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<mods:title id="0A6CEC7AB9DF839746F10523D91C6A47">A taxonomic revision of the genus Arcynopteryx Klapálek, 1904 (Plecoptera, Perlodidae)</mods:title>
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<taxonomicName id="C1B94D3CFFF0897DFF2D1EF8A0C2FF17" ID-CoL="GC98" authority="McLachlan, 1872" authorityName="McLachlan" authorityYear="1872" box="[151,657,151,177]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF0897DFF2D1EF8A0C2FF17" bold="true" box="[151,657,151,177]" pageId="1" pageNumber="2">
<emphasis id="34CDEAADFFF0897DFF2D1EF8A3CDFF17" bold="true" box="[151,414,151,177]" italics="true" pageId="1" pageNumber="2">Arcynopteryx dichroa</emphasis>
(
<bibRefCitation id="62284B4EFFF0897DFE171EF8A0DBFF17" author="McLachlan" box="[429,648,151,177]" pageId="1" pageNumber="2" refString="McLachlan, R. (1872) Perlides. In: Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps &amp; McLachlan. Annales de la Societe entomologique de Belgique (Bruxelles), 15, 47 - 55." type="journal article" year="1872">McLachlan, 1872</bibRefCitation>
)
</emphasis>
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</heading>
</paragraph>
<paragraph id="060636BFFFF0897DFF2D1ED5A34AFF74" blockId="1.[151,657,151,210]" box="[151,281,186,210]" pageId="1" pageNumber="2">
(
<figureCitation id="9E822A3AFFF0897DFF251ED5A343FF74" box="[159,272,186,210]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 8" captionStart-2="FIGURES 9 – 12" captionStartId-0="2.[151,260,1441,1464]" captionStartId-1="3.[151,264,927,950]" captionStartId-2="3.[151,264,1946,1969]" captionTargetBox-0="[264,1317,602,1408]" captionTargetBox-1="[165,1404,210,889]" captionTargetBox-2="[264,1319,1032,1898]" captionTargetId-0="figure@2.[264,1323,593,1417]" captionTargetId-1="figure@3.[151,1436,193,906]" captionTargetId-2="figure@3.[264,1323,1032,1922]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." captionText-1="FIGURES 7 – 8. Arcynopteryx dichroa, type: 7. Male abdominal tip, dorsolateral, pinned. 8. Same, dorsal, cleaned. (Photograph by D. Goodger)." captionText-2="FIGURES 9 – 12. Arcynopteryx dichroa, egg: 9. Habitus: anchor, chorion structure, row of micropyles, lateral. 10. Collar, lateral. 11. Micropylar orifices and mounds, dorsal view. 12. Chorion structure, dorsal view." httpUri-0="https://zenodo.org/record/210961/files/figure.png" httpUri-1="https://zenodo.org/record/210962/files/figure.png" httpUri-2="https://zenodo.org/record/210963/files/figure.png" pageId="1" pageNumber="2">Figs 1–12</figureCitation>
)
</paragraph>
<paragraph id="060636BFFFF0897DFF2D1F6DA30FFE56" blockId="1.[151,1436,257,496]" pageId="1" pageNumber="2">
<bibRefCitation id="62284B4EFFF0897DFF2D1F6DA310FEBE" author="McLachlan" box="[151,323,258,280]" pageId="1" pageNumber="2" refString="McLachlan, R. (1872) Perlides. In: Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps &amp; McLachlan. Annales de la Societe entomologique de Belgique (Bruxelles), 15, 47 - 55." type="journal article" year="1872">McLachlan 1872</bibRefCitation>
: 52−53, pl. I, figs 4−4a, 5−5b (
<taxonomicName id="C1B94D3CFFF0897DFD3A1F6CA104FEBE" box="[640,855,258,280]" class="Insecta" family="Perlodidae" genus="Dictyopteryx" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF0897DFD3A1F6CA104FEBE" box="[640,855,258,280]" italics="true" pageId="1" pageNumber="2">Dictyopteryx dichroa</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFF0897DFCD61F6DA657FEBE" author="Klapalek" box="[876,1028,258,280]" pageId="1" pageNumber="2" refString="Klapalek, F. (1912) Plecopteres I. Fam. Perlodidae. Collections Zoologiques du Baron Edmond de Selys-Longchamps. Bruxelles, 4, 1 - 66." type="journal article" year="1912">Klapálek 1912</bibRefCitation>
: 14–16, fig. 8 (
<taxonomicName id="C1B94D3CFFF0897DFB261F6CA7DFFEBE" box="[1180,1420,259,280]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFF0897DFB261F6CA7DFFEBE" box="[1180,1420,259,280]" italics="true" pageId="1" pageNumber="2">Arcynopteryx compacta</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFF0897DFF7D1F4EA318FE91" author="Brinck" box="[199,331,289,311]" pageId="1" pageNumber="2" refString="Brinck, P. (1949) Studies on Swedish stoneflies (Plecoptera). Opuscula Entomologica, 11, Lund, 250 pp." type="book" year="1949">Brinck 1949</bibRefCitation>
: 58−60, figs 4 A–F (
<taxonomicName id="C1B94D3CFFF0897DFD971F4DA173FE91" box="[557,800,290,311]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFF0897DFD971F4DA173FE91" box="[557,800,290,311]" italics="true" pageId="1" pageNumber="2">Arcynopteryx compacta</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFF0897DFC801F4EA1ECFE91" author="Brinck" box="[826,959,289,311]" pageId="1" pageNumber="2" refString="Brinck, P. (1956) Reproductive system and mating in Plecoptera. Opuscula Entomologica, 21, 57 - 127." type="journal article" year="1956">Brinck 1956</bibRefCitation>
: 66–71, figs 6A–D (
<taxonomicName id="C1B94D3CFFF0897DFB231F4DA7DFFE91" box="[1177,1420,290,311]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFF0897DFB231F4DA7DFFE91" box="[1177,1420,290,311]" italics="true" pageId="1" pageNumber="2">Arcynopteryx compacta</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFF0897DFF7D1F2FA335FEF0" author="Zhiltzova" box="[199,358,320,342]" pageId="1" pageNumber="2" refString="Zhiltzova, L. (1966) The stoneflies (Plecoptera) of the European part of USSR (without Caucasus). Entomological Review, 45, 525 - 549 (in Russian)." type="journal article" year="1966">Zhiltzova 1966</bibRefCitation>
: 539 (
<taxonomicName id="C1B94D3CFFF0897DFE101F2EA0CFFEF0" box="[426,668,321,342]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFF0897DFE101F2EA0CFFEF0" box="[426,668,321,342]" italics="true" pageId="1" pageNumber="2">Arcynopteryx compacta</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFF0897DFD0E1F2FA176FEF0" author="Illies" box="[692,805,320,342]" pageId="1" pageNumber="2" refString="Illies, J. (1966) Katalog der rezenten Plecoptera. Das Tierreich, Berlin, 82, 632 s." type="journal article" year="1966">Illies 1966</bibRefCitation>
: 353 (
<taxonomicName id="C1B94D3CFFF0897DFCD31F2EA614FEF0" box="[873,1095,320,342]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF0897DFCD31F2EA614FEF0" box="[873,1095,320,342]" italics="true" pageId="1" pageNumber="2">Arcynopteryx dichroa</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFF0897DFBE41F2FA6A8FEF3" author="Kimmins" box="[1118,1275,320,342]" pageId="1" pageNumber="2" refString="Kimmins, D. (1970) A list of the type-specimens of Plecoptera and Megaloptera in the British Museum (Natural History). Bulletin of the British Museum (Natural History) Entomology, 24, 335 - 361." type="journal article" year="1970">Kimmins 1970</bibRefCitation>
: 340 (
<taxonomicName id="C1B94D3CFFF0897DFA851F2EA31DFED3" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF0897DFA851F2EA31DFED3" italics="true" pageId="1" pageNumber="2">Arcynopteryx dichroa</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFF0897DFEDF1F31A3B7FED2" author="Zwick" box="[357,484,350,372]" pageId="1" pageNumber="2" refString="Zwick, P. (1973) Insecta: Plecoptera. Phylogenetisches System und Katalog. Das Tierreich. Walter de Gruyter, Berlin, 94, ixxxii + 465 pp." type="book" year="1973">Zwick 1973</bibRefCitation>
: 224 (
<taxonomicName id="C1B94D3CFFF0897DFD9E1F30A145FED3" box="[548,790,351,373]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFF0897DFD9E1F30A145FED3" box="[548,790,351,373]" italics="true" pageId="1" pageNumber="2">Arcynopteryx compacta</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFF0897DFC971F31A64DFED2" author="Stark" box="[813,1054,350,372]" pageId="1" pageNumber="2" refString="Stark, B. &amp; Szczytko, W. (1988) Egg morphology and phylogeny in Arcynopterygini (Plecoptera: Perlodidae). Journal of the Kansas Entomological Society, 61, 143 - 160." type="journal article" year="1988">Stark &amp; Szczytko 1988</bibRefCitation>
: 156 (
<taxonomicName id="C1B94D3CFFF0897DFBDA1F30A702FED3" box="[1120,1361,351,373]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFF0897DFBDA1F30A702FED3" box="[1120,1361,351,373]" italics="true" pageId="1" pageNumber="2">Arcynopteryx compacta</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFF0897DFAD21F30A300FE34" author="Kondratieff" pageId="1" pageNumber="2" refString="Kondratieff, B. C. (2004) Perlodidae-Perlodinae (The Springflies). In: Stark B. S. &amp; Armitage B. J. (Eds.), Stoneflies (Plecoptera) of Eastern North America. Vol. 2. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Ohio Biological Survey, pp. 149 - 180." type="book" year="2004">Kondratieff 2004</bibRefCitation>
: 166, figs 8.1–8.3 (
