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<document id="911DAE74A790B872D76110447AEFD388" ID-DOI="10.5281/zenodo.193866" ID-GBIF-Dataset="d4514350-31c1-48f4-a3ac-3c768b6b5e65" ID-ISSN="1175-5326" ID-Zenodo-Dep="193866" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460174872716" checkinUser="plazi" docAuthor="Ledford, Joel M. &amp; Griswold, Charles E." docDate="2010" docId="881A879DFF83FFC8FF5DF96E1F39F8D9" docLanguage="en" docName="zt02391p032.pdf" docOrigin="Zootaxa 2391" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Darkoneta reddelli Ledford &amp; Griswold, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="7423FFE5FF9AFFD3FFCAFFCC1B3EFFC0" masterDocTitle="A study of the subfamily Archoleptonetinae (Araneae, Leptonetidae) with a review of the morphology and relationships for the Leptonetidae" masterLastPageNumber="32" masterPageNumber="1" pageNumber="26" updateTime="1698236069209" updateUser="plazi">
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<mods:title id="F2B14FDB86C411CDFB36D86A671C203B">A study of the subfamily Archoleptonetinae (Araneae, Leptonetidae) with a review of the morphology and relationships for the Leptonetidae</mods:title>
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<mods:namePart id="8C09CC03BDFFD7E19BE384683CA53775">Ledford, Joel M.</mods:namePart>
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<mods:namePart id="3395FCCAB4F485110753DACD56E3B23C">Griswold, Charles E.</mods:namePart>
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<treatment id="881A879DFF83FFC8FF5DF96E1F39F8D9" ID-DOI="http://doi.org/10.5281/zenodo.6204734" ID-GBIF-Taxon="119383534" ID-Zenodo-Dep="6204734" LSID="urn:lsid:plazi:treatment:881A879DFF83FFC8FF5DF96E1F39F8D9" httpUri="http://treatment.plazi.org/id/881A879DFF83FFC8FF5DF96E1F39F8D9" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
<subSubSection id="48A96500FF83FFCAFF5DF96E1AB2F91F" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="000C368BFF83FFCAFF5DF96E1AD8F97C" blockId="25.[151,486,1698,1759]" box="[151,486,1698,1724]" pageId="25" pageNumber="26">
<heading id="5B4481E7FF83FFCAFF5DF96E1AD8F97C" bold="true" box="[151,486,1698,1724]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<emphasis id="32C7EA99FF83FFCAFF5DF96E1AD8F97C" bold="true" box="[151,486,1698,1724]" pageId="25" pageNumber="26">
<emphasis id="32C7EA99FF83FFCAFF5DF96E1ABFF97C" bold="true" box="[151,385,1698,1724]" italics="true" pageId="25" pageNumber="26">
<taxonomicName id="C7B34D08FF83FFCAFF5DF96E1A44F97C" box="[151,378,1698,1724]" class="Arachnida" family="Leptonetidae" genus="Darkoneta" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="reddelli" status="sp. nov.">Darkoneta reddelli</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="29F457E2FF83FFCAFE42F96E1AD8F97C" box="[392,486,1698,1724]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="000C368BFF83FFCAFF5DF9091AB2F91F" blockId="25.[151,486,1698,1759]" box="[151,396,1733,1759]" pageId="25" pageNumber="26">
(
