treatments-xml/data/0B/1B/87/0B1B8791FFCF0610F8EEEC777C755A63.xml

<document id="A9F3D9FB427A9C6C57760AAF83039EF2" ID-ISSN="0024-4082" ID-ZooBank="74C07292-2BD6-4E3E-B68D-B144B81BBD83" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629585783401" checkinUser="felipe" docAuthor="Bonifácio, Paulo &amp; Menot, Lénaïck" docDate="2019" docId="0B1B8791FFCF0610F8EEEC777C755A63" docLanguage="en" docName="ZoolJLinnSoc.185.555–635.pdf" docOrigin="Zoological Journal of the Linnean Society 185" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Macellicephaloides moustachu Bonifácio &amp; Menot 2019, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="611" masterDocId="F722FFE9FFFA0628F820E81A7D025E58" masterDocTitle="New genera and species from the Equatorial Pacific provide phylogenetic insights into deep-sea Polynoidae (Annelida)" masterLastPageNumber="635" masterPageNumber="555" pageNumber="608" updateTime="1699081480826" updateUser="plazi">
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<mods:title id="76B752686DDD1B09B7C10D0A13B30ED6">New genera and species from the Equatorial Pacific provide phylogenetic insights into deep-sea Polynoidae (Annelida)</mods:title>
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<mods:namePart id="8D5D579FC34D685EFB2C3638B39D5786">Bonifácio, Paulo</mods:namePart>
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<mods:namePart id="EA5F983DF4A9D3D7B5E9D3D5B9807CDE">Menot, Lénaïck</mods:namePart>
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<mods:title id="B174AAD9EF63397872C841EF3BC239C7">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="6F6792D0B4A9FECE3C67A3B976F848A3">2019</mods:date>
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<heading id="D84581EBFFCF061DF8EEEC777FDD5ADD" box="[206,735,1133,1157]" centered="true" fontSize="9" level="2" pageId="53" pageNumber="608" reason="2">
<taxonomicName id="44B24D04FFCF061DF8EEEC777F775ADB" ID-CoL="6R2GB" authority="Bonifácio &amp; Menot, 2019" authorityName="Bonifácio &amp; Menot" authorityYear="2019" box="[206,629,1133,1156]" class="Polychaeta" family="Polynoidae" genus="Macellicephaloides" kingdom="Animalia" order="Phyllodocida" pageId="53" pageNumber="627" phylum="Annelida" rank="species" species="moustachu" status="sp. nov.">MACELLICEPHALOIDES MOUSTACHU</taxonomicName>
<emphasis id="B1C6EA95FFCF061DFA5DEC6A7FDD5ADD" bold="true" box="[637,735,1133,1157]" pageId="53" pageNumber="608">
<taxonomicNameLabel id="AAF557EEFFCF061DFA5DEC6A7FDD5ADD" box="[637,735,1133,1157]" pageId="53" pageNumber="608" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="CBA8650CFFCF0610F964EC8E7C755A63" lastPageId="56" lastPageNumber="611" pageId="53" pageNumber="608" type="description">
<paragraph id="830D3687FFCF061DF964EC8E7F6B5AF5" blockId="53.[324,617,1172,1197]" box="[324,617,1172,1197]" pageId="53" pageNumber="608">
(
<figureCitation id="1B892A02FFCF061DF96CEC8E7F325AF3" box="[332,560,1172,1197]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="53" pageNumber="608">FIG. 16A–J; TABLES</figureCitation>
1, 2)
</paragraph>
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<emphasis id="B1C6EA95FFCF061DF883ECC77C5B5AAA" box="[163,345,1245,1266]" italics="true" pageId="53" pageNumber="608">Type material:</emphasis>
<materialsCitation id="33DA3CDAFFCF061DF951ECC77F795B93" collectingDate="2015-04-04" collectionCode="IOM" county="Clarion-Clipperton Fracture Zone" elevation="0" latitude="11.043533" location="Equatorial Eastern Pacific Ocean" longLatPrecision="1" longitude="-119.65853" municipality="Eastern Pacific Ocean" pageId="53" pageNumber="608" specimenCode="IFR421" specimenCount="1" typeStatus="holotype">
<typeStatus id="5C098825FFCF061DF951ECC77CE15AAB" box="[369,483,1245,1267]" pageId="53" pageNumber="608" type="holotype">Holotype</typeStatus>
, MNHN-IA-TYPE 1832 (
<specimenCode id="D3149EFCFFCF061DF88AECE67C015B4A" box="[170,259,1276,1298]" collectionCode="IFR" pageId="53" pageNumber="608">IFR421</specimenCode>
), complete, length 
<quantity id="444A9B62FFCF061DF9F1ECE17F365B49" box="[465,564,1275,1297]" metricMagnitude="-3" metricUnit="m" metricValue="2.69" pageId="53" pageNumber="608" unit="mm" value="2.69">2.69 mm</quantity>
, width 
<quantity id="444A9B62FFCF061DFAA4ECE17FE55B49" box="[644,743,1275,1297]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="53" pageNumber="608" unit="mm" value="0.66">0.66 mm</quantity>
, 15 segments (including tentacular segment), 
<location id="866D605CFFCF061DFAAEED007CAF5B16" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DFAAEED007CAF5B16" county="Clarion-Clipperton Fracture Zone" latitude="11.043533" longLatPrecision="1" longitude="-119.65853" municipality="Eastern Pacific Ocean" name="Equatorial Eastern Pacific Ocean" pageId="53" pageNumber="608">
Equatorial 
<collectingMunicipality id="6369ACFDFFCF061DF883ED207CAF5B16" box="[163,429,1337,1359]" pageId="53" pageNumber="608">Eastern Pacific Ocean</collectingMunicipality>
</location>
, 
<collectingCounty id="6A6C4E0BFFCF061DF999ED237DDF5B36" pageId="53" pageNumber="608">Clarion-Clipperton Fracture Zone</collectingCounty>
, 
<collectionCode id="E5A3AE42FFCF061DF8CAED427C225B35" box="[234,288,1368,1389]" country="Russia" httpUri="http://grbio.org/cool/9zah-8jyz" name="Institute of Oceanology, Academy of Sciences" pageId="53" pageNumber="608">IOM</collectionCode>
license area, station 99, collected 
<date id="F70C1047FFCF061DFA99ED427DE25BD5" pageId="53" pageNumber="608" value="2015-04-04">
<collectingDate id="E748E9AFFFCF061DFA99ED427DE25BD5" pageId="53" pageNumber="608" value="2015-04-04">4 April 2015</collectingDate>
</date>
, epibenthic sledge epi-net, start 
<geoCoordinate id="E6865040FFCF061DFA5EED6D7E055BD5" box="[638,775,1398,1421]" degrees="11" direction="north" minutes="2.296" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="11.038267">11°2.296′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DF883ED8C7C475BF3" box="[163,325,1429,1452]" degrees="119" direction="west" minutes="40.825" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-119.68042">119°40.825′W</geoCoordinate>
, end 
<geoCoordinate id="E6865040FFCF061DF9A5ED8C7F0A5BF3" box="[389,520,1429,1452]" degrees="11" direction="north" minutes="2.612" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="11.043533">11°2.612′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFA35ED8C7FBA5BF3" box="[533,696,1429,1452]" degrees="119" direction="west" minutes="39.512" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-119.65853">119°39.512′W</geoCoordinate>
, 
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<quantity id="444A9B62FFCF061DFAE4ED8C7DFA5B92" metricMagnitude="3" metricUnit="m" metricValue="4.4" metricValueMax="4.401999999999999" metricValueMin="4.398" pageId="53" pageNumber="608" unit="m" value="4400.0" valueMax="4402.0" valueMin="4398.0">4398– 4402 m</quantity>
depth, 
<quantity id="444A9B62FFCF061DF970EDAF7CA75B92" box="[336,421,1461,1483]" metricMagnitude="3" metricUnit="m" metricValue="2.529" pageId="53" pageNumber="608" unit="m" value="2529.0">2529 m</quantity>
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trawling distance
</materialsCitation>
. 