<taxonomicName id="C1B94D3CFFF0897DFD971F11A171FE35" box="[557,802,382,403]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFF0897DFD971F11A171FE35" box="[557,802,382,403]" italics="true" pageId="1" pageNumber="2">Arcynopteryx compacta</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFF0897DFC841F12A61EFE35" author="Stewart" box="[830,1101,381,403]" pageId="1" pageNumber="2" refString="Stewart, K. &amp; Oswood, M. (2006) The stoneflies (Plecoptera) of Alaska and western Canada. The Caddis Press, Columbus, Ohio. pp. 1 - 325." type="book chapter" year="2006">Stewart &amp; Oswood 2006</bibRefCitation>
: 182 (
<taxonomicName id="C1B94D3CFFF0897DFB2D1F11A7DFFE35" box="[1175,1420,382,403]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFF0897DFB2D1F11A7DFFE35" box="[1175,1420,382,403]" italics="true" pageId="1" pageNumber="2">Arcynopteryx compacta</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFF0897DFF7D1FF4A3BCFE17" author="Teslenko" box="[199,495,411,433]" pageId="1" pageNumber="2" refString="Teslenko, V. &amp; Zhiltzova, L. (2009) Keys to the stoneflies (Insecta, Plecoptera) of Russia and adjacent countries. Imagines and larvae. Dalnauka, Vladivostok, 1 - 382 pp (in Russian)." type="book chapter" year="2009">Teslenko &amp; Zhiltzova 2009</bibRefCitation>
: 26, figs. 118−121 (
<taxonomicName id="C1B94D3CFFF0897DFD771FF3A191FE14" box="[717,962,412,434]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFF0897DFD771FF3A191FE14" box="[717,962,412,434]" italics="true" pageId="1" pageNumber="2">Arcynopteryx compacta</emphasis>
</taxonomicName>
). The additional synonyms are presented in Brinck’s (1949), Ricker’s (1952), Illies’s (1955) papers, on the 
<taxonomicName id="C1B94D3CFFF0897DFCE11FD5A195FE76" box="[859,966,442,464]" class="Insecta" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
species file website (DeWalt, 
<bibRefCitation id="62284B4EFFF0897DFABF1FD5A302FE56" author="Neu-Becker" pageId="1" pageNumber="2" refString="DeWalt, R., Neu-Becker, U. &amp; Steuber, G. (2012) Plecoptera Species File Online. Version 1.1 / 4.0, http: // Plecoptera. Species- File. org" type="book" year="2012">Neu-Becker &amp; Stueber 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4EA36534FFF0897DFF2D1C74A0D0F982" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="060636BFFFF0897DFF2D1C74A0F0FCF2" blockId="1.[151,1437,539,2004]" pageId="1" pageNumber="2">
<emphasis id="34CDEAADFFF0897DFF2D1C74A342FD92" bold="true" box="[151,273,539,564]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
<taxonomicName id="C1B94D3CFFF0897DFEA21C73A3DEFD92" box="[280,397,540,564]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF0897DFEA21C73A3DEFD92" box="[280,397,540,564]" italics="true" pageId="1" pageNumber="2">A. dichroa</emphasis>
</taxonomicName>
can be distinguished by a shape of the hemitergal lobes which bear lobe apices directed posteriorly, apical margins prolonged and rounded, well sclerotized (
<figureCitation id="9E822A3AFFF0897DFC261C50A1B7FDF1" box="[924,996,575,599]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
); the hemitergal lobes are in contact mesoanteriorly (
<figureCitation id="9E822A3AFFF0897DFEEA1C0BA3CAFDDA" box="[336,409,612,636]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Figs 2</figureCitation>
&amp; 
<figureCitation id="9E822A3AFFF0897DFE7A1C0BA383FDDA" box="[448,464,612,636]" captionStart="FIGURES 7 – 8" captionStartId="3.[151,264,927,950]" captionTargetBox="[165,1404,210,889]" captionTargetId="figure@3.[151,1436,193,906]" captionTargetPageId="3" captionText="FIGURES 7 – 8. Arcynopteryx dichroa, type: 7. Male abdominal tip, dorsolateral, pinned. 8. Same, dorsal, cleaned. (Photograph by D. Goodger)." httpUri="https://zenodo.org/record/210962/files/figure.png" pageId="1" pageNumber="2">7</figureCitation>
). Each hemitergal lobe bears a knob erected above a membranous patch close to the anterior hemitergal margin (
<figureCitation id="9E822A3AFFF0897DFE751CE8A062FD39" box="[463,561,647,671]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Figs 2, 3</figureCitation>
&amp; 
<figureCitation id="9E822A3AFFF0897DFDEE1CE8A037FD39" box="[596,612,647,671]" captionStart="FIGURES 7 – 8" captionStartId="3.[151,264,927,950]" captionTargetBox="[165,1404,210,889]" captionTargetId="figure@3.[151,1436,193,906]" captionTargetPageId="3" captionText="FIGURES 7 – 8. Arcynopteryx dichroa, type: 7. Male abdominal tip, dorsolateral, pinned. 8. Same, dorsal, cleaned. (Photograph by D. Goodger)." httpUri="https://zenodo.org/record/210962/files/figure.png" pageId="1" pageNumber="2">8</figureCitation>
). Epiproct is very distinct from that of the other 
<taxonomicName id="C1B94D3CFFF0897DFBC51CE7A746FD06" box="[1151,1301,648,672]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="34CDEAADFFF0897DFBC51CE7A746FD06" box="[1151,1301,648,672]" italics="true" pageId="1" pageNumber="2">Arcynopteryx</emphasis>
</taxonomicName>
species, the stylet of the epiproct resembles a strong, long, fine bristle directed upward and forward (
<figureCitation id="9E822A3AFFF0897DFB1E1CC3A6BFFD62" box="[1188,1260,684,708]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Fig. 5</figureCitation>
). The shape of female subgenital plate is variable: the posterior margin of the subgenital plate has a shallow notch that separates two small lobes (
<figureCitation id="9E822A3AFFF0897DFEE21C9BA3CEFCAA" box="[344,413,756,780]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Fig. 6</figureCitation>
); sometimes there are two shallow notches with three small lobes. Egg is distinguished by chorionic structure which is presented by a pattern of hexagonal follicle cell impressions (FCI’s), flat floors which contain often 12 shallow punctations (
<figureCitation id="9E822A3AFFF0897DFDF81D53A0C5FCF2" box="[578,662,828,852]" captionStart="FIGURES 9 – 12" captionStartId="3.[151,264,1946,1969]" captionTargetBox="[264,1319,1032,1898]" captionTargetId="figure@3.[264,1323,1032,1922]" captionTargetPageId="3" captionText="FIGURES 9 – 12. Arcynopteryx dichroa, egg: 9. Habitus: anchor, chorion structure, row of micropyles, lateral. 10. Collar, lateral. 11. Micropylar orifices and mounds, dorsal view. 12. Chorion structure, dorsal view." httpUri="https://zenodo.org/record/210963/files/figure.png" pageId="1" pageNumber="2">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="060636BFFFF0897DFF731D30A0D0F982" blockId="1.[151,1437,539,2004]" pageId="1" pageNumber="2">
<emphasis id="34CDEAADFFF0897DFF731D30A324FCDE" bold="true" box="[201,375,863,888]" pageId="1" pageNumber="2">Adult habitus.</emphasis>
The head, pronotum and abdomen are brown, the meso- and metanotum dark brown. Head (
<figureCitation id="9E822A3AFFF0897DFF251DEBA2B2FC3A" box="[159,225,900,924]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
) with wide, transverse, M-shaped dark brown band between antennal bases, delimited by epicranial suture posteriorly. In front of the distinct M-shaped band, an indistinct darkish spot projects onto the clypeus; the frontoclypeus is broadly truncated and pale laterally; the tentorial pits are dark (
<figureCitation id="9E822A3AFFF0897DFC651DA3A671FC42" box="[991,1058,972,996]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
). The interocellar area exhibits a pale spot rounded anteriorly that does not reach the median ocellus, a pale spot continues to the medial surface of the occiput. Two tentorial pits in front of the lateral ocelli and two small oval patches laterally to the lateral ocelli are pale. Behind each compound eye is a posterolateral spot with brown callosities (
<figureCitation id="9E822A3AFFF0897DFB851A57A6D3FBF6" box="[1087,1152,1080,1104]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
). The antennae and palpi are brownish; the basal antennal segments are brown. The submental gills are long and thin. The pronotum is the same width as the head width under the compound eyes, brownish, quadrangular, with rounded angles; the lateral margins are straight; a broad, median yellow band is present, slightly wider in its posterior third (
<figureCitation id="9E822A3AFFF0897DFB651ACBA772FB1A" box="[1247,1313,1188,1212]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
). The pronotal rugosities are dark brown. The lateral fields are slightly darker than the median stripe, and the pronotum callosities are sometimes indistinct. The arms of the mesosternal ridge meet the anterior corners of the furcal pits. The abdomen is covered by colorless hairs, pronounced on the abdominal terga posterolaterally. The legs are brown; the distal end of the femur and the basal part of the tibia are dark brown. The cerci are longer than the abdomen, with long brownish hairs; the basal cercal segments are brown. The distal half of the apical cercal segments is dark brown. The females are macropterous. The males usually have very shortened wings, or their wings reach to the end of the abdomen or slightly past it. The forewing is long, narrow, and transparent, with brown veins and a pale yellow C vein. The venation includes an irregular net near the apex, sometimes consisting of three rows of cells. The hind wing anal area is large, and A2, A4 and A5 are forked. Brachypterous and long-winged specimens occurred together at the same sampling site.
