity>
bsl: [FAO37]
<collectingCountry id="55FDA1E5FF08FF07FD04F874FD40F82C" box="[651,703,1992,2017]" name="Italy" pageId="150" pageNumber="151">Italy</collectingCountry>
, Foggia Province, Tremiti Islands, southern area of San Domino
<collectingCountry id="55FDA1E5FF09FF06FF46FF2BFEEFFF7D" box="[201,272,151,176]" name="Iceland" pageId="151" pageNumber="152">Island</collectingCountry>
, in the proximity of Punta Zio Cesare, submarine cave Grotta delle Viole, external cave’s entrance, coarse organogenic detritus.
<emphasis id="1F9E3D67FF09FF06FD83FF00FD1DFF18" bold="true" box="[524,738,188,213]" pageId="151" pageNumber="152">
D’Addabbo
<bibRefCitation id="497B9C84FF09FF06FD2FFF00FD1DFF18" box="[672,738,188,213]" pageId="151" pageNumber="152" refString="D'Addabbo Gallo, M., Grimaldi de Zio, S. &amp; Sanduli, R. (2001) Heterotardigrada of two submarine caves in S. Domino Island (Tremiti Islands) in the Mediterranean Sea with the description of two species of Stygarctidae. Zoologischer Anzeiger, 240, 361 - 369. http: // dx. doi. org / 10.1078 / 0044 - 5231 - 00043" type="journal article">Gallo</bibRefCitation>
</emphasis>
<emphasis id="1F9E3D67FF09FF06FD65FF01FCDCFF18" box="[746,803,188,213]" italics="true" pageId="151" pageNumber="152">et al.</emphasis>
<emphasis id="1F9E3D67FF09FF06FCA5FF00FE76FF35" bold="true" pageId="151" pageNumber="152">
(2001), Grimaldi de Zio &amp; D’Addabbo
<bibRefCitation id="497B9C84FF09FF06FA8EFF00FA69FF18" author="Gallo" box="[1281,1430,188,213]" pageId="151" pageNumber="152" refString="D'Addabbo Gallo, M., Grimaldi de Zio, S. &amp; Sanduli, R. (2001) Heterotardigrada of two submarine caves in S. Domino Island (Tremiti Islands) in the Mediterranean Sea with the description of two species of Stygarctidae. Zoologischer Anzeiger, 240, 361 - 369. http: // dx. doi. org / 10.1078 / 0044 - 5231 - 00043" type="journal article" year="2001">Gallo (2001)</bibRefCitation>
, Grimaldi de
<bibRefCitation id="497B9C84FF09FF06FEEEFF63FE76FF35" box="[353,393,223,248]" pageId="151" pageNumber="152" refString="Hansen, J. G., D'Addabbo Gallo, M. &amp; Grimaldi De Zio, S. (2003) A comparison of morphological characters within the genus Rhomboarctus (Tardigrada: Heterotardigrada) with the description of two new species. Zoologischer Anzeiger, (242) 1, 83 - 96. http: // dx. doi. org / 10.1078 / 0044 - 5231 - 00089" type="journal article">Zio</bibRefCitation>
</emphasis>
<emphasis id="1F9E3D67FF09FF06FE1EFF5DFE36FF35" box="[401,457,223,248]" italics="true" pageId="151" pageNumber="152">et al.</emphasis>
<emphasis id="1F9E3D67FF09FF06FE5FFF63FDE5FF35" bold="true" box="[464,538,223,248]" pageId="151" pageNumber="152">(2003)</emphasis>
</paragraph>
<paragraph id="2D55E175FF09FF06FF18FEB8FECBFED0" blockId="151.[151,1436,151,717]" box="[151,308,260,285]" pageId="151" pageNumber="152">
<emphasis id="1F9E3D67FF09FF06FF18FEB8FECBFED0" bold="true" box="[151,308,260,285]" pageId="151" pageNumber="152">Alboran Sea:</emphasis>
</paragraph>
<paragraph id="2D55E175FF09FF06FF18FE94FD04FEA8" blockId="151.[151,1436,151,717]" pageId="151" pageNumber="152">
•
<emphasis id="1F9E3D67FF09FF06FF48FE9BFE04FE8D" box="[199,507,295,320]" italics="true" pageId="151" pageNumber="152">
<geoCoordinate id="48DE87B2FF09FF06FF48FE9BFEDCFE8D" box="[199,291,295,320]" direction="north" orientation="latitude" pageId="151" pageNumber="152" precision="925" value="35.183334">35°11′N</geoCoordinate>
,
<geoCoordinate id="48DE87B2FF09FF06FEA0FE9BFE6EFE8D" box="[303,401,295,320]" direction="west" orientation="longitude" pageId="151" pageNumber="152" precision="925" value="-2.4333334">02°26′W</geoCoordinate>
; 0 m bsl:
</emphasis>
<emphasis id="1F9E3D67FF09FF06FD8CFE9BFD94FE8D" bold="true" box="[515,619,295,320]" pageId="151" pageNumber="152">[FAO37]</emphasis>
<collectingCountry id="55FDA1E5FF09FF06FDFDFE94FD25FEF2" box="[626,730,296,319]" name="Morocco" pageId="151" pageNumber="152">Morocco</collectingCountry>
, Chafarinas Islands, Isable II
<collectingCountry id="55FDA1E5FF09FF06FBA5FE9BFB91FE8D" box="[1066,1134,295,320]" name="Iceland" pageId="151" pageNumber="152">Island</collectingCountry>
[
<collectingCountry id="55FDA1E5FF09FF06FBF2FE9BFB57FE8D" box="[1149,1192,295,320]" name="Iceland" pageId="151" pageNumber="152">Isla</collectingCountry>
Isabel II], midlittoral pool.
