156 lines
16 KiB
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156 lines
16 KiB
XML
<document id="DD371AAFEA4AF0367A39E0007687767F" ID-CLB-Dataset="28501" ID-DOI="http://dx.doi.org/10.3897/zookeys.826.26488" ID-GBIF-Dataset="9d62af83-e4f5-423b-8e12-7dd567f69921" ID-PMC="PMC6405737" ID-Pensoft-Pub="1313-2970-826-1" ID-PubMed="30858752" ID-ZooBank="B1550A3AE54744509A44BC4366D5E110" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-826-1" ModsDocOrigin="ZooKeys 826" ModsDocTitle="Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records" checkinTime="1551446249571" checkinUser="pensoft" docAuthor="Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z." docDate="2019" docId="7AD0E12BCD9E76B9E7821622C1A9B876" docLanguage="en" docName="ZooKeys 826: 1-343" docOrigin="ZooKeys 826" docSource="http://dx.doi.org/10.3897/zookeys.826.26488" docTitle="Dichotomius (Dichotomius) satanas subsp. angustus Luederwaldt 1923" docType="treatment" docVersion="7" lastPageNumber="122" masterDocId="FFECFFBAFFFC2D25D00B9843FFD67749" masterDocTitle="Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records" masterLastPageNumber="343" masterPageNumber="1" pageNumber="122" updateTime="1701379963039" updateUser="ExternalLinkService">
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<mods:title id="B922AF44830A087EF597D446665FA3A2">Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records</mods:title>
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<mods:namePart id="CCA552A455313188A8343FC14E156CFD">Chamorro, William</mods:namePart>
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<mods:namePart id="C0809EEED488B59D2FBBC4D7CFBC4A95">Marin-Armijos, Diego</mods:namePart>
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<mods:namePart id="07F8C9E7A4C7EA0D9BBBB6D4357400D3">senjo, Angelico</mods:namePart>
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<mods:namePart id="56D0C6AC7087C1F81C19BEF4126E46D5">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="98993EC03749B8677D6E74FC56B8E500">text</mods:typeOfResource>
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<mods:date id="44D37692497A0A48F85FCC4F990B2FD0">2019</mods:date>
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<mods:detail id="A44FB721B23D1F56539E1C7014023614" type="volume">
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<mods:number id="2E6FF980F367C6E6443AB8974D289B05">826</mods:number>
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<mods:classification id="DF481E2F3A8F3C15D8728FC75389D5AA">journal article</mods:classification>
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<mods:identifier id="5F75DB86ECCA42ED37492B67AB347251" type="DOI">http://dx.doi.org/10.3897/zookeys.826.26488</mods:identifier>
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<treatment id="7AD0E12BCD9E76B9E7821622C1A9B876" ID-GBIF-Taxon="154127762" LSID="urn:lsid:plazi:treatment:7AD0E12BCD9E76B9E7821622C1A9B876" httpUri="http://treatment.plazi.org/id/7AD0E12BCD9E76B9E7821622C1A9B876" lastPageNumber="122" pageId="121" pageNumber="122">
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<subSubSection id="CB5D9E55C3CFB85B2B0051397D787A48" pageId="121" pageNumber="122" type="nomenclature">
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<paragraph id="E3C3BF4F23F64478AA855D726D9F8DC6" pageId="121" pageNumber="122">
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<taxonomicName id="F4B12CF155CF930ECB5414C71721537D" ID-CoL="7ZS9C" authority="Luederwaldt, 1923" authorityName="Luederwaldt" authorityYear="1923" class="Insecta" family="Scarabaeidae" genus="Dichotomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichotomius (Dichotomius) satanas" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="subSpecies" species="satanas" subGenus="Dichotomius" subSpecies="angustus">Dichotomius (Dichotomius) satanas angustus (Luederwaldt, 1923)</taxonomicName>
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Plate 28A
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</paragraph>
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</subSubSection>
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<subSubSection id="1B11B6B93E8A67B69B49B266632B4009" pageId="121" pageNumber="122" type="reference_group">
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<paragraph id="49A2B4F1B42B8542AC15F36A7C85967B" pageId="121" pageNumber="122">
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<taxonomicName id="6C3F322DDC5E5D7AC586F290692A02A1" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Pinotus satanas var. angustus" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="variety" species="satanas" variety="angustus">Pinotus satanas var. angustus</taxonomicName>
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Luederwaldt, 1923: 10 (original description. Type locality: Ecuador Sarayacu; Ost-Kordillere, Sued-Ecuador [= eastern Cordillera, southern Ecuador], 300 m; Canolos [= Canelos] Ost-Ecuador [= eastern Cordillera, Ecuador], 700 m; Macas Ost-Ecuador [= eastern Cordillera, Ecuador]; Loja-Sabanilla Ost-Ecuador [= eastern Cordillera, Ecuador]; Bannos-Mirador [=
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<normalizedToken id="2003998D49F85D81DBB5CECD998C0C87" originalValue="Baños-Mirador">Banos-Mirador</normalizedToken>
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] Ost-Ecuador [= eastern Cordillera, Ecuador], 14-1600 m).