</paragraph>
</subSubSection>
<subSubSection id="4EA36534FFF0897EFF73185FA185F87F" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
<paragraph id="060636BFFFF0897EFF73185FA1BEFDB1" blockId="1.[151,1437,539,2004]" lastBlockId="2.[151,1436,151,535]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<emphasis id="34CDEAADFFF0897DFF73185FA355F9EE" bold="true" box="[201,262,1584,1608]" pageId="1" pageNumber="2">Male</emphasis>
. Body length 
<quantity id="C1419B5AFFF0897DFE16185FA005F9EE" box="[428,598,1584,1608]" metricMagnitude="-2" metricUnit="m" metricValue="1.4849999999999999" metricValueMax="1.95" metricValueMin="1.02" pageId="1" pageNumber="2" unit="mm" value="14.85" valueMax="19.5" valueMin="10.2">10.2−19.5 mm</quantity>
, forewing of full-winged male 12.0− 
<quantity id="C1419B5AFFF0897DFBB9185FA638F9EE" box="[1027,1131,1584,1608]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="2" unit="mm" value="12.7">12.7 mm</quantity>
, wingspan 25.4–27.0 mm; forewing of male with shortened wings 4.3−6.0 mm, wingspan 
<quantity id="C1419B5AFFF0897DFCE3183BA1ACF9CA" box="[857,1023,1620,1644]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" metricValueMax="1.35" metricValueMin="1.01" pageId="1" pageNumber="2" unit="mm" value="11.8" valueMax="13.5" valueMin="10.1">10.1–13.5 mm</quantity>
. Abdominal tergum 9 exhibits a thin, transversal, membranous, pale median line and two pairs of small pale spots: one pair of spots is close to the line medially; the second pair of spots is situated close to the anterior margin of tergum 9 (
<figureCitation id="9E822A3AFFF0897DFBED18F3A6CFF912" box="[1111,1180,1692,1716]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
). The posterior margin has a medial arcuate notch, which runs ½ of the length of tergum 9, and two submedial, transversely elongated, and rounded swellings. These swellings are covered by small stout setae close to the notch and by long fine colorless hairs posteriolaterally (
<figureCitation id="9E822A3AFFF0897DFE221968A38DF8B9" box="[408,478,1799,1823]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Figs 2</figureCitation>
&amp; 
<figureCitation id="9E822A3AFFF0897DFDB81968A041F8B9" box="[514,530,1799,1823]" captionStart="FIGURES 7 – 8" captionStartId="3.[151,264,927,950]" captionTargetBox="[165,1404,210,889]" captionTargetId="figure@3.[151,1436,193,906]" captionTargetPageId="3" captionText="FIGURES 7 – 8. Arcynopteryx dichroa, type: 7. Male abdominal tip, dorsolateral, pinned. 8. Same, dorsal, cleaned. (Photograph by D. Goodger)." httpUri="https://zenodo.org/record/210962/files/figure.png" pageId="1" pageNumber="2">7</figureCitation>
). Sternum 9 is scoop-shaped, extended backward and curved upward. Tergum 10 is divided into two hemiterga (
<figureCitation id="9E822A3AFFF0897DFE4D1943A06DF8E2" box="[503,574,1836,1860]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
). The hemitergal lobes in a dorsal view are wide, with lobe apices directed posteriorly, apical margins prolonged and rounded, well sclerotized; the lobes are in contact mesoanteriorly (
<figureCitation id="9E822A3AFFF0897DFAF51920A2F6F82A" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Figs 2, 3</figureCitation>
&amp; 
<figureCitation id="9E822A3AFFF0897DFF7D191BA284F82A" box="[199,215,1908,1932]" captionStart="FIGURES 7 – 8" captionStartId="3.[151,264,927,950]" captionTargetBox="[165,1404,210,889]" captionTargetId="figure@3.[151,1436,193,906]" captionTargetPageId="3" captionText="FIGURES 7 – 8. Arcynopteryx dichroa, type: 7. Male abdominal tip, dorsolateral, pinned. 8. Same, dorsal, cleaned. (Photograph by D. Goodger)." httpUri="https://zenodo.org/record/210962/files/figure.png" pageId="1" pageNumber="2">8</figureCitation>
). Their mesal edges are membranous and resemble prolonged small white oval patches. Each hemitergal lobe bears a knob erected above a membranous patch close to the anterior hemitergal margin (
<figureCitation id="9E822A3AFFF0897DFB3E19F8A69FF809" box="[1156,1228,1943,1967]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Figs 2</figureCitation>
&amp; 
<figureCitation id="9E822A3AFFF0897DFB4B19F8A752F809" box="[1265,1281,1943,1967]" captionStart="FIGURES 7 – 8" captionStartId="3.[151,264,927,950]" captionTargetBox="[165,1404,210,889]" captionTargetId="figure@3.[151,1436,193,906]" captionTargetPageId="3" captionText="FIGURES 7 – 8. Arcynopteryx dichroa, type: 7. Male abdominal tip, dorsolateral, pinned. 8. Same, dorsal, cleaned. (Photograph by D. Goodger)." httpUri="https://zenodo.org/record/210962/files/figure.png" pageId="1" pageNumber="2">8</figureCitation>
). The knob is ovular, membranous ventrally, sclerotized and rounded dorsally, and covered by a few small, stout setae (
<figureCitation id="9E822A3AFFF0897DFA8219D3A72CF872" box="[1336,1407,1980,2004]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="1" pageNumber="2">Figs 3</figureCitation>
&amp; 
<figureCitation id="9E822A3AFFF3897EFF2D1EF8A2F4FF09" box="[151,167,151,175]" captionStart="FIGURES 7 – 8" captionStartId="3.[151,264,927,950]" captionTargetBox="[165,1404,210,889]" captionTargetId="figure@3.[151,1436,193,906]" captionTargetPageId="3" captionText="FIGURES 7 – 8. Arcynopteryx dichroa, type: 7. Male abdominal tip, dorsolateral, pinned. 8. Same, dorsal, cleaned. (Photograph by D. Goodger)." httpUri="https://zenodo.org/record/210962/files/figure.png" pageId="2" pageNumber="3">8</figureCitation>
). The cowl is membranous, folded, resembles a deep pouch between and under the hemitergal lobes and is attached around the base of the epiproct and the internal basal anchor (
<figureCitation id="9E822A3AFFF3897EFC071ED3A656FF72" box="[957,1029,188,212]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="2" pageNumber="3">Figs 5</figureCitation>
&amp; 
<figureCitation id="9E822A3AFFF3897EFB911ED3A668FF72" box="[1067,1083,188,212]" captionStart="FIGURES 7 – 8" captionStartId="3.[151,264,927,950]" captionTargetBox="[165,1404,210,889]" captionTargetId="figure@3.[151,1436,193,906]" captionTargetPageId="3" captionText="FIGURES 7 – 8. Arcynopteryx dichroa, type: 7. Male abdominal tip, dorsolateral, pinned. 8. Same, dorsal, cleaned. (Photograph by D. Goodger)." httpUri="https://zenodo.org/record/210962/files/figure.png" pageId="2" pageNumber="3">8</figureCitation>
). The dorsolateral edges of the cowl are supported by flat and darkly sclerotized paragenital plates. Otherwise, in a lateral view, the lever arm of the epiproct is arcuate and is found at the bottom of the deep cowl; and terminates in a basal sclerite ventrally, the top of which serves as a place for the attachment of the loop of the stylet and sclerotized bands (
<figureCitation id="9E822A3AFFF3897EFB681F48A744FE99" box="[1234,1303,295,319]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
). The stylet of the epiproct resembles a strong, long, fine bristle directed upward and forward; the basal plate of the loop of the stylet is thin (
<figureCitation id="9E822A3AFFF3897EFE8F1F00A32FFE21" box="[309,380,367,391]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
). Two lateral sclerotized bands are narrowed at the base, and each band has a deep rounded notch on the inner edge in the last third of its length. In a dorsal view, the everted aedeagus is large and membranous, with a pair of lateral lobes at dorsolateral margins (
<figureCitation id="9E822A3AFFF3897EFCAA1FD8A106FE69" box="[784,853,439,463]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
). The lobes may not be fully everted. One large prolonged lobe is narrowed to the apex. Fine, erect, clear spinules are visible ventrally and dorsally on the prolonged lobe. The swollen rounded apex of the aedeagus is without spinules (
<figureCitation id="9E822A3AFFF3897EFC211F90A1B3FDB1" box="[923,992,511,535]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
).