<materialsCitation id="33DA3CDAFFCF061DFAA6EDAF7CE2589A" collectingDate="2015-04-07" collectionCode="GSR" county="Clarion-Clipperton Fracture Zone" elevation="0" latitude="13.877033" location="Equatorial Eastern Pacific Ocean" longLatPrecision="1" longitude="-123.23772" municipality="Eastern Pacific Ocean" pageId="53" pageNumber="608" specimenCode="IFR520-1" specimenCount="1" typeStatus="paratype">
<typeStatus id="5C098825FFCF061DFAA6EDAF7FED5B92" box="[646,751,1461,1482]" pageId="53" pageNumber="608" type="paratype">Paratype</typeStatus>
1, MNHN-IA-TYPE 1833 (
<specimenCode id="D3149EFCFFCF061DF9E0EDCE7F335BB2" box="[448,561,1492,1514]" collectionCode="IFR" pageId="53" pageNumber="608">IFR520-1</specimenCode>
), complete, length 
<quantity id="444A9B62FFCF061DF883EDE97C125850" box="[163,272,1523,1544]" metricMagnitude="-3" metricUnit="m" metricValue="1.97" pageId="53" pageNumber="608" unit="mm" value="1.97">1.97 mm</quantity>
, width 
<quantity id="444A9B62FFCF061DF94FEDE97CDF5850" box="[367,477,1523,1545]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="53" pageNumber="608" unit="mm" value="0.52">0.52 mm</quantity>
, 15 segments (including tentacular segment), 
<location id="866D605CFFCF061DF996EE087DEE581E" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DF996EE087DEE581E" county="Clarion-Clipperton Fracture Zone" latitude="13.877033" longLatPrecision="1" longitude="-123.23772" municipality="Eastern Pacific Ocean" name="Equatorial Eastern Pacific Ocean" pageId="53" pageNumber="608">
Equatorial 
<collectingMunicipality id="6369ACFDFFCF061DFA69EE087DEE581E" pageId="53" pageNumber="608">Eastern Pacific Ocean</collectingMunicipality>
</location>
, 
<collectingCounty id="6A6C4E0BFFCF061DF8D6EE2B7F75581E" box="[246,631,1585,1607]" pageId="53" pageNumber="608">Clarion-Clipperton Fracture Zone</collectingCounty>
, 
<collectionCode id="E5A3AE42FFCF061DFAA2EE2B7FB5581E" box="[642,695,1585,1606]" pageId="53" pageNumber="608">GSR</collectionCode>
license area, station 117, collected 
<date id="F70C1047FFCF061DF9CCEE4A7F80583D" box="[492,642,1615,1637]" pageId="53" pageNumber="608" value="2015-04-07">
<collectingDate id="E748E9AFFFCF061DF9CCEE4A7F80583D" box="[492,642,1615,1637]" pageId="53" pageNumber="608" value="2015-04-07">7 April 2015</collectingDate>
</date>
, epibenthic sledge supra-net, start 
<geoCoordinate id="E6865040FFCF061DF9E0EE747F5658DC" box="[448,596,1645,1668]" degrees="13" direction="north" minutes="52.317" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="13.87195">13°52.317′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFA42EE747E0B58DC" box="[610,777,1645,1668]" degrees="123" direction="west" minutes="15.442" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-123.25737">123°15.442′W</geoCoordinate>
, end 
<geoCoordinate id="E6865040FFCF061DF8F5EE977C6758FB" box="[213,357,1676,1699]" degrees="13" direction="north" minutes="52.622" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="13.877033">13°52.622′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DF951EE977F1058FB" box="[369,530,1676,1699]" degrees="123" direction="west" minutes="14.263" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-123.23772">123°14.263′W</geoCoordinate>
, 
<elevation id="089FD1B4FFCF061DFA3DEE977DFE589A" isEnumeration="true" pageId="53" pageNumber="608">
<quantity id="444A9B62FFCF061DFA3DEE977FB858FB" box="[541,698,1677,1699]" metricMagnitude="3" metricUnit="m" metricValue="4.5095" metricValueMax="4.521" metricValueMin="4.498" pageId="53" pageNumber="608" unit="m" value="4509.5" valueMax="4521.0" valueMin="4498.0">4498–4521 m</quantity>
depth, 
<quantity id="444A9B62FFCF061DF883EEB67DFE589A" box="[163,252,1708,1730]" metricMagnitude="3" metricUnit="m" metricValue="3.129" pageId="53" pageNumber="608" unit="m" value="3129.0">3129 m</quantity>
</elevation>
trawling distance
</materialsCitation>
. 
<materialsCitation id="33DA3CDAFFCF061DF9CEEEB77E9C5C48" collectingDate="2015-04-07" collectingDateMax="2015-04-15" collectingDateMin="2015-04-07" collectionCode="GSR" country="Clipperton Island" county="Fracture Zone" elevation="0" latitude="13.877033" location="Equatorial Eastern Pacific Ocean" longLatPrecision="1" longitude="-123.23772" municipality="Eastern Pacific Ocean" pageId="53" pageNumber="608" specimenCode="TYPE 1834, IFR449, IFR529-1" specimenCount="1" typeStatus="paratype">
<typeStatus id="5C098825FFCF061DF9CEEEB77F59589A" box="[494,603,1709,1730]" pageId="53" pageNumber="608" type="paratype">Paratype</typeStatus>
2, MNHN-IA- 
<specimenCode id="D3149EFCFFCF061DF883EED67C2C58B8" box="[163,302,1739,1761]" collectionCode="TYPE" pageId="53" pageNumber="608">TYPE 1834</specimenCode>
(
<specimenCode id="D3149EFCFFCF061DF960EED17C9D58B9" box="[320,415,1739,1761]" collectionCode="IFR" pageId="53" pageNumber="608">IFR449</specimenCode>
b), complete, length 
<quantity id="444A9B62FFCF061DFABAEED17E0458B9" box="[666,774,1739,1761]" metricMagnitude="-3" metricUnit="m" metricValue="1.41" pageId="53" pageNumber="608" unit="mm" value="1.41">1.41 mm</quantity>
, width 
<quantity id="444A9B62FFCF061DF8D1EEF07C5E58A7" box="[241,348,1770,1792]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="53" pageNumber="608" unit="mm" value="0.46">0.46 mm</quantity>
, 15 segments (including tentacular segment), 
<location id="866D605CFFCF061DF93AEF137FA25946" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DF93AEF137FA25946" box="[282,672,1801,1823]" country="Clipperton Island" county="Fracture Zone" latitude="13.877033" longLatPrecision="1" longitude="-123.23772" municipality="Eastern Pacific Ocean" name="Equatorial Eastern Pacific Ocean" pageId="53" pageNumber="608">
Equatorial 
<location id="866D605CFFCF061DF9BEEF137FA25946" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DF9BEEF137FA25946" box="[414,672,1801,1823]" country="Clipperton Island" county="Fracture Zone" latitude="13.877033" longLatPrecision="1" longitude="-123.23772" municipality="Eastern Pacific Ocean" name="Eastern Pacific Ocean" pageId="53" pageNumber="608">Eastern Pacific Ocean</location>
</location>
, Clarion- 
<collectingCountry id="FBA57617FFCF061DF883EF327C285966" box="[163,298,1832,1854]" name="Clipperton Island" pageId="53" pageNumber="608">Clipperton</collectingCountry>
<location id="866D605CFFCF061DF916EF327CEC5965" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DF916EF327CEC5965" box="[310,494,1832,1853]" country="Clipperton Island" county="Fracture Zone" latitude="13.877033" longLatPrecision="1" longitude="-123.23772" municipality="Eastern Pacific Ocean" name="Fracture Zone" pageId="53" pageNumber="608">Fracture Zone</location>
, 
<location id="866D605CFFCF061DF9DEEF327F5F5966" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DF9DEEF327F5F5966" box="[510,605,1832,1854]" country="Clipperton Island" county="Fracture Zone" latitude="13.877033" longLatPrecision="1" longitude="-123.23772" municipality="Eastern Pacific Ocean" name="Ifremer" pageId="53" pageNumber="608">Ifremer</location>
license area, station 158, collected 
<date id="F70C1047FFCF061DF9E2EF5D7F775905" box="[450,629,1863,1885]" pageId="53" pageNumber="608" value="2015-04-15">
<collectingDate id="E748E9AFFFCF061DF9E2EF5D7F775905" box="[450,629,1863,1885]" pageId="53" pageNumber="608" value="2015-04-15">15 April 2015</collectingDate>
</date>
, epibenthic sledge supra-net, start 
<geoCoordinate id="E6865040FFCF061DFC6CE8DC79CC5E84" box="[1100,1230,197,220]" degrees="14" direction="north" minutes="3.411" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="14.05685">14°3.411′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFCFAE8DC786F5E84" box="[1242,1389,197,220]" degrees="130" direction="west" minutes="7.989" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-130.13315">130°7.989′W</geoCoordinate>
, end 
<geoCoordinate id="E6865040FFCF061DFB1BE8FF7EBD5EA2" box="[827,959,228,251]" degrees="14" direction="north" minutes="3.813" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="14.06355">14°3.813′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFBECE8FF79635EA2" box="[972,1121,228,251]" degrees="130" direction="west" minutes="6.481" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-130.10802">130°6.481′W</geoCoordinate>
, 
<elevation id="089FD1B4FFCF061DFC4EE8FF7E535F41" isEnumeration="true" pageId="53" pageNumber="608">
<quantity id="444A9B62FFCF061DFC4EE8FF780C5EA2" box="[1134,1294,229,251]" metricMagnitude="3" metricUnit="m" metricValue="4.962" metricValueMax="4.978" metricValueMin="4.946" pageId="53" pageNumber="608" unit="m" value="4962.0" valueMax="4978.0" valueMin="4946.0">4946–4978 m</quantity>
depth, 
<quantity id="444A9B62FFCF061DFD4AE8FF7E535F41" metricMagnitude="3" metricUnit="m" metricValue="3.789" pageId="53" pageNumber="608" unit="m" value="3789.0">3789 m</quantity>
</elevation>
trawling distance. 
