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<document id="FA6D6F33D3847DEDF99C7DD896FD2030" ID-CLB-Dataset="31172" ID-DOI="10.11646/zootaxa.4363.4.12" ID-GBIF-Dataset="5e7426db-618e-4be0-818a-0f1b4e11539b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1114811" ID-ZooBank="90765905-FD57-4412-BE16-01DF1E04E6D5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513152347109" checkinUser="plazi" docAuthor="Nurinsiyah, Ayu Savitri &amp; Hausdorf, Bernhard" docDate="2017" docId="373B87F8FFF9001FACAC7C22498EFD5E" docLanguage="en" docName="zootaxa.4363.4.11.pdf" docOrigin="Zootaxa 4363 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dicharax Kobelt &amp; Mollendorff 1900, n. sp." docType="treatment" docVersion="9" lastPageNumber="591" masterDocId="CB02FF80FFF9001DAC3B793C4870FF87" masterDocTitle="Dicharax (?) candrakirana n. sp. (Gastropoda: Cyclophoridae) from Sempu Island, Indonesia" masterLastPageNumber="591" masterPageNumber="589" pageNumber="589" updateTime="1698509516600" updateUser="ExternalLinkService">
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<mods:title id="FA9BA448E31C369D5810C024DF6C7130">Dicharax (?) candrakirana n. sp. (Gastropoda: Cyclophoridae) from Sempu Island, Indonesia</mods:title>
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<mods:namePart id="9AA48989834A943063ED438449581E73">Nurinsiyah, Ayu Savitri</mods:namePart>
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<mods:namePart id="CF043BC21A99D868E20B48514D26793F">Hausdorf, Bernhard</mods:namePart>
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<mods:date id="95340F210BFCB52C3448DE4919C036C8">2017</mods:date>
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<mods:number id="483F0C869B5B0877B3ABA12559625BC9">4363</mods:number>
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<treatment id="373B87F8FFF9001FACAC7C22498EFD5E" ID-DOI="http://doi.org/10.5281/zenodo.6029286" ID-GBIF-Taxon="137679986" ID-Zenodo-Dep="6029286" LSID="urn:lsid:plazi:treatment:373B87F8FFF9001FACAC7C22498EFD5E" httpUri="http://treatment.plazi.org/id/373B87F8FFF9001FACAC7C22498EFD5E" lastPageId="2" lastPageNumber="591" pageId="0" pageNumber="589">
<subSubSection id="F7886565FFF9001DACAC7C224A57FADC" pageId="0" pageNumber="589" type="nomenclature">
<paragraph id="BF2D36EEFFF9001DACAC7C224A1DFABF" blockId="0.[151,1436,1206,2027]" box="[151,621,1310,1336]" pageId="0" pageNumber="589">
<heading id="E4658182FFF9001DACAC7C224A1DFABF" bold="true" box="[151,621,1310,1336]" fontSize="11" level="1" pageId="0" pageNumber="589" reason="1">
<taxonomicName id="78924D6DFFF9001DACAC7C224A1DFABF" ID-CoL="7P2XZ" authority="Kobelt &amp; Mollendorff, 1900" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[151,621,1310,1336]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF9001DACAC7C224A1DFABF" bold="true" box="[151,621,1310,1336]" pageId="0" pageNumber="589">
<emphasis id="8DE6EAFCFFF9001DACAC7C224979FABF" bold="true" box="[151,265,1310,1336]" italics="true" pageId="0" pageNumber="589">Dicharax</emphasis>
<bibRefCitation id="DB034B1FFFF9001DAD2B7C224A1DFABF" author="Kobelt" box="[272,621,1310,1336]" pageId="0" pageNumber="589" refString="Kobelt, W. &amp; Mollendorff, O. von (1900) Zur Systematik der Pneumonopomen. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft, 32, 186." type="journal article" year="1900">Kobelt &amp; Möllendorff, 1900</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="BF2D36EEFFF9001DACAC7C7D4A57FADC" blockId="0.[151,1436,1206,2027]" box="[151,551,1345,1371]" pageId="0" pageNumber="589">
<emphasis id="8DE6EAFCFFF9001DACAC7C7D4A57FADC" bold="true" box="[151,551,1345,1371]" pageId="0" pageNumber="589">
<taxonomicName id="78924D6DFFF9001DACAC7C7D4979FADC" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[151,265,1345,1371]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF9001DACAC7C7D4979FADC" bold="true" box="[151,265,1345,1371]" italics="true" pageId="0" pageNumber="589">Dicharax</emphasis>
</taxonomicName>
(?)
<taxonomicName id="78924D6DFFF9001DAD017C7D49AFFADC" authority="Nurinsiyah &amp; Hausdorf, 2017" authorityName="Nurinsiyah &amp; Hausdorf" authorityYear="2017" box="[314,479,1345,1371]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="species" species="candrakirana" status="sp. nov.">
<emphasis id="8DE6EAFCFFF9001DAD017C7D49AFFADC" bold="true" box="[314,479,1345,1371]" italics="true" pageId="0" pageNumber="589">candrakirana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96D55787FFF9001DADDE7C7D4A57FADC" box="[485,551,1345,1371]" pageId="0" pageNumber="589" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="F7886565FFF9001DACAC7C584B5DF89C" pageId="0" pageNumber="589" type="diagnosis">
<paragraph id="BF2D36EEFFF9001DACAC7C584D60FA47" blockId="0.[151,1436,1206,2027]" pageId="0" pageNumber="589">
<emphasis id="8DE6EAFCFFF9001DACAC7C58497CFAFC" bold="true" box="[151,268,1380,1403]" pageId="0" pageNumber="589">Diagnosis.</emphasis>
<taxonomicName id="78924D6DFFF9001DAD237C5F490CFAFC" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[280,380,1379,1403]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF9001DAD237C5F490CFAFC" box="[280,380,1379,1403]" italics="true" pageId="0" pageNumber="589">Dicharax</emphasis>
</taxonomicName>
(?)