</paragraph>
<caption id="52C66637FFF3897EFF2D1BCEA720FA52" httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="2" pageNumber="3" targetBox="[264,1317,602,1408]" targetPageId="2">
<paragraph id="060636BFFFF3897EFF2D1BCEA720FA52" blockId="2.[151,1436,1441,1525]" pageId="2" pageNumber="3">
<emphasis id="34CDEAADFFF3897EFF2D1BCEA366FA11" bold="true" box="[151,309,1441,1464]" pageId="2" pageNumber="3">FIGURES 1–6.</emphasis>
<taxonomicName id="C1B94D3CFFF3897EFE861BCDA05DFA11" box="[316,526,1441,1463]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF3897EFE861BCDA05DFA11" box="[316,526,1441,1463]" italics="true" pageId="2" pageNumber="3">Arcynopteryx dichroa</emphasis>
</taxonomicName>
: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3–5 = 1.0.
</paragraph>
</caption>
<paragraph id="060636BFFFF3897EFF73184DA0FFF8AF" blockId="2.[151,1436,1569,2009]" pageId="2" pageNumber="3">
<emphasis id="34CDEAADFFF3897EFF73184DA373F99C" bold="true" box="[201,288,1570,1594]" pageId="2" pageNumber="3">Female</emphasis>
is macropterous, larger and darker than male. Body length 11.0− 
<quantity id="C1419B5AFFF3897EFBB7184EA626F99F" box="[1037,1141,1569,1593]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="2" pageNumber="3" unit="mm" value="20.5">20.5 mm</quantity>
, forewing 11.0– 
<quantity id="C1419B5AFFF3897EFA8A184EA7CBF99F" box="[1328,1432,1569,1593]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="2" pageNumber="3" unit="mm" value="18.4">18.4 mm</quantity>
, wingspan 24.0− 
<quantity id="C1419B5AFFF3897EFEF6182BA3E6F9FA" box="[332,437,1604,1628]" metricMagnitude="-2" metricUnit="m" metricValue="3.92" pageId="2" pageNumber="3" unit="mm" value="39.2">39.2 mm</quantity>
. Sternum 8 bears two brown spots close to the anterior margin (
<figureCitation id="9E822A3AFFF3897EFB2D182BA68EF9FA" box="[1175,1245,1604,1628]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
). The subgenital plate is wide, relatively short, and pale mesoanteriorly, extending laterally from the sides of sternum 8 and reaching almost half the length of sternum 9 (
<figureCitation id="9E822A3AFFF3897EFD8F18E6A028F907" box="[565,635,1673,1697]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
). The posterior margin of the subgenital plate has a shallow notch that separates two small lobes. The subgenital plate covered with small, brownish setae. The shape of the subgenital plate is variable. Sometimes there are two shallow notches with three small lobes. Sternum 9 is pale medially, with two brown circular spots mesolaterally (
<figureCitation id="9E822A3AFFF3897EFDE0189EA0CDF8AF" box="[602,670,1777,1801]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="060636BFFFF3897EFF73197AA185F87F" blockId="2.[151,1436,1569,2009]" pageId="2" pageNumber="3">
<emphasis id="34CDEAADFFF3897EFF73197AA2ABF88B" bold="true" box="[201,248,1813,1837]" pageId="2" pageNumber="3">Egg</emphasis>
is ovular and circular in cross-section, 370 X 238 µm. Anchor mushroom-shaped covers the collar completely (
<figureCitation id="9E822A3AFFF3897EFF481958A364F8E9" box="[242,311,1847,1871]" captionStart="FIGURES 9 – 12" captionStartId="3.[151,264,1946,1969]" captionTargetBox="[264,1319,1032,1898]" captionTargetId="figure@3.[264,1323,1032,1922]" captionTargetPageId="3" captionText="FIGURES 9 – 12. Arcynopteryx dichroa, egg: 9. Habitus: anchor, chorion structure, row of micropyles, lateral. 10. Collar, lateral. 11. Micropylar orifices and mounds, dorsal view. 12. Chorion structure, dorsal view." httpUri="https://zenodo.org/record/210963/files/figure.png" pageId="2" pageNumber="3">Fig. 9</figureCitation>
). The collar is stalked, its rim flanged and irregularly incised (
<figureCitation id="9E822A3AFFF3897EFC481958A615F8E9" box="[1010,1094,1847,1871]" captionStart="FIGURES 9 – 12" captionStartId="3.[151,264,1946,1969]" captionTargetBox="[264,1319,1032,1898]" captionTargetId="figure@3.[264,1323,1032,1922]" captionTargetPageId="3" captionText="FIGURES 9 – 12. Arcynopteryx dichroa, egg: 9. Habitus: anchor, chorion structure, row of micropyles, lateral. 10. Collar, lateral. 11. Micropylar orifices and mounds, dorsal view. 12. Chorion structure, dorsal view." httpUri="https://zenodo.org/record/210963/files/figure.png" pageId="2" pageNumber="3">Fig. 10</figureCitation>
). The sides of collar has irregular meshwork and projections; shoulder is low (
<figureCitation id="9E822A3AFFF3897EFD7A1936A147F8D7" box="[704,788,1881,1905]" captionStart="FIGURES 9 – 12" captionStartId="3.[151,264,1946,1969]" captionTargetBox="[264,1319,1032,1898]" captionTargetId="figure@3.[264,1323,1032,1922]" captionTargetPageId="3" captionText="FIGURES 9 – 12. Arcynopteryx dichroa, egg: 9. Habitus: anchor, chorion structure, row of micropyles, lateral. 10. Collar, lateral. 11. Micropylar orifices and mounds, dorsal view. 12. Chorion structure, dorsal view." httpUri="https://zenodo.org/record/210963/files/figure.png" pageId="2" pageNumber="3">Fig. 10</figureCitation>
). The chorion is covered with hexagonal FCI’s (
<figureCitation id="9E822A3AFFF3897EFA8D1936A2E5F832" captionStart="FIGURES 9 – 12" captionStartId="3.[151,264,1946,1969]" captionTargetBox="[264,1319,1032,1898]" captionTargetId="figure@3.[264,1323,1032,1922]" captionTargetPageId="3" captionText="FIGURES 9 – 12. Arcynopteryx dichroa, egg: 9. Habitus: anchor, chorion structure, row of micropyles, lateral. 10. Collar, lateral. 11. Micropylar orifices and mounds, dorsal view. 12. Chorion structure, dorsal view." httpUri="https://zenodo.org/record/210963/files/figure.png" pageId="2" pageNumber="3">Figs 9 &amp; 12</figureCitation>
); the FCI walls are slightly raised with thin, shallow furrows; flat floors often contain 12 shallow punctations (
<figureCitation id="9E822A3AFFF3897EFF2519F0A2A7F811" box="[159,244,1951,1975]" captionStart="FIGURES 9 – 12" captionStartId="3.[151,264,1946,1969]" captionTargetBox="[264,1319,1032,1898]" captionTargetId="figure@3.[264,1323,1032,1922]" captionTargetPageId="3" captionText="FIGURES 9 – 12. Arcynopteryx dichroa, egg: 9. Habitus: anchor, chorion structure, row of micropyles, lateral. 10. Collar, lateral. 11. Micropylar orifices and mounds, dorsal view. 12. Chorion structure, dorsal view." httpUri="https://zenodo.org/record/210963/files/figure.png" pageId="2" pageNumber="3">Fig. 12</figureCitation>
). Row of micropyles subequatorial; orifices small without lips, some set on low micropylar mounds are occasionally surrounded by rosettes (
<figureCitation id="9E822A3AFFF3897EFD8C19AEA0EEF87F" box="[566,701,1985,2009]" captionStart="FIGURES 9 – 12" captionStartId="3.[151,264,1946,1969]" captionTargetBox="[264,1319,1032,1898]" captionTargetId="figure@3.[264,1323,1032,1922]" captionTargetPageId="3" captionText="FIGURES 9 – 12. Arcynopteryx dichroa, egg: 9. Habitus: anchor, chorion structure, row of micropyles, lateral. 10. Collar, lateral. 11. Micropylar orifices and mounds, dorsal view. 12. Chorion structure, dorsal view." httpUri="https://zenodo.org/record/210963/files/figure.png" pageId="2" pageNumber="3">Figs 9 &amp; 11</figureCitation>
). Eclosion line is absent.
