547 lines
75 KiB
XML
547 lines
75 KiB
XML
<document id="5924FCE1D07C1830558BE9695C7F70D1" ID-ISSN="0024-4082" ID-ZooBank="74C07292-2BD6-4E3E-B68D-B144B81BBD83" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629585783401" checkinUser="felipe" docAuthor="Bonifácio, Paulo & Menot, Lénaïck" docDate="2019" docId="0B1B8791FFCB061CFB5BEA6679585BF5" docLanguage="en" docName="ZoolJLinnSoc.185.555–635.pdf" docOrigin="Zoological Journal of the Linnean Society 185" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Macellicephala parvafauces Bonifácio & Menot 2019, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="607" masterDocId="F722FFE9FFFA0628F820E81A7D025E58" masterDocTitle="New genera and species from the Equatorial Pacific provide phylogenetic insights into deep-sea Polynoidae (Annelida)" masterLastPageNumber="635" masterPageNumber="555" pageNumber="604" updateTime="1699081480826" updateUser="plazi">
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<mods:titleInfo id="332274F0F37DD2A1056C9E773A384658">
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<mods:title id="5E8E68A0B8068F267AE84D3B653592BE">New genera and species from the Equatorial Pacific provide phylogenetic insights into deep-sea Polynoidae (Annelida)</mods:title>
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<mods:namePart id="EB495491378B47B9B423863B5C6C990D">Bonifácio, Paulo</mods:namePart>
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<mods:roleTerm id="1A77AB70744B7EA69587776E25F8AB27">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="340432A013C5731524EC734BA30A64D3">Menot, Lénaïck</mods:namePart>
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<mods:typeOfResource id="EB2F137B103A2925BA7CD4F5244701B2">text</mods:typeOfResource>
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<mods:title id="B6E055C43CBB53981A0839F56A228B3E">Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part id="EE2B225CA2F7591DD5C9AC02B7E2A9FA">
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<mods:date id="6AF1C4D95A8B8DB2BCC2DC1E2D498CE9">2019</mods:date>
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<mods:detail id="80232E47305A0398DD0275AD19EC2001" type="volume">
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<mods:number id="14A0F467C9A47D0F53BF4C11C4697AF3">185</mods:number>
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<mods:classification id="2A37042BD6808B4AEE02FDAA8366C9BC">journal article</mods:classification>
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<mods:identifier id="8F26D38DAC8C88CE9627D64C3517EE00" type="ISSN">0024-4082</mods:identifier>
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</mods:mods>
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<treatment id="0B1B8791FFCB061CFB5BEA6679585BF5" LSID="urn:lsid:plazi:treatment:0B1B8791FFCB061CFB5BEA6679585BF5" httpUri="http://treatment.plazi.org/id/0B1B8791FFCB061CFB5BEA6679585BF5" lastPageId="52" lastPageNumber="607" pageId="49" pageNumber="604">
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<subSubSection id="CBA8650CFFCB0619FB5BEA6678615CCB" box="[891,1379,635,659]" pageId="49" pageNumber="604" type="nomenclature">
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<paragraph id="830D3687FFCB0619FB5BEA6678615CCB" blockId="49.[891,1379,635,659]" box="[891,1379,635,659]" pageId="49" pageNumber="604">
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<heading id="D84581EBFFCB0619FB5BEA6678615CCB" box="[891,1379,635,659]" centered="true" fontSize="9" level="2" pageId="49" pageNumber="604" reason="2">
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<taxonomicName id="44B24D04FFCB0619FB5BEA6679FA5CCA" ID-CoL="3WXTX" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[891,1272,636,659]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="49" pageNumber="628" phylum="Annelida" rank="species" species="parvafauces" status="sp. nov.">MACELLICEPHALA PARVAFAUCES</taxonomicName>
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<emphasis id="B1C6EA95FFCB0619FD20EA6578615CCB" bold="true" box="[1280,1379,635,659]" pageId="49" pageNumber="604">
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<taxonomicNameLabel id="AAF557EEFFCB0619FD20EA6578615CCB" box="[1280,1379,635,659]" pageId="49" pageNumber="604" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CBA8650CFFCB061CFBF0EAB879585BF5" lastPageId="52" lastPageNumber="607" pageId="49" pageNumber="604" type="description">
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<paragraph id="830D3687FFCB0619FBF0EAB8780F5CE3" blockId="49.[976,1293,674,699]" box="[976,1293,674,699]" pageId="49" pageNumber="604">
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(
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<figureCitation id="1B892A02FFCB0619FBF9EAB879BB5CE2" box="[985,1209,674,699]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="49" pageNumber="604">FIG. 15A–I; TABLES</figureCitation>
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1, 2, 4)
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||
</paragraph>
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||
<paragraph id="830D3687FFCB0619FB1BEAF178325AD7" blockId="49.[827,1446,746,1167]" pageId="49" pageNumber="604">
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<emphasis id="B1C6EA95FFCB0619FB1BEAF17EF75CA7" box="[827,1013,746,768]" italics="true" pageId="49" pageNumber="604">Type material:</emphasis>
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<materialsCitation id="33DA3CDAFFCB0619FC23EAF07E9D5D8F" collectingDate="2015-04-20" collectingDateMax="2015-04-21" collectingDateMin="2015-04-20" collectionCode="IFR, APEI" county="Kiel" elevation="4949" latitude="18.802166" location="Equatorial Eastern Pacific Ocean" longLatPrecision="13" longitude="-128.30333" municipality="Clarion-Clipperton Fracture Zone" pageId="49" pageNumber="604" specimenCode="IFR602, APEI#3" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="5C098825FFCB0619FC23EAF079745D58" box="[1027,1142,746,768]" pageId="49" pageNumber="604" type="holotype">Holotype</typeStatus>
|
||
, MNHN-IA-TYPE 1830 (
|
||
<specimenCode id="D3149EFCFFCB0619FB62EB137E995D47" box="[834,923,777,799]" collectionCode="IFR" pageId="49" pageNumber="604">IFR602</specimenCode>
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), complete, length
|
||
<quantity id="444A9B62FFCB0619FC44EB1379D15D47" box="[1124,1235,777,799]" metricMagnitude="-2" metricUnit="m" metricValue="1.182" pageId="49" pageNumber="604" unit="mm" value="11.82">11.82 mm</quantity>
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, width
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<quantity id="444A9B62FFCB0619FD3FEB1378825D47" box="[1311,1408,777,799]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="49" pageNumber="604" unit="mm" value="1.5">1.50 mm</quantity>
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||
, 18 segments (including tentacular segment),
|
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<location id="866D605CFFCB0619FD06EB3279475D04" LSID="urn:lsid:plazi:treatment:0B1B8791FFCB061CFB5BEA6679585BF5:866D605CFFCB0619FD06EB3279475D04" county="Kiel" latitude="18.802166" longLatPrecision="13" longitude="-128.30333" municipality="Clarion-Clipperton Fracture Zone" name="Equatorial Eastern Pacific Ocean" pageId="49" pageNumber="604">
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||
Equatorial
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||
<location id="866D605CFFCB0619FB1BEB5D79475D04" LSID="urn:lsid:plazi:treatment:0B1B8791FFCB061CFB5BEA6679585BF5:866D605CFFCB0619FB1BEB5D79475D04" box="[827,1093,838,860]" county="Kiel" latitude="18.802166" longLatPrecision="13" longitude="-128.30333" municipality="Clarion-Clipperton Fracture Zone" name="Eastern Pacific Ocean" pageId="49" pageNumber="604">Eastern Pacific Ocean</location>
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||
</location>
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,
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||
<collectingMunicipality id="6369ACFDFFCB0619FC71EB5C7E745D23" pageId="49" pageNumber="604">Clarion-Clipperton Fracture Zone</collectingMunicipality>
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||
,
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||
<specimenCode id="D3149EFCFFCB0619FBA2EB7F7EDD5D22" box="[898,991,869,890]" collectionCode="APEI" country="Canada" httpUri="http://grbio.org/cool/pg9t-riai" name="Agriculture Canada Research Station" pageId="49" pageNumber="604">APEI#3</specimenCode>
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||
nodules, station 189, collected
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||
<date id="F70C1047FFCB0619FD7CEB7F7EBB5DC2" pageId="49" pageNumber="604" value="2015-04-20" valueMax="2015-04-21" valueMin="2015-04-20">
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<collectingDate id="E748E9AFFFCB0619FD7CEB7F7EBB5DC2" pageId="49" pageNumber="604" value="2015-04-20" valueMax="2015-04-21" valueMin="2015-04-20">20–21 April 2015</collectingDate>
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||
</date>
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||
, ROV
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||
<collectingCounty id="6A6C4E0BFFCB0619FC24EB9E79365DC2" box="[1028,1076,900,922]" pageId="49" pageNumber="604">Kiel</collectingCounty>
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6000, biobox, start
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||
<geoCoordinate id="E6865040FFCB0619FD3DEB9E78A25DC1" box="[1309,1440,899,922]" degrees="18" direction="north" minutes="47.80" orientation="latitude" pageId="49" pageNumber="604" precision="9" value="18.796667">18°47.80′N</geoCoordinate>
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,
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<geoCoordinate id="E6865040FFCB0619FB1BEBB87EC85DE0" box="[827,970,929,952]" degrees="128" direction="west" minutes="18.53" orientation="longitude" pageId="49" pageNumber="604" precision="9" value="-128.30884">128°18.53′W</geoCoordinate>
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, end
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<geoCoordinate id="E6865040FFCB0619FC25EBB979815DE0" box="[1029,1155,929,952]" degrees="18" direction="north" minutes="48.13" orientation="latitude" pageId="49" pageNumber="604" precision="9" value="18.802166">18°48.13′N</geoCoordinate>
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,
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<geoCoordinate id="E6865040FFCB0619FCAEEBB9781F5DE0" box="[1166,1309,929,952]" degrees="128" direction="west" minutes="18.20" orientation="longitude" pageId="49" pageNumber="604" precision="9" value="-128.30333">128°18.20′W</geoCoordinate>
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,
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<elevation id="089FD1B4FFCB0619FD07EBB87E9D5D8F" metricMagnitude="3" metricUnit="m" metricValue="4.9485" metricValueMax="4.964" metricValueMin="4.933" pageId="49" pageNumber="604" unit="m" value="4948.5" valueMax="4964.0" valueMin="4933.0">
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<quantity id="444A9B62FFCB0619FD07EBB87E535D8F" metricMagnitude="3" metricUnit="m" metricValue="4.9485" metricValueMax="4.964" metricValueMin="4.933" pageId="49" pageNumber="604" unit="m" value="4948.5" valueMax="4964.0" valueMin="4933.0">4933–4964 m</quantity>
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depth
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||
</elevation>
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||
</materialsCitation>
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.