<taxonomicName id="78924D6DFFF9001DAD887C5F4A34FAFC" authority="Nurinsiyah &amp; Hausdorf, 2017" authorityName="Nurinsiyah &amp; Hausdorf" authorityYear="2017" box="[435,580,1379,1403]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="species" species="candrakirana" status="sp. nov.">
<emphasis id="8DE6EAFCFFF9001DAD887C5F4A34FAFC" box="[435,580,1379,1403]" italics="true" pageId="0" pageNumber="589">candrakirana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96D55787FFF9001DAE6A7C584AE2FAFC" box="[593,658,1380,1403]" pageId="0" pageNumber="589" rank="species">
<emphasis id="8DE6EAFCFFF9001DAE6A7C584AE2FAFC" bold="true" box="[593,658,1380,1403]" pageId="0" pageNumber="589">n. sp.</emphasis>
</taxonomicNameLabel>
is characterized by a small, depressed conical shell, with a smooth protoconch and an irregularly striated teleoconch, with spiral striae especially around the umbilicus, and with a short sutural tube about a quarter of a whorl posterior to the aperture from which 5–9 radial microtunnels extend.
</paragraph>
<paragraph id="BF2D36EEFFF9001DACFC7CF04B88F934" blockId="0.[151,1436,1206,2027]" pageId="0" pageNumber="589">
<emphasis id="8DE6EAFCFFF9001DACFC7CF04911FA64" bold="true" box="[199,353,1484,1507]" pageId="0" pageNumber="589">
Shell (
<figureCitation id="27A92A6BFFF9001DAD347CF04922FA64" box="[271,338,1484,1507]" captionStart="FIGURE 1" captionStartId="1.[151,244,1085,1107]" captionTargetBox="[178,1422,319,1073]" captionTargetId="figure@1.[178,1422,319,1073]" captionTargetPageId="1" captionText="FIGURE 1. Dicharax (?) candrakirana n. sp. Indonesia: Malang, Sempu Island, at entrance of Kelabang Cave (holotype MZB 19025).Scale bar = 1mm." httpUri="https://zenodo.org/record/1114817/files/figure.png" pageId="0" pageNumber="589">Fig. 1</figureCitation>
).
</emphasis>
Depressed conical; with 3 whorls separated by a deep suture. Protoconch low, smooth. Teleoconch irregularly striated, especially at the bottom side around the umbilicus with spiral striae, less distinct spiral striae are present also on the upper side close to the suture. Color whitish; subtranslucent. Last whorl rounded, with an indistinct constriction about a quarter of a whorl posterior to the aperture; posterior to the constriction with a short (
<quantity id="786A9B0BFFF9001DA9377F084DEBF9CC" box="[1292,1435,1588,1611]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="0" pageNumber="589" unit="mm" value="0.275" valueMax="0.3" valueMin="0.25">0.25–0.3 mm</quantity>
long) sutural tube; below the tube with 5–9 radial microtunnels. Aperture almost circular, not crenulated; upper insertion of the peristome descending towards aperture; insertions of the peristome connected by a thick callus; peristome expanded, but not reflexed, strongly thickened inside. Umbilicus wide, eccentric.
</paragraph>
<paragraph id="BF2D36EEFFF9001DACFC7F824B5DF89C" blockId="0.[151,1436,1206,2027]" pageId="0" pageNumber="589">
Measurements (n = 8): shell diameter D: 2.0–
<quantity id="786A9B0BFFF9001DAE917F824A8DF952" box="[682,765,1726,1749]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="0" pageNumber="589" unit="mm" value="2.2">2.2 mm</quantity>
, mean 2.1 +
<quantity id="786A9B0BFFF9001DAFBE7F824B97F952" box="[901,999,1726,1749]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="0" pageNumber="589" unit="mm" value="0.05">0.05 mm</quantity>
; shell height H:
<quantity id="786A9B0BFFF9001DA8A87F824D53F952" box="[1171,1315,1726,1749]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="0" pageNumber="589" unit="mm" value="1.275" valueMax="1.3" valueMin="1.25">1.25–1.3 mm</quantity>
, mean 1.25 +
<quantity id="786A9B0BFFF9001DAC947FDD4963F97F" box="[175,275,1761,1784]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="0" pageNumber="589" unit="mm" value="0.05">0.05 mm</quantity>
; aperture (outer peristome) diameter da:
<quantity id="786A9B0BFFF9001DAEE87FDD4B16F97F" box="[723,870,1761,1784]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="9.0" metricValueMin="8.5" pageId="0" pageNumber="589" unit="mm" value="0.875" valueMax="0.9" valueMin="0.85">0.85–0.9 mm</quantity>
, mean 0.85 + 0.0 mm; aperture height ha:
<quantity id="786A9B0BFFF9001DA9047FDD48B3F89C" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="9.0" metricValueMin="8.5" pageId="0" pageNumber="589" unit="mm" value="0.875" valueMax="0.9" valueMin="0.85">0.85–0.9 mm</quantity>
, mean 0.85 + 0.0 mm; D/H: 1.63–1.71, mean 1.67 + 0.03.