</paragraph>
</subSubSection>
<caption id="52C66637FFF2897FFF2D1DF0A36DFC72" httpUri="https://zenodo.org/record/210962/files/figure.png" pageId="3" pageNumber="4" targetBox="[165,1404,210,889]" targetPageId="3">
<paragraph id="060636BFFFF2897FFF2D1DF0A36DFC72" blockId="3.[151,1436,927,980]" pageId="3" pageNumber="4">
<emphasis id="34CDEAADFFF2897FFF2D1DF0A369FC13" bold="true" box="[151,314,927,950]" pageId="3" pageNumber="4">FIGURES 7–8.</emphasis>
<taxonomicName id="C1B94D3CFFF2897FFE851DCFA049FC13" box="[319,538,927,949]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF2897FFE851DCFA049FC13" box="[319,538,927,949]" italics="true" pageId="3" pageNumber="4">Arcynopteryx dichroa</emphasis>
</taxonomicName>
, type: 7. Male abdominal tip, dorsolateral, pinned. 8. Same, dorsal, cleaned. (Photograph by D. Goodger).
</paragraph>
</caption>
<caption id="52C66637FFF2897FFF2D19F5A672F869" httpUri="https://zenodo.org/record/210963/files/figure.png" pageId="3" pageNumber="4" targetBox="[264,1319,1032,1898]" targetPageId="3">
<paragraph id="060636BFFFF2897FFF2D19F5A672F869" blockId="3.[151,1436,1946,1999]" pageId="3" pageNumber="4">
<emphasis id="34CDEAADFFF2897FFF2D19F5A31BF809" bold="true" box="[151,328,1946,1969]" pageId="3" pageNumber="4">FIGURES 9–12.</emphasis>
<taxonomicName id="C1B94D3CFFF2897FFEF519F4A07FF816" box="[335,556,1946,1968]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF2897FFEF519F4A07FF816" box="[335,556,1946,1968]" italics="true" pageId="3" pageNumber="4">Arcynopteryx dichroa</emphasis>
</taxonomicName>
, egg: 9. Habitus: anchor, chorion structure, row of micropyles, lateral. 10. Collar, lateral. 11. Micropylar orifices and mounds, dorsal view. 12. Chorion structure, dorsal view.
</paragraph>
</caption>
<subSubSection id="4EA36534FFF58978FF731EF8A016FE52" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="060636BFFFF58978FF731EF8A016FE52" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="5">
<emphasis id="34CDEAADFFF58978FF731EF8A3F8FF16" bold="true" box="[201,427,151,176]" pageId="4" pageNumber="5">Material examined</emphasis>
. 
<typeStatus id="D902881DFFF58978FE021EF7A07BFF16" box="[440,552,152,176]" pageId="4" pageNumber="5" type="lectotype">Lectotype</typeStatus>
male (pinned), 
<taxonomicName id="C1B94D3CFFF58978FD631EF6A6D1FF09" authority="McLachlan" authorityName="McLachlan" box="[729,1154,151,176]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FD631EF6A102FF16" box="[729,849,152,176]" italics="true" pageId="4" pageNumber="5">D. dichroa</emphasis>
McLachlan (
<emphasis id="34CDEAADFFF58978FC5D1EF6A62BFF16" box="[999,1144,153,176]" italics="true" pageId="4" pageNumber="5">Dictyopteryx</emphasis>
)
</taxonomicName>
, [McL. label] / Sibir. orient. (Maa(c)k) / 
<taxonomicName id="C1B94D3CFFF58978FEE81ED3A3F5FF72" box="[338,422,188,212]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">dichroa</taxonomicName>
McL. / 
<taxonomicName id="C1B94D3CFFF58978FDBE1ED2A0BCFF72" box="[516,751,188,212]" class="Insecta" family="Perlodidae" genus="Dictiopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FDBE1ED2A0BCFF72" box="[516,751,188,212]" italics="true" pageId="4" pageNumber="5">Dictiopteryx dichroa</emphasis>
</taxonomicName>
McL. The type-series in 
<collectionCode id="60A8AE7AFFF58978FBAB1ED2A636FF72" box="[1041,1125,189,212]" pageId="4" pageNumber="5">BMNH</collectionCode>
includes one male and one female 
<typeStatus id="D902881DFFF58978FF501EB0A3DFFF51" box="[234,396,223,247]" pageId="4" pageNumber="5" type="paralectotype">paralectotypes</typeStatus>
(
<bibRefCitation id="62284B4EFFF58978FE231EB0A015FF51" author="Kimmins" box="[409,582,223,247]" pageId="4" pageNumber="5" refString="Kimmins, D. (1970) A list of the type-specimens of Plecoptera and Megaloptera in the British Museum (Natural History). Bulletin of the British Museum (Natural History) Entomology, 24, 335 - 361." type="journal article" year="1970">Kimmins 1970</bibRefCitation>
). 