<location id="866D605CFFCF061DFC61E91E79B75F41" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DFC61E91E79B75F41" box="[1089,1205,258,281]" country="Clipperton Island" county="Fracture Zone" latitude="13.877033" longLatPrecision="1" longitude="-123.23772" municipality="Eastern Pacific Ocean" name="Paratype'" pageId="53" pageNumber="608">Paratype′</location>
3, MNHN-IA-TYPE 1835 (
<specimenCode id="D3149EFCFFCF061DFBA0E9387EEF5F60" box="[896,1005,290,312]" collectionCode="IFR" pageId="53" pageNumber="608">IFR529-1</specimenCode>
), incomplete for SEM, length 
<quantity id="444A9B62FFCF061DFD1BE939789D5F60" box="[1339,1439,291,312]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="53" pageNumber="608" unit="mm" value="1.24">1.24 mm</quantity>
, width 
<quantity id="444A9B62FFCF061DFBA6E95B7EEF5F0F" box="[902,1005,321,343]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="53" pageNumber="608" unit="mm" value="0.4">0.40 mm</quantity>
, nine segments (including tentacular segment), 
<location id="866D605CFFCF061DFB92E97A783A5F2D" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DFB92E97A783A5F2D" box="[946,1336,352,374]" country="Clipperton Island" county="Fracture Zone" latitude="13.877033" longLatPrecision="1" longitude="-123.23772" municipality="Eastern Pacific Ocean" name="Equatorial Eastern Pacific Ocean" pageId="53" pageNumber="608">
Equatorial 
<collectingMunicipality id="6369ACFDFFCF061DFC16E97B783A5F2D" box="[1078,1336,352,374]" pageId="53" pageNumber="608">Eastern Pacific Ocean</collectingMunicipality>
</location>
, Clarion- 
<collectingCountry id="FBA57617FFCF061DFB1BE9657EBB5FCD" box="[827,953,383,405]" name="Clipperton Island" pageId="53" pageNumber="608">Clipperton</collectingCountry>
<collectingCounty id="6A6C4E0BFFCF061DFBE1E96579685FCC" box="[961,1130,383,404]" pageId="53" pageNumber="608">Fracture Zone</collectingCounty>
, 
<collectionCode id="E5A3AE42FFCF061DFC57E96579AE5FCC" box="[1143,1196,383,404]" pageId="53" pageNumber="608">GSR</collectionCode>
license area, station 117, collected 
<date id="F70C1047FFCF061DFBFEE984796D5FEC" box="[990,1135,414,436]" pageId="53" pageNumber="608" value="2015-04-07">
<collectingDate id="E748E9AFFFCF061DFBFEE984796D5FEC" box="[990,1135,414,436]" pageId="53" pageNumber="608" value="2015-04-07">7 April 2015</collectingDate>
</date>
, epibenthic sledge epi-net, start 
<geoCoordinate id="E6865040FFCF061DFB5FE9A679115F8A" box="[895,1043,443,466]" degrees="13" direction="north" minutes="52.317" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="13.87195">13°52.317′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFC02E9A679C55F8A" box="[1058,1223,443,466]" degrees="123" direction="west" minutes="15.442" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-123.25737">123°15.442′W</geoCoordinate>
, end 
<geoCoordinate id="E6865040FFCF061DFD2AE9A6789D5F8A" box="[1290,1439,443,466]" degrees="13" direction="north" minutes="52.622" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="13.877033">13°52.622′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFB1BE9C17EDE5FA9" box="[827,988,474,497]" degrees="123" direction="west" minutes="14.263" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-123.23772">123°14.263′W</geoCoordinate>
, 
<elevation id="089FD1B4FFCF061DFBC8E9C178345FA9" box="[1000,1334,475,497]" isEnumeration="true" pageId="53" pageNumber="608">
<quantity id="444A9B62FFCF061DFBC8E9C179875FA9" box="[1000,1157,475,497]" metricMagnitude="3" metricUnit="m" metricValue="4.5095" metricValueMax="4.521" metricValueMin="4.498" pageId="53" pageNumber="608" unit="m" value="4509.5" valueMax="4521.0" valueMin="4498.0">4498–4521 m</quantity>
depth, 
<quantity id="444A9B62FFCF061DFCC0E9C678345FA9" box="[1248,1334,476,497]" metricMagnitude="3" metricUnit="m" metricValue="3.129" pageId="53" pageNumber="608" unit="m" value="3129.0">3129 m</quantity>
</elevation>
trawling distance
</materialsCitation>
. 
<materialsCitation id="33DA3CDAFFCF061DFB8AE9E1789C5C48" box="[938,1438,506,528]" collectionCode="NHMUK" country="Clipperton Island" pageId="53" pageNumber="608" specimenCode="NHMUK 2018.25351, IFR208" specimenCount="1" typeStatus="paratype">
<typeStatus id="5C098825FFCF061DFB8AE9E179115C48" box="[938,1043,507,528]" pageId="53" pageNumber="608" type="paratype">Paratype</typeStatus>
4, 
<specimenCode id="D3149EFCFFCF061DFC17E9E1782E5C48" box="[1079,1324,506,528]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="53" pageNumber="608" type="Museum">NHMUK 2018.25351</specimenCode>
(
<specimenCode id="D3149EFCFFCF061DFD1CE9E078945C48" box="[1340,1430,506,528]" collectionCode="IFR" pageId="53" pageNumber="608">IFR208</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="33DA3CDAFFCF061DFB1BEA037E715CD3" collectionCode="IFR" country="Clipperton Island" location="Eastern Pacific Ocean" pageId="53" pageNumber="608" specimenCount="1" typeStatus="paratype">
complete, length 
<quantity id="444A9B62FFCF061DFC02EA0379945C76" box="[1058,1174,537,558]" metricMagnitude="-3" metricUnit="m" metricValue="2.19" pageId="53" pageNumber="608" unit="mm" value="2.19">2.19 mm</quantity>
, width 
<quantity id="444A9B62FFCF061DFCDFEA0378715C76" box="[1279,1395,537,559]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="53" pageNumber="608" unit="mm" value="0.52">0.52 mm</quantity>
, 15 segments (including tentacular segment), Equatorial 
<location id="866D605CFFCF061DFB1BEA4D79475C34" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DFB1BEA4D79475C34" box="[827,1093,599,621]" country="Clipperton Island" name="Eastern Pacific Ocean" pageId="53" pageNumber="608">Eastern Pacific Ocean</location>
, Clarion-Clipperton Fracture Zone
</materialsCitation>
, 
<materialsCitation id="33DA3CDAFFCF061DFB5EEA6C78115CB0" collectingDate="2015-03-21" collectionCode="BGR" country="Clipperton Island" elevation="0" latitude="11.836333" longLatPrecision="13" longitude="-116.974335" pageId="53" pageNumber="608" specimenCount="1" typeStatus="paratype">
<collectionCode id="E5A3AE42FFCF061DFB5EEA6C7EB75CD3" box="[894,949,630,651]" country="Germany" httpUri="http://grbio.org/cool/r4mz-yr8h" name="Bundesanstalt fur Geowissenschaften und Rohstoffe" pageId="53" pageNumber="608">BGR</collectionCode>
license area, station 20, collected 
<date id="F70C1047FFCF061DFD17EA6C7E7A5CF2" pageId="53" pageNumber="608" value="2015-03-21">
<collectingDate id="E748E9AFFFCF061DFD17EA6C7E7A5CF2" pageId="53" pageNumber="608" value="2015-03-21">21 March 2015</collectingDate>
</date>
, epibenthic sledge epi-net, start 
<geoCoordinate id="E6865040FFCF061DFD36EA8E789D5CF2" box="[1302,1439,659,682]" degrees="11" direction="north" minutes="50.15" orientation="latitude" pageId="53" pageNumber="608" precision="9" value="11.835834">11°50.15′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFB1BEAA97ED65C91" box="[827,980,690,713]" degrees="117" direction="west" minutes="58.49" orientation="longitude" pageId="53" pageNumber="608" precision="9" value="-117.97483">117°58.49′W</geoCoordinate>
, end 
<geoCoordinate id="E6865040FFCF061DFC3AEAA979A05C91" box="[1050,1186,690,713]" degrees="11" direction="north" minutes="50.18" orientation="latitude" pageId="53" pageNumber="608" precision="9" value="11.836333">11°50.18′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFC91EAA978495C91" box="[1201,1355,690,713]" degrees="116" direction="west" minutes="58.46" orientation="longitude" pageId="53" pageNumber="608" precision="9" value="-116.974335">116°58.46′W</geoCoordinate>
, 4144– 
<elevation id="089FD1B4FFCF061DFB1BEAC8793F5CBF" box="[827,1085,722,744]" isEnumeration="true" pageId="53" pageNumber="608">
<quantity id="444A9B62FFCF061DFB1BEAC87E925CBF" box="[827,912,722,743]" metricMagnitude="3" metricUnit="m" metricValue="4.093" pageId="53" pageNumber="608" unit="m" value="4093.0">4093 m</quantity>
depth, 
<quantity id="444A9B62FFCF061DFBC8EAC8793F5CBF" box="[1000,1085,722,744]" metricMagnitude="3" metricUnit="m" metricValue="2.7689999999999997" pageId="53" pageNumber="608" unit="m" value="2769.0">2769 m</quantity>
</elevation>
trawling distance
</materialsCitation>
. 