</paragraph>
</subSubSection>
<subSubSection id="F7886565FFF9001CACFC7E1A4DE3FE90" lastPageId="1" lastPageNumber="590" pageId="0" pageNumber="589" type="discussion">
<paragraph id="BF2D36EEFFF9001CACFC7E1A4DE3FE90" blockId="0.[151,1436,1206,2027]" lastBlockId="1.[151,1436,152,279]" lastPageId="1" lastPageNumber="590" pageId="0" pageNumber="589">
<emphasis id="8DE6EAFCFFF9001DACFC7E1A4945F8BA" bold="true" box="[199,309,1830,1853]" pageId="0" pageNumber="589">Remarks.</emphasis>
<bibRefCitation id="DB034B1FFFF9001DAD077E1A49A3F8BA" author="Kobelt" box="[316,467,1830,1853]" pageId="0" pageNumber="589" refString="Kobelt, W. (1902) Cyclophoridae. In: Schulze, F. E. (Ed.), Das Tierreich. Vol. 16. R. Friedlander und Sohn, Berlin, pp. I - XXXIX + 1 - 662, 1 map." type="book chapter" year="1902">Kobelt (1902)</bibRefCitation>
classified
<taxonomicName id="78924D6DFFF9001DAE7E7E194AD9F8BA" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[581,681,1829,1853]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF9001DAE7E7E194AD9F8BA" box="[581,681,1829,1853]" italics="true" pageId="0" pageNumber="589">Dicharax</emphasis>
</taxonomicName>
and
<taxonomicName id="78924D6DFFF9001DAEE57E194C36F8BA" authority="Mollendorff, 1897" authorityName="Mollendorff" authorityYear="1897" box="[734,1094,1829,1853]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF9001DAEE57E194B0BF8BA" box="[734,891,1829,1853]" italics="true" pageId="0" pageNumber="589">Chamalycaeus</emphasis>
<bibRefCitation id="DB034B1FFFF9001DAFB97E1A4C36F8BA" author="Mollendorff" box="[898,1094,1830,1853]" pageId="0" pageNumber="589" refString="Mollendorff, O. von (1897) Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft, 29, 89 - 97." type="journal article" year="1897">Möllendorff, 1897</bibRefCitation>
</taxonomicName>
as subgenera of
<taxonomicName id="78924D6DFFF9001DA8C07E1948BCF8E7" authority="Gray, 1850" authorityName="Gray" authorityYear="1850" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF9001DA8C07E194D2BF8BA" box="[1275,1371,1829,1853]" italics="true" pageId="0" pageNumber="589">Alycaeus</emphasis>
<bibRefCitation id="DB034B1FFFF9001DA95A7E1A48BCF8E7" author="Gray" pageId="0" pageNumber="589" refString="Gray, J. E. (1850) Nomenclature of molluscous animals and shells in the collection of the British Museum. Part I. Cyclophoridae. Spottiswoodes and Shaw, London, 68 pp." type="book" year="1850">Gray, 1850</bibRefCitation>
</taxonomicName>
and distinguished
<taxonomicName id="78924D6DFFF9001DADAC7E74498BF8E7" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[407,507,1864,1888]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF9001DADAC7E74498BF8E7" box="[407,507,1864,1888]" italics="true" pageId="0" pageNumber="589">Dicharax</emphasis>
</taxonomicName>
from
<taxonomicName id="78924D6DFFF9001DAE047E744AACF8E7" authorityName="Mollendorff" authorityYear="1897" box="[575,732,1864,1888]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF9001DAE047E744AACF8E7" box="[575,732,1864,1888]" italics="true" pageId="0" pageNumber="589">Chamalycaeus</emphasis>
</taxonomicName>
by the position of the constriction and the presence of a swelling near the aperture. In contrast,
<bibRefCitation id="DB034B1FFFF9001DADC67E504B56F804" box="[509,806,1899,1923]" pageId="0" pageNumber="589" refString="Pall-Gergely, B., Hunyadi, A., Do, D. S., Naggs, F. &amp; Asami, T. (2017) Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa, 4331 (1), 1 - 124. https: // doi. org / 10.11646 / zootaxa. 4331.1.1" type="journal article">
Páll-Gergely
<emphasis id="8DE6EAFCFFF9001DAEA87E504AA1F804" box="[659,721,1899,1923]" italics="true" pageId="0" pageNumber="589">et al.</emphasis>
(
<docYear id="A7F4DA15FFF9001DAEDC7E504B6FF804" box="[743,799,1900,1923]" pageId="0" pageNumber="589">2017</docYear>
)
</bibRefCitation>
classified these two taxa as genera and distinguished
<taxonomicName id="78924D6DFFF9001DACAC7EB14944F822" authorityName="Mollendorff" authorityYear="1897" box="[151,308,1933,1957]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF9001DACAC7EB14944F822" box="[151,308,1933,1957]" italics="true" pageId="0" pageNumber="589">Chamalycaeus</emphasis>
</taxonomicName>
from
<taxonomicName id="78924D6DFFF9001DADBA7EB14996F822" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[385,486,1933,1957]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF9001DADBA7EB14996F822" box="[385,486,1933,1957]" italics="true" pageId="0" pageNumber="589">Dicharax</emphasis>
</taxonomicName>
based on the presence of elevated spiral striations on the teleoconch. Whereas
<taxonomicName id="78924D6DFFF9001DA9B97EB34958F84F" authority="Nurinsiyah &amp; Hausdorf, 2017" authorityName="Nurinsiyah &amp; Hausdorf" authorityYear="2017" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="species" species="candrakirana" status="sp. nov.">
<emphasis id="8DE6EAFCFFF9001DA9B97EB34958F84F" italics="true" pageId="0" pageNumber="589">D. candrakirana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96D55787FFF9001DAD147E8D491CF84F" box="[303,364,1969,1992]" pageId="0" pageNumber="589" rank="species">