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</paragraph>
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<paragraph id="8A86CE88113CE33C01DD8C6F0D945077" pageId="121" pageNumber="122">
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<taxonomicName id="218F1B35840551B673DB464E8A7D8660" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Pinotus satanas var. angustus" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="variety" species="satanas" variety="angustus">Pinotus satanas var. angustus</taxonomicName>
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:
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<bibRefCitation id="E9CE76C321553691C1D3671AFE1C20A9" author="Luederwaldt, H" journalOrPublisher="Revista do Museu Paulista" pageId="272" pageNumber="273" pagination="1 - 174" title="As especies brasileiras do genero Pinotus, com algumas consideractes tambem sobra outras especies." volume="16" year="1929">Luederwaldt 1929</bibRefCitation>
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: 36 (characters in key).
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</paragraph>
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<paragraph id="0028B25164306FA46A5F1266B5B65CF1" pageId="121" pageNumber="122">
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<taxonomicName id="9186BEDC885E9A7DA80A02E9041BE5D0" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinotus angustus" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="angustus">Pinotus angustus</taxonomicName>
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:
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<bibRefCitation id="DA72A47D9AE402AF6E517F7A40296A41" author="Blackwelder, RE" journalOrPublisher="National Museum Bulletin" pageId="262" pageNumber="263" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
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: 206 (list of species from Latin America);
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<bibRefCitation id="8AEF20172C06A2ACABC75C420346BF07" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
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: 320-321 (cited for Ecuador).
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</paragraph>
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<paragraph id="C80945BCE48C9419C32732F705D6BD90" pageId="121" pageNumber="122">
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<taxonomicName id="992995E480E11D411A5F6EC181C71B43" class="Insecta" family="Scarabaeidae" genus="Dichotomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichotomius (Dichotomius) satanas" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="subSpecies" species="satanas" subGenus="Dichotomius" subSpecies="angustus">Dichotomius (Dichotomius) satanas angustus</taxonomicName>
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:
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<bibRefCitation id="E7F1F317F3F0F7B094A407411D806025" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
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: 95 (cited for Ecuador).
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</paragraph>
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</subSubSection>
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<subSubSection id="7A8BFC021EB80EA6C4A3D0AE37055697" pageId="121" pageNumber="122" type="type specimens">
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<paragraph id="0BA206A3384D9B4B4F2727DA7261E0B5" pageId="121" pageNumber="122">Type specimens.</paragraph>
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<paragraph id="3A4D43FFB700E8C5DB4AA62B3D9DD845" pageId="121" pageNumber="122">
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<taxonomicName id="8EDE60B91B672D2761BFCFD2594B47AF" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Pinotus satanas var. angustus" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="variety" species="satanas" variety="angustus">Pinotus satanas var. angustus</taxonomicName>
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Luederwaldt, 1923. Two syntypes examined deposited at the MZUSP. Lectotype to be designated in a future work on this species group.