<collectingCountry id="7EAE762FFFF58978FDE21EB0A0F4FF51" box="[600,679,223,247]" name="Russia" pageId="4" pageNumber="5">Russia</collectingCountry>
: Altai Mountains, 
<specimenCount id="10BFFD36FFF58978FCCF1EB0A18FFF51" box="[885,988,223,247]" pageId="4" pageNumber="5" type="male">12 males</specimenCount>
, 
<specimenCount id="10BFFD36FFF58978FC5C1EB0A606FF51" box="[998,1109,223,247]" pageId="4" pageNumber="5" type="female">2 females</specimenCount>
, Chuya River, near Iodro settlement, 
<date id="7207107FFFF58978FF461F6BA32EFEBA" box="[252,381,260,284]" pageId="4" pageNumber="5" value="1989-05-18">18.05.1989</date>
, coll. E. Makarchenko; Khabarovskiy Region, 
<specimenCount id="10BFFD36FFF58978FC1B1F6BA658FEBA" box="[929,1035,260,284]" pageId="4" pageNumber="5" type="male">10 males</specimenCount>
, 
<specimenCount id="10BFFD36FFF58978FBA01F6BA6DEFEBA" box="[1050,1165,260,284]" pageId="4" pageNumber="5" type="female">2 females</specimenCount>
, Khor River, Ussuri R. Basin, Amur R. Basin, 
<date id="7207107FFFF58978FE271F48A04DFE99" box="[413,542,295,319]" pageId="4" pageNumber="5" value="1961-05-31">31.05.1961</date>
, coll. I. Levanidova; Tuva: 
<specimenCount id="10BFFD36FFF58978FCEF1F48A1FDFE99" box="[853,942,295,319]" pageId="4" pageNumber="5" type="male">3 males</specimenCount>
, 
<specimenCount id="10BFFD36FFF58978FC001F48A679FE99" box="[954,1066,295,319]" pageId="4" pageNumber="5" type="female">2 females</specimenCount>
, Kara-Hol‘ Lake, Baj-Taiginskyi District, 
<date id="7207107FFFF58978FF401F23A329FEC2" box="[250,378,332,356]" pageId="4" pageNumber="5" value="2003-06-23">23.06.2003</date>
, coll. M. Zasypkina; Kamchatka Peninsula, 
<specimenCount id="10BFFD36FFF58978FCD41F23A19BFEC2" box="[878,968,332,356]" pageId="4" pageNumber="5" type="male">6 males</specimenCount>
, 
<specimenCount id="10BFFD36FFF58978FC6E1F23A617FEC2" box="[980,1092,332,356]" pageId="4" pageNumber="5" type="female">3 females</specimenCount>
, Avacha R., below Krutaya R. mouth, 
<date id="7207107FFFF58978FF571F00A33EFE21" box="[237,365,367,391]" pageId="4" pageNumber="5" value="1969-06-20">20.06.1969</date>
, coll. I. Levanidova; Chukotka Peninsula, 
<specimenCount id="10BFFD36FFF58978FCF31F00A1C4FE21" box="[841,919,367,391]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
, Levaya Rechka, 
<quantity id="C1419B5AFFF58978FBE41F00A6E0FE21" box="[1118,1203,367,391]" metricMagnitude="5" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="5" unit="km" value="105.0">105 km</quantity>
on the road between Egvekinot and Iultin settlements, 3.07.1973, coll. I. Chereshnev; 
<specimenCount id="10BFFD36FFF58978FCCA1FFBA1EDFE0A" box="[880,958,404,428]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
, 
<specimenCount id="10BFFD36FFF58978FC731FFBA67EFE0A" box="[969,1069,404,428]" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
, Pravaya Rechka, 
<quantity id="C1419B5AFFF58978FB411FFBA703FE0A" box="[1275,1360,404,428]" metricMagnitude="5" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="5" unit="km" value="105.0">105 km</quantity>
on the road between Egvekinot and Iultin settlements, 
<date id="7207107FFFF58978FD701FD8A118FE69" box="[714,843,439,463]" pageId="4" pageNumber="5" value="1973-10-07">10.07.1973</date>
, coll. I. Chereshnev; 
<specimenCount id="10BFFD36FFF58978FBF61FD8A6CCFE69" box="[1100,1183,439,463]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
, Lena River, Lenskie Stolby, 1.06.1974, coll. I. Levanidova.
</paragraph>
</subSubSection>
<subSubSection id="4EA36534FFF58978FF731F90A094FCB2" pageId="4" pageNumber="5" type="distribution">
<paragraph id="060636BFFFF58978FF731F90A094FCB2" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="5">
<emphasis id="34CDEAADFFF58978FF731F90A331FDBE" bold="true" box="[201,354,511,536]" pageId="4" pageNumber="5">Distribution.</emphasis>
<emphasis id="34CDEAADFFF58978FED71C6FA3BEFDBE" box="[365,493,512,536]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="C1B94D3CFFF58978FED71C6FA3BAFDBE" box="[365,489,512,536]" class="Insecta" family="Perlodidae" genus="Dictiopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">A. dichroa</taxonomicName>
,
</emphasis>
is a Holarctic circumpolar species primarily of the northern latitudes of Europe, Asia, and North 
<collectingCountry id="7EAE762FFFF58978FEEE1C4BA3EBFD9A" box="[340,440,548,572]" name="United States of America" pageId="4" pageNumber="5">America</collectingCountry>
, occurring in streams, but also lakes throughout its range. The species inhabits northern Europe (
<bibRefCitation id="62284B4EFFF58978FF4D1C28A3FBFDF9" author="Zhiltzova" box="[247,424,583,607]" pageId="4" pageNumber="5" refString="Zhiltzova, L. (1966) The stoneflies (Plecoptera) of the European part of USSR (without Caucasus). Entomological Review, 45, 525 - 549 (in Russian)." type="journal article" year="1966">Zhiltzova 1966</bibRefCitation>
, 
<bibRefCitation id="62284B4EFFF58978FE0E1C28A0DBFDF9" author="Lillehammer" box="[436,648,583,607]" pageId="4" pageNumber="5" refString="Lillehammer, A. (1974) Norwegian stoneflies I. Analysis of the variations in morphological and structural characters used in taxonomy. Norsk Entomologisk Tidsskrift, 21, 59 - 107." type="journal article" year="1974">Lillehammer 1974</bibRefCitation>
, 
<bibRefCitation id="62284B4EFFF58978FD2E1C28A11CFDF9" author="Loskutova" box="[660,847,583,607]" pageId="4" pageNumber="5" refString="Loskutova, O. (2006) Fauna of the European North-East of Russia. Stoneflies. 9. Nauka Publishing House, Sankt-Petersburg, 224 pp (in Russian)." type="book" year="2006">Loskutova 2006</bibRefCitation>
), in Central and Southwest Europe 
<taxonomicName id="C1B94D3CFFF58978FB591C27A70AFDC6" box="[1251,1369,584,608]" class="Insecta" family="Perlodidae" genus="Dictiopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FB591C27A70AFDC6" box="[1251,1369,584,608]" italics="true" pageId="4" pageNumber="5">A. dichroa</emphasis>
</taxonomicName>
has a disjunct boreomontane distribution with Pleistocene relict populations (
<bibRefCitation id="62284B4EFFF58978FC701C03A61BFD22" author="Illies" box="[970,1096,620,644]" pageId="4" pageNumber="5" refString="Illies, J. (1955) Steinfliegen oder Plecoptera. Die Tierwelt Deutschlands, 43, 150 s." type="journal article" year="1955">Illies 1955</bibRefCitation>
, 
<bibRefCitation id="62284B4EFFF58978FBEC1C03A6B7FD22" author="Zwick" box="[1110,1252,620,644]" pageId="4" pageNumber="5" refString="Zwick, P. (2004) Key to the West Palaearctic genera of stoneflies (Plecoptera) in the larval stage. Limnologica, 34, 315 - 348." type="journal article" year="2004">Zwick 2004</bibRefCitation>
). The species is widespread in Asia including 
<collectingCountry id="7EAE762FFFF58978FE5E1CE0A007FD01" box="[484,596,655,679]" name="Mongolia" pageId="4" pageNumber="5">Mongolia</collectingCountry>
, Siberia and the Russian Far East. In North 
<collectingCountry id="7EAE762FFFF58978FB861CE0A6CFFD01" box="[1084,1180,655,679]" name="United States of America" pageId="4" pageNumber="5">America</collectingCountry>
<taxonomicName id="C1B94D3CFFF58978FB191CFFA74BFD0E" box="[1187,1304,656,680]" class="Insecta" family="Perlodidae" genus="Dictiopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FB191CFFA74BFD0E" box="[1187,1304,656,680]" italics="true" pageId="4" pageNumber="5">A. dichroa</emphasis>
</taxonomicName>
also occurs in alpine zones of Central Rocky Mountains (
<bibRefCitation id="62284B4EFFF58978FD191CDBA167FD6A" author="Ricker" box="[675,820,692,716]" pageId="4" pageNumber="5" refString="Ricker, W. (1964) Distribution of Canadian stoneflies. Gewasser und Abwasser, 34 / 35, 50 - 71." type="journal article" year="1964">Ricker 1964</bibRefCitation>