<emphasis id="8DE6EAFCFFF9001DAD147E8D491CF84F" bold="true" box="[303,364,1969,1992]" pageId="0" pageNumber="589">n. sp.</emphasis>
</taxonomicNameLabel>
would be classified in
<taxonomicName id="78924D6DFFF9001DAE5D7E8C4B73F84F" authorityName="Mollendorff" authorityYear="1897" box="[614,771,1968,1992]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF9001DAE5D7E8C4B73F84F" box="[614,771,1968,1992]" italics="true" pageId="0" pageNumber="589">Chamalycaeus</emphasis>
</taxonomicName>
according to the diagnosis of
<bibRefCitation id="DB034B1FFFF9001DA8727E8D4C93F84F" author="Kobelt" box="[1097,1251,1969,1992]" pageId="0" pageNumber="589" refString="Kobelt, W. (1902) Cyclophoridae. In: Schulze, F. E. (Ed.), Das Tierreich. Vol. 16. R. Friedlander und Sohn, Berlin, pp. I - XXXIX + 1 - 662, 1 map." type="book chapter" year="1902">Kobelt (1902)</bibRefCitation>
, its classification is more problematic with regard to the classification of
<bibRefCitation id="DB034B1FFFF9001DAECE7EE84C7FF86C" box="[757,1039,2003,2027]" pageId="0" pageNumber="589" refString="Pall-Gergely, B., Hunyadi, A., Do, D. S., Naggs, F. &amp; Asami, T. (2017) Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa, 4331 (1), 1 - 124. https: // doi. org / 10.11646 / zootaxa. 4331.1.1" type="journal article">
Páll-Gergely
<emphasis id="8DE6EAFCFFF9001DAFBE7EE84BCDF86C" box="[901,957,2003,2027]" italics="true" pageId="0" pageNumber="589">et al.</emphasis>
(2017)
</bibRefCitation>
. There are distinct spiral lines at the bottom side around the umbilicus and less distinct spiral lines on the upper side close to the suture. However, the spiral lines are not elevated ridges as in
<taxonomicName id="78924D6DFFF8001CADC579854AEBFF56" authorityName="Mollendorff" authorityYear="1897" box="[510,667,185,209]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="590" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF8001CADC579854AEBFF56" box="[510,667,185,209]" italics="true" pageId="1" pageNumber="590">Chamalycaeus</emphasis>
</taxonomicName>
.
<bibRefCitation id="DB034B1FFFF8001CAE9379864BC8FF56" box="[680,952,185,209]" pageId="1" pageNumber="590" refString="Pall-Gergely, B., Hunyadi, A., Do, D. S., Naggs, F. &amp; Asami, T. (2017) Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa, 4331 (1), 1 - 124. https: // doi. org / 10.11646 / zootaxa. 4331.1.1" type="journal article">
Páll-Gergely
<emphasis id="8DE6EAFCFFF8001CAF0D79864B1CFF56" box="[822,876,185,209]" italics="true" pageId="1" pageNumber="590">et al.</emphasis>
(2017)
</bibRefCitation>
found spiral striations also in a population of
<taxonomicName id="78924D6DFFF8001CACAC79E04AB8FF73" authority="(Bavay &amp; Dautzenberg, 1912)" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[151,712,220,244]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="590" phylum="Mollusca" rank="species" species="depressus">
<emphasis id="8DE6EAFCFFF8001CACAC79E0491EFF73" box="[151,366,220,244]" italics="true" pageId="1" pageNumber="590">Dicharax depressus</emphasis>
(Bavay &amp; Dautzenberg, 1912)
</taxonomicName>
of
<collectingCountry id="C785767EFFF8001CAECF79E14B21FF73" box="[756,849,221,244]" name="Vietnam" pageId="1" pageNumber="590">Vietnam</collectingCountry>
, which they nevertheless classified in
<taxonomicName id="78924D6DFFF8001CA93879E04D17FF73" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[1283,1383,220,244]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="590" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF8001CA93879E04D17FF73" box="[1283,1383,220,244]" italics="true" pageId="1" pageNumber="590">Dicharax</emphasis>
</taxonomicName>
. We follow a recommendation of B. Páll-Gergely (pers. comm.) and classify the new species also preliminarily in
<taxonomicName id="78924D6DFFF8001CA91C79C34DFCFE90" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[1319,1420,255,279]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="590" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF8001CA91C79C34DFCFE90" box="[1319,1420,255,279]" italics="true" pageId="1" pageNumber="590">Dicharax</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="EBED6666FFF8001CACAC7D0149E7FBF7" httpUri="https://zenodo.org/record/1114817/files/figure.png" pageId="1" pageNumber="590" targetBox="[178,1422,319,1073]" targetPageId="1">
<paragraph id="BF2D36EEFFF8001CACAC7D0149E7FBF7" blockId="1.[151,1436,1085,1136]" pageId="1" pageNumber="590">
<emphasis id="8DE6EAFCFFF8001CACAC7D014960FBD4" bold="true" box="[151,272,1085,1107]" pageId="1" pageNumber="590">FIGURE 1.</emphasis>
<taxonomicName id="78924D6DFFF8001CAD227D014904FBD5" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[281,372,1085,1106]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="590" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF8001CAD227D014904FBD5" box="[281,372,1085,1106]" italics="true" pageId="1" pageNumber="590">Dicharax</emphasis>
</taxonomicName>
(?)