<figureCitation id="98882A0EFF83FFCAFF55F9091A3CF91F" box="[159,258,1733,1759]" captionStart="FIGURES 29 34" captionStartId="17.[151,273,1856,1880]" captionTargetBox="[194,1387,194,1828]" captionTargetId="figure@17.[194,1387,194,1828]" captionTargetPageId="17" captionText="FIGURES 29 34. Archoleptonetinae general morphology. 29. Darkoneta quetzal sp. nov., male carapace, dorsal, arrows to stridulatory picks; 30. Darkoneta quetzal sp. nov., stridulatory structures, dorsal; 31. Archoleptoneta schusteri, female, Burdell Mountain, Marin Co., CA, patella III, dorsal, showing gland; 32. Darkoneta reddelli sp. nov., juvenile carapace, dorsal; 33. Archoleptoneta schusteri, female, Burdell Mountain, Marin Co., CA, carapace, posterior; 34. Darkoneta quetzal sp. nov., male abdomen, ventral, showing gonopore anteriad of epigastric furrow." httpUri="https://zenodo.org/record/193872/files/figure.png" pageId="25" pageNumber="26">Figs. 32</figureCitation>
,
<figureCitation id="98882A0EFF83FFCAFEC4F9091A10F91F" box="[270,302,1733,1759]" captionStart="FIGURES 43 48" captionStartId="19.[151,269,1864,1888]" captionTargetBox="[151,1436,567,1834]" captionTargetId="figure@19.[151,1441,567,1834]" captionTargetPageId="19" captionText="FIGURES 43 48. Archoleptonetinae spinnerets and calamistra, SEM. 43. Archoleptoneta schusteri, female, spinnerets, ventral, Burdell Mountain, Marin County, CA, USA; 44. A. schusteri, female, cribellum, Burdell Mountain; 45. A. schusteri, female, calamistrum, right metatarsus IV, Burdell Mountain; 46. Darkoneta stridulans, female, spinnerets, ventral, Chiriqui, Panama; 47. A. schusteri, female, cribellum, spigots, Burdell Mountain; 48. A. schusteri, female, calamistrum, right metatarsus IV, Burdell Mountain." httpUri="https://zenodo.org/record/193874/files/figure.png" pageId="25" pageNumber="26">46</figureCitation>
,
<figureCitation id="98882A0EFF83FFCAFEF0F9091ABAF91F" box="[314,388,1733,1759]" captionStart="FIGURES 91 96" captionStartId="26.[151,270,1906,1930]" captionTargetBox="[151,1435,679,1884]" captionTargetId="figure@26.[151,1436,679,1884]" captionTargetPageId="26" captionText="FIGURES 91 96. Microstructure views of Darkoneta reddelli sp. nov., male right palp, Cueva de Tasalolpan, Mexico. 91. Retrolateral; 92. Ventral; 93. Prolateral; 94. Bulb, ventral; 95. Embolus, ventral; 96. Accessory sclerites, prolateral. E, embolus; MS, median sclerite; PRS, prolateral sclerite; RLS, retrolateral sclerite." httpUri="https://zenodo.org/record/193882/files/figure.png" pageId="25" pageNumber="26">9196</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="48A96500FF83FFCAFF5DF8C31E6CF85A" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="000C368BFF83FFCAFF5DF8C31A32F88F" blockId="25.[151,1436,1807,2020]" pageId="25" pageNumber="26">
<emphasis id="32C7EA99FF83FFCAFF5DF8C31A6AF8E9" bold="true" box="[151,340,1807,1833]" pageId="25" pageNumber="26">
<typeStatus id="DF088829FF83FFCAFF5DF8C31BE8F8E9" box="[151,214,1807,1833]" pageId="25" pageNumber="26">Type</typeStatus>
material.
</emphasis>
Male
<typeStatus id="DF088829FF83FFCAFE69F8DC1930F8EA" box="[419,526,1808,1834]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
from Cueva de Tasalolpan,
<quantity id="C74B9B6EFF83FFCAFCA2F8DC1898F8EA" box="[872,934,1808,1834]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="km" value="5.0">5 km</quantity>
southwest of Cuetzalan, Puebla,
<collectingCountry id="78A4761BFF83FFCAFA8AF8DC1EA2F8EA" box="[1344,1436,1808,1834]" name="Mexico" pageId="25" pageNumber="26">Mexico</collectingCountry>
(
<collectionCode id="66A2AE4EFF83FFCAFF55F8F91A3EF88F" LSID="urn:lsid:biocol.org:col:34925" box="[159,256,1845,1871]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH</collectionCode>
).