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||
<materialsCitation id="33DA3CDAFFCB0619FB8AEBD878A05A28" collectingDate="2015-04-09" collectingDateMax="2015-04-10" collectingDateMin="2015-04-09" collectionCode="GSR" county="Kiel" latitude="13.873167" location="Equatorial Eastern Pacific Ocean" longLatPrecision="13" longitude="-123.2505" municipality="Clarion-Clipperton Fracture Zone" pageId="49" pageNumber="604" specimenCode="IFR403" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="5C098825FFCB0619FB8AEBD879145D8F" box="[938,1046,962,983]" pageId="49" pageNumber="604" type="paratype">Paratype</typeStatus>
|
||
, MNHN-IA-TYPE 1831 (
|
||
<specimenCode id="D3149EFCFFCB0619FD1CEBDB78955D8F" box="[1340,1431,961,983]" collectionCode="IFR" pageId="49" pageNumber="604">IFR403</specimenCode>
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), complete, length
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<quantity id="444A9B62FFCB0619FBD9EBFA795E5DAD" box="[1017,1116,992,1013]" metricMagnitude="-3" metricUnit="m" metricValue="8.02" pageId="49" pageNumber="604" unit="mm" value="8.02">8.02 mm</quantity>
|
||
, width
|
||
<quantity id="444A9B62FFCB0619FC8BEBFA780C5DAD" box="[1195,1294,992,1014]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="49" pageNumber="604" unit="mm" value="1.05">1.05 mm</quantity>
|
||
, 18 segments (including tentacular segment),
|
||
<location id="866D605CFFCB0619FC9CEBE47ED45A6A" LSID="urn:lsid:plazi:treatment:0B1B8791FFCB061CFB5BEA6679585BF5:866D605CFFCB0619FC9CEBE47ED45A6A" county="Kiel" latitude="13.873167" longLatPrecision="13" longitude="-123.2505" municipality="Clarion-Clipperton Fracture Zone" name="Equatorial Eastern Pacific Ocean" pageId="49" pageNumber="604">
|
||
Equatorial
|
||
<location id="866D605CFFCB0619FD64EBE57ED45A6A" LSID="urn:lsid:plazi:treatment:0B1B8791FFCB061CFB5BEA6679585BF5:866D605CFFCB0619FD64EBE57ED45A6A" county="Kiel" latitude="13.873167" longLatPrecision="13" longitude="-123.2505" municipality="Clarion-Clipperton Fracture Zone" name="Eastern Pacific Ocean" pageId="49" pageNumber="604">Eastern Pacific Ocean</location>
|
||
</location>
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||
,
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||
<collectingMunicipality id="6369ACFDFFCB0619FBC1EC0778615A6B" box="[993,1379,1053,1075]" pageId="49" pageNumber="604">Clarion-Clipperton Fracture Zone</collectingMunicipality>
|
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,
|
||
<collectionCode id="E5A3AE42FFCB0619FD4EEC0778A15A6A" box="[1390,1443,1053,1074]" pageId="49" pageNumber="604">GSR</collectionCode>
|
||
license area, station 131, collected
|
||
<date id="F70C1047FFCB0619FCFEEC26789C5A0A" box="[1246,1438,1084,1106]" pageId="49" pageNumber="604" value="2015-04-09" valueMax="2015-04-10" valueMin="2015-04-09">
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<collectingDate id="E748E9AFFFCB0619FCFEEC26789C5A0A" box="[1246,1438,1084,1106]" pageId="49" pageNumber="604" value="2015-04-09" valueMax="2015-04-10" valueMin="2015-04-09">9–10 April 2015</collectingDate>
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</date>
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, ROV
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<collectingCounty id="6A6C4E0BFFCB0619FB58EC407EA55A28" box="[888,935,1114,1136]" pageId="49" pageNumber="604">Kiel</collectingCounty>
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6000, biobox, start
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,
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<geoCoordinate id="E6865040FFCB0619FD31EC4078A05A28" box="[1297,1442,1113,1136]" degrees="123" direction="west" minutes="15.03" orientation="longitude" pageId="49" pageNumber="604" precision="9" value="-123.2505">123°15.03′W</geoCoordinate>
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, end
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<geoCoordinate id="E6865040FFCB0619FB4BEC637EEB5AD7" box="[875,1001,1144,1167]" degrees="13" direction="north" minutes="52.44" orientation="latitude" pageId="49" pageNumber="604" precision="9" value="13.874">13°52.44′N</geoCoordinate>
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,
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<geoCoordinate id="E6865040FFCB0619FBD4EC6379815AD7" box="[1012,1155,1144,1167]" degrees="123" direction="west" minutes="14.88" orientation="longitude" pageId="49" pageNumber="604" precision="9" value="-123.248">123°14.88′W</geoCoordinate>
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,
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<quantity id="444A9B62FFCB0619FCADEC6379E35AD7" box="[1165,1249,1145,1167]" metricMagnitude="3" metricUnit="m" metricValue="4.478" pageId="49" pageNumber="604" unit="m" value="4478.0">4478 m</quantity>
|
||
depth.
|
||
</paragraph>
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<paragraph id="830D3687FFCB0619FB1BECA27EEC5BDE" blockId="49.[827,1444,1208,1874]" pageId="49" pageNumber="604">
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<emphasis id="B1C6EA95FFCB0619FB1BECA278A65A95" box="[827,1444,1208,1229]" italics="true" pageId="49" pageNumber="604">
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Description (based on
|
||
<typeStatus id="5C098825FFCB0619FC49ECA279D65A95" box="[1129,1236,1208,1229]" pageId="49" pageNumber="604" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="5C098825FFCB0619FD00ECA278965A95" box="[1312,1428,1208,1229]" pageId="49" pageNumber="604" type="paratype">paratype</typeStatus>
|
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):
|
||
</emphasis>
|
||
<typeStatus id="5C098825FFCB0619FB1BECCC7EA05AB4" box="[827,930,1238,1260]" pageId="49" pageNumber="604" type="holotype">Holotype</typeStatus>
|
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complete,
|
||
<quantity id="444A9B62FFCB0619FC3DECCD798D5AB4" box="[1053,1167,1239,1260]" metricMagnitude="-2" metricUnit="m" metricValue="1.182" pageId="49" pageNumber="604" unit="mm" value="11.82">11.82 mm</quantity>
|
||
long and
|
||
<quantity id="444A9B62FFCB0619FD22ECCC78675AB4" box="[1282,1381,1238,1260]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="49" pageNumber="604" unit="mm" value="1.5">1.50 mm</quantity>
|
||
wide for 18 segments, slightly dorsoventrally flattened, not tapering posteriorly; live specimen whitish anteriorly and slightly brownish posteriorly; ethanol-preserved specimen pale yellow throughout (
|
||
<figureCitation id="1B892A02FFCB0619FCEFED4B78535B3E" box="[1231,1361,1361,1383]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="49" pageNumber="604">Fig. 15A, C</figureCitation>
|
||
). Body surface smooth.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFCB0619FB73ED94783258AD" blockId="49.[827,1444,1208,1874]" pageId="49" pageNumber="604">
|
||
Prostomium bilobed, about as wide as long, lobes pronounced anteriorly, truncated; small, oval frontal filaments present, inserted at innermost margin of prostomial lobes; median notch between prostomial lobes moderately wide and deep (
|
||
<figureCitation id="1B892A02FFCB0619FCE7EE1378295847" box="[1223,1323,1545,1567]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="49" pageNumber="604">Fig. 15D</figureCitation>
|
||
); eyes absent. Median antenna present, lateral antennae absent; ceratophore of median antenna large, cylindrical, short (shorter than anterior margin of prostomial lobes), inserted near median notch, style missing. Palps smooth, tapering, short (reaching segment 4;
|
||
<figureCitation id="1B892A02FFCB0619FB1BEEDB7E9E588F" box="[827,924,1729,1751]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="49" pageNumber="604">Fig. 15D</figureCitation>