<taxonomicName id="78924D6DFFF8001CAD997D014A55FBD5" authority="Nurinsiyah &amp; Hausdorf, 2017" authorityName="Nurinsiyah &amp; Hausdorf" authorityYear="2017" box="[418,549,1085,1106]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="590" phylum="Mollusca" rank="species" species="candrakirana" status="sp. nov.">
<emphasis id="8DE6EAFCFFF8001CAD997D014A55FBD5" box="[418,549,1085,1106]" italics="true" pageId="1" pageNumber="590">candrakirana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96D55787FFF8001CAE157D014A18FBD4" box="[558,616,1085,1107]" pageId="1" pageNumber="590" rank="species">
<emphasis id="8DE6EAFCFFF8001CAE157D014A18FBD4" bold="true" box="[558,616,1085,1107]" pageId="1" pageNumber="590">n. sp.</emphasis>
</taxonomicNameLabel>
Indonesia: Malang, Sempu Island, at entrance of Kelabang Cave (holotype MZB 19025). Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="EBED6666FFF8001CACAC7EFC488BF874" httpUri="https://zenodo.org/record/1114823/files/figure.png" pageId="1" pageNumber="590" targetBox="[194,1399,1176,1979]" targetPageId="1">
<paragraph id="BF2D36EEFFF8001CACAC7EFC488BF874" blockId="1.[151,1436,1984,2035]" pageId="1" pageNumber="590">
<emphasis id="8DE6EAFCFFF8001CACAC7EFC497FF851" bold="true" box="[151,271,1984,2006]" pageId="1" pageNumber="590">FIGURE 2.</emphasis>
Type locality of
<taxonomicName id="78924D6DFFF8001CAD847EFD4A6AF851" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[447,538,1985,2006]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="590" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFF8001CAD847EFD4A6AF851" box="[447,538,1985,2006]" italics="true" pageId="1" pageNumber="590">Dicharax</emphasis>
</taxonomicName>
(?)
<taxonomicName id="78924D6DFFF8001CAE7D7EFD4AB9F851" authority="Nurinsiyah &amp; Hausdorf, 2017" authorityName="Nurinsiyah &amp; Hausdorf" authorityYear="2017" box="[582,713,1985,2006]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="590" phylum="Mollusca" rank="species" species="candrakirana" status="sp. nov.">
<emphasis id="8DE6EAFCFFF8001CAE7D7EFD4AB9F851" box="[582,713,1985,2006]" italics="true" pageId="1" pageNumber="590">candrakirana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96D55787FFF8001CAEE97EFC4B7BF851" box="[722,779,1984,2006]" pageId="1" pageNumber="590" rank="species">
<emphasis id="8DE6EAFCFFF8001CAEE97EFC4B7BF851" bold="true" box="[722,779,1984,2006]" pageId="1" pageNumber="590">n. sp.</emphasis>
</taxonomicNameLabel>
: limestone rocks at entrance of Kelabang Cave on Sempu Island, Indonesia.
</paragraph>
</caption>
<subSubSection id="F7886565FFFB001FACFC79AB4B65FEBE" pageId="2" pageNumber="591" type="reference_group">
<paragraph id="BF2D36EEFFFB001FACFC79AB4B65FEBE" blockId="2.[151,1436,151,729]" pageId="2" pageNumber="591">
<taxonomicName id="78924D6DFFFB001FACFC79AB495CFF28" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[199,300,151,175]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="591" phylum="Mollusca" rank="genus">
<emphasis id="8DE6EAFCFFFB001FACFC79AB495CFF28" box="[199,300,151,175]" italics="true" pageId="2" pageNumber="591">Dicharax</emphasis>
</taxonomicName>
(?)