</paragraph>
<paragraph id="000C368BFF83FFCAFF0EF8961E6CF85A" blockId="25.[151,1436,1807,2020]" pageId="25" pageNumber="26">
<emphasis id="32C7EA99FF83FFCAFF0EF8961931F8B4" bold="true" box="[196,527,1882,1908]" pageId="25" pageNumber="26">Other material examined.</emphasis>
<collectingCountry id="78A4761BFF83FFCAFDDDF8961949F8B4" box="[535,631,1882,1908]" name="Mexico" pageId="25" pageNumber="26">Mexico</collectingCountry>
: Puebla: Cueva de Tasalolpan,
<quantity id="C74B9B6EFF83FFCAFC25F8961F12F8B4" box="[1007,1068,1882,1908]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="km" value="5.0">5 km</quantity>
southwest Cuetzalan,
<geoCoordinate id="6587504CFF83FFCAFAF6F8961EA6F8B4" box="[1340,1432,1882,1908]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="20.02">20.02N</geoCoordinate>
,
<geoCoordinate id="6587504CFF83FFCAFF5DF84C1BCBF85A" box="[151,245,1920,1946]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="555" value="-97.54">97.54W</geoCoordinate>
,
<date id="740D104BFF83FFCAFECAF84C1AB7F85A" box="[256,393,1920,1946]" pageId="25" pageNumber="26" value="1976-12-22">22.xii.1976</date>
, J. Reddell, A. Grubbs, C. Soileau, D. McKenzie,
<specimenCount id="16B5FD02FF83FFCAFC1CF84C1F74F85A" box="[982,1098,1920,1946]" pageId="25" pageNumber="26" type="female">2 females</specimenCount>
,
<specimenCount id="16B5FD02FF83FFCAFB9CF84C1FE8F85A" box="[1110,1238,1920,1946]" pageId="25" pageNumber="26" type="juvenile">6 juveniles</specimenCount>
(
<collectionCode id="66A2AE4EFF83FFCAFB2FF84C1E7BF85A" LSID="urn:lsid:biocol.org:col:34925" box="[1253,1349,1920,1946]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="48A96500FF83FFC9FF0EF8681AE6FF72" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="000C368BFF83FFC9FF0EF8681AE6FF72" blockId="25.[151,1436,1807,2020]" lastBlockId="26.[151,1437,152,626]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="32C7EA99FF83FFCAFF0EF8681A7DF87E" bold="true" box="[196,323,1956,1982]" pageId="25" pageNumber="26">Diagnosis.</emphasis>
Pale, eyeless troglobite (
<figureCitation id="98882A0EFF83FFCAFDADF8691982F87F" box="[615,700,1957,1983]" captionStart="FIGURES 29 34" captionStartId="17.[151,273,1856,1880]" captionTargetBox="[194,1387,194,1828]" captionTargetId="figure@17.[194,1387,194,1828]" captionTargetPageId="17" captionText="FIGURES 29 34. Archoleptonetinae general morphology. 29. Darkoneta quetzal sp. nov., male carapace, dorsal, arrows to stridulatory picks; 30. Darkoneta quetzal sp. nov., stridulatory structures, dorsal; 31. Archoleptoneta schusteri, female, Burdell Mountain, Marin Co., CA, patella III, dorsal, showing gland; 32. Darkoneta reddelli sp. nov., juvenile carapace, dorsal; 33. Archoleptoneta schusteri, female, Burdell Mountain, Marin Co., CA, carapace, posterior; 34. Darkoneta quetzal sp. nov., male abdomen, ventral, showing gonopore anteriad of epigastric furrow." httpUri="https://zenodo.org/record/193872/files/figure.png" pageId="25" pageNumber="26">Fig. 32</figureCitation>
) separated from other
<taxonomicName id="C7B34D08FF83FFCAFC09F8691F7EF87E" box="[963,1088,1957,1982]" class="Arachnida" family="Leptonetidae" genus="Darkoneta" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="32C7EA99FF83FFCAFC09F8691F7EF87E" box="[963,1088,1957,1982]" italics="true" pageId="25" pageNumber="26">Darkoneta</emphasis>
</taxonomicName>
species by having males with a stout embolus that extends 1/2 the length of the tarsus (
<figureCitation id="98882A0EFF83FFCAFCF8F806188DF824" box="[818,947,1994,2020]" captionStart="FIGURES 91 96" captionStartId="26.[151,270,1906,1930]" captionTargetBox="[151,1435,679,1884]" captionTargetId="figure@26.[151,1436,679,1884]" captionTargetPageId="26" captionText="FIGURES 91 96. Microstructure views of Darkoneta reddelli sp. nov., male right palp, Cueva de Tasalolpan, Mexico. 91. Retrolateral; 92. Ventral; 93. Prolateral; 94. Bulb, ventral; 95. Embolus, ventral; 96. Accessory sclerites, prolateral. E, embolus; MS, median sclerite; PRS, prolateral sclerite; RLS, retrolateral sclerite." httpUri="https://zenodo.org/record/193882/files/figure.png" pageId="25" pageNumber="26">Fig. 92, 95</figureCitation>