|
||
). Trilobed facial tubercle present, median tubercle larger than lateral ones (
|
||
<figureCitation id="1B892A02FFCB0619FC82EEFA782258AD" box="[1186,1312,1760,1782]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="49" pageNumber="604">Fig. 15B, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="830D3687FFCB0619FB73EEE478A0590A" blockId="49.[827,1444,1208,1874]" pageId="49" pageNumber="604">Tentacular segment with short lobes, inserted laterally and slightly ventral to prostomium; without acicula or chaetae; tentaculophores large, cylindrical,</paragraph>
|
||
<caption id="D7CD660FFFC8061AF8AAE9837ECF5FF7" box="[138,973,409,431]" pageId="50" pageNumber="605" startId="50.[138,202,409,431]">
|
||
<paragraph id="830D3687FFC8061AF8AAE9837ECF5FF7" blockId="50.[138,973,409,431]" box="[138,973,409,431]" pageId="50" pageNumber="605">
|
||
<emphasis id="B1C6EA95FFC8061AF8AAE9837DE65FF7" bold="true" box="[138,228,409,431]" pageId="50" pageNumber="605">Table 4.</emphasis>
|
||
Diagnostic characters for all valid species in the genus
|
||
<taxonomicName id="44B24D04FFC8061AFB0FE9837ECF5FF7" authorityName="McIntosh" authorityYear="1885" box="[815,973,409,431]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="genus">Macellicephala</taxonomicName>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="830D3687FFC8061AF937E9C97A1D5849" pageId="50" pageNumber="605">
|
||
<table id="F1B2C427FFC8F9D7F8AAE9C97A215818" box="[138,1827,467,1600]" gridcols="12" gridrows="12" pageId="50" pageNumber="605">
|
||
<tr id="3D8234C5FFC8F9D7F8AAE9C97A215FA5" box="[138,1827,467,509]" gridrow="0" pageId="50" pageNumber="605" rowspan-0="1">
|
||
<th id="7E535DB9FFC8F9D7F937E9C97C9A5FA5" box="[279,408,467,509]" gridcol="1" gridrow="0" pageId="50" pageNumber="605">
|
||
<taxonomicName id="44B24D04FFC8061AF937E9C97C835FBD" baseAuthorityName="Moore" baseAuthorityYear="1910" box="[279,385,467,485]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="species" species="aciculata">
|
||
<emphasis id="B1C6EA95FFC8061AF937E9C97C835FBD" box="[279,385,467,485]" italics="true" pageId="50" pageNumber="605">M. aciculata</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="7E535DB9FFC8F9D7F986E9C97F2A5FA5" box="[422,552,467,509]" gridcol="2" gridrow="0" pageId="50" pageNumber="605">
|
||
<taxonomicName id="44B24D04FFC8061AF987E9C97F0D5FBD" authorityName="Stop-Bowitz" authorityYear="1948" box="[423,527,467,485]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="species" species="atlantica">
|
||
<emphasis id="B1C6EA95FFC8061AF987E9C97F0D5FBD" box="[423,527,467,485]" italics="true" pageId="50" pageNumber="605">M. atlantica</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="7E535DB9FFC8F9D7FA16E9C97FB05FA5" box="[566,690,467,509]" gridcol="3" gridrow="0" pageId="50" pageNumber="605">
|
||
<taxonomicName id="44B24D04FFC8061AFA16E9C97F9D5FBD" authorityName="Wu & Wang" authorityYear="1987" box="[566,671,467,485]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="species" species="australis">
|
||
<emphasis id="B1C6EA95FFC8061AFA16E9C97F9D5FBD" box="[566,671,467,485]" italics="true" pageId="50" pageNumber="605">M. australis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="7E535DB9FFC8F9D7FAE0E9C97EC15FA5" box="[704,963,467,509]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="50" pageNumber="605">
|
||
<emphasis id="B1C6EA95FFC8061AFAE1E9C97EBB5FBD" box="[705,953,467,485]" italics="true" pageId="50" pageNumber="605">
|
||
<taxonomicName id="44B24D04FFC8061AFAE1E9C97E505FBD" authorityName="Bonifácio & Menot" authorityYear="2019" box="[705,850,467,485]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="species" species="clarionensis">
|
||
<emphasis id="B1C6EA95FFC8061AFAE1E9C97E505FBD" bold="true" box="[705,850,467,485]" italics="true" pageId="50" pageNumber="605">M. clarionensis</emphasis>
|
||
</taxonomicName>
|
||
M. galapa-
|
||
</emphasis>
|
||
<emphasis id="B1C6EA95FFC8061AFAE1E9F17E0A5FA5" bold="true" box="[705,776,491,509]" pageId="50" pageNumber="605">sp. nov.</emphasis>
|
||
<emphasis id="B1C6EA95FFC8061AFB7CE9F17E935FA5" box="[860,913,491,509]" italics="true" pageId="50" pageNumber="605">gensis</emphasis>
|
||
</th>
|
||
<th id="7E535DB9FFC8F9D7FBF8E9C979555FA5" box="[984,1111,467,509]" gridcol="6" gridrow="0" pageId="50" pageNumber="605">
|
||
<taxonomicName id="44B24D04FFC8061AFBF9E9C9793A5FBD" authorityName="Reyss, 1971 Pettibone" authorityYear="1976" box="[985,1080,467,485]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="species" species="laubieri">
|
||
<emphasis id="B1C6EA95FFC8061AFBF9E9C9793A5FBD" box="[985,1080,467,485]" italics="true" pageId="50" pageNumber="605">M. laubieri</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="7E535DB9FFC8F9D7FC48E9C979E95FA5" box="[1128,1259,467,509]" gridcol="7" gridrow="0" pageId="50" pageNumber="605">
|
||
<taxonomicName id="44B24D04FFC8061AFC48E9C979DD5FBD" authorityName="Uschakov" authorityYear="1957" box="[1128,1247,467,485]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="species" species="longipalpa">
|
||
<emphasis id="B1C6EA95FFC8061AFC48E9C979DD5FBD" box="[1128,1247,467,485]" italics="true" pageId="50" pageNumber="605">M. longipalpa</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="7E535DB9FFC8F9D7FCD7E9C978765FA5" box="[1271,1396,467,509]" gridcol="8" gridrow="0" pageId="50" pageNumber="605">
|
||
<taxonomicName id="44B24D04FFC8061AFCD8E9C978605FBD" baseAuthorityName="McIntosh" baseAuthorityYear="1885" box="[1272,1378,467,485]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="species" species="mirabilis">
|
||
<emphasis id="B1C6EA95FFC8061AFCD8E9C978605FBD" box="[1272,1378,467,485]" italics="true" pageId="50" pageNumber="605">M. mirabilis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="7E535DB9FFC8F9D7FDA0E9C97B105FA5" box="[1408,1554,467,509]" gridcol="9" gridrow="0" pageId="50" pageNumber="605">
|
||
<emphasis id="B1C6EA95FFC8061AFDA0E9C978C55FA5" bold="true" pageId="50" pageNumber="605">
|
||
<taxonomicName id="44B24D04FFC8061AFDA0E9C97B105FBD" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[1408,1554,467,485]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="species" species="parvafauces" status="sp. nov.">
|
||
<emphasis id="B1C6EA95FFC8061AFDA0E9C97B105FBD" box="[1408,1554,467,485]" italics="true" pageId="50" pageNumber="605">M. parvafauces</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AAF557EEFFC8061AFDA0E9F178C55FA5" box="[1408,1479,491,509]" pageId="50" pageNumber="605" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<th id="7E535DB9FFC8F9D7FE3FE9C97B995FA5" box="[1567,1691,467,509]" gridcol="10" gridrow="0" pageId="50" pageNumber="605">
|
||
<taxonomicName id="44B24D04FFC8061AFE3FE9C97B855FBD" baseAuthorityName="Moore" baseAuthorityYear="1910" box="[1567,1671,467,485]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="species" species="remigata">
|
||
<emphasis id="B1C6EA95FFC8061AFE3FE9C97B855FBD" box="[1567,1671,467,485]" italics="true" pageId="50" pageNumber="605">M. remigata</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="7E535DB9FFC8F9D7FE86E9C97A215FA5" box="[1702,1827,467,509]" gridcol="11" gridrow="0" pageId="50" pageNumber="605">
|
||
<taxonomicName id="44B24D04FFC8061AFE87E9C97A055FBD" baseAuthorityName="Levinsen" baseAuthorityYear="1886" box="[1703,1799,467,485]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="species" species="violacea">
|
||
<emphasis id="B1C6EA95FFC8061AFE87E9C97A055FBD" box="[1703,1799,467,485]" italics="true" pageId="50" pageNumber="605">M. violacea</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="3D8234C5FFC8F9D7F8AAEA057A215C11" box="[138,1827,543,585]" gridrow="1" pageId="50" pageNumber="605">