<materialsCitation id="33DA3CDAFFCF061DFD3EEAC879C85D5F" collectionCode="NHMUK" country="Clipperton Island" pageId="53" pageNumber="608" specimenCode="NHMUK 2018.25352, IFR538-3" specimenCount="1" typeStatus="paratype">
<typeStatus id="5C098825FFCF061DFD3EEAC878855CBF" box="[1310,1415,722,743]" pageId="53" pageNumber="608" type="paratype">Paratype</typeStatus>
5, 
<specimenCode id="D3149EFCFFCF061DFB1BEAEB79385D5F" box="[827,1082,753,775]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="53" pageNumber="608" type="Museum">NHMUK 2018.25352</specimenCode>
(
<specimenCode id="D3149EFCFFCF061DFC6CEAEB79C05D5F" box="[1100,1218,753,775]" collectionCode="IFR" pageId="53" pageNumber="608">IFR538-3</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="33DA3CDAFFCF061DFCF9EAEB780D5D3B" collectionCode="IFR" country="Clipperton Island" location="Eastern Pacific Ocean" pageId="53" pageNumber="608" specimenCount="1" typeStatus="paratype">
complete, length 
<quantity id="444A9B62FFCF061DFB1BEB0A7EAA5D7D" box="[827,936,784,805]" metricMagnitude="-3" metricUnit="m" metricValue="2.13" pageId="53" pageNumber="608" unit="mm" value="2.13">2.13 mm</quantity>
, width 
<quantity id="444A9B62FFCF061DFC27EB1579775D7D" box="[1031,1141,783,805]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="53" pageNumber="608" unit="mm" value="0.51">0.51 mm</quantity>
, 14 segments (including tentacular segment), Equatorial 
<location id="866D605CFFCF061DFCC1EB357E865D3A" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DFCC1EB357E865D3A" country="Clipperton Island" name="Eastern Pacific Ocean" pageId="53" pageNumber="608">Eastern Pacific Ocean</location>
, Clarion-Clipperton Fracture Zone
</materialsCitation>
, 
<materialsCitation id="33DA3CDAFFCF061DFD3AEB5779235D86" collectingDate="2015-04-10" collectionCode="GSR" country="Clipperton Island" elevation="0" latitude="13.8521" longLatPrecision="1" longitude="-123.23552" pageId="53" pageNumber="608" specimenCount="1" typeStatus="paratype">
<collectionCode id="E5A3AE42FFCF061DFD3AEB57784D5D3A" box="[1306,1359,845,866]" pageId="53" pageNumber="608">GSR</collectionCode>
license area, station 133, collected 
<date id="F70C1047FFCF061DFC5CEB76781F5DDA" box="[1148,1309,876,898]" pageId="53" pageNumber="608" value="2015-04-10">
<collectingDate id="E748E9AFFFCF061DFC5CEB76781F5DDA" box="[1148,1309,876,898]" pageId="53" pageNumber="608" value="2015-04-10">10 April 2015</collectingDate>
</date>
, epibenthic sledge supra-net, start 
<geoCoordinate id="E6865040FFCF061DFC1EEB9179C85DF8" box="[1086,1226,905,929]" degrees="13" direction="north" minutes="50.751" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="13.84585">13°50.751′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFCF4EB9178725DF8" box="[1236,1392,905,929]" degrees="123" direction="west" minutes="15.649" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-123.26082">123°15.649′W</geoCoordinate>
, end 
<geoCoordinate id="E6865040FFCF061DFB1BEBB37ECA5DE7" box="[827,968,936,959]" degrees="13" direction="north" minutes="51.126" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="13.8521">13°51.126′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFBF3EBB079735DE7" box="[979,1137,936,959]" degrees="123" direction="west" minutes="14.131" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-123.23552">123°14.131′W</geoCoordinate>
, 4516– 
<elevation id="089FD1B4FFCF061DFCE1EBB37E535D86" isEnumeration="true" pageId="53" pageNumber="608">
<quantity id="444A9B62FFCF061DFCE1EBB378175DE7" box="[1217,1301,937,959]" metricMagnitude="3" metricUnit="m" metricValue="4.427" pageId="53" pageNumber="608" unit="m" value="4427.0">4427 m</quantity>
depth, 
<quantity id="444A9B62FFCF061DFD4BEBB07E535D86" metricMagnitude="3" metricUnit="m" metricValue="2.289" pageId="53" pageNumber="608" unit="m" value="2289.0">2289 m</quantity>
</elevation>
trawling distance
</materialsCitation>
.
</paragraph>
<paragraph id="830D3687FFCF061EFB1BEC1D7DF55F42" blockId="53.[827,1444,1031,1885]" lastBlockId="54.[145,762,197,282]" lastPageId="54" lastPageNumber="609" pageId="53" pageNumber="608">
<emphasis id="B1C6EA95FFCF061DFB1BEC1D794E5A44" box="[827,1100,1031,1052]" italics="true" pageId="53" pageNumber="608">Additional material:</emphasis>
<materialsCitation id="33DA3CDAFFCF061EFC48EC1D7DF15F42" collectingDate="2015-04-04" collectingDateMax="2015-04-15" collectingDateMin="2015-04-04" collectionCode="IOM" collectorName="Specimen &amp; Equatorial Eastern Pacific Ocean &amp; Clarion-Clipperton Fracture Zone &amp; Ifremer &amp; Paulo Bonifácio &amp; Fracture Zone" country="Clipperton Island" county="Eastern Pacific Ocean" elevation="0" lastPageId="54" lastPageNumber="609" latitude="14.06355" location="Equatorial Eastern Pacific Ocean" longLatPrecision="1" longitude="-130.10802" municipality="Eastern Pacific Ocean" pageId="53" pageNumber="608" specimenCode="IFR422, IFR423, IFR682, IFR460" specimenCount="1">
Specimen 1, MNHN-IA- PNT 79 (
<specimenCode id="D3149EFCFFCF061DFB87EC3C79015A64" box="[935,1027,1062,1084]" collectionCode="IFR" pageId="53" pageNumber="608">IFR422</specimenCode>
), complete, length 
<quantity id="444A9B62FFCF061DFCC7EC3C78535A63" box="[1255,1361,1062,1084]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="53" pageNumber="608" unit="mm" value="2.75">2.75 mm</quantity>
, width 
<quantity id="444A9B62FFCF061DFB1BEC5F7E9D5A02" box="[827,927,1093,1115]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="53" pageNumber="608" unit="mm" value="0.62">0.62 mm</quantity>
, 16 segments (including tentacular segment), 
<location id="866D605CFFCF061DFB1BEC7E79BC5A21" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DFB1BEC7E79BC5A21" box="[827,1214,1124,1146]" country="Clipperton Island" county="Eastern Pacific Ocean" latitude="14.06355" longLatPrecision="1" longitude="-130.10802" municipality="Eastern Pacific Ocean" name="Equatorial Eastern Pacific Ocean" pageId="53" pageNumber="608">
Equatorial 
<location id="866D605CFFCF061DFB9EEC7E79BC5A21" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DFB9EEC7E79BC5A21" box="[958,1214,1124,1146]" country="Clipperton Island" county="Eastern Pacific Ocean" latitude="14.06355" longLatPrecision="1" longitude="-130.10802" municipality="Eastern Pacific Ocean" name="Eastern Pacific Ocean" pageId="53" pageNumber="608">Eastern Pacific Ocean</location>
</location>
, 
<location id="866D605CFFCF061DFCE9EC7E7EDE5AC0" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DFCE9EC7E7EDE5AC0" country="Clipperton Island" county="Eastern Pacific Ocean" latitude="14.06355" longLatPrecision="1" longitude="-130.10802" municipality="Eastern Pacific Ocean" name="Clarion-Clipperton Fracture Zone" pageId="53" pageNumber="608">Clarion-Clipperton Fracture Zone</location>
, 
<collectionCode id="E5A3AE42FFCF061DFBC6EC9979195AC0" box="[998,1051,1155,1176]" country="Russia" httpUri="http://grbio.org/cool/9zah-8jyz" name="Institute of Oceanology, Academy of Sciences" pageId="53" pageNumber="608">IOM</collectionCode>
license area, station 99, collected 
<date id="F70C1047FFCF061DFDB5EC997EB65AEF" pageId="53" pageNumber="608" value="2015-04-04">
<collectingDate id="E748E9AFFFCF061DFDB5EC997EB65AEF" pageId="53" pageNumber="608" value="2015-04-04">4 April 2015</collectingDate>
</date>
, epibenthic sledge epi-net, start 
<geoCoordinate id="E6865040FFCF061DFD01ECBB78A25AEF" box="[1313,1440,1184,1207]" degrees="11" direction="north" minutes="2.296" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="11.038267">11°2.296′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFB1BECDA7EDF5A8D" box="[827,989,1215,1238]" degrees="119" direction="west" minutes="40.825" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-119.68042">119°40.825′W</geoCoordinate>