, 
<bibRefCitation id="62284B4EFFF58978FCFB1CDBA631FD6A" author="Stewart" box="[833,1122,692,716]" pageId="4" pageNumber="5" refString="Stewart, K. &amp; Oswood, M. (2006) The stoneflies (Plecoptera) of Alaska and western Canada. The Caddis Press, Columbus, Ohio. pp. 1 - 325." type="book chapter" year="2006">Stewart &amp; Oswood 2006</bibRefCitation>
). Populations of 
<taxonomicName id="C1B94D3CFFF58978FA9E1CDBA7CFFD6A" box="[1316,1436,692,716]" class="Insecta" family="Perlodidae" genus="Dictiopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FA9E1CDBA7CFFD6A" box="[1316,1436,692,716]" italics="true" pageId="4" pageNumber="5">A. dichroa</emphasis>
</taxonomicName>
have been recorded from Alaska, Alberta, British 
<collectingCountry id="7EAE762FFFF58978FD651CB8A103FD49" box="[735,848,727,751]" name="United States of America" pageId="4" pageNumber="5">Columbia</collectingCountry>
, Montana, Colorado, Maine, New Hampshire, Saskatchewan and Wyoming (
<bibRefCitation id="62284B4EFFF58978FE781C93A0E8FCB2" author="Stewart" box="[450,699,764,788]" pageId="4" pageNumber="5" refString="Stewart, K. W. &amp; Stark, B. P. (2002) Nymphs of North American Stonefly Genera (Plecoptera). 2 nd edition, The Caddis Press, Columbus, Ohio, pp. 1 - 510." type="book chapter" year="2002">Stewart &amp; Stark 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4EA36534FFF58978FF731D4FA301F9F6" pageId="4" pageNumber="5" type="discussion">
<paragraph id="060636BFFFF58978FF731D4FA0E7FC4A" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="5">
<emphasis id="34CDEAADFFF58978FF731D4FA36EFC9E" bold="true" box="[201,317,800,824]" pageId="4" pageNumber="5">Remarks.</emphasis>
<taxonomicName id="C1B94D3CFFF58978FEFD1D4FA06DFC9E" box="[327,574,800,824]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FEFD1D4FA06DFC9E" box="[327,574,800,824]" italics="true" pageId="4" pageNumber="5">Arcynopteryx dichroa</emphasis>
</taxonomicName>
has been described based on male and female specimens as 
<taxonomicName id="C1B94D3CFFF58978FAB11D4EA39AFCFA" authority="McLachlan 1872" authorityName="McLachlan" authorityYear="1872" class="Insecta" family="Perlodidae" genus="Dictyopteryx" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FAB11D4EA2BEFCFA" italics="true" pageId="4" pageNumber="5">Dictyopteryx dichroa</emphasis>
(
<bibRefCitation id="62284B4EFFF58978FF461D2BA392FCFA" author="McLachlan" box="[252,449,836,860]" pageId="4" pageNumber="5" refString="McLachlan, R. (1872) Perlides. In: Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps &amp; McLachlan. Annales de la Societe entomologique de Belgique (Bruxelles), 15, 47 - 55." type="journal article" year="1872">McLachlan 1872</bibRefCitation>
)
</taxonomicName>
. The description has omitted the date, locality and the structural details of the epiproct complex. However, the stylet of the epiproct was noted and illustrated (
<bibRefCitation id="62284B4EFFF58978FC641D08A76DFCD9" author="McLachlan" box="[990,1342,871,896]" pageId="4" pageNumber="5" refString="McLachlan, R. (1872) Perlides. In: Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps &amp; McLachlan. Annales de la Societe entomologique de Belgique (Bruxelles), 15, 47 - 55." type="journal article" year="1872">
McLachlan 1872, pl. I. 
<figureCitation id="9E822A3AFFF58978FB4C1D08A76DFCD9" box="[1270,1342,871,895]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,260,1441,1464]" captionTargetBox="[264,1317,602,1408]" captionTargetId="figure@2.[264,1323,593,1417]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Arcynopteryx dichroa: 1. Head and pronotum of male. 2. Male abdominal tip, dorsal. 3. Male left hemitergal lobe, dorsal. 4. Aedeagus, dorsal. 5. Cowl: sclerites of the internal basal anchor, lever arm, the paragenital plate, hemitergal lobe with knob, stylet and loop of the epiproct, and two lateral sclerotized bands, lateral. 6. Female abdominal tip, ventral. Scale (mm): 3 – 5 = 1.0." httpUri="https://zenodo.org/record/210961/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
</bibRefCitation>
). 
<taxonomicName id="C1B94D3CFFF58978FAEC1D06A314FC02" class="Insecta" family="Perlodidae" genus="Dictyopteryx" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FAEC1D06A314FC02" italics="true" pageId="4" pageNumber="5">Dictyopteryx dichroa</emphasis>
</taxonomicName>
<typeStatus id="D902881DFFF58978FEF41DE2A32DFC03" box="[334,382,909,933]" pageId="4" pageNumber="5">type</typeStatus>
is well associated with newly acquired material in the shape of the hemitergal lobes and knobs on them. These characters suggest that the species 
<taxonomicName id="C1B94D3CFFF58978FD5E1DDEA10DFC6E" box="[740,862,944,968]" class="Insecta" family="Perlodidae" genus="Dictyopteryx" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FD5E1DDEA10DFC6E" box="[740,862,944,968]" italics="true" pageId="4" pageNumber="5">D. dichroa</emphasis>
</taxonomicName>
belongs to 
<taxonomicName id="C1B94D3CFFF58978FC521DDFA766FC61" authority="Klapalek, 1904" authorityName="Klapalek" authorityYear="1904" box="[1000,1333,943,968]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="34CDEAADFFF58978FC521DDFA62DFC6E" box="[1000,1150,944,968]" italics="true" pageId="4" pageNumber="5">Arcynopteryx</emphasis>
<bibRefCitation id="62284B4EFFF58978FB3D1DC0A766FC61" author="Klapalek" box="[1159,1333,943,967]" pageId="4" pageNumber="5" refString="Klapalek, F. (1904) Evropske druhy celede Dictyopterygidae. Rozpravy C eske Akademie (Praha), 2, 1 - 10." type="journal article" year="1904">Klapálek, 1904</bibRefCitation>
</taxonomicName>
with the name 
<taxonomicName id="C1B94D3CFFF58978FF601DBBA0E3FC4A" authority="McLachlan, 1872" authorityName="McLachlan" authorityYear="1872" box="[218,688,980,1004]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FF601DBBA39EFC4A" box="[218,461,980,1004]" italics="true" pageId="4" pageNumber="5">Arcynopteryx dichroa</emphasis>
(
<bibRefCitation id="62284B4EFFF58978FE661DBBA0F4FC4A" author="McLachlan" box="[476,679,980,1004]" pageId="4" pageNumber="5" refString="McLachlan, R. (1872) Perlides. In: Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps &amp; McLachlan. Annales de la Societe entomologique de Belgique (Bruxelles), 15, 47 - 55." type="journal article" year="1872">McLachlan, 1872</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="060636BFFFF58978FF731D98A6E1FB62" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="5">
Klapalek designated 
<taxonomicName id="C1B94D3CFFF58978FE0E1D97A1CEFBA9" authority="McLachlan, 1872" authorityName="McLachlan" authorityYear="1872" box="[436,925,1015,1040]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFF58978FE0E1D97A0EDFBB6" box="[436,702,1016,1040]" italics="true" pageId="4" pageNumber="5">Arcynopteryx compacta</emphasis>
(
<bibRefCitation id="62284B4EFFF58978FD761D98A1C7FBA9" author="McLachlan" box="[716,916,1015,1039]" pageId="4" pageNumber="5" refString="McLachlan, R. (1872) Perlides. In: Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps &amp; McLachlan. Annales de la Societe entomologique de Belgique (Bruxelles), 15, 47 - 55." type="journal article" year="1872">McLachlan, 1872</bibRefCitation>
)
</taxonomicName>
as 
<typeStatus id="D902881DFFF58978FC781D96A1A1FBB7" box="[962,1010,1017,1041]" pageId="4" pageNumber="5">type</typeStatus>
species of genus 
<emphasis id="34CDEAADFFF58978FB021D97A705FBB6" box="[1208,1366,1016,1040]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="C1B94D3CFFF58978FB021D97A701FBB6" box="[1208,1362,1016,1040]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Arcynopteryx</taxonomicName>
.