<emphasis id="1F9E3D67FF09FF06FE87FEF0FD04FEA8" bold="true" box="[264,763,332,357]" pageId="151" pageNumber="152">
Villora-Moreno &amp; Grimaldi de
<bibRefCitation id="497B9C84FF09FF06FD0EFEF0FD04FEA8" author="Zio" box="[641,763,332,357]" pageId="151" pageNumber="152" refString="Villora-Moreno, S. &amp; Grimaldi de Zio, S. (1996) New records of marine Tardigrada in the Mediterranean Sea. Zoological Journal of the Linnean Society, 116, 149 - 166. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1996. tb 02340. x" type="journal article" year="1996">Zio (1996)</bibRefCitation>
</emphasis>
</paragraph>
<paragraph id="2D55E175FF09FF06FF18FECCFC87FE60" blockId="151.[151,1436,151,717]" pageId="151" pageNumber="152">
•
<emphasis id="1F9E3D67FF09FF06FF48FED3FDF1FE45" box="[199,526,367,392]" italics="true" pageId="151" pageNumber="152">
<geoCoordinate id="48DE87B2FF09FF06FF48FED3FEDCFE45" box="[199,291,367,392]" direction="north" orientation="latitude" pageId="151" pageNumber="152" precision="925" value="35.183334">35°11′N</geoCoordinate>
,
<geoCoordinate id="48DE87B2FF09FF06FEBBFED3FE6AFE45" box="[308,405,367,392]" direction="west" orientation="longitude" pageId="151" pageNumber="152" precision="925" value="-2.4166667">02°25′W</geoCoordinate>
; 0 m bsl:
</emphasis>
<emphasis id="1F9E3D67FF09FF06FD95FED3FD7EFE45" bold="true" box="[538,641,367,392]" pageId="151" pageNumber="152">[FAO37]</emphasis>
<collectingCountry id="55FDA1E5FF09FF06FD01FECCFD09FE4A" box="[654,758,368,391]" name="Morocco" pageId="151" pageNumber="152">Morocco</collectingCountry>
, Chafarinas Islands, Rey Francisco
<collectingCountry id="55FDA1E5FF09FF06FB2EFED3FB1AFE45" box="[1185,1253,367,392]" name="Iceland" pageId="151" pageNumber="152">Island</collectingCountry>
[
<collectingCountry id="55FDA1E5FF09FF06FB76FED3FADBFE45" box="[1273,1316,367,392]" name="Iceland" pageId="151" pageNumber="152">Isla</collectingCountry>
del Rey], midlittoral pool.
<emphasis id="1F9E3D67FF09FF06FE09FE28FC87FE60" bold="true" box="[390,888,404,429]" pageId="151" pageNumber="152">
Villora-Moreno &amp; Grimaldi de
<bibRefCitation id="497B9C84FF09FF06FD71FE28FC87FE60" author="Zio" box="[766,888,404,429]" pageId="151" pageNumber="152" refString="Villora-Moreno, S. &amp; Grimaldi de Zio, S. (1996) New records of marine Tardigrada in the Mediterranean Sea. Zoological Journal of the Linnean Society, 116, 149 - 166. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1996. tb 02340. x" type="journal article" year="1996">Zio (1996)</bibRefCitation>
</emphasis>
</paragraph>
<paragraph id="2D55E175FF09FF06FF18FE0BFEABFE1D" blockId="151.[151,1436,151,717]" box="[151,340,439,464]" pageId="151" pageNumber="152">
<emphasis id="1F9E3D67FF09FF06FF18FE0BFEABFE1D" bold="true" box="[151,340,439,464]" pageId="151" pageNumber="152">Atlantic Ocean:</emphasis>
</paragraph>
<paragraph id="2D55E175FF09FF06FF18FE61FB10FDD5" blockId="151.[151,1436,151,717]" pageId="151" pageNumber="152">
•
<emphasis id="1F9E3D67FF09FF06FF48FE60FE07FE38" box="[199,504,476,501]" italics="true" pageId="151" pageNumber="152">
<geoCoordinate id="48DE87B2FF09FF06FF48FE60FED9FE38" box="[199,294,476,501]" direction="north" orientation="latitude" pageId="151" pageNumber="152" precision="925" value="43.35">43°21′N</geoCoordinate>
,
<geoCoordinate id="48DE87B2FF09FF06FEBFFE60FE6DFE38" box="[304,402,476,501]" direction="west" orientation="longitude" pageId="151" pageNumber="152" precision="925" value="-8.25">08°15′W</geoCoordinate>
; 0 m bsl:
</emphasis>
<emphasis id="1F9E3D67FF09FF06FE71FE60FCE6FE38" bold="true" box="[510,793,476,501]" pageId="151" pageNumber="152">
[FAO27]
<typeStatus id="F2515FD7FF09FF06FDE2FE60FD57FE38" box="[621,680,476,501]" pageId="151" pageNumber="152">Type</typeStatus>
Locality:
</emphasis>
<collectingCountry id="55FDA1E5FF09FF06FCAFFE60FC9CFE38" box="[800,867,476,501]" name="Spain" pageId="151" pageNumber="152">Spain</collectingCountry>
, Coruña Province, near La Coruña, Sada, mid-tidal zone on sandy beach, barnacles (
<emphasis id="1F9E3D67FF09FF06FDB6FE43FCEBFDD5" box="[569,788,511,536]" italics="true" pageId="151" pageNumber="152">Eliminius modestus</emphasis>
) from stone.