, end 
<geoCoordinate id="E6865040FFCF061DFC3DECDA79A25A8D" box="[1053,1184,1215,1238]" degrees="11" direction="north" minutes="2.612" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="11.043533">11°2.612′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFC8DECDA78525A8D" box="[1197,1360,1215,1238]" degrees="119" direction="west" minutes="39.512" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-119.65853">119°39.512′W</geoCoordinate>
, 
<elevation id="089FD1B4FFCF061DFD7CECDA79395AAC" isEnumeration="true" pageId="53" pageNumber="608">
<quantity id="444A9B62FFCF061DFD7CECDA7E8D5AAC" metricMagnitude="3" metricUnit="m" metricValue="4.4" metricValueMax="4.401999999999999" metricValueMin="4.398" pageId="53" pageNumber="608" unit="m" value="4400.0" valueMax="4402.0" valueMin="4398.0">4398– 4402 m</quantity>
depth, 
<quantity id="444A9B62FFCF061DFBC6ECC579395AAC" box="[998,1083,1247,1269]" metricMagnitude="3" metricUnit="m" metricValue="2.529" pageId="53" pageNumber="608" unit="m" value="2529.0">2529 m</quantity>
</elevation>
trawling distance. Specimen 2, MNHN-IA-PNT 80 (
<specimenCode id="D3149EFCFFCF061DFC66ECE479AE5B4C" box="[1094,1196,1278,1300]" collectionCode="IFR" pageId="53" pageNumber="608">IFR423</specimenCode>
), complete, length 
<quantity id="444A9B62FFCF061DFB1BED077EAA5B6A" box="[827,936,1309,1330]" metricMagnitude="-3" metricUnit="m" metricValue="1.84" pageId="53" pageNumber="608" unit="mm" value="1.84">1.84 mm</quantity>
, width 
<quantity id="444A9B62FFCF061DFC27ED0779775B6A" box="[1031,1141,1309,1330]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="53" pageNumber="608" unit="mm" value="0.44">0.44 mm</quantity>
, 15 segments (including tentacular segment), Equatorial 
<location id="866D605CFFCF061DFCC1ED267E865B37" LSID="urn:lsid:plazi:treatment:0B1B8791FFCF0610F8EEEC777C755A63:866D605CFFCF061DFCC1ED267E865B37" country="Clipperton Island" county="Eastern Pacific Ocean" latitude="14.06355" longLatPrecision="1" longitude="-130.10802" municipality="Eastern Pacific Ocean" name="Eastern Pacific Ocean" pageId="53" pageNumber="608">Eastern Pacific Ocean</location>
, Clarion-Clipperton Fracture Zone, 
<collectionCode id="E5A3AE42FFCF061DFD3AED40784D5B37" box="[1306,1359,1370,1391]" country="Russia" httpUri="http://grbio.org/cool/9zah-8jyz" name="Institute of Oceanology, Academy of Sciences" pageId="53" pageNumber="608">IOM</collectionCode>
license area, station 99, collected 
<date id="F70C1047FFCF061DFC5DED6378155BD7" box="[1149,1303,1401,1423]" pageId="53" pageNumber="608" value="2015-04-04">
<collectingDate id="E748E9AFFFCF061DFC5DED6378155BD7" box="[1149,1303,1401,1423]" pageId="53" pageNumber="608" value="2015-04-04">4 April 2015</collectingDate>
</date>
, epibenthic sledge epi-net, start 
<geoCoordinate id="E6865040FFCF061DFC15ED8279BB5BF5" box="[1077,1209,1430,1454]" degrees="11" direction="north" minutes="2.296" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="11.038267">11°2.296′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFCE7ED8D78695BF5" box="[1223,1387,1430,1453]" degrees="119" direction="west" minutes="40.825" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-119.68042">119°40.825′W</geoCoordinate>
, end 
<geoCoordinate id="E6865040FFCF061DFB1BEDAC7EBE5B94" box="[827,956,1461,1484]" degrees="11" direction="north" minutes="2.612" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="11.043533">11°2.612′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFBE9EDAC796B5B94" box="[969,1129,1461,1484]" degrees="119" direction="west" minutes="39.512" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-119.65853">119°39.512′W</geoCoordinate>
, 
<elevation id="089FD1B4FFCF061DFC55EDAD7E535BB3" isEnumeration="true" pageId="53" pageNumber="608">
<quantity id="444A9B62FFCF061DFC55EDAD78135B94" box="[1141,1297,1463,1484]" metricMagnitude="3" metricUnit="m" metricValue="4.4" metricValueMax="4.401999999999999" metricValueMin="4.398" pageId="53" pageNumber="608" unit="m" value="4400.0" valueMax="4402.0" valueMin="4398.0">4398–4402 m</quantity>
depth, 
<quantity id="444A9B62FFCF061DFD4BEDAC7E535BB3" metricMagnitude="3" metricUnit="m" metricValue="2.529" pageId="53" pageNumber="608" unit="m" value="2529.0">2529 m</quantity>
</elevation>
trawling distance. 
<collectorName id="2E475351FFCF061DFC13EDCF79A15BB3" box="[1075,1187,1493,1515]" pageId="53" pageNumber="608">Specimen</collectorName>
3, MNHN-IA-PNT 81 (
<specimenCode id="D3149EFCFFCF061DFB62EDEE7E995852" box="[834,923,1524,1546]" collectionCode="IFR" pageId="53" pageNumber="608">IFR682</specimenCode>
), complete, length 
<quantity id="444A9B62FFCF061DFC49EDEE79CE5852" box="[1129,1228,1524,1546]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="53" pageNumber="608" unit="mm" value="1.11">1.11 mm</quantity>
, width 
<quantity id="444A9B62FFCF061DFD3CEDEE787D5852" box="[1308,1407,1524,1546]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="53" pageNumber="608" unit="mm" value="0.3">0.30 mm</quantity>
, 11 segments (including tentacular segment), 
<collectorName id="2E475351FFCF061DFD06EE097947581F" pageId="53" pageNumber="608">
Equatorial 
<collectingMunicipality id="6369ACFDFFCF061DFB1BEE287947581F" box="[827,1093,1586,1608]" pageId="53" pageNumber="608">Eastern Pacific Ocean</collectingMunicipality>
</collectorName>
, 
<collectorName id="2E475351FFCF061DFC72EE287E77583E" pageId="53" pageNumber="608">Clarion-Clipperton Fracture Zone</collectorName>
, 
<collectorName id="2E475351FFCF061DFBA2EE4A7EDE583E" box="[898,988,1616,1638]" pageId="53" pageNumber="608">Ifremer</collectorName>
license area, station 158, collected 
<date id="F70C1047FFCF061DFDA7EE4A7EB658DD" pageId="53" pageNumber="608" value="2015-04-15">
<collectingDate id="E748E9AFFFCF061DFDA7EE4A7EB658DD" pageId="53" pageNumber="608" value="2015-04-15">15 April 2015</collectingDate>
</date>
, epibenthic sledge epi-net, start 
<geoCoordinate id="E6865040FFCF061DFD01EE7578A258DD" box="[1313,1440,1646,1669]" degrees="14" direction="north" minutes="3.411" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="14.05685">14°3.411′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFB1BEE947EC858FC" box="[827,970,1677,1700]" degrees="130" direction="west" minutes="7.989" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-130.13315">130°7.989′W</geoCoordinate>
, end 
<geoCoordinate id="E6865040FFCF061DFC25EE94798158FC" box="[1029,1155,1677,1700]" degrees="14" direction="north" minutes="3.813" orientation="latitude" pageId="53" pageNumber="608" precision="1" value="14.06355">14°3.813′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCF061DFCAEEE94781F58FC" box="[1166,1309,1677,1700]" degrees="130" direction="west" minutes="6.481" orientation="longitude" pageId="53" pageNumber="608" precision="1" value="-130.10802">130°6.481′W</geoCoordinate>
, 
<elevation id="089FD1B4FFCF061DFD07EE947EF9589A" isEnumeration="true" pageId="53" pageNumber="608">
<quantity id="444A9B62FFCF061DFD07EE947E53589A" metricMagnitude="3" metricUnit="m" metricValue="4.962" metricValueMax="4.978" metricValueMin="4.946" pageId="53" pageNumber="608" unit="m" value="4962.0" valueMax="4978.0" valueMin="4946.0">4946–4978 m</quantity>
depth, 
<quantity id="444A9B62FFCF061DFB87EEB77EF9589A" box="[935,1019,1709,1730]" metricMagnitude="3" metricUnit="m" metricValue="3.789" pageId="53" pageNumber="608" unit="m" value="3789.0">3789 m</quantity>
</elevation>
trawling distance. 