|
||
<th id="7E535DB9FFC8F9D7F8AAEA057C0B5C11" box="[138,265,543,585]" gridcol="0" gridrow="1" pageId="50" pageNumber="605">Prostomial shape</th>
|
||
<td id="7E535DB9FFC8F9D7F937EA057C9A5C11" box="[279,408,543,585]" gridcol="1" gridrow="1" pageId="50" pageNumber="605">?</td>
|
||
<td id="7E535DB9FFC8F9D7F986EA057F2A5C11" box="[422,552,543,585]" gridcol="2" gridrow="1" pageId="50" pageNumber="605">Subretangular</td>
|
||
<td id="7E535DB9FFC8F9D7FA16EA057FB05C11" box="[566,690,543,585]" gridcol="3" gridrow="1" pageId="50" pageNumber="605">Oval</td>
|
||
<td id="7E535DB9FFC8F9D7FAE0EA057E525C11" box="[704,848,543,585]" gridcol="4" gridrow="1" pageId="50" pageNumber="605">Subretangular</td>
|
||
<td id="7E535DB9FFC8F9D7FB7CEA057EC15C11" box="[860,963,543,585]" gridcol="5" gridrow="1" pageId="50" pageNumber="605">Oval</td>
|
||
<td id="7E535DB9FFC8F9D7FBF8EA0579555C11" box="[984,1111,543,585]" gridcol="6" gridrow="1" pageId="50" pageNumber="605">Subretangular</td>
|
||
<td id="7E535DB9FFC8F9D7FC48EA0579E95C11" box="[1128,1259,543,585]" gridcol="7" gridrow="1" pageId="50" pageNumber="605">Subretangular</td>
|
||
<td id="7E535DB9FFC8F9D7FCD7EA0578765C11" box="[1271,1396,543,585]" gridcol="8" gridrow="1" pageId="50" pageNumber="605">Oval</td>
|
||
<td id="7E535DB9FFC8F9D7FDA0EA057B105C11" box="[1408,1554,543,585]" gridcol="9" gridrow="1" pageId="50" pageNumber="605">Subretangular</td>
|
||
<td id="7E535DB9FFC8F9D7FE3FEA057B995C11" box="[1567,1691,543,585]" gridcol="10" gridrow="1" pageId="50" pageNumber="605">Oval</td>
|
||
<td id="7E535DB9FFC8F9D7FE86EA057A215C11" box="[1702,1827,543,585]" gridcol="11" gridrow="1" pageId="50" pageNumber="605">Oval</td>
|
||
</tr>
|
||
<tr id="3D8234C5FFC8F9D7F8AAEA487A215C24" box="[138,1827,594,636]" gridrow="2" pageId="50" pageNumber="605">
|
||
<th id="7E535DB9FFC8F9D7F8AAEA487C0B5C24" box="[138,265,594,636]" gridcol="0" gridrow="2" pageId="50" pageNumber="605">Frontal/facial tubercle</th>
|
||
<td id="7E535DB9FFC8F9D7F937EA487C9A5C24" box="[279,408,594,636]" gridcol="1" gridrow="2" pageId="50" pageNumber="605">Absent</td>
|
||
<td id="7E535DB9FFC8F9D7F986EA487F2A5C24" box="[422,552,594,636]" gridcol="2" gridrow="2" pageId="50" pageNumber="605">Absent</td>
|
||
<td id="7E535DB9FFC8F9D7FA16EA487FB05C24" box="[566,690,594,636]" gridcol="3" gridrow="2" pageId="50" pageNumber="605">?</td>
|
||
<td id="7E535DB9FFC8F9D7FAE0EA487E525C24" box="[704,848,594,636]" gridcol="4" gridrow="2" pageId="50" pageNumber="605">Present</td>
|
||
<td id="7E535DB9FFC8F9D7FB7CEA487EC15C24" box="[860,963,594,636]" gridcol="5" gridrow="2" pageId="50" pageNumber="605">Present</td>
|
||
<td id="7E535DB9FFC8F9D7FBF8EA4879555C24" box="[984,1111,594,636]" gridcol="6" gridrow="2" pageId="50" pageNumber="605">Absent</td>
|
||
<td id="7E535DB9FFC8F9D7FC48EA4879E95C24" box="[1128,1259,594,636]" gridcol="7" gridrow="2" pageId="50" pageNumber="605">Absent</td>
|
||
<td id="7E535DB9FFC8F9D7FCD7EA4878765C24" box="[1271,1396,594,636]" gridcol="8" gridrow="2" pageId="50" pageNumber="605">Absent</td>
|
||
<td id="7E535DB9FFC8F9D7FDA0EA487B105C24" box="[1408,1554,594,636]" gridcol="9" gridrow="2" pageId="50" pageNumber="605">Present</td>
|
||
<td id="7E535DB9FFC8F9D7FE3FEA487B995C24" box="[1567,1691,594,636]" gridcol="10" gridrow="2" pageId="50" pageNumber="605">Present</td>
|
||
<td id="7E535DB9FFC8F9D7FE86EA487A215C24" box="[1702,1827,594,636]" gridcol="11" gridrow="2" pageId="50" pageNumber="605">Present</td>
|
||
</tr>
|
||
<tr id="3D8234C5FFC8F9D7F8AAEA9F7A215CF7" box="[138,1827,645,687]" gridrow="3" pageId="50" pageNumber="605">
|
||
<th id="7E535DB9FFC8F9D7F8AAEA9F7C0B5CF7" box="[138,265,645,687]" gridcol="0" gridrow="3" pageId="50" pageNumber="605">Frontal filaments</th>
|
||
<td id="7E535DB9FFC8F9D7F937EA9F7C9A5CF7" box="[279,408,645,687]" gridcol="1" gridrow="3" pageId="50" pageNumber="605">Present</td>
|
||
<td id="7E535DB9FFC8F9D7F986EA9F7F2A5CF7" box="[422,552,645,687]" gridcol="2" gridrow="3" pageId="50" pageNumber="605">?</td>
|
||
<td id="7E535DB9FFC8F9D7FA16EA9F7FB05CF7" box="[566,690,645,687]" gridcol="3" gridrow="3" pageId="50" pageNumber="605">Present</td>
|
||
<td id="7E535DB9FFC8F9D7FAE0EA9F7E525CF7" box="[704,848,645,687]" gridcol="4" gridrow="3" pageId="50" pageNumber="605">Present</td>
|
||
<td id="7E535DB9FFC8F9D7FB7CEA9F7EC15CF7" box="[860,963,645,687]" gridcol="5" gridrow="3" pageId="50" pageNumber="605">Absent</td>
|
||
<td id="7E535DB9FFC8F9D7FBF8EA9F79555CF7" box="[984,1111,645,687]" gridcol="6" gridrow="3" pageId="50" pageNumber="605">Absent</td>
|
||
<td id="7E535DB9FFC8F9D7FC48EA9F79E95CF7" box="[1128,1259,645,687]" gridcol="7" gridrow="3" pageId="50" pageNumber="605">Absent</td>
|
||
<td id="7E535DB9FFC8F9D7FCD7EA9F78765CF7" box="[1271,1396,645,687]" gridcol="8" gridrow="3" pageId="50" pageNumber="605">Present</td>
|
||
<td id="7E535DB9FFC8F9D7FDA0EA9F7B105CF7" box="[1408,1554,645,687]" gridcol="9" gridrow="3" pageId="50" pageNumber="605">Present</td>
|
||
<td id="7E535DB9FFC8F9D7FE3FEA9F7B995CF7" box="[1567,1691,645,687]" gridcol="10" gridrow="3" pageId="50" pageNumber="605">Present</td>
|
||
<td id="7E535DB9FFC8F9D7FE86EA9F7A215CF7" box="[1702,1827,645,687]" gridcol="11" gridrow="3" pageId="50" pageNumber="605">Present</td>
|
||
</tr>
|
||
<tr id="3D8234C5FFC8F9D7F8AAEAA27A215D4A" box="[138,1827,696,786]" gridrow="4" pageId="50" pageNumber="605">
|
||
<th id="7E535DB9FFC8F9D7F8AAEAA27C0B5D4A" box="[138,265,696,786]" gridcol="0" gridrow="4" pageId="50" pageNumber="605">Dorsal tuber- cles on non- elytrigerous segments</th>
|
||
<td id="7E535DB9FFC8F9D7F937EAA27C9A5D4A" box="[279,408,696,786]" gridcol="1" gridrow="4" pageId="50" pageNumber="605">Inconspicuous</td>
|
||
<td id="7E535DB9FFC8F9D7F986EAA27F2A5D4A" box="[422,552,696,786]" gridcol="2" gridrow="4" pageId="50" pageNumber="605">Inconspicuous</td>
|
||
<td id="7E535DB9FFC8F9D7FA16EAA27FB05D4A" box="[566,690,696,786]" gridcol="3" gridrow="4" pageId="50" pageNumber="605">Inconspicuous</td>
|
||
<td id="7E535DB9FFC8F9D7FAE0EAA27E525D4A" box="[704,848,696,786]" gridcol="4" gridrow="4" pageId="50" pageNumber="605">Prominent</td>
|
||
<td id="7E535DB9FFC8F9D7FB7CEAA27EC15D4A" box="[860,963,696,786]" gridcol="5" gridrow="4" pageId="50" pageNumber="605">Prominent</td>
|
||
<td id="7E535DB9FFC8F9D7FBF8EAA279555D4A" box="[984,1111,696,786]" gridcol="6" gridrow="4" pageId="50" pageNumber="605">Inconspicuous</td>
|
||
<td id="7E535DB9FFC8F9D7FC48EAA279E95D4A" box="[1128,1259,696,786]" gridcol="7" gridrow="4" pageId="50" pageNumber="605">Inconspicuous</td>
|
||
<td id="7E535DB9FFC8F9D7FCD7EAA278765D4A" box="[1271,1396,696,786]" gridcol="8" gridrow="4" pageId="50" pageNumber="605">Inconspicuous</td>
|
||
<td id="7E535DB9FFC8F9D7FDA0EAA27B105D4A" box="[1408,1554,696,786]" gridcol="9" gridrow="4" pageId="50" pageNumber="605">Prominent</td>
|
||
<td id="7E535DB9FFC8F9D7FE3FEAA27B995D4A" box="[1567,1691,696,786]" gridcol="10" gridrow="4" pageId="50" pageNumber="605">Prominent</td>
|
||
<td id="7E535DB9FFC8F9D7FE86EAA27A215D4A" box="[1702,1827,696,786]" gridcol="11" gridrow="4" pageId="50" pageNumber="605">Prominent</td>
|
||
</tr>
|
||
<tr id="3D8234C5FFC8F9D7F8AAEB017A215D05" box="[138,1827,795,861]" gridrow="5" pageId="50" pageNumber="605">
|
||
<th id="7E535DB9FFC8F9D7F8AAEB017C0B5D05" box="[138,265,795,861]" gridcol="0" gridrow="5" pageId="50" pageNumber="605">Shape of dorsal tubercles</th>
|
||
<td id="7E535DB9FFC8F9D7F937EB017C9A5D05" box="[279,408,795,861]" gridcol="1" gridrow="5" pageId="50" pageNumber="605">Bulbous</td>
|
||
<td id="7E535DB9FFC8F9D7F986EB017F2A5D05" box="[422,552,795,861]" gridcol="2" gridrow="5" pageId="50" pageNumber="605">–</td>