<bibRefCitation id="497B9C84FF09FF06FC24FE43FB10FDD5" author="Kristensen" box="[939,1263,511,536]" pageId="151" pageNumber="152" refString="Kristensen, R. M. &amp; Hallas, T. E. (1980) The tidal genus Echiniscoides and its variability, with erection of Echiniscoididae fam. n. (Tardigrada). Zoologica Scripta, 9, 113 - 127. http: // dx. doi. org / 10.1111 / j. 1463 - 6409.1980. tb 00657. x" type="journal article" year="1980">
<emphasis id="1F9E3D67FF09FF06FC24FE43FB10FDD5" bold="true" box="[939,1263,511,536]" pageId="151" pageNumber="152">Kristensen &amp; Hallas (1980)</emphasis>
</bibRefCitation>
</paragraph>
<paragraph id="2D55E175FF09FF06FF18FD98FABBFDF0" blockId="151.[151,1436,151,717]" box="[151,1348,548,573]" pageId="151" pageNumber="152">
<emphasis id="1F9E3D67FF09FF06FF18FD98FD5EFDF0" bold="true" box="[151,673,548,573]" pageId="151" pageNumber="152">Record numbers (Sea/Ocean classification):</emphasis>
Adriatic Sea: 1, Alboran Sea: 2, Atlantic Ocean: 1;
<emphasis id="1F9E3D67FF09FF06FB67FD98FABBFDF0" bold="true" box="[1256,1348,548,573]" pageId="151" pageNumber="152">total: 4.</emphasis>
</paragraph>
<paragraph id="2D55E175FF09FF06FF18FDFBFC46FDAD" blockId="151.[151,1436,151,717]" box="[151,953,583,608]" pageId="151" pageNumber="152">
<emphasis id="1F9E3D67FF09FF06FF18FDFBFDA3FDAD" bold="true" box="[151,604,583,608]" pageId="151" pageNumber="152">Record numbers (FAO classification):</emphasis>
FAO27: 1, FAO37: 3;
<emphasis id="1F9E3D67FF09FF06FCD2FDFBFC46FDAD" bold="true" box="[861,953,583,608]" pageId="151" pageNumber="152">total: 4.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="65F0B2FEFF09FF06FF18FDD0FC37FD01" pageId="151" pageNumber="152" type="discussion">
<paragraph id="2D55E175FF09FF06FF18FDD0FC37FD01" blockId="151.[151,1436,151,717]" pageId="151" pageNumber="152">
<emphasis id="1F9E3D67FF09FF06FF18FDD0FEF2FD48" bold="true" box="[151,269,620,645]" pageId="151" pageNumber="152">Remarks:</emphasis>
The original description of this subspecies was from the Atlantic coast of
<collectingCountry id="55FDA1E5FF09FF06FBEDFDD0FB5DFD48" box="[1122,1186,620,645]" name="Spain" pageId="151" pageNumber="152">Spain</collectingCountry>
from barnacles in the mid-tidal zone. Later references include warm waters of the Mediterranean, where it has been collected in sand down to
<quantity id="EA124C90FF09FF06FEA4FD08FE9BFD01" box="[299,356,692,716]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="151" pageNumber="152" unit="m" value="10.0">10 m</quantity>
bsl (see also remarks for
<taxonomicName id="EAEA9AF6FF09FF06FD0CFD08FC47FD00" box="[643,952,692,717]" class="Heterotardigrada" family="Echiniscoididae" genus="Echiniscoides" kingdom="Animalia" order="Echiniscoidea" pageId="151" pageNumber="152" phylum="Tardigrada" rank="subSpecies" species="sigismundi" subSpecies="sigismundi">
<emphasis id="1F9E3D67FF09FF06FD0CFD08FC47FD00" box="[643,952,692,717]" italics="true" pageId="151" pageNumber="152">Echiniscoides s. sigismundi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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