) and a proventrally curved and thickened prolateral sclerite (
<figureCitation id="98882A0EFF80FFC9FEB9FF541AF5FF72" box="[371,459,152,178]" captionStart="FIGURES 91 96" captionStartId="26.[151,270,1906,1930]" captionTargetBox="[151,1435,679,1884]" captionTargetId="figure@26.[151,1436,679,1884]" captionTargetPageId="26" captionText="FIGURES 91 96. Microstructure views of Darkoneta reddelli sp. nov., male right palp, Cueva de Tasalolpan, Mexico. 91. Retrolateral; 92. Ventral; 93. Prolateral; 94. Bulb, ventral; 95. Embolus, ventral; 96. Accessory sclerites, prolateral. E, embolus; MS, median sclerite; PRS, prolateral sclerite; RLS, retrolateral sclerite." httpUri="https://zenodo.org/record/193882/files/figure.png" pageId="26" pageNumber="27">Fig. 96</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="48A96500FF80FFC9FF0EFF711806FDB2" pageId="26" pageNumber="27" type="description">
<paragraph id="000C368BFF80FFC9FF0EFF711BF0FF3C" blockId="26.[151,1437,152,626]" pageId="26" pageNumber="27">
<emphasis id="32C7EA99FF80FFC9FF0EFF711A97FF17" bold="true" box="[196,425,189,215]" pageId="26" pageNumber="27">Description. Male</emphasis>
(
<typeStatus id="DF088829FF80FFC9FE73FF711917FF17" box="[441,553,189,215]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
). Total length 0.87. Specimen pale, eyeless; missing patella-tarsus for all legs.
</paragraph>
<paragraph id="000C368BFF80FFC9FF0EFEC418F2FEAC" blockId="26.[151,1437,152,626]" pageId="26" pageNumber="27">Carapace 0.38 long, 0.32 wide, height at fovea 0.56 x carapace width; clypeus 0.04 high, chelicerae 0.21 long, fang furrow with three small teeth on retromargin and a single large proximal tooth. Sternum 0.26 long, 0.28 wide; labium.0 5 long,.0 6 wide; palpal coxae.12 long,.0 7 wide.</paragraph>
<paragraph id="000C368BFF80FFC9FF0EFEB41A71FE1C" blockId="26.[151,1437,152,626]" pageId="26" pageNumber="27">Femora elongate and covered with fine setae. Leg measurements (Femur only): I: 0.9; II: 0.58; III: 0.73; IV: 0.93; pedipalpus: 0.25 + 0.09 + 0.11 + 0.25 = [0.7]. Femur I 2.8 x carapace width, palpal femur 0.78 x carapace width.</paragraph>
<paragraph id="000C368BFF80FFC9FF0EFE2419C8FD8C" blockId="26.[151,1437,152,626]" pageId="26" pageNumber="27">Palpal bulb 0.15 long, 0.08 wide; Prolateral sclerite enlarged and with a retrolateral bend; median sclerite long and thin, tapering retrolaterally; retrolateral sclerite sharply tapering retrolaterally; embolus short, enlarged, and extending 1/2 the length of the tarsus.</paragraph>
<paragraph id="000C368BFF80FFC9FF0EFD941806FDB2" blockId="26.[151,1437,152,626]" box="[196,824,600,626]" pageId="26" pageNumber="27">Abdomen pale, without pattern, 0.49 long, 0.34 wide.</paragraph>
</subSubSection>
<caption id="54CC6603FF80FFC9FF5DF8BE18C8F809" httpUri="https://zenodo.org/record/193882/files/figure.png" pageId="26" pageNumber="27" targetBox="[151,1435,679,1884]" targetPageId="26">
<paragraph id="000C368BFF80FFC9FF5DF8BE18C8F809" blockId="26.[151,1437,1906,1993]" pageId="26" pageNumber="27">
<emphasis id="32C7EA99FF80FFC9FF5DF8BE1A60F84A" bold="true" box="[151,350,1906,1930]" pageId="26" pageNumber="27">FIGURES 9196.</emphasis>
Microstructure views of
<taxonomicName id="C7B34D08FF80FFC9FDA4F8BE1804F849" box="[622,826,1906,1929]" class="Arachnida" family="Leptonetidae" genus="Darkoneta" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="reddelli" status="sp. nov.">
<emphasis id="32C7EA99FF80FFC9FDA4F8BE1804F849" box="[622,826,1906,1929]" italics="true" pageId="26" pageNumber="27">Darkoneta reddelli</emphasis>
</taxonomicName>
<emphasis id="32C7EA99FF80FFC9FC8BF8BE18A6F84A" bold="true" box="[833,920,1906,1930]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="29F457E2FF80FFC9FC8BF8BE18A6F84A" box="[833,920,1906,1930]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male right palp, Cueva de Tasalolpan, Mexico. 91. Retrolateral; 92. Ventral; 93. Prolateral; 94. Bulb, ventral; 95. Embolus, ventral; 96. Accessory sclerites, prolateral. E, embolus; MS, median sclerite; PRS, prolateral sclerite; RLS, retrolateral sclerite.