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</paragraph>
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<paragraph id="81ED3A807FF3A40B2C18786D7CF27902" pageId="121" pageNumber="122">
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Syntype (♂): "O.ECUADOR / Canelos / F. Ohs. 23. 12. 05 [p]", "COTIPO [p, black margin, red label]", "Pinotus ♂ / angustus
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<normalizedToken id="A91DDFC0D13DA74A337348C6FE8ABC4E" originalValue="Lüd">Lued</normalizedToken>
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. /
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<normalizedToken id="064B5AEBA35B47C61CA4C439C594B7BE" originalValue="Lüd">Lued</normalizedToken>
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. det. 22. [hw]", "17228 [p]".
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</paragraph>
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<paragraph id="2FA8A4A2F3D5A6E4C8658D02DFAA3DF3" pageId="121" pageNumber="122">Syntype (♂): "Sud-Ecuador, / Ostcordill 3000 m. / F. Ohaus 19-x- [hw]", "17226 [p]", "COTIPO [p, black margin, red label]".</paragraph>
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</subSubSection>
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<subSubSection id="01D0A925B1E50B1744A07D05655CAB72" pageId="121" pageNumber="122" type="distribution">
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<paragraph id="0FBEA258F7232C1572D10D53E3062B1E" pageId="121" pageNumber="122">Distribution.</paragraph>
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<paragraph id="C9BC4CA57E8E0EC909A8D7F8B2E32E4A" pageId="121" pageNumber="122">Colombia and Ecuador.</paragraph>
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</subSubSection>
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<subSubSection id="A96AFB38A2E6B760D06D90724059B30A" pageId="121" pageNumber="122" type="records examined">
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<paragraph id="00718519AFFFCACDDFBA1AE5A1F15768" pageId="121" pageNumber="122">Records examined.</paragraph>
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<paragraph id="2290EFBD70A9D0F8D4181B45EE6CEFD4" pageId="121" pageNumber="122">
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LOJA: without specific locality (1 specimen MZUSP). NAPO: Archidona (1 specimen CEMT);
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<normalizedToken id="ADC5AAA732785829ED0CE4B1B1925CD8" originalValue="Cabañas">Cabanas</normalizedToken>
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San Isidro, 2 km NW de Cosanga, 2150 m (2 specimens CEMT); Cosanga Yanayacu Biost, 2150 m (3 specimens MECN);
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<normalizedToken id="5544BEA6D024861E02B7C86EBB6EA663" originalValue="Río">Rio</normalizedToken>
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<normalizedToken id="BA014D374F6D1A41D7EAC6647F04B883" originalValue="Hollín">Hollin</normalizedToken>
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, 1100 m (1 specimen CEMT). PASTAZA: Canelos (1 specimen MZUSP). TUNGURAHUA:
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<normalizedToken id="6D83ED91EB13E8DEF31A2AE0E2A20742" originalValue="Baños">Banos</normalizedToken>
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El Topo, 1590 m (24 specimens CEMT).
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<normalizedToken id="F5E5D7EA53AD68BE0577B30CE93C2E48" originalValue="SUCUMBÍOS">SUCUMBIOS</normalizedToken>
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: La
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<normalizedToken id="EF5FE9EDB0CA90F40884557526357A60" originalValue="Sofía">Sofia</normalizedToken>
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, 1800 m (2 specimens MUTPL); Sebundoy, 2200 m (2 specimens MECN). ZAMORA CHINCHIPE: Bombuscaru, Parque Nacional Podocarpus, 1150 m (2 specimens MECN); Chito
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<normalizedToken id="DD22DC9272AA0B6321767680004A2A3F" originalValue="Río">Rio</normalizedToken>
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Sangolas, 1800 m (4 specimens MUTPL); San Andres, 1850 m (4 specimens CEMT). UNDETERMINED PROVINCE: Ostcordill [= without specific locality], 3000 m (1 specimen MZUSP).