<taxonomicName id="78924D6DFFFB001FAD6579AB499FFF28" authority="Nurinsiyah &amp; Hausdorf, 2017" authorityName="Nurinsiyah &amp; Hausdorf" authorityYear="2017" box="[350,495,151,175]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="591" phylum="Mollusca" rank="species" species="candrakirana" status="sp. nov.">
<emphasis id="8DE6EAFCFFFB001FAD6579AB499FFF28" box="[350,495,151,175]" italics="true" pageId="2" pageNumber="591">candrakirana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96D55787FFFB001FADC179A44A4AFF28" box="[506,570,152,175]" pageId="2" pageNumber="591" rank="species">
<emphasis id="8DE6EAFCFFFB001FADC179A44A4AFF28" bold="true" box="[506,570,152,175]" pageId="2" pageNumber="591">n. sp.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="78924D6DFFFB001FAEE879AB4C13FF28" authority="(Martens, 1864)" authorityName="Martens" authorityYear="1864" baseAuthorityName="Martens" baseAuthorityYear="1864" box="[723,1123,151,175]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="591" phylum="Mollusca" rank="species" species="longituba">
<emphasis id="8DE6EAFCFFFB001FAEE879AB4BD8FF28" box="[723,936,151,175]" italics="true" pageId="2" pageNumber="591">Dicharax longituba</emphasis>
(
<bibRefCitation id="DB034B1FFFFB001FAF8179A44C2BFF28" author="Martens" box="[954,1115,152,175]" pageId="2" pageNumber="591" refString="Martens, E. von (1864) Uber neue Cyclostomaceen und Helicinen aus dem indischen Archipel. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 1864, 116 - 121." type="journal article" year="1864">Martens, 1864</bibRefCitation>
)
</taxonomicName>
of
<collectingRegion id="7D56F80CFFFB001FA8B679A44CCCFF28" box="[1165,1212,152,175]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="591">Java</collectingRegion>
and
<collectingRegion id="7D56F80CFFFB001FA8C279A44D22FF28" box="[1273,1362,152,175]" country="Indonesia" name="Sumatera Utara" pageId="2" pageNumber="591">Sumatra</collectingRegion>
in the smaller shell with a shorter sutural tube, an only indistinct constriction without swelling, the lack of ribbing and the presence of spiral striae at the bottom side around the umbilicus. It differs from
<taxonomicName id="78924D6DFFFB001FAFDD79E048A5FE90" authority="(Mollendorff, 1897)" authorityName="Mollendorff" authorityYear="1897" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="591" phylum="Mollusca" rank="species" species="fruhstorferi">
<emphasis id="8DE6EAFCFFFB001FAFDD79E04D74FF73" box="[998,1284,220,244]" italics="true" pageId="2" pageNumber="591">Chamalycaeus fruhstorferi</emphasis>
(
<bibRefCitation id="DB034B1FFFFB001FA92979E148BEFE90" author="Mollendorff" pageId="2" pageNumber="591" refString="Mollendorff, O. von (1897) Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft, 29, 89 - 97." type="journal article" year="1897">Möllendorff, 1897</bibRefCitation>
)
</taxonomicName>
of
<collectingRegion id="7D56F80CFFFB001FACC5783C495DFE90" box="[254,301,256,279]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="591">Java</collectingRegion>
and
<taxonomicName id="78924D6DFFFB001FAD5C79C34B57FE90" authority="Vermeulen, 1996" authorityName="Vermeulen" authorityYear="1996" box="[359,807,255,279]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="591" phylum="Mollusca" rank="species" species="kessneri">
<emphasis id="8DE6EAFCFFFB001FAD5C79C34A15FE90" box="[359,613,255,279]" italics="true" pageId="2" pageNumber="591">Chamalycaeus kessneri</emphasis>
<bibRefCitation id="DB034B1FFFFB001FAE54783C4B57FE90" author="Vermeulen" box="[623,807,256,279]" pageId="2" pageNumber="591" refString="Vermeulen, J. J. (1996) Notes on terrestrial molluscs of Java, Bali, and Nusa Penida. Basteria, 59, 149 - 162." type="journal article" year="1996">Vermeulen, 1996</bibRefCitation>
</taxonomicName>
of
<collectingRegion id="7D56F80CFFFB001FAF6A783C4B0DFE90" box="[849,893,256,279]" country="Indonesia" name="Bali" pageId="2" pageNumber="591">Bali</collectingRegion>
and Nusa Penida in the smaller shell with fewer whorls and the lack of ribbing and elevated spiral striations.
</paragraph>
</subSubSection>
<subSubSection id="F7886565FFFB001FACFC78794B32FDC8" pageId="2" pageNumber="591" type="materials_examined">
<paragraph id="BF2D36EEFFFB001FACFC78794CCDFD8E" blockId="2.[151,1436,151,729]" pageId="2" pageNumber="591">
<materialsCitation id="0FFA3CB3FFFB001FACFC78794AC8FE26" ID-GBIF-Occurrence="1705513545" collectingDate="2014-06-09" collectionCode="MZB" collectorName="A. S. Nurinsiyah &amp; Fauzia" country="Indonesia" county="Malang" elevation="44" latitude="-8.449445" location="Kelabang Cave" longLatPrecision="21" longitude="112.69111" municipality="Sempu Island" pageId="2" pageNumber="591" specimenCount="2" stateProvince="East Java" typeStatus="holotype">
<emphasis id="8DE6EAFCFFFB001FACFC78794900FEDB" bold="true" box="[199,368,325,348]" pageId="2" pageNumber="591">
<typeStatus id="6029884CFFFB001FACFC7879488FFEDB" box="[199,255,325,348]" pageId="2" pageNumber="591">Type</typeStatus>
material.
</emphasis>
<typeStatus id="6029884CFFFB001FAD4278794991FEDB" box="[377,481,325,348]" pageId="2" pageNumber="591" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="C785767EFFFB001FADD678794A27FEDB" box="[493,599,325,348]" name="Indonesia" pageId="2" pageNumber="591">Indonesia</collectingCountry>
,
<collectingRegion id="7D56F80CFFFB001FAE5D78794AA0FEDB" box="[614,720,325,348]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="591">East Java</collectingRegion>
:
<collectingCounty id="564C4E62FFFB001FAEE578794B42FEDB" box="[734,818,325,348]" pageId="2" pageNumber="591">Malang</collectingCounty>
,
<collectingMunicipality id="5F49AC94FFFB001FAF7B78794BA8FEDB" box="[832,984,325,348]" pageId="2" pageNumber="591">Sempu Island</collectingMunicipality>
, limestone rocks in lowland rainforest at entrance of
<location id="BA4D6035FFFB001FAD2C785449CDFEF8" LSID="urn:lsid:plazi:treatment:373B87F8FFF9001FACAC7C22498EFD5E:BA4D6035FFFB001FAD2C785449CDFEF8" box="[279,445,360,383]" country="Indonesia" county="Malang" latitude="-8.449445" longLatPrecision="21" longitude="112.69111" municipality="Sempu Island" name="Kelabang Cave" pageId="2" pageNumber="591" stateProvince="East Java">Kelabang Cave</location>
,
<elevation id="34BFD1DDFFFB001FADF178544A4AFEF8" box="[458,570,360,383]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="591" unit="m" value="44.0">
<quantity id="786A9B0BFFFB001FADF178544A71FEF8" box="[458,513,360,383]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="591" unit="m" value="44.0">44 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="DAA65029FFFB001FAE7378544AC3FEF8" box="[584,691,360,383]" direction="south" orientation="latitude" pageId="2" pageNumber="591" precision="15" value="-8.449445">8°26'58&quot;S</geoCoordinate>
<geoCoordinate id="DAA65029FFFB001FAE8078544B32FEF8" box="[699,834,360,383]" direction="east" orientation="longitude" pageId="2" pageNumber="591" precision="15" value="112.69111">112°41'28&quot;E</geoCoordinate>
(
<collectionCode id="D983AE2BFFFB001FAF6A78544BFCFEF8" box="[849,908,360,383]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="591">MZB</collectionCode>
19025, leg.