|
||
<td id="7E535DB9FFC8F9D7FA16EB017FB05D05" box="[566,690,795,861]" gridcol="3" gridrow="5" pageId="50" pageNumber="605">–</td>
|
||
<td id="7E535DB9FFC8F9D7FAE0EB017E525D05" box="[704,848,795,861]" gridcol="4" gridrow="5" pageId="50" pageNumber="605">Conical</td>
|
||
<td id="7E535DB9FFC8F9D7FB7CEB017EC15D05" box="[860,963,795,861]" gridcol="5" gridrow="5" pageId="50" pageNumber="605">Conical</td>
|
||
<td id="7E535DB9FFC8F9D7FBF8EB0179555D05" box="[984,1111,795,861]" gridcol="6" gridrow="5" pageId="50" pageNumber="605">–</td>
|
||
<td id="7E535DB9FFC8F9D7FC48EB0179E95D05" box="[1128,1259,795,861]" gridcol="7" gridrow="5" pageId="50" pageNumber="605">–</td>
|
||
<td id="7E535DB9FFC8F9D7FCD7EB0178765D05" box="[1271,1396,795,861]" gridcol="8" gridrow="5" pageId="50" pageNumber="605">–</td>
|
||
<td id="7E535DB9FFC8F9D7FDA0EB017B105D05" box="[1408,1554,795,861]" gridcol="9" gridrow="5" pageId="50" pageNumber="605">Conical</td>
|
||
<td id="7E535DB9FFC8F9D7FE3FEB017B995D05" box="[1567,1691,795,861]" gridcol="10" gridrow="5" pageId="50" pageNumber="605">Bulbous</td>
|
||
<td id="7E535DB9FFC8F9D7FE86EB017A215D05" box="[1702,1827,795,861]" gridcol="11" gridrow="5" pageId="50" pageNumber="605">Conical</td>
|
||
</tr>
|
||
<tr id="3D8234C5FFC8F9D7F8AAEB7C7A215DC8" box="[138,1827,870,912]" gridrow="6" pageId="50" pageNumber="605">
|
||
<th id="7E535DB9FFC8F9D7F8AAEB7C7C0B5DC8" box="[138,265,870,912]" gridcol="0" gridrow="6" pageId="50" pageNumber="605">Dorsal body surface</th>
|
||
<td id="7E535DB9FFC8F9D7F937EB7C7C9A5DC8" box="[279,408,870,912]" gridcol="1" gridrow="6" pageId="50" pageNumber="605">Papillated</td>
|
||
<td id="7E535DB9FFC8F9D7F986EB7C7F2A5DC8" box="[422,552,870,912]" gridcol="2" gridrow="6" pageId="50" pageNumber="605">Smooth</td>
|
||
<td id="7E535DB9FFC8F9D7FA16EB7C7FB05DC8" box="[566,690,870,912]" gridcol="3" gridrow="6" pageId="50" pageNumber="605">?</td>
|
||
<td id="7E535DB9FFC8F9D7FAE0EB7C7E525DC8" box="[704,848,870,912]" gridcol="4" gridrow="6" pageId="50" pageNumber="605">Smooth</td>
|
||
<td id="7E535DB9FFC8F9D7FB7CEB7C7EC15DC8" box="[860,963,870,912]" gridcol="5" gridrow="6" pageId="50" pageNumber="605">?</td>
|
||
<td id="7E535DB9FFC8F9D7FBF8EB7C79555DC8" box="[984,1111,870,912]" gridcol="6" gridrow="6" pageId="50" pageNumber="605">Smooth</td>
|
||
<td id="7E535DB9FFC8F9D7FC48EB7C79E95DC8" box="[1128,1259,870,912]" gridcol="7" gridrow="6" pageId="50" pageNumber="605">Papillated</td>
|
||
<td id="7E535DB9FFC8F9D7FCD7EB7C78765DC8" box="[1271,1396,870,912]" gridcol="8" gridrow="6" pageId="50" pageNumber="605">Papillated</td>
|
||
<td id="7E535DB9FFC8F9D7FDA0EB7C7B105DC8" box="[1408,1554,870,912]" gridcol="9" gridrow="6" pageId="50" pageNumber="605">Smooth</td>
|
||
<td id="7E535DB9FFC8F9D7FE3FEB7C7B995DC8" box="[1567,1691,870,912]" gridcol="10" gridrow="6" pageId="50" pageNumber="605">Smooth</td>
|
||
<td id="7E535DB9FFC8F9D7FE86EB7C7A215DC8" box="[1702,1827,870,912]" gridcol="11" gridrow="6" pageId="50" pageNumber="605">Smooth</td>
|
||
</tr>
|
||
<tr id="3D8234C5FFC8F9D7F8AAEB807A215D9C" box="[138,1827,922,964]" gridrow="7" pageId="50" pageNumber="605">
|
||
<th id="7E535DB9FFC8F9D7F8AAEB807C0B5D9C" box="[138,265,922,964]" gridcol="0" gridrow="7" pageId="50" pageNumber="605">Ventral body surface</th>
|
||
<td id="7E535DB9FFC8F9D7F937EB807C9A5D9C" box="[279,408,922,964]" gridcol="1" gridrow="7" pageId="50" pageNumber="605">Smooth</td>
|
||
<td id="7E535DB9FFC8F9D7F986EB807F2A5D9C" box="[422,552,922,964]" gridcol="2" gridrow="7" pageId="50" pageNumber="605">Smooth</td>
|
||
<td id="7E535DB9FFC8F9D7FA16EB807FB05D9C" box="[566,690,922,964]" gridcol="3" gridrow="7" pageId="50" pageNumber="605">?</td>
|
||
<td id="7E535DB9FFC8F9D7FAE0EB807E525D9C" box="[704,848,922,964]" gridcol="4" gridrow="7" pageId="50" pageNumber="605">Smooth</td>
|
||
<td id="7E535DB9FFC8F9D7FB7CEB807EC15D9C" box="[860,963,922,964]" gridcol="5" gridrow="7" pageId="50" pageNumber="605">?</td>
|
||
<td id="7E535DB9FFC8F9D7FBF8EB8079555D9C" box="[984,1111,922,964]" gridcol="6" gridrow="7" pageId="50" pageNumber="605">Smooth</td>
|
||
<td id="7E535DB9FFC8F9D7FC48EB8079E95D9C" box="[1128,1259,922,964]" gridcol="7" gridrow="7" pageId="50" pageNumber="605">Papillated</td>
|
||
<td id="7E535DB9FFC8F9D7FCD7EB8078765D9C" box="[1271,1396,922,964]" gridcol="8" gridrow="7" pageId="50" pageNumber="605">Smooth</td>
|
||
<td id="7E535DB9FFC8F9D7FDA0EB807B105D9C" box="[1408,1554,922,964]" gridcol="9" gridrow="7" pageId="50" pageNumber="605">Smooth</td>
|
||
<td id="7E535DB9FFC8F9D7FE3FEB807B995D9C" box="[1567,1691,922,964]" gridcol="10" gridrow="7" pageId="50" pageNumber="605">Smooth</td>
|
||
<td id="7E535DB9FFC8F9D7FE86EB807A215D9C" box="[1702,1827,922,964]" gridcol="11" gridrow="7" pageId="50" pageNumber="605">Smooth</td>
|
||
</tr>
|
||
<tr id="3D8234C5FFC8F9D7F8AAEBD77A215A7F" box="[138,1827,973,1063]" gridrow="8" pageId="50" pageNumber="605">
|
||
<th id="7E535DB9FFC8F9D7F8AAEBD77C0B5A7F" box="[138,265,973,1063]" gridcol="0" gridrow="8" pageId="50" pageNumber="605">First seg- ment with nephridial papillae</th>
|
||
<td id="7E535DB9FFC8F9D7F937EBD77C9A5A7F" box="[279,408,973,1063]" gridcol="1" gridrow="8" pageId="50" pageNumber="605">Segment 10</td>
|
||
<td id="7E535DB9FFC8F9D7F986EBD77F2A5A7F" box="[422,552,973,1063]" gridcol="2" gridrow="8" pageId="50" pageNumber="605">?</td>
|
||
<td id="7E535DB9FFC8F9D7FA16EBD77FB05A7F" box="[566,690,973,1063]" gridcol="3" gridrow="8" pageId="50" pageNumber="605">Segment 10</td>
|
||
<td id="7E535DB9FFC8F9D7FAE0EBD77E525A7F" box="[704,848,973,1063]" gridcol="4" gridrow="8" pageId="50" pageNumber="605">Segment 10</td>
|
||
<td id="7E535DB9FFC8F9D7FB7CEBD77EC15A7F" box="[860,963,973,1063]" gridcol="5" gridrow="8" pageId="50" pageNumber="605">?</td>
|
||
<td id="7E535DB9FFC8F9D7FBF8EBD779555A7F" box="[984,1111,973,1063]" gridcol="6" gridrow="8" pageId="50" pageNumber="605">Segment 10</td>
|
||
<td id="7E535DB9FFC8F9D7FC48EBD779E95A7F" box="[1128,1259,973,1063]" gridcol="7" gridrow="8" pageId="50" pageNumber="605">Segment 10</td>
|
||
<td id="7E535DB9FFC8F9D7FCD7EBD778765A7F" box="[1271,1396,973,1063]" gridcol="8" gridrow="8" pageId="50" pageNumber="605">Segment 10</td>
|
||
<td id="7E535DB9FFC8F9D7FDA0EBD77B105A7F" box="[1408,1554,973,1063]" gridcol="9" gridrow="8" pageId="50" pageNumber="605">Segment 10</td>
|
||
<td id="7E535DB9FFC8F9D7FE3FEBD77B995A7F" box="[1567,1691,973,1063]" gridcol="10" gridrow="8" pageId="50" pageNumber="605">Segment 9</td>
|
||
<td id="7E535DB9FFC8F9D7FE86EBD77A215A7F" box="[1702,1827,973,1063]" gridcol="11" gridrow="8" pageId="50" pageNumber="605">Segment 10</td>
|
||
</tr>
|
||
<tr id="3D8234C5FFC8F9D7F8AAEC2A7A215AE2" box="[138,1827,1072,1210]" gridrow="9" pageId="50" pageNumber="605">
|
||
<th id="7E535DB9FFC8F9D7F8AAEC2A7C0B5AE2" box="[138,265,1072,1210]" gridcol="0" gridrow="9" pageId="50" pageNumber="605">Notochaetae develop- ment com- pared to neurochae- tae</th>
|
||
<td id="7E535DB9FFC8F9D7F937EC2A7C9A5AE2" box="[279,408,1072,1210]" gridcol="1" gridrow="9" pageId="50" pageNumber="605">Notochaetae stouter than neurochae- tae</td>
|
||
<td id="7E535DB9FFC8F9D7F986EC2A7F2A5AE2" box="[422,552,1072,1210]" gridcol="2" gridrow="9" pageId="50" pageNumber="605">Notochaetae stouter than neurochae- tae</td>
|
||
<td id="7E535DB9FFC8F9D7FA16EC2A7FB05AE2" box="[566,690,1072,1210]" gridcol="3" gridrow="9" pageId="50" pageNumber="605">Notochaetae stouter than neuro- chaetae</td>
|
||
<td id="7E535DB9FFC8F9D7FAE0EC2A7E525AE2" box="[704,848,1072,1210]" gridcol="4" gridrow="9" pageId="50" pageNumber="605">Notochaetae stouter than neurochaetae</td>
|
||