<collectorName id="2E475351FFCF061DFCF7EEB77847589B" box="[1239,1349,1709,1731]" pageId="53" pageNumber="608">Specimen</collectorName>
4, 
<collectorName id="2E475351FFCF061DFD48EEB77897589A" box="[1384,1429,1709,1730]" name="Paulo Bonifácio" pageId="53" pageNumber="608">P.B.</collectorName>
’s collection (
<specimenCode id="D3149EFCFFCF061DFBE0EED6791D58BA" box="[960,1055,1740,1762]" collectionCode="IFR" pageId="53" pageNumber="608">IFR460</specimenCode>
b), incomplete, length 
<quantity id="444A9B62FFCF061DFD12EED6789C58B9" box="[1330,1438,1740,1761]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="53" pageNumber="608" unit="mm" value="1.4">1.40 mm</quantity>
, width 
<quantity id="444A9B62FFCF061DFBA8EEF17EF35958" box="[904,1009,1771,1792]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="53" pageNumber="608" unit="mm" value="0.42">0.42 mm</quantity>
, ten segments (including tentacular segment), 
<collectorName id="2E475351FFCF061DFB92EF13783A5946" box="[946,1336,1801,1823]" pageId="53" pageNumber="608">
Equatorial 
<collectingCounty id="6A6C4E0BFFCF061DFC16EF10783A5946" box="[1078,1336,1801,1823]" pageId="53" pageNumber="608">Eastern Pacific Ocean</collectingCounty>
</collectorName>
, Clarion- 
<collectingCountry id="FBA57617FFCF061DFB1BEF327EC05966" box="[827,962,1832,1854]" name="Clipperton Island" pageId="53" pageNumber="608">Clipperton</collectingCountry>
<collectorName id="2E475351FFCF061DFBEEEF3379845966" box="[974,1158,1833,1854]" pageId="53" pageNumber="608">Fracture Zone</collectorName>
, 
<collectorName id="2E475351FFCF061DFCB6EF3279F75966" box="[1174,1269,1832,1854]" pageId="53" pageNumber="608">Ifremer</collectorName>
license area, station 158, collected 
<date id="F70C1047FFCF061DFC13EF5D79D35905" box="[1075,1233,1863,1885]" pageId="53" pageNumber="608" value="2015-04-15">
<collectingDate id="E748E9AFFFCF061DFC13EF5D79D35905" box="[1075,1233,1863,1885]" pageId="53" pageNumber="608" value="2015-04-15">15 April 2015</collectingDate>
</date>
, epibenthic sledge epi-net, start 
<geoCoordinate id="E6865040FFCC061EF908E8DC7CA55E84" box="[296,423,197,220]" degrees="14" direction="north" minutes="3.411" orientation="latitude" pageId="54" pageNumber="609" precision="1" value="14.05685">14°3.411′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCC061EF990E8DC7F3D5E84" box="[432,575,197,220]" degrees="130" direction="west" minutes="7.989" orientation="longitude" pageId="54" pageNumber="609" precision="1" value="-130.13315">130°7.989′W</geoCoordinate>
, end 
<geoCoordinate id="E6865040FFCC061EFA57E8DC7FF45E84" box="[631,758,197,220]" degrees="14" direction="north" minutes="3.813" orientation="latitude" pageId="54" pageNumber="609" precision="1" value="14.06355">14°3.813′N</geoCoordinate>
, 
<geoCoordinate id="E6865040FFCC061EF8B1E8FF7C255EA2" box="[145,295,228,251]" degrees="130" direction="west" minutes="6.481" orientation="longitude" pageId="54" pageNumber="609" precision="1" value="-130.10802">130°6.481′W</geoCoordinate>
, 
<elevation id="089FD1B4FFCC061EF914E8FF7F885EA2" box="[308,650,229,251]" isEnumeration="true" pageId="54" pageNumber="609">
<quantity id="444A9B62FFCC061EF914E8FF7CD65EA2" box="[308,468,229,251]" metricMagnitude="3" metricUnit="m" metricValue="4.962" metricValueMax="4.978" metricValueMin="4.946" pageId="54" pageNumber="609" unit="m" value="4962.0" valueMax="4978.0" valueMin="4946.0">4946–4978 m</quantity>
depth, 
<quantity id="444A9B62FFCC061EFA11E8FF7F885EA2" box="[561,650,229,250]" metricMagnitude="3" metricUnit="m" metricValue="3.789" pageId="54" pageNumber="609" unit="m" value="3789.0">3789 m</quantity>
</elevation>
trawling distance
</materialsCitation>
.
</paragraph>
<paragraph id="830D3687FFCC061EF8B1E9597FC45C48" blockId="54.[145,762,322,1877]" pageId="54" pageNumber="609">
<emphasis id="B1C6EA95FFCC061EF8B1E9597F855F0F" box="[145,647,322,344]" italics="true" pageId="54" pageNumber="609">Description(basedonholotypeandparatypes):</emphasis>
<typeStatus id="5C098825FFCC061EFAB2E9587FFA5F00" box="[658,760,322,344]" pageId="54" pageNumber="609" type="holotype">Holotype</typeStatus>
complete, 
<quantity id="444A9B62FFCC061EF92EE97B7C7B5F2E" box="[270,377,353,375]" metricMagnitude="-3" metricUnit="m" metricValue="2.69" pageId="54" pageNumber="609" unit="mm" value="2.69">2.69 mm</quantity>
long and 
<quantity id="444A9B62FFCC061EF9D8E97B7F615F2E" box="[504,611,353,375]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="54" pageNumber="609" unit="mm" value="0.66">0.66 mm</quantity>
wide for 15 segments (including tentacular segment), slightly dorsoventrally flattened, slightly tapering posteriorly; live specimen bluish; ethanol-preserved pale brown (
<figureCitation id="1B892A02FFCC061EF8B9E9C67DFE5FA9" box="[153,252,475,498]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig. 16A</figureCitation>
), with brownish bands subbasally on palps and medially on dorsal cirri; prostomium whitish.
</paragraph>
<paragraph id="830D3687FFCC061EF889EA037C785B14" blockId="54.[145,762,322,1877]" pageId="54" pageNumber="609">
Prostomium bilobed but continuously fused, wider than long, lobes not pronounced anteriorly but extending to ventral side (
<figureCitation id="1B892A02FFCC061EF9A7EA4C7F0D5C34" box="[391,527,598,620]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig. 16A–D</figureCitation>
); frontal filaments small, inserted anteriorly (
<figureCitation id="1B892A02FFCC061EF9E7EA6F7F455CD3" box="[455,583,629,651]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig. 16A–C</figureCitation>
); median notch between prostomial lobes absent; eyes absent. Median antenna present, lateral antennae absent; ceratophore of median antenna cylindrical, short (shorter than distance between ceratophore insertion and posterior margin of prostomium), inserted anteriormedially on prostomium; style missing in 
<typeStatus id="5C098825FFCC061EF9D8EB377F625D1B" box="[504,608,813,835]" pageId="54" pageNumber="609" type="holotype">holotype</typeStatus>
; in 
<typeStatus id="5C098825FFCC061EFAAEEB377FFA5D1A" box="[654,760,813,834]" pageId="54" pageNumber="609" type="paratype">paratype</typeStatus>
(MNHN-IA-TYPE 1833) style smooth, tapering into thin tips, long (reaching segment 5). Palps smooth, tapering, long (reaching segment 4–5), inserted ventrally, directed ventroposteriorly (
<figureCitation id="1B892A02FFCC061EFA40EBBD7FEB5DE5" box="[608,745,935,957]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig. 16C, D</figureCitation>
). Pharynx not everted in 
<typeStatus id="5C098825FFCC061EF9BBEBDC7CFC5D84" box="[411,510,966,988]" pageId="54" pageNumber="609" type="holotype">holotype</typeStatus>
; dissected in 
<typeStatus id="5C098825FFCC061EFAB3EBDD7FFB5D84" box="[659,761,967,988]" pageId="54" pageNumber="609" type="paratype">paratype</typeStatus>
(MNHN-IA-TYPE 1834), with pharyngeal papillae not possible to count; two pairs of jaws, dorsal jaw fused, each dorsal jaw with two main blunt teeth, innermost one with a smaller, pointed secondary tooth, margin smooth (
<figureCitation id="1B892A02FFCC061EF8D9EC457C5C5A2D" box="[249,350,1119,1141]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig. 16E</figureCitation>
); each ventral jaw with one main blunt tooth, outer margin serrated (four to five small teeth), with two smaller secondary teeth, pointed on each side, margin smooth and short (
<figureCitation id="1B892A02FFCC061EFA12ECA17F8D5A89" box="[562,655,1211,1233]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig. 16F</figureCitation>
). Bilobed facial tubercle, very small, located between palps and upper lip (
<figureCitation id="1B892A02FFCC061EF928ECE37C6B5B57" box="[264,361,1273,1295]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig. 16D</figureCitation>
). In most specimens, bilobed facial tubercle difficult to see but made visible using forceps to move palps apart.
</paragraph>
<paragraph id="830D3687FFCC061EF889ED4F7CE958E3" blockId="54.[145,762,322,1877]" pageId="54" pageNumber="609">
Tentacular segment very reduced, fused to prostomium, with very short lobe, inserted ventrolaterally to prostomium;withoutaciculaorchaetae;tentaculophores small but distinct, dorsal tentaculophores slightly smaller than ventral tentaculophores; styles missing in 
<typeStatus id="5C098825FFCC061EF8B1EDF47DF4585C" box="[145,246,1518,1540]" pageId="54" pageNumber="609" type="holotype">holotype</typeStatus>
; in 
<typeStatus id="5C098825FFCC061EF901EDF47C8B585B" box="[289,393,1518,1539]" pageId="54" pageNumber="609" type="paratype">paratype</typeStatus>
(MNHN-IA-TYPE 1833), styles smooth, tapering into thin tips, short; dorsal tentacular style shorter than ventral tentacular style (reaching segment 4). Second segment with elytrophores, subbiramous parapodia, with chaetae and ventral cirri. Segments 3 and 4 medioventrally inflated, forming a truncated structure (
<figureCitation id="1B892A02FFCC061EF95DEEBC7CDE58E4" box="[381,476,1702,1724]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig. 16C</figureCitation>
).