<collectorName id="12675338FFFB001FA82F78544CB1FEF8" box="[1044,1217,360,383]" pageId="2" pageNumber="591">A.S. Nurinsiyah</collectorName>
&amp; H.
<collectorName id="12675338FFFB001FA93C78544D3EFEF8" box="[1287,1358,360,383]" pageId="2" pageNumber="591">Fauzia</collectorName>
<collectingDate id="DB68E9C6FFFB001FA96D785448A0FE26" pageId="2" pageNumber="591" value="2014-06-09">
<date id="CB2C102EFFFB001FA96D785448A0FE26" pageId="2" pageNumber="591" value="2014-06-09">9 June 2014</date>
</collectingDate>
; measurements: D =
<quantity id="786A9B0BFFFB001FAD8778B64A63FE26" box="[444,531,394,417]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="591" unit="mm" value="2.2">2.2 mm</quantity>
, H =
<quantity id="786A9B0BFFFB001FAE6C78B64AC0FE26" box="[599,688,394,417]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="591" unit="mm" value="1.3">1.3 mm</quantity>
)
</materialsCitation>
.
<materialsCitation id="0FFA3CB3FFFB001FAEFF78B64CCDFD8E" ID-GBIF-Occurrence="1705513544" collectionCode="MZB" collectorName="Malang &amp; Jembatan Anakan Hill" country="Indonesia" county="Malang" elevation="44" latitude="-8.451388" location="Kelabang Cave" longLatPrecision="21" longitude="112.69111" municipality="Sempu Island" pageId="2" pageNumber="591" specimenCode="ZMH 133203, ZMH 133204" specimenCount="1" stateProvince="East Java" typeStatus="paratype">
<typeStatus id="6029884CFFFB001FAEFF78B64B40FE26" box="[708,816,394,417]" pageId="2" pageNumber="591" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="C785767EFFFB001FAF0678B64BD7FE26" box="[829,935,394,417]" name="Indonesia" pageId="2" pageNumber="591">Indonesia</collectingCountry>
,
<collectingRegion id="7D56F80CFFFB001FAF8E78B64C6FFE26" box="[949,1055,394,417]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="591">East Java</collectingRegion>
:
<collectingCounty id="564C4E62FFFB001FA81678B64CF1FE26" box="[1069,1153,394,417]" pageId="2" pageNumber="591">Malang</collectingCounty>
,
<collectingMunicipality id="5F49AC94FFFB001FA8B478B64D56FE26" box="[1167,1318,394,417]" pageId="2" pageNumber="591">Sempu Island</collectingMunicipality>
, limestone rocks at entrance of
<location id="BA4D6035FFFB001FAD4478914A59FE43" LSID="urn:lsid:plazi:treatment:373B87F8FFF9001FACAC7C22498EFD5E:BA4D6035FFFB001FAD4478914A59FE43" box="[383,553,429,452]" country="Indonesia" county="Malang" latitude="-8.451388" longLatPrecision="21" longitude="112.69111" municipality="Sempu Island" name="Kelabang Cave" pageId="2" pageNumber="591" stateProvince="East Java">Kelabang Cave</location>
,
<elevation id="34BFD1DDFFFB001FAE0378914ADFFE43" box="[568,687,429,452]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="591" unit="m" value="44.0">
<quantity id="786A9B0BFFFB001FAE0378914A02FE43" box="[568,626,429,452]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="591" unit="m" value="44.0">44 m</quantity>
a.s.l.
</elevation>
,
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(
<collectionCode id="D983AE2BFFFB001FAFF578914C79FE43" box="[974,1033,429,452]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="591">MZB</collectionCode>
19026/1;
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/2); Malang,
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, summit of Kembar Satu Hill,
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a.s.l.
</elevation>
,
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(
<collectionCode id="D983AE2BFFFB001FA81478EC4C1AFE60" box="[1071,1130,464,487]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="591">MZB</collectionCode>
19027/2;
<specimenCode id="EF349E95FFFB001FA8DB78EC4D0EFE60" box="[1248,1406,464,487]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="2" pageNumber="591">ZMH 133204</specimenCode>
/1);
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,
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,
<collectorName id="12675338FFFB001FADAE78CE4AF5FD8E" box="[405,645,498,521]" pageId="2" pageNumber="591">Jembatan Anakan Hill</collectorName>
,
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<quantity id="786A9B0BFFFB001FAEAB78CE4AB6FD8E" box="[656,710,498,521]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="591" unit="m" value="46.0">46 m</quantity>
a.s.l.