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</paragraph>
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</subSubSection>
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<subSubSection id="67FC1C0A4629ACAD0F60E17039E7D5C6" pageId="121" pageNumber="122" type="literature records">
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<paragraph id="C511FACB56511E3565FED74768166396" pageId="121" pageNumber="122">Literature records.</paragraph>
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<paragraph id="F45A0D9E9D1865A4D4FFD5AB40945285" pageId="121" pageNumber="122">
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LOJA: Sabanilla [= El Tambo, ZAMORA CHINCHIPE] (
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<bibRefCitation id="6FEA3955EEDB59E01DF4057B723EEFF9" author="Luederwaldt, H" journalOrPublisher="Separata da Revista do Museu Paulista" pageId="272" pageNumber="273" pagination="1 - 13" title="Neue Pinotus Arten." volume="14" year="1923">Luederwaldt 1923</bibRefCitation>
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: 11). MORONA SANTIAGO: Macas: (
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<bibRefCitation id="B70D2EC43828895C65AF318072B15EF4" author="Luederwaldt, H" journalOrPublisher="Separata da Revista do Museu Paulista" pageId="272" pageNumber="273" pagination="1 - 13" title="Neue Pinotus Arten." volume="14" year="1923">Luederwaldt 1923</bibRefCitation>
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: 11). PASTAZA: Canolos [= Canelos], 700 m (
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<bibRefCitation id="9D48EF0C8817397E32CDAE4F66CF5AB7" author="Luederwaldt, H" journalOrPublisher="Separata da Revista do Museu Paulista" pageId="272" pageNumber="273" pagination="1 - 13" title="Neue Pinotus Arten." volume="14" year="1923">Luederwaldt 1923</bibRefCitation>
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: 11); Sarayacu (
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<bibRefCitation id="9FB53056B24559C469183D1D815E2F9E" author="Luederwaldt, H" journalOrPublisher="Separata da Revista do Museu Paulista" pageId="272" pageNumber="273" pagination="1 - 13" title="Neue Pinotus Arten." volume="14" year="1923">Luederwaldt 1923</bibRefCitation>
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: 11). TUNGURAHUA: Bannos-Mirador [=
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<normalizedToken id="2682F075418A50F667706DDEB07D9B6F" originalValue="Baños">Banos</normalizedToken>
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Mirador], 1400-1600 m. (
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<bibRefCitation id="7DA7A8049559BF961EBA8E8A84B8DF2D" author="Luederwaldt, H" journalOrPublisher="Separata da Revista do Museu Paulista" pageId="272" pageNumber="273" pagination="1 - 13" title="Neue Pinotus Arten." volume="14" year="1923">Luederwaldt 1923</bibRefCitation>
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: 11). UNDETERMINED PROVINCE: Ost-Kordillere, Sued-Ecuador, 3000 m (
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<bibRefCitation id="91AF7F4E53223FD1684A4A5D5494A451" author="Luederwaldt, H" journalOrPublisher="Separata da Revista do Museu Paulista" pageId="272" pageNumber="273" pagination="1 - 13" title="Neue Pinotus Arten." volume="14" year="1923">Luederwaldt 1923</bibRefCitation>
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: 11).
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</paragraph>
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</subSubSection>
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<subSubSection id="A7EA776107407AFF78A3C135C4DF4CDC" pageId="121" pageNumber="122" type="temporal data">
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<paragraph id="4EAB81E5C972714DB199AD3012A81D29" pageId="121" pageNumber="122">Temporal data.</paragraph>
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<paragraph id="DE73D30008653329827DA683B42332FD" pageId="121" pageNumber="122">Collected in January, February, May, July, October, September, November, and December.</paragraph>
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</subSubSection>
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<subSubSection id="B645B1C68A25439E317531086CBE9C99" pageId="121" pageNumber="122" type="remarks">
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<paragraph id="69438C239F1B8163ECA887BED24E9902" pageId="121" pageNumber="122">Remarks.</paragraph>
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<paragraph id="DAA5133354A0AB94C0428B41B048B679" pageId="121" pageNumber="122">Inhabits the evergreen foothill forests of the Amazon region from 700-1150 m a.s.l. In the Andean region, it was recorded in the evergreen montane forests and montane cloud forests from 1600-2200 m a.s.l. Collected with pitfall traps baited with human feces.</paragraph>
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</subSubSection>
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</treatment>
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</document> |