</paragraph>
</caption>
<caption id="54CC6603FF81FFC8FF5DFA521EA9FA36" httpUri="https://zenodo.org/record/193883/files/figure.png" pageId="27" pageNumber="28" targetBox="[151,1435,194,1417]" targetPageId="27">
<paragraph id="000C368BFF81FFC8FF5DFA521EA9FA36" blockId="27.[151,1437,1438,1526]" pageId="27" pageNumber="28">
<emphasis id="32C7EA99FF81FFC8FF5DFA521A49FA76" bold="true" box="[151,375,1438,1462]" pageId="27" pageNumber="28">FIGURES 97102.</emphasis>
Microstructure views of
<taxonomicName id="C7B34D08FF81FFC8FD56FA531884FA76" box="[668,954,1439,1462]" class="Arachnida" family="Leptonetidae" genus="Archoleptoneta" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="schusteri">
<emphasis id="32C7EA99FF81FFC8FD56FA531884FA76" box="[668,954,1439,1462]" italics="true" pageId="27" pageNumber="28">Archoleptoneta schusteri</emphasis>
</taxonomicName>
, male right palp, Soquel, Santa Cruz Co., California, USA. 97. Retrolateral; 98. Ventral; 99. Prolateral; 100. Bulb, prolateral; 101. Accessory sclerites, ventral; 102. Accessory sclerites, prolateral. E, embolus; MS, median sclerite; PRS, prolateral sclerite; RLS, retrolateral sclerite.
</paragraph>
</caption>
<subSubSection id="48A96500FF81FFC8FF0EF9D21A14F969" pageId="27" pageNumber="28" type="discussion">
<paragraph id="000C368BFF81FFC8FF0EF9D21A14F969" blockId="27.[151,1436,1566,1817]" pageId="27" pageNumber="28">
<emphasis id="32C7EA99FF81FFC8FF0EF9D21A03F9F8" bold="true" box="[196,317,1566,1592]" pageId="27" pageNumber="28">Remarks.</emphasis>
This series of specimens includes eight individuals, two of which are adult males. Most of the specimens are missing leg segments and their abdomens. While a female individual is almost certainly a part of the series, the absence of abdomens makes it impossible to identify a mature specimen, therefore no female is described.
</paragraph>
</subSubSection>
<subSubSection id="48A96500FF81FFC8FF0EF9781961F934" pageId="27" pageNumber="28" type="etymology">
<paragraph id="000C368BFF81FFC8FF0EF9781961F934" blockId="27.[151,1436,1566,1817]" pageId="27" pageNumber="28">
<emphasis id="32C7EA99FF81FFC8FF0EF9781A6FF90E" bold="true" box="[196,337,1716,1742]" pageId="27" pageNumber="28">Etymology.</emphasis>
This species is named in honor of James Reddell, collector of this and many other species of leptonetids throughout North
<collectingCountry id="78A4761BFF81FFC8FE38F9161965F934" box="[498,603,1754,1780]" name="United States of America" pageId="27" pageNumber="28">America</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="48A96500FF81FFC8FF0EF9321F39F8D9" box="[196,1031,1790,1817]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="000C368BFF81FFC8FF0EF9321F39F8D9" blockId="27.[151,1436,1566,1817]" box="[196,1031,1790,1817]" pageId="27" pageNumber="28">
<emphasis id="32C7EA99FF81FFC8FF0EF9321A5AF8D8" bold="true" box="[196,356,1790,1816]" pageId="27" pageNumber="28">Distribution.</emphasis>
Known only from Cueva de Tasalolpan, Puebla,
<collectingCountry id="78A4761BFF81FFC8FC6FF9331F3DF8D9" box="[933,1027,1791,1817]" name="Mexico" pageId="27" pageNumber="28">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>