<td id="7E535DB9FFC8F9D7FB7CEC2A7EC15AE2" box="[860,963,1072,1210]" gridcol="5" gridrow="9" pageId="50" pageNumber="605">About the same</td>
|
||
<td id="7E535DB9FFC8F9D7FBF8EC2A79555AE2" box="[984,1111,1072,1210]" gridcol="6" gridrow="9" pageId="50" pageNumber="605">About the same</td>
|
||
<td id="7E535DB9FFC8F9D7FC48EC2A79E95AE2" box="[1128,1259,1072,1210]" gridcol="7" gridrow="9" pageId="50" pageNumber="605">Notochaetae subequal to or more slender than neurochae- tae</td>
|
||
<td id="7E535DB9FFC8F9D7FCD7EC2A78765AE2" box="[1271,1396,1072,1210]" gridcol="8" gridrow="9" pageId="50" pageNumber="605">Notochaetae stouter than neuro- chaetae</td>
|
||
<td id="7E535DB9FFC8F9D7FDA0EC2A7B105AE2" box="[1408,1554,1072,1210]" gridcol="9" gridrow="9" pageId="50" pageNumber="605">About the same</td>
|
||
<td id="7E535DB9FFC8F9D7FE3FEC2A7B995AE2" box="[1567,1691,1072,1210]" gridcol="10" gridrow="9" pageId="50" pageNumber="605">Notochaetae slightly stouter than neuro- chaetae</td>
|
||
<td id="7E535DB9FFC8F9D7FE86EC2A7A215AE2" box="[1702,1827,1072,1210]" gridcol="11" gridrow="9" pageId="50" pageNumber="605">Notochaetae stouter than neuro- chaetae</td>
|
||
</tr>
|
||
<tr id="3D8234C5FFC8F9D7F8AAECD97A215B15" box="[138,1827,1219,1357]" gridrow="10" pageId="50" pageNumber="605">
|
||
<th id="7E535DB9FFC8F9D7F8AAECD97C0B5B15" box="[138,265,1219,1357]" gridcol="0" gridrow="10" pageId="50" pageNumber="605">Notochaetae</th>
|
||
<td id="7E535DB9FFC8F9D7F937ECD97C9A5B15" box="[279,408,1219,1357]" gridcol="1" gridrow="10" pageId="50" pageNumber="605">With two rows of spines in one side</td>
|
||
<td id="7E535DB9FFC8F9D7F986ECD97F2A5B15" box="[422,552,1219,1357]" gridcol="2" gridrow="10" pageId="50" pageNumber="605">With two rows of spines in one side, blunt tips</td>
|
||
<td id="7E535DB9FFC8F9D7FA16ECD97FB05B15" box="[566,690,1219,1357]" gridcol="3" gridrow="10" pageId="50" pageNumber="605">With spinous rows, blunt tips</td>
|
||
<td id="7E535DB9FFC8F9D7FAE0ECD97E525B15" box="[704,848,1219,1357]" gridcol="4" gridrow="10" pageId="50" pageNumber="605">With faint spinous rows, pointed tips</td>
|
||
<td id="7E535DB9FFC8F9D7FB7CECD97EC15B15" box="[860,963,1219,1357]" gridcol="5" gridrow="10" pageId="50" pageNumber="605">Widely- spaced spinous rows, rounded tips</td>
|
||
<td id="7E535DB9FFC8F9D7FBF8ECD979555B15" box="[984,1111,1219,1357]" gridcol="6" gridrow="10" pageId="50" pageNumber="605">With faint spinous rows, blunt tips</td>
|
||
<td id="7E535DB9FFC8F9D7FC48ECD979E95B15" box="[1128,1259,1219,1357]" gridcol="7" gridrow="10" pageId="50" pageNumber="605">Smooth or with faint spinous rows, blunt tips</td>
|
||
<td id="7E535DB9FFC8F9D7FCD7ECD978765B15" box="[1271,1396,1219,1357]" gridcol="8" gridrow="10" pageId="50" pageNumber="605">Smooth, blunt tips</td>
|
||
<td id="7E535DB9FFC8F9D7FDA0ECD97B105B15" box="[1408,1554,1219,1357]" gridcol="9" gridrow="10" pageId="50" pageNumber="605">With faint spinous rows, pointed tips</td>
|
||
<td id="7E535DB9FFC8F9D7FE3FECD97B995B15" box="[1567,1691,1219,1357]" gridcol="10" gridrow="10" pageId="50" pageNumber="605">Smooth or with distinct spinous rows, blunt tips</td>
|
||
<td id="7E535DB9FFC8F9D7FE86ECD97A215B15" box="[1702,1827,1219,1357]" gridcol="11" gridrow="10" pageId="50" pageNumber="605">Smooth or with faint spinous rows, blunt tips</td>
|
||
</tr>
|
||
<tr id="3D8234C5FFC8F9D7F8AAED4C7A215818" box="[138,1827,1366,1600]" gridrow="11" pageId="50" pageNumber="605">
|
||
<th id="7E535DB9FFC8F9D7F8AAED4C7C0B5818" box="[138,265,1366,1600]" gridcol="0" gridrow="11" pageId="50" pageNumber="605">Neurochaetae</th>
|
||
<td id="7E535DB9FFC8F9D7F937ED4C7C9A5818" box="[279,408,1366,1600]" gridcol="1" gridrow="11" pageId="50" pageNumber="605">With spinous pocket on both margins, tapered to blunt tips</td>
|
||
<td id="7E535DB9FFC8F9D7F986ED4C7F2A5818" box="[422,552,1366,1600]" gridcol="2" gridrow="11" pageId="50" pageNumber="605">With spinous pocket on both margins, tapered to blunt tips</td>
|
||
<td id="7E535DB9FFC8F9D7FA16ED4C7FB05818" box="[566,690,1366,1600]" gridcol="3" gridrow="11" pageId="50" pageNumber="605">With spines on both margins, hooked tips</td>
|
||
<td id="7E535DB9FFC8F9D7FAE0ED4C7E525818" box="[704,848,1366,1600]" gridcol="4" gridrow="11" pageId="50" pageNumber="605">With spinous pocket on both margins, minute biden- tate hooked tips</td>
|
||
<td id="7E535DB9FFC8F9D7FB7CED4C7EC15818" box="[860,963,1366,1600]" gridcol="5" gridrow="11" pageId="50" pageNumber="605">With spines on both margins, tapered rounded tips</td>
|
||
<td id="7E535DB9FFC8F9D7FBF8ED4C79555818" box="[984,1111,1366,1600]" gridcol="6" gridrow="11" pageId="50" pageNumber="605">With spinous pocket on both margins, tapered blunt tips</td>
|
||
<td id="7E535DB9FFC8F9D7FC48ED4C79E95818" box="[1128,1259,1366,1600]" gridcol="7" gridrow="11" pageId="50" pageNumber="605">With spines or spinous pocket on both mar- gins, blunt tips</td>
|
||
<td id="7E535DB9FFC8F9D7FCD7ED4C78765818" box="[1271,1396,1366,1600]" gridcol="8" gridrow="11" pageId="50" pageNumber="605">With spinous pocket on both mar- gins, blunt tips</td>
|
||
<td id="7E535DB9FFC8F9D7FDA0ED4C7B105818" box="[1408,1554,1366,1600]" gridcol="9" gridrow="11" pageId="50" pageNumber="605">Type 1, with spinous pocket on both mar- gins, pointed tips; type 2, with spinous pocket more centrally, hooked tips</td>
|
||
<td id="7E535DB9FFC8F9D7FE3FED4C7B995818" box="[1567,1691,1366,1600]" gridcol="10" gridrow="11" pageId="50" pageNumber="605">With spines on both margins, pointed tips</td>
|
||
<td id="7E535DB9FFC8F9D7FE86ED4C7A215818" box="[1702,1827,1366,1600]" gridcol="11" gridrow="11" pageId="50" pageNumber="605">Smooth or with faint spinous rows in one side, abruptly to slightly hooked tips</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="830D3687FFC8061AF8AAEE717C4D58CA" blockId="50.[138,1837,1643,1682]" pageId="50" pageNumber="605">
|
||
References for species are provided in
|
||
<tableCitation id="CE30033CFFC8061AF9FBEE717F195825" box="[475,539,1643,1661]" captionStart="Table 2" captionStartId="7.[163,227,196,217]" captionText="Table 2. List of taxa included in the different datasets: only molecular data (MDS), molecular and morphological data (CDS) and only morphological data" pageId="50" pageNumber="605">Table 2</tableCitation>
|
||
. The descriptions of the valid species
|
||
<taxonomicName id="44B24D04FFC8061AFB4BEE7179175825" authorityName="Levenstein" authorityYear="1978" box="[875,1045,1643,1661]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="species" species="alia">
|
||
<emphasis id="B1C6EA95FFC8061AFB4BEE7179175825" box="[875,1045,1643,1661]" italics="true" pageId="50" pageNumber="605">Macellicephala alia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="44B24D04FFC8061AFC63EE7178165825" authorityName="Levenstein" authorityYear="1975" box="[1091,1300,1643,1661]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="50" pageNumber="605" phylum="Annelida" rank="species" species="tricornis">
|
||
<emphasis id="B1C6EA95FFC8061AFC63EE7178165825" box="[1091,1300,1643,1661]" italics="true" pageId="50" pageNumber="605">Macellicephala tricornis</emphasis>
|
||
</taxonomicName>
|
||
were not included (see Taxa selection for detail). ‘?’ indicates uncertain information.
|
||
</paragraph>
|
||
<caption id="D7CD660FFFC9061BF883EEBD7EB25908" pageId="51" pageNumber="606" startId="51.[163,242,1703,1725]" targetBox="[162,1443,158,1653]" targetPageId="51">
|
||
<paragraph id="830D3687FFC9061BF883EEBD7EB25908" blockId="51.[163,1443,1702,1872]" pageId="51" pageNumber="606">
|
||
<emphasis id="B1C6EA95FFC9061BF883EEBD7FCE58E4" bold="true" box="[163,716,1702,1725]" pageId="51" pageNumber="606">
|
||
Figure 15.