</paragraph>
<paragraph id="830D3687FFCC061EF889EEDF7CF35940" blockId="54.[145,762,322,1877]" pageId="54" pageNumber="609">Eight pairs of knob-like elytrophores present on segments 2, 4, 5, 7, 9, 11, 13 and 15 (all elytra missing); on segment 2, elytrophores larger.</paragraph>
<paragraph id="830D3687FFCC061EF889EF3B796F5F6F" blockId="54.[145,762,322,1877]" lastBlockId="54.[809,1426,197,1016]" pageId="54" pageNumber="609">
Cirrigerous segments with large, cylindrical, elongate dorsal cirrophores (
<figureCitation id="1B892A02FFCC061EF9CEEF257F56590D" box="[494,596,1855,1877]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig. 16G</figureCitation>
); dorsodistal, pointed projection (
<figureCitation id="1B892A02FFCC061EFC35E8DF79795E83" box="[1045,1147,197,219]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig. 16G</figureCitation>
) on cirrophores; styles smooth, tapering into long thin tips, long (much longer than tip of neuroacicular lobe), mostly directed posteriorly; dorsal tubercle absent.
</paragraph>
<paragraph id="830D3687FFCC061EFB61E925798A5C13" blockId="54.[809,1426,197,1016]" pageId="54" pageNumber="609">
Ventral cirri smooth, tapering, present from segment 2 to last segment; inserted basally on neuropodia of segment 2, style missing in 
<typeStatus id="5C098825FFCC061EFCADE96779F05FCB" box="[1165,1266,381,403]" pageId="54" pageNumber="609" type="holotype">holotype</typeStatus>
; in 
<typeStatus id="5C098825FFCC061EFD3DE96778935FCA" box="[1309,1425,381,402]" pageId="54" pageNumber="609" type="paratype">paratypes</typeStatus>
(MNHN-IA-TYPE 1833 and MNHN-IA-TYPE 1835), style short (shorter than tip of neuroacicular lobe); in subsequent segments inserted distally on neuropodia, styles short on segments 3 and 4 (shorter than tip of neuroacicular lobe) and long from segment 5 (longer than tip of neuroacicular lobe).
</paragraph>
<paragraph id="830D3687FFCC061EFB61EA4979805D25" blockId="54.[809,1426,197,1016]" pageId="54" pageNumber="609">
Parapodia subbiramous, notopodia very reduced, much shorter than neuropodia(
<figureCitation id="1B892A02FFCC061EFC5DEA6879D55CD0" box="[1149,1239,626,648]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig.16G</figureCitation>
).Notopodia with inconspicuous to very short acicular lobe, notoacicula slender, tip of notoacicula not penetrating epidermis. Neuropodia large, subtriangular, tapering into long acicular lobe, tip of neuroacicula not penetrating epidermis. Notochaetae absent. Neurochaetae moderate in number (nine to 22 observed), short to long, very narrow, distally with spines along both margins, with pointed tips (
<figureCitation id="1B892A02FFCC061EFBD7EB7D79715D25" box="[1015,1139,871,893]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig. 16H–J</figureCitation>
).
</paragraph>
<paragraph id="830D3687FFCC061EFB61EB9C78905DA0" blockId="54.[809,1426,197,1016]" pageId="54" pageNumber="609">
Nephridial papillae present on segments 7–14, small, bulbous. Pygidium rounded, with minute papilla ventrally; slightly enclosed by last segment; with terminal anus (
<figureCitation id="1B892A02FFCC061EFBF6EBF879315DA0" box="[982,1075,994,1016]" captionStart="Figure 16" captionStartId="55.[163,241,1682,1704]" captionTargetBox="[163,1443,195,1641]" captionTargetId="figure-56@55.[702,973,768,960]" captionTargetPageId="55" captionText="Figure 16. Macellicephaloides moustachu sp. nov., holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection." pageId="54" pageNumber="609">Fig. 16A</figureCitation>
). Anal cirri lost, scars not seen.
</paragraph>
<paragraph id="830D3687FFCC061EFB09EC3B7E835B74" blockId="54.[809,1425,1057,1324]" pageId="54" pageNumber="609">
<emphasis id="B1C6EA95FFCC061EFB09EC3B79615A6E" box="[809,1123,1057,1078]" italics="true" pageId="54" pageNumber="609">Morphological variation:</emphasis>
The specimens vary in the number of segments from 11 (MNHN-IA-PNT 81) to 16 (MNHN-IA-PNT 79) but share many of the morphological characters described above except for the number of pairs of elytra and the presence of nephridial papillae. The specimen with 11 segments presents five pairs of elytrophores and no nephridial papillae; these characters are probably linked to growth.
</paragraph>
<paragraph id="830D3687FFCC061EFB09ED4F799B5BBD" blockId="54.[809,1425,1365,1509]" pageId="54" pageNumber="609">
<emphasis id="B1C6EA95FFCC061EFB09ED4F79CC5B32" box="[809,1230,1365,1386]" italics="true" pageId="54" pageNumber="609">Methyl Green staining pattern:</emphasis>
The 
<typeStatus id="5C098825FFCC061EFD3CED4F788C5B32" box="[1308,1422,1365,1386]" pageId="54" pageNumber="609" type="paratype">paratype</typeStatus>
(NHMUK 2018.25351) stains slightly but uniformly all along the body (not shown). Two densely stained spots are observed anteriorly on prostomium, located close to the median ceratophore.
</paragraph>
<paragraph id="830D3687FFCC0610FB09EE147F5A5F2C" blockId="54.[809,1425,1550,1879]" lastBlockId="56.[145,761,197,372]" lastPageId="56" lastPageNumber="611" pageId="54" pageNumber="609">
<emphasis id="B1C6EA95FFCC061EFB09EE147EA0587B" box="[809,930,1550,1571]" italics="true" pageId="54" pageNumber="609">Remarks:</emphasis>
Among the eight species described in 
<taxonomicName id="44B24D04FFCC061EFB79EE377971581A" authorityName="Uschakov" authorityYear="1955" box="[857,1139,1581,1602]" class="Polychaeta" family="Polynoidae" genus="Macellicephaloides" kingdom="Animalia" order="Phyllodocida" pageId="54" pageNumber="609" phylum="Annelida" rank="genus">
<emphasis id="B1C6EA95FFCC061EFB79EE377971581A" box="[857,1139,1581,1602]" italics="true" pageId="54" pageNumber="609">Macellicephaloides</emphasis>
</taxonomicName>
only one species, 
<taxonomicName id="44B24D04FFCC061EFB09EE5178895838" authority="Levenstein, 1983" authorityName="Levenstein" authorityYear="1983" box="[809,1419,1611,1633]" class="Polychaeta" family="Polynoidae" genus="Macellicephaloides" kingdom="Animalia" order="Phyllodocida" pageId="54" pageNumber="609" phylum="Annelida" rank="species" species="improvisa">
<emphasis id="B1C6EA95FFCC061EFB09EE5179A15839" box="[809,1187,1611,1633]" italics="true" pageId="54" pageNumber="609">Macellicephaloides improvisa</emphasis>
<bibRefCitation id="E7234B76FFCC061EFC8FEE5178895838" author="Levenstein RYA" box="[1199,1419,1611,1633]" pageId="54" pageNumber="609" pagination="1419 - 1421" refId="ref54816" refString="Levenstein RYA. 1983. Macellicephaloides improvisa sp. n. (Polychaeta, Polynoidae) from the Kuril-Kamchatka Trough. Zoologicheskii Zhurnal 62: 1419 - 1421." type="journal article" year="1983">Levenstein, 1983</bibRefCitation>
</taxonomicName>
, shows notopodia shorter than neuropodia, with thin notoacicula (
<bibRefCitation id="E7234B76FFCC061EFBE9EE93799B58C7" author="Pettibone MH" box="[969,1177,1673,1695]" pageId="54" pageNumber="609" pagination="154 - 168" refId="ref56575" refString="Pettibone MH. 1989 b. Polynoidae and Sigalionidae (Polychaeta) from the Guaymas Basin, with descriptions of two new species, and additional records from hydrothermal vents of the Galapagos Rift, 21 ° N, and seep-sites in the Gulf of Mexico (Florida and Louisiana). Proceedings of the Biological Society of Washington 102: 154 - 168." type="journal article" year="1989">Pettibone, 1989b</bibRefCitation>
). However, the new species 
<taxonomicName id="44B24D04FFCC061EFBA2EEBD79E158E4" authority="Bonifácio &amp; Menot, 2019" authorityName="Bonifácio &amp; Menot" authorityYear="2019" box="[898,1251,1703,1724]" class="Polychaeta" family="Polynoidae" genus="Macellicephaloides" kingdom="Animalia" order="Phyllodocida" pageId="54" pageNumber="609" phylum="Annelida" rank="species" species="moustachu" status="sp. nov.">
<emphasis id="B1C6EA95FFCC061EFBA2EEBD79E158E4" box="[898,1251,1703,1724]" italics="true" pageId="54" pageNumber="609">Macellicephaloides moustachu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAF557EEFFCC061EFCCBEEB2783D58E5" box="[1259,1343,1704,1725]" pageId="54" pageNumber="609" rank="species">sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="44B24D04FFCC061EFB4AEEDC79CF5884" authorityName="Levenstein" authorityYear="1983" box="[874,1229,1734,1756]" class="Polychaeta" family="Polynoidae" genus="Macellicephaloides" kingdom="Animalia" order="Phyllodocida" pageId="54" pageNumber="609" phylum="Annelida" rank="species" species="improvisa">
<emphasis id="B1C6EA95FFCC061EFB4AEEDC79CF5884" box="[874,1229,1734,1756]" italics="true" pageId="54" pageNumber="609">Macellicephaloides improvisa</emphasis>
</taxonomicName>
in having up to 16 segments and elongate cirrophores with a distal pointed projection, whereas 
<taxonomicName id="44B24D04FFCC061EFCBEEF197EA15960" authorityName="Levenstein" authorityYear="1983" class="Polychaeta" family="Polynoidae" genus="Macellicephaloides" kingdom="Animalia" order="Phyllodocida" pageId="54" pageNumber="609" phylum="Annelida" rank="species" species="improvisa">
<emphasis id="B1C6EA95FFCC061EFCBEEF197EA15960" italics="true" pageId="54" pageNumber="609">Macellicephaloides improvisa</emphasis>
</taxonomicName>
has 18 segments and short cirrophores without a distal pointed projection. 