</emphasis>
However, he was mistaken and misidentified, when provided (
<bibRefCitation id="62284B4EFFF58978FCA71A73A620FB92" author="Klapalek" box="[797,1139,1052,1077]" pageId="4" pageNumber="5" refString="Klapalek, F. (1912) Plecopteres I. Fam. Perlodidae. Collections Zoologiques du Baron Edmond de Selys-Longchamps. Bruxelles, 4, 1 - 66." type="journal article" year="1912">Klapálek 1912, fig. 8, page 13</bibRefCitation>
) an illustration of “ 
<taxonomicName id="C1B94D3CFFF58978FAEB1A73A3BFFBFE" authority="McLachlan" authorityName="McLachlan" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFF58978FAEB1A73A30CFBFE" italics="true" pageId="4" pageNumber="5">Arcynopteryx compacta</emphasis>
McLachlan
</taxonomicName>
”, which was actually 
<taxonomicName id="C1B94D3CFFF58978FD5A1A2FA1A0FBFE" authority="McLachlan" authorityName="McLachlan" box="[736,1011,1088,1112]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FD5A1A2FA105FBFE" box="[736,854,1088,1112]" italics="true" pageId="4" pageNumber="5">A. dichroa</emphasis>
(McLachlan)
</taxonomicName>
, thus both taxonomic species actually were involved in the misidentification (International Code of Zoological Nomenclature 1999). The 
<typeStatus id="D902881DFFF58978FB4E1A0AA777FBDB" box="[1268,1316,1125,1149]" pageId="4" pageNumber="5">type</typeStatus>
species of 
<taxonomicName id="C1B94D3CFFF58978FF2D1AE7A37EFB06" box="[151,301,1160,1184]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="34CDEAADFFF58978FF2D1AE7A37EFB06" box="[151,301,1160,1184]" italics="true" pageId="4" pageNumber="5">Arcynopteryx</emphasis>
</taxonomicName>
is fixed (under Article 70.3.2 of the Code) as 
<taxonomicName id="C1B94D3CFFF58978FC911AE7A6AEFB06" authority="McLachlan, 1872" authorityName="McLachlan" authorityYear="1872" box="[811,1277,1160,1184]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FC911AE7A64EFB06" box="[811,1053,1160,1184]" italics="true" pageId="4" pageNumber="5">Arcynopteryx dichroa</emphasis>
(
<bibRefCitation id="62284B4EFFF58978FB911AE7A6A6FB06" author="McLachlan" box="[1067,1269,1160,1184]" pageId="4" pageNumber="5" refString="McLachlan, R. (1872) Perlides. In: Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps &amp; McLachlan. Annales de la Societe entomologique de Belgique (Bruxelles), 15, 47 - 55." type="journal article" year="1872">McLachlan, 1872</bibRefCitation>
)
</taxonomicName>
, misidentified as 
<taxonomicName id="C1B94D3CFFF58978FF0F1AC3A0CDFB62" authority="McLachlan, 1872" authorityName="McLachlan" authorityYear="1872" box="[181,670,1196,1220]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFF58978FF0F1AC3A3ECFB62" box="[181,447,1196,1220]" italics="true" pageId="4" pageNumber="5">Arcynopteryx compacta</emphasis>
(
<bibRefCitation id="62284B4EFFF58978FE771AC3A0C6FB62" author="McLachlan" box="[461,661,1196,1220]" pageId="4" pageNumber="5" refString="McLachlan, R. (1872) Perlides. In: Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps &amp; McLachlan. Annales de la Societe entomologique de Belgique (Bruxelles), 15, 47 - 55." type="journal article" year="1872">McLachlan, 1872</bibRefCitation>
)
</taxonomicName>
in the original designation by 
<bibRefCitation id="62284B4EFFF58978FC4E1AC3A6FEFB62" author="Klapalek" box="[1012,1197,1196,1220]" pageId="4" pageNumber="5" refString="Klapalek, F. (1912) Plecopteres I. Fam. Perlodidae. Collections Zoologiques du Baron Edmond de Selys-Longchamps. Bruxelles, 4, 1 - 66." type="journal article" year="1912">Klapalek (1912)</bibRefCitation>
.
</paragraph>
<paragraph id="060636BFFFF58978FF731ABFA6F0FAF2" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="5">
The shape of the female subgenital plate of 
<taxonomicName id="C1B94D3CFFF58978FD0E1ABFA1F4FB4E" box="[692,935,1232,1256]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="34CDEAADFFF58978FD0E1ABFA1F4FB4E" box="[692,935,1232,1256]" italics="true" pageId="4" pageNumber="5">Arcynopteryx dichroa</emphasis>
</taxonomicName>
is variable (
<bibRefCitation id="62284B4EFFF58978FB8A1ABFA692FB4E" author="Brinck" box="[1072,1217,1232,1256]" pageId="4" pageNumber="5" refString="Brinck, P. (1949) Studies on Swedish stoneflies (Plecoptera). Opuscula Entomologica, 11, Lund, 250 pp." type="book" year="1949">Brinck 1949</bibRefCitation>
, 
<bibRefCitation id="62284B4EFFF58978FB761ABFA716FB4E" author="Illies" box="[1228,1349,1232,1256]" pageId="4" pageNumber="5" refString="Illies, J. (1955) Steinfliegen oder Plecoptera. Die Tierwelt Deutschlands, 43, 150 s." type="journal article" year="1955">Illies 1955</bibRefCitation>
, Zhiltsova 1964, 
<bibRefCitation id="62284B4EFFF58978FEB11A9AA3CFFAAA" author="Rauser" box="[267,412,1268,1292]" pageId="4" pageNumber="5" refString="Rauser, J. (1968) Plecoptera. Ergebnisse der zoologischen Forschungen von Dr. Z. Kazsab in der Mongolia. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden, 34, 398 s." type="journal article" year="1968">Rauser 1968</bibRefCitation>
). In the East Palaearctic and Nearctic Regions the females without associated males are difficult to separate from those of 
<taxonomicName id="C1B94D3CFFF58978FE401B77A018FA96" box="[506,587,1304,1328]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="34CDEAADFFF58978FE401B77A018FA96" box="[506,587,1304,1328]" italics="true" pageId="4" pageNumber="5">Skwala</emphasis>
</taxonomicName>
or 
<taxonomicName id="C1B94D3CFFF58978FDCB1B77A0B3FA96" box="[625,736,1304,1328]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="polaris">
<emphasis id="34CDEAADFFF58978FDCB1B77A0B3FA96" box="[625,736,1304,1328]" italics="true" pageId="4" pageNumber="5">A. polaris</emphasis>
</taxonomicName>
, since their subgenital pates are similar, and both genera have the arms of mesosternal ridge meet anterior corners of furcal pits (
<bibRefCitation id="62284B4EFFF58978FCC11B53A6C4FAF2" author="Stewart" box="[891,1175,1340,1364]" pageId="4" pageNumber="5" refString="Stewart, K. &amp; Oswood, M. (2006) The stoneflies (Plecoptera) of Alaska and western Canada. The Caddis Press, Columbus, Ohio. pp. 1 - 325." type="book chapter" year="2006">Stewart &amp; Oswood 2006</bibRefCitation>
).
</paragraph>
<paragraph id="060636BFFFF58978FF731B0FA301F9F6" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="5">
Wide variations in body size, wing length and color pattern are observed throughout the distribution of the species (
<bibRefCitation id="62284B4EFFF58978FF6B1BEBA31FFA3A" author="Illies" box="[209,332,1412,1436]" pageId="4" pageNumber="5" refString="Illies, J. (1955) Steinfliegen oder Plecoptera. Die Tierwelt Deutschlands, 43, 150 s." type="journal article" year="1955">Illies 1955</bibRefCitation>
, 
<bibRefCitation id="62284B4EFFF58978FEEC1BEBA07CFA3A" author="Lillehammer" box="[342,559,1412,1436]" pageId="4" pageNumber="5" refString="Lillehammer, A. (1988) Stoneflies (Plecoptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica. Brill E. J. &amp; Scandinavian Science Press Ltd., Leiden, 165 pp." type="book" year="1988">Lillehammer, 1988</bibRefCitation>
). For example, 
<bibRefCitation id="62284B4EFFF58978FD661BEAA1D3FA3A" author="Despax" box="[732,896,1412,1436]" pageId="4" pageNumber="5" refString="Despax, R. (1951) Plecopteres. Faune de France, 55, 280 pp." type="book" year="1951">Despax (1951)</bibRefCitation>
described three forms of the species with different wing lengths from the Pyrenees. In the population from Kara-Hol Lake in Asia (Tuva, Siberia, 
<collectingCountry id="7EAE762FFFF58978FB671BC7A77FFA66" box="[1245,1324,1448,1472]" name="Russia" pageId="4" pageNumber="5">Russia</collectingCountry>
), the head and pronotum of males are generally pale, with an indistinct brownish pattern. On the Chukotian and Kamchatka peninsulas (the Far East of 
<collectingCountry id="7EAE762FFFF58978FE711B9FA049F9AE" box="[459,538,1520,1544]" name="Russia" pageId="4" pageNumber="5">Russia</collectingCountry>
), the stoneflies have a dark brown body pattern, and the males are usually brachypterous. Both the males and the females from the Ussuri River (Amur R. Basin) have long wings (
<bibRefCitation id="62284B4EFFF58978FB43187BA316F9F6" author="Levanidova" pageId="4" pageNumber="5" refString="Levanidova, I. &amp; Zhiltzova, L. (1979) An annotated list of the stoneflies (Plecoptera) of the Soviet Far East. Internationale Revue der Gesamten Hydrobiologie, 64, 551 - 576." type="journal article" year="1979">Levanidova &amp; Zhiltzova 1979</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>