</elevation>
,
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<geoCoordinate id="DAA65029FFFB001FAF4578CE4C75FD8E" box="[894,1029,498,521]" direction="east" orientation="longitude" pageId="2" pageNumber="591" precision="15" value="112.69111">112°41'28&quot;E</geoCoordinate>
(
<collectionCode id="D983AE2BFFFB001FA82878CE4C3EFD8E" box="[1043,1102,498,521]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="591">MZB</collectionCode>
19028/1).
</materialsCitation>
</paragraph>
<paragraph id="BF2D36EEFFFB001FACFC7B294B32FDC8" blockId="2.[151,1436,151,729]" pageId="2" pageNumber="591">
<materialsCitation id="0FFA3CB3FFFB001FACFC7B294B32FDC8" ID-GBIF-Occurrence="1705513543" country="Indonesia" county="Malang" elevation="44" latitude="-8.449445" location="Kelabang Cave" longLatPrecision="21" longitude="112.69111" municipality="Sempu Island" pageId="2" pageNumber="591" specimenCount="1" stateProvince="East Java" typeStatus="holotype">
<emphasis id="8DE6EAFCFFFB001FACFC7B2949B7FDAB" bold="true" box="[199,455,533,556]" pageId="2" pageNumber="591">
<typeStatus id="6029884CFFFB001FACFC7B29488FFDAB" box="[199,255,533,556]" pageId="2" pageNumber="591">Type</typeStatus>
locality (
<figureCitation id="27A92A6BFFFB001FAD497B2949CAFDAB" box="[370,442,533,556]" captionStart="FIGURE 2" captionStartId="1.[151,244,1984,2006]" captionTargetBox="[194,1399,1176,1979]" captionTargetId="figure@1.[194,1399,1176,1980]" captionTargetPageId="1" captionText="FIGURE 2. Type locality of Dicharax (?) candrakirana n. sp.: limestone rocks at entrance of Kelabang Cave on Sempu Island, Indonesia." httpUri="https://zenodo.org/record/1114823/files/figure.png" pageId="2" pageNumber="591">Fig. 2</figureCitation>
).
</emphasis>
<collectingCountry id="C785767EFFFB001FADEF7B294A4EFDAB" box="[468,574,533,556]" name="Indonesia" pageId="2" pageNumber="591">Indonesia</collectingCountry>
,
<collectingRegion id="7D56F80CFFFB001FAE757B294ACBFDAB" box="[590,699,533,556]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="591">East Java</collectingRegion>
:
<collectingCounty id="564C4E62FFFB001FAEF07B294B6FFDAB" box="[715,799,533,556]" pageId="2" pageNumber="591">Malang</collectingCounty>
,
<collectingMunicipality id="5F49AC94FFFB001FAF147B294BBAFDAB" box="[815,970,533,556]" pageId="2" pageNumber="591">Sempu Island</collectingMunicipality>
, limestone rocks in lowland rainforest at entrance of
<location id="BA4D6035FFFB001FAD2F7B0449CAFDC8" LSID="urn:lsid:plazi:treatment:373B87F8FFF9001FACAC7C22498EFD5E:BA4D6035FFFB001FAD2F7B0449CAFDC8" box="[276,442,568,591]" country="Indonesia" county="Malang" latitude="-8.449445" longLatPrecision="21" longitude="112.69111" municipality="Sempu Island" name="Kelabang Cave" pageId="2" pageNumber="591" stateProvince="East Java">Kelabang Cave</location>
,
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<quantity id="786A9B0BFFFB001FADFE7B04498CFDC8" box="[453,508,568,591]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="591" unit="m" value="44.0">44 m</quantity>
a.s.l.
</elevation>
,
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<geoCoordinate id="DAA65029FFFB001FAE887B044B4DFDC8" box="[691,829,568,591]" direction="east" orientation="longitude" pageId="2" pageNumber="591" precision="15" value="112.69111">112°41'28&quot;E</geoCoordinate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7886565FFFB001FACFC7B664976FD13" pageId="2" pageNumber="591" type="distribution">
<paragraph id="BF2D36EEFFFB001FACFC7B664976FD13" blockId="2.[151,1436,151,729]" pageId="2" pageNumber="591">
<emphasis id="8DE6EAFCFFFB001FACFC7B664929FDF6" bold="true" box="[199,345,602,625]" pageId="2" pageNumber="591">Distribution.</emphasis>
The new species is known only from limestone rocks in lowland rainforest on Sempu Island in
<collectingCountry id="C785767EFFFB001FACAC7B414972FD13" box="[151,258,637,660]" name="Indonesia" pageId="2" pageNumber="591">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="F7886565FFFB001FACFC7B9C498EFD5E" pageId="2" pageNumber="591" type="etymology">
<paragraph id="BF2D36EEFFFB001FACFC7B9C498EFD5E" blockId="2.[151,1436,151,729]" pageId="2" pageNumber="591">
<emphasis id="8DE6EAFCFFFB001FACFC7B9C4937FD30" bold="true" box="[199,327,672,695]" pageId="2" pageNumber="591">Etymology.</emphasis>
The species is named after Candra Kirana, a beautiful princess in the famous Indonesian folk tale ‘Keong Mas’, the ‘Golden Snail’.
</paragraph>
</subSubSection>
</treatment>
</document>
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