<taxonomicName id="44B24D04FFCC0610FC98EF5B7C125E82" authority="Bonifácio &amp; Menot, 2019" authorityName="Bonifácio &amp; Menot" authorityYear="2019" class="Polychaeta" family="Polynoidae" genus="Macellicephaloides" kingdom="Animalia" lastPageId="56" lastPageNumber="611" order="Phyllodocida" pageId="54" pageNumber="609" phylum="Annelida" rank="species" species="moustachu" status="sp. nov.">
<emphasis id="B1C6EA95FFCC0610FC98EF5B7C125E82" italics="true" lastPageId="56" lastPageNumber="611" pageId="54" pageNumber="609">Macellicephaloides moustachu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAF557EEFFC20610F938E8DC7C6F5E83" box="[280,365,198,219]" pageId="56" pageNumber="611" rank="species">sp. nov.</taxonomicNameLabel>
is a relatively common species in the CCFZ, with&gt; 
<specimenCount id="95B4FD0EFFC20610F97EE8FE7CF45EA2" box="[350,502,228,250]" pageId="56" pageNumber="611" type="generic">38 specimens</specimenCount>
sampled in four out of the five areas. Unfortunately, only the 18S gene could be sequenced; DNA amplification of the 16S and 
<emphasis id="B1C6EA95FFC20610FAE9E93B7FFA5F6E" box="[713,760,289,310]" italics="true" pageId="56" pageNumber="611">COI</emphasis>
genes was unsuccessful. Consequently, identification of damaged specimens was challenging.
</paragraph>
<caption id="D7CD660FFFCD061FF883EE887913592E" pageId="55" pageNumber="610" startId="55.[163,241,1682,1704]" targetBox="[163,1443,195,1641]" targetPageId="55">
<paragraph id="830D3687FFCD061FF883EE887913592E" blockId="55.[163,1443,1682,1910]" pageId="55" pageNumber="610">
<emphasis id="B1C6EA95FFCD061FF883EE887FE158F0" bold="true" box="[163,739,1682,1704]" pageId="55" pageNumber="610">
Figure 16. 
<taxonomicName id="44B24D04FFCD061FF902EE887F8A58F0" authority="Bonifácio &amp; Menot, 2019" authorityName="Bonifácio &amp; Menot" authorityYear="2019" box="[290,648,1682,1704]" class="Polychaeta" family="Polynoidae" genus="Macellicephaloides" kingdom="Animalia" order="Phyllodocida" pageId="55" pageNumber="610" phylum="Annelida" rank="species" species="moustachu" status="sp. nov.">
<emphasis id="B1C6EA95FFCD061FF902EE887F8A58F0" bold="true" box="[290,648,1682,1704]" italics="true" pageId="55" pageNumber="610">Macellicephaloides moustachu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAF557EEFFCD061FFAAEEE897FE158F0" box="[654,739,1683,1704]" pageId="55" pageNumber="610" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype MNHN-IA-TYPE 1832 (A–C, G–I), paratype 2 MNHN-IA- TYPE 1834 (D–F) and paratype 3 MNHN-IA-TYPE 1835 (J). A, dorsal view of a preserved complete specimen. B, anterior end, dorsal view, chaetae omitted. C, anterior end, ventral view, chaetae omitted. D, anterior end, ventral view, chaetae omitted. E, inner view of dorsal side of jaws from dissected pharynx, papillae omitted. F, inner view of ventral side of jaws from dissected pharynx, papillae omitted. G, right parapodia, posterior view (ventral cirri lost), segment 6. H, neurochaeta with spines along both margins, frontal view, segment 6. I, neurochaeta with spines along both margins, frontolateral view, segment 6. J, scanning electron micrograph of neurochaeta; up side is the distal part of chaeta. Abbreviations: fil, frontal filament; ft, frontal tubercles; inf, inflated truncate structure; pp, pointed projection.
</paragraph>
</caption>
<paragraph id="830D3687FFC20610F8B1E9877F025C57" blockId="56.[145,761,413,527]" pageId="56" pageNumber="611">
<emphasis id="B1C6EA95FFC20610F8B1E9877C105FEA" box="[145,274,413,434]" italics="true" pageId="56" pageNumber="611">Etymology:</emphasis>
The species name came from the French word ‘ 
<taxonomicName id="44B24D04FFC20610F8F3E9A67C4D5F89" authorityName="Bonifácio &amp; Menot" authorityYear="2019" box="[211,335,444,465]" class="Polychaeta" family="Polynoidae" genus="Macellicephaloides" kingdom="Animalia" order="Phyllodocida" pageId="56" pageNumber="611" phylum="Annelida" rank="species" species="moustachu">
<emphasis id="B1C6EA95FFC20610F8F3E9A67C4D5F89" box="[211,335,444,465]" italics="true" pageId="56" pageNumber="611">moustachu</emphasis>
</taxonomicName>
’ meaning ‘with a moustache’. It refers to the palps directed ventrally, giving the impression that the worm has a moustache.
</paragraph>
<paragraph id="830D3687FFC20610F8B1EA227FBD5CD3" blockId="56.[145,761,568,651]" pageId="56" pageNumber="611">
<emphasis id="B1C6EA95FFC20610F8B1EA227C325C15" box="[145,304,568,589]" italics="true" pageId="56" pageNumber="611">Genetic data:</emphasis>
DNA sequencing for this species was successful only for 18S but not for 
<emphasis id="B1C6EA95FFC20610FA37EA4D7F445C34" box="[535,582,599,620]" italics="true" pageId="56" pageNumber="611">COI</emphasis>
or 16S, sharing 100% of genetic material between the specimens.
</paragraph>
<paragraph id="830D3687FFC20610F8B1EAAE7FF85D7D" blockId="56.[145,762,692,806]" pageId="56" pageNumber="611">
<emphasis id="B1C6EA95FFC20610F8B1EAAE7C2B5C91" box="[145,297,692,713]" italics="true" pageId="56" pageNumber="611">Distribution:</emphasis>
Based on the material examined (
<specimenCount id="95B4FD0EFFC20610FAF2EAAE7C0A5CB0" pageId="56" pageNumber="611" type="generic">ten specimens</specimenCount>
), this species has a wide distribution within Clarion-Clipperton Fracture Zone, being sampled in BGR, IOM (
<typeStatus id="5C098825FFC20610F930EB0A7C435D7D" box="[272,321,784,805]" pageId="56" pageNumber="611">type</typeStatus>
locality), GSR and Ifremer license areas.
</paragraph>
<paragraph id="830D3687FFC20610F8B1EB547C755A63" blockId="56.[145,761,846,1083]" pageId="56" pageNumber="611">
<emphasis id="B1C6EA95FFC20610F8B1EB547C715D3C" box="[145,371,846,868]" italics="true" pageId="56" pageNumber="611">Reproductive notes:</emphasis>
<typeStatus id="5C098825FFC20610F95FEB557CEB5D3C" box="[383,489,847,868]" pageId="56" pageNumber="611" type="paratype">Paratype</typeStatus>
MNHN-IA-TYPE 1833 seems to be brooding dorsally; two large interconnected pouches were observed linked to the body through an anterior small hole on the notopodia of segment 10. The sac directly connected to the body is surrounded by a membrane and has visible small ova inside; the second sac shows an agglomerate of small ova without a visible membrane.
</paragraph>
</subSubSection>
</treatment>
</document>