|
||
<taxonomicName id="44B24D04FFC9061BF903EEBC7F6C58E5" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[291,622,1702,1725]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="51" pageNumber="606" phylum="Annelida" rank="species" species="parvafauces" status="sp. nov.">
|
||
<emphasis id="B1C6EA95FFC9061BF903EEBC7F6C58E5" bold="true" box="[291,622,1702,1725]" italics="true" pageId="51" pageNumber="606">Macellicephala parvafauces</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AAF557EEFFC9061BFA55EEBD7FCE58E4" box="[629,716,1703,1724]" pageId="51" pageNumber="606" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10
|
||
<emphasis id="B1C6EA95FFC9061BFBD7EEF8790058AF" box="[1015,1026,1762,1783]" italics="true" pageId="51" pageNumber="606">–</emphasis>
|
||
12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="830D3687FFCE061CF8B1E8DF7C7C5FC3" blockId="52.[145,762,197,1885]" pageId="52" pageNumber="607">
|
||
equal sized (
|
||
<figureCitation id="1B892A02FFCE061CF90AE8DF7C8D5E83" box="[298,399,197,219]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="52" pageNumber="607">Fig. 15D</figureCitation>
|
||
); styles missing. Pharynx not everted in
|
||
<typeStatus id="5C098825FFCE061CF92AE8FF7C6F5EA3" box="[266,365,229,251]" pageId="52" pageNumber="607" type="holotype">holotype</typeStatus>
|
||
; dissected in
|
||
<typeStatus id="5C098825FFCE061CFA24E8FC7F685EA3" box="[516,618,230,251]" pageId="52" pageNumber="607" type="paratype">paratype</typeStatus>
|
||
(MNHN-IA- TYPE 1831), with nine pairs of rounded to subtriangular, equal-sized distal papillae; two pairs of small jaws with main fang, margin smooth (
|
||
<figureCitation id="1B892A02FFCE061CFA23E95F7F635F03" box="[515,609,325,347]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="52" pageNumber="607">Fig. 15E</figureCitation>
|
||
). Second segment with elytrophores, subbiramous parapodia, chaetae and ventral cirri.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFCE061CF889E9BF7CB45C24" blockId="52.[145,762,197,1885]" pageId="52" pageNumber="607">Nine pairs of large, bulbous elytrophores present on segments 2, 4, 5, 7, 9, 11, 13, 15 and 17; elytron dropped from segment 13 but still attached on segment 15, both elytra in poor condition; moderately large (partly covering preceding segment and subsequent two segments), surface smooth; posterior margin papillated, with large, short papillae.</paragraph>
|
||
<paragraph id="830D3687FFCE061CF889EA9C7C4C5D63" blockId="52.[145,762,197,1885]" pageId="52" pageNumber="607">
|
||
Cirrigerous segments with large, bulbous dorsal cirrophores, inserted subdistally on notopodia; style smooth, tapering, long (reaching three or four subsequent segments, much longer than tip of neuroacicular lobe); dorsal tubercles distinct, conical, small (
|
||
<figureCitation id="1B892A02FFCE061CF8FEEB3C7C3D5D64" box="[222,319,806,828]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="52" pageNumber="607">Fig. 15D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="830D3687FFCE061CF889EB5C7C7E5A64" blockId="52.[145,762,197,1885]" pageId="52" pageNumber="607">Ventral cirri smooth, tapering, present from segment 2 to last segment; inserted basally on neuropodia of segment 2, style long (longer than tip of neuroacicular lobe); in subsequent segments inserted medially on neuropodia (in middle body slightly subdistally on neuropodia), style short (shorter than tip of neuroacicular lobe), style longer than neuropodia lobe in last segment.</paragraph>
|
||
<paragraph id="830D3687FFCE061CF889EC5C7DFD58E5" blockId="52.[145,762,197,1885]" pageId="52" pageNumber="607">
|
||
Parapodia subbiramous; notopodia shorter than neuropodia (
|
||
<figureCitation id="1B892A02FFCE061CF93FEC7C7C795A24" box="[287,379,1126,1148]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="52" pageNumber="607">Fig. 15F</figureCitation>
|
||
). Dorsal ridges absent. Notopodia subtriangular, tapering into long acicular lobe, tip of notoacicula not penetrating epidermis. Neuropodia large, subtriangular, tapering into long acicular lobe, tip of neuroacicula not penetrating epidermis. Notochaetae moderate in number (15–20 observed), short to long, distally straight to slightly curved, with very faint spinous rows, with pointed tips (
|
||
<figureCitation id="1B892A02FFCE061CFAA8ED5D7FE85B05" box="[648,746,1351,1373]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="52" pageNumber="607">Fig. 15G</figureCitation>
|
||
); some with slightly hooked tip; notochaetae as stout as neurochaetae. Neurochaetae of
|
||
<specimenCount id="95B4FD0EFFCE061CFA2AED9D7F825BC4" box="[522,640,1415,1436]" pageId="52" pageNumber="607" type="generic" typeStatus="types">two types</specimenCount>
|
||
: (1) upper group, moderate in number (~20 observed), long to very long, distally flattened to concave, with very faint spinous rows along both margins, with pointed tips (
|
||
<figureCitation id="1B892A02FFCE061CF8B9EE1D7DFF5845" box="[153,253,1543,1565]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="52" pageNumber="607">Fig. 15H</figureCitation>
|
||
), laterally appearing with hooked tips; and (2) middle and lower groups, moderate in number (~20 observed) long to short, distally flattened to concave with very faint spinous rows on both sides but distally not touching the margins, with minute hooked tips (
|
||
<figureCitation id="1B892A02FFCE061CF8B9EEBD7DF258E5" box="[153,240,1703,1725]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="52" pageNumber="607">Fig. 15I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="830D3687FFCE061CF889EEDD7CE15905" blockId="52.[145,762,197,1885]" pageId="52" pageNumber="607">
|
||
Nephridial papillae on segments 10, 11 and 12, small, bulbous (
|
||
<figureCitation id="1B892A02FFCE061CF96AEEFD7CA958A5" box="[330,427,1767,1789]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="52" pageNumber="607">Fig. 15C</figureCitation>
|
||
). In last segment, notopodia and neuropodia of similar size. Pygidium rounded, not enclosed by last segment; with dorsal anus (
|
||
<figureCitation id="1B892A02FFCE061CFAABEF3D7FE95965" box="[651,747,1831,1853]" captionStart="Figure 15" captionStartId="51.[163,242,1703,1725]" captionTargetBox="[162,1443,158,1653]" captionTargetPageId="51" captionText="Figure 15. Macellicephala parvafauces sp. nov., holotype MNHN-IA-TYPE 1830 (A, B, D, F–I) and paratype MNHN- IA-TYPE 1831 (C, E). A, dorsal view of a preserved complete specimen. B, anterior end, ventral view of a preserved specimen. C, ventral view of a live specimen with nephridial papillae on segments 10–12. D, anterior end, dorsal view, chaetae omitted. E, inner view of half side of dissected pharynx with papillae. F, right parapodia, posterior view, segment 7. G, notochaeta with very faint spinous rows, segment 7. H, upper neurochaeta, segment 7. I, lower neurochaeta, segment 7. Abbreviations: dt, dorsal tubercle; ft, facial tubercle; np, nephridial papilla." pageId="52" pageNumber="607">Fig. 15A</figureCitation>
|
||
). Anal cirri lost, scars not seen.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFCE061CFB09E8DC79195FCB" blockId="52.[809,1425,198,404]" pageId="52" pageNumber="607">
|
||
<emphasis id="B1C6EA95FFCE061CFB09E8DC79445E83" box="[809,1094,198,219]" italics="true" pageId="52" pageNumber="607">Morphological variation:</emphasis>
|
||
<materialsCitation id="33DA3CDAFFCE061CFC72E8DC79155FCB" collectionCode="CCFZ" pageId="52" pageNumber="607" specimenCount="2" typeStatus="holotype">
|
||
The specimens found in the
|
||
<collectionCode id="E5A3AE42FFCE061CFB09E8FF7E6F5EA2" box="[809,877,229,250]" pageId="52" pageNumber="607">CCFZ</collectionCode>
|
||
are highly similar. The only difference between them concerns the chaetae. In the
|
||
<typeStatus id="5C098825FFCE061CFC84E91978045F41" box="[1188,1286,259,281]" pageId="52" pageNumber="607" type="holotype">holotype</typeStatus>
|
||
, the chaetae are slightly wilted, giving an impression of flatness, whereas in the
|
||
<typeStatus id="5C098825FFCE061CFBC1E95B79485F0E" box="[993,1098,321,342]" pageId="52" pageNumber="607" type="paratype">paratype</typeStatus>
|
||
the chaetae have the usual appearance. This could be an artefact of the fixation/ conservation process
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFCE061CFB09E9A678145D7C" blockId="52.[809,1425,444,804]" pageId="52" pageNumber="607">
|
||
<emphasis id="B1C6EA95FFCE061CFB09E9A6780E5F8A" box="[809,1292,444,466]" italics="true" pageId="52" pageNumber="607">
|
||
Remarks:
|
||
<taxonomicName id="44B24D04FFCE061CFB93E9A6780E5F8A" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[947,1292,444,466]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="52" pageNumber="607" phylum="Annelida" rank="species" species="parvafauces" status="sp. nov.">Macellicephala parvafauces</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="AAF557EEFFCE061CFD37E9A778705F8A" box="[1303,1394,445,466]" pageId="52" pageNumber="607" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is closer to
|
||
<taxonomicName id="44B24D04FFCE061CFBAEE9C1785D5FA9" authority="(Levinsen, 1886)" baseAuthorityName="Levinsen" baseAuthorityYear="1886" box="[910,1375,475,497]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="52" pageNumber="607" phylum="Annelida" rank="species" species="violacea">
|
||
<emphasis id="B1C6EA95FFCE061CFBAEE9C1799E5FA8" box="[910,1180,475,496]" italics="true" pageId="52" pageNumber="607">Macellicephala violacea</emphasis>
|
||
(
|
||
<bibRefCitation id="E7234B76FFCE061CFC8BE9C178555FA9" author="Levinsen GMR" box="[1195,1367,475,497]" pageId="52" pageNumber="607" pagination="287 - 303" refId="ref54845" refString="Levinsen GMR. 1886. Kara-Havets Ledorme (Annulata). In: Lutken CF, ed. Dijmphna-Togtets Zoologisk-botaniske Udbytte. Kobenhaven: J. Hagerup, 287 - 303." type="book chapter" year="1886">Levinsen, 1886</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="44B24D04FFCE061CFB09E9E078225C57" authority="Wu & Wang, 1987" authorityName="Wu & Wang" authorityYear="1987" box="[809,1312,506,527]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="52" pageNumber="607" phylum="Annelida" rank="species" species="australis">
|
||
<emphasis id="B1C6EA95FFCE061CFB09E9E0794A5C57" box="[809,1096,506,527]" italics="true" pageId="52" pageNumber="607">Macellicephala australis</emphasis>
|
||
<bibRefCitation id="E7234B76FFCE061CFC6EE9E078225C57" author="Wu BL & Wang YH" box="[1102,1312,506,527]" pageId="52" pageNumber="607" pagination="23 - 29" refId="ref58344" refString="Wu BL, Wang YH. 1987. Two new species of Polychaeta from South Ocean. Acta Zootaxonomica Sinica 12: 23 - 29." type="journal article" year="1987">Wu & Wang, 1987</bibRefCitation>
|
||
</taxonomicName>
|
||
, having a smooth body surface and neurochaetae with hooked tips (
|
||
<tableCitation id="CE30033CFFCE061CFB46EA2D7EBF5C14" box="[870,957,567,589]" captionStart="Table 4" captionStartId="50.[138,202,409,431]" captionText="Table 4. Diagnostic characters for all valid species in the genus Macellicephala" pageId="52" pageNumber="607">Table 4</tableCitation>
|
||
). However, the styles of the dorsal cirri in
|
||
<taxonomicName id="44B24D04FFCE061CFB67EA4C79585C33" baseAuthorityName="Levinsen" baseAuthorityYear="1886" box="[839,1114,598,619]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="52" pageNumber="607" phylum="Annelida" rank="species" species="violacea">
|
||
<emphasis id="B1C6EA95FFCE061CFB67EA4C79585C33" box="[839,1114,598,619]" italics="true" pageId="52" pageNumber="607">Macellicephala violacea</emphasis>
|
||
</taxonomicName>
|
||
are shorter than the tip of neuropodial lobe, whereas in
|
||
<taxonomicName id="44B24D04FFCE061CFC57EA6F78935CD2" authorityName="Wu & Wang" authorityYear="1987" box="[1143,1425,629,650]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="52" pageNumber="607" phylum="Annelida" rank="species" species="australis">
|
||
<emphasis id="B1C6EA95FFCE061CFC57EA6F78935CD2" box="[1143,1425,629,650]" italics="true" pageId="52" pageNumber="607">Macellicephala australis</emphasis>
|
||
</taxonomicName>
|
||
they are longer, as in
|
||
<taxonomicName id="44B24D04FFCE061CFC1CEA8978925CF0" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[1084,1424,659,680]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="52" pageNumber="607" phylum="Annelida" rank="species" species="parvafauces" status="sp. nov.">
|
||
<emphasis id="B1C6EA95FFCE061CFC1CEA8978925CF0" box="[1084,1424,659,680]" italics="true" pageId="52" pageNumber="607">Macellicephala parvafauces</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AAF557EEFFCE061CFB09EAA87E835C9F" box="[809,897,690,711]" pageId="52" pageNumber="607" rank="species">sp. nov.</taxonomicNameLabel>
|
||
This new species also has prostomial lobes anteriorly truncated and conical dorsal tubercles, whereas
|
||
<taxonomicName id="44B24D04FFCE061CFBADEAF579A65D5C" authorityName="Wu & Wang" authorityYear="1987" box="[909,1188,751,772]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" kingdom="Animalia" order="Phyllodocida" pageId="52" pageNumber="607" phylum="Annelida" rank="species" species="australis">
|
||
<emphasis id="B1C6EA95FFCE061CFBADEAF579A65D5C" box="[909,1188,751,772]" italics="true" pageId="52" pageNumber="607">Macellicephala australis</emphasis>
|
||
</taxonomicName>
|
||
has prostomial lobes anteriorly rounded and no dorsal tubercles.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFCE061CFB09EB5779DB5DF8" blockId="52.[809,1425,845,928]" pageId="52" pageNumber="607">
|
||
<emphasis id="B1C6EA95FFCE061CFB09EB577EA95D3A" box="[809,939,845,866]" italics="true" pageId="52" pageNumber="607">Etymology:</emphasis>
|
||
The species name came from the Latin ‘
|
||
<emphasis id="B1C6EA95FFCE061CFB0EEB767EC75DD8" box="[814,965,875,897]" italics="true" pageId="52" pageNumber="607">parvă fauces</emphasis>
|
||
’ meaning ‘small jaws’. It refers to the relatively smaller jaws of this species.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFCE061CFB09EBD37E825A20" blockId="52.[809,1425,969,1144]" pageId="52" pageNumber="607">
|
||
<emphasis id="B1C6EA95FFCE061CFB09EBD37EC45D86" box="[809,966,969,990]" italics="true" pageId="52" pageNumber="607">Genetic data:</emphasis>
|
||
DNA sequencing for this species was successful for
|
||
<emphasis id="B1C6EA95FFCE061CFBF6EBF279055DA5" box="[982,1031,1000,1021]" italics="true" pageId="52" pageNumber="607">COI</emphasis>
|
||
, 16S and 18S. The
|
||
<typeStatus id="5C098825FFCE061CFCD5EBFD78595DA5" box="[1269,1371,999,1021]" pageId="52" pageNumber="607" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="5C098825FFCE061CFB09EC1D7E975A44" box="[809,917,1031,1052]" pageId="52" pageNumber="607" type="paratype">paratype</typeStatus>
|
||
shared 98.6% of genetic material in
|
||
<emphasis id="B1C6EA95FFCE061CFD40EC1C78935A43" box="[1376,1425,1030,1051]" italics="true" pageId="52" pageNumber="607">COI</emphasis>
|
||
and 100% in 16S or 18S. The average K2P distance for intraspecific variation was 1.4% for
|
||
<emphasis id="B1C6EA95FFCE061CFCD0EC5E781D5A01" box="[1264,1311,1092,1113]" italics="true" pageId="52" pageNumber="607">COI</emphasis>
|
||
and 0.0% for 16S.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFCE061CFB09ECBB784D5B4A" blockId="52.[809,1425,1185,1299]" pageId="52" pageNumber="607">
|
||
<emphasis id="B1C6EA95FFCE061CFB09ECBB7EC35AEE" box="[809,961,1185,1206]" italics="true" pageId="52" pageNumber="607">Distribution:</emphasis>
|
||
Based on the material examined (
|
||
<specimenCount id="95B4FD0EFFCE061CFD46ECBB7EA35A8E" pageId="52" pageNumber="607" type="generic">two specimens</specimenCount>
|
||
), this species has a wide distribution within the Clarion-Clipperton Fracture Zone, being sampled in APEI#3 (
|
||
<typeStatus id="5C098825FFCE061CFB8EECE77EE25B4A" box="[942,992,1277,1298]" pageId="52" pageNumber="607">type</typeStatus>
|
||
locality) and GSR license areas.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFCE061CFB09ED2679585BF5" blockId="52.[809,1425,1340,1454]" pageId="52" pageNumber="607">
|
||
<emphasis id="B1C6EA95FFCE061CFB09ED267EF25B09" box="[809,1008,1340,1361]" italics="true" pageId="52" pageNumber="607">Ecological notes:</emphasis>
|
||
The
|
||
<typeStatus id="5C098825FFCE061CFC1BED2679A25B0A" box="[1083,1184,1340,1362]" pageId="52" pageNumber="607" type="holotype">holotype</typeStatus>
|
||
was extracted from a sponge,
|
||
<taxonomicName id="44B24D04FFCE061CFBBDED407E635BD6" authority="Tabachnick & Levi, 2004" authorityName="Tabachnick & Levi" authorityYear="2004" class="Hexactinellida" family="Euplectellidae" genus="Corbitella" kingdom="Animalia" order="Lyssacinosida" pageId="52" pageNumber="607" phylum="Porifera" rank="species" species="discasterosa">
|
||
<emphasis id="B1C6EA95FFCE061CFBBDED4079A55B37" box="[925,1191,1370,1391]" italics="true" pageId="52" pageNumber="607">Corbitella discasterosa</emphasis>
|
||
<bibRefCitation id="E7234B76FFCE061CFC8EED407E635BD6" author="Tabachnick KR & Levi C" pageId="52" pageNumber="607" pagination="11 - 71" refId="ref57759" refString="Tabachnick KR, Levi C. 2004. Lyssacinosida du Pacifique sudouest (Porifera: Hexactinellida). In: Marshall BA, Richer de Forges B, eds. Tropical Deep-Sea Benthos 23. Paris: Memoires du Museum national d'Histoire naturelle 191: 11 - 71." type="journal article" year="2004">Tabachnick & Lévi, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="E7234B76FFCE061CFB52ED6379625BD6" author="Kersken D & Janussen D & Arbizu PM" box="[882,1120,1401,1423]" pageId="52" pageNumber="607" refId="ref54317" refString="Kersken D, Janussen D, Arbizu PM. 2018. Deep-sea glass sponges (Hexactinellida) from polymetallic nodule fields in the Clarion-Clipperton Fracture Zone (CCFZ), northeastern Pacific: Part II - Hexasterophora. Marine Biodiversity. doi: 10.1007 / s 12526 - 018 - 0880 - y." type="book" year="2018">
|
||
Kersken
|
||
<emphasis id="B1C6EA95FFCE061CFBFEED60791B5BD6" box="[990,1049,1401,1423]" italics="true" pageId="52" pageNumber="607">et al.</emphasis>
|
||
, 2018
|
||
</bibRefCitation>
|
||
), indicating a commensal life mode with this sponge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |