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<document id="F44410B6D2321E882792E93430602B05" ID-DOI="10.11865/zs.2023101" ID-ISSN="2095-6827" ID-Zenodo-Dep="10941307" ID-ZooBank="7A57D6D4-279E-4A4F-9D9B-2ADF461B5E28" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712576969620" checkinUser="felipe" docAuthor="Lin, Yejie, Li, Shuqiang &amp; Pham, Dinh-Sac" docDate="2023" docId="431B4B17FF8DFF9AFF7ED560FD39FDFC" docLanguage="en" docName="ZoolSyst.48.1.1-99.pdf" docOrigin="Zoological Systematics 48 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8283969" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Latouchia wenruni Lin &amp; Li 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="13" masterDocId="BF22336FFF86FF96FFF6D010FFFAFFC0" masterDocTitle="Taxonomic notes on some spider species (Arachnida: Araneae) from China and Vietnam" masterLastPageNumber="99" masterPageNumber="1" pageNumber="12" updateTime="1712578185271" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C57032B7F498D48E9D25220C80CB3BB8">Taxonomic notes on some spider species (Arachnida: Araneae) from China and Vietnam</mods:title>
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<mods:namePart id="33243A5C8E4FDE5215EB79FF7EC09001">Lin, Yejie</mods:namePart>
<mods:affiliation id="4015F37CE48D8D918629B29BF0B309CF">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart id="8BE0BB822D920F1B20E82ED29BD63B9D">Li, Shuqiang</mods:namePart>
<mods:affiliation id="AA73D8E8F5647DFB9E7FA52304818A19">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart id="FE0B9245A3D1829865E471EF8E198137">Pham, Dinh-Sac</mods:namePart>
<mods:affiliation id="FDBAA128AC6B792E0A92A5E008918A0E">Vietnam National Museum of Nature (VNMN), Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam Corresponding author, E-mail: phamdinhsac @ vnmn. vast. vn</mods:affiliation>
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<mods:date id="C24FA064FEB2DD625572A02AF312B3DC">2023</mods:date>
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<treatment id="431B4B17FF8DFF9AFF7ED560FD39FDFC" ID-DOI="http://doi.org/10.5281/zenodo.10941531" ID-Zenodo-Dep="10941531" LSID="urn:lsid:plazi:treatment:431B4B17FF8DFF9AFF7ED560FD39FDFC" httpUri="http://treatment.plazi.org/id/431B4B17FF8DFF9AFF7ED560FD39FDFC" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="83A8A98AFF8DFF9DFF7ED560FD41FA48" box="[136,699,1392,1416]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="CB0DFA01FF8DFF9DFF7ED560FD41FA48" blockId="11.[136,699,1392,1416]" box="[136,699,1392,1416]" pageId="11" pageNumber="12">
<heading id="90454D6DFF8DFF9DFF7ED560FD41FA48" box="[136,699,1392,1416]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="12" reason="2">
<emphasis id="F9C62613FF8DFF9DFF7ED560FDD0FA48" bold="true" box="[136,554,1392,1416]" pageId="11" pageNumber="12">
<taxonomicName id="0CB28182FF8DFF9DFF7ED560FE33FA48" authority="Lin &amp; Li" authorityName="Lin &amp; Li" authorityYear="2023" box="[136,457,1392,1416]" class="Arachnida" family="Halonoproctidae" genus="Latouchia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="wenruni" status="sp. nov.">
<emphasis id="F9C62613FF8DFF9DFF7ED560FEA6FA48" bold="true" box="[136,348,1392,1416]" italics="true" pageId="11" pageNumber="12">Latouchia wenruni</emphasis>
Lin &amp; Li
</taxonomicName>
,
<taxonomicNameLabel id="E2F59B68FF8DFF9DFE22D560FDD0FA48" box="[468,554,1392,1416]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="5389E684FF8DFF9DFDB1D560FD49FA48" box="[583,691,1392,1416]" captionStart-0="Figure 9" captionStart-1="Figure 10" captionStartId-0="12.[155,218,2026,2048]" captionStartId-1="13.[155,218,2026,2048]" captionTargetBox-0="[189,1392,633,1980]" captionTargetBox-1="[189,1399,252,1991]" captionTargetId-0="figure-194@12.[189,1399,633,1987]" captionTargetId-1="figure-48@13.[189,1399,252,1991]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Figure 9. Latouchia wenruni Lin &amp; Li, sp. nov., holotype male, left palp. A. Ventral view; B. Retrolateral view; C. Embolus tip, dorsal view.Abbreviations: AE—apophysis of embolic tip; E—embolus. Scale bars: A–B=0.5mm; C=0.1mm." captionText-1="Figure 10. Latouchia wenruni Lin &amp; Li, sp. nov., holotype male (B) and paratype female (A, C). A. Vulva, dorsal view; B–C. Habitus, dorsal view. Abbreviations: SH—spermathecal head; Sta—stalk. Scale bars: A=0.3mm; B–C=2.0mm." figureDoi-0="http://doi.org/10.5281/zenodo.10941337" figureDoi-1="http://doi.org/10.5281/zenodo.10941342" httpUri-0="https://zenodo.org/record/10941337/files/figure.png" httpUri-1="https://zenodo.org/record/10941342/files/figure.png" pageId="11" pageNumber="12">Figs 9–10</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="83A8A98AFF8DFF9AFF4BD5B7FD39FDFC" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="etymology">
<paragraph id="CB0DFA01FF8DFF9DFF4BD5B7FEE0FA22" blockId="11.[136,1452,1447,2061]" pageId="11" pageNumber="12">
Etymology. The species is named after Mr. Run Wen, the collector of the
<typeStatus id="140944A3FF8DFF9DFC02D5B7FBABFA7F" box="[1012,1105,1447,1471]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
and
<typeStatus id="140944A3FF8DFF9DFB7DD5B8FB0FFA00" box="[1163,1269,1448,1472]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
; noun (name) in genitive case.
</paragraph>
<paragraph id="CB0DFA01FF8DFF9DFF4BD5FDFF28F94F" blockId="11.[136,1452,1447,2061]" pageId="11" pageNumber="12">
Diagnosis. The male of the new species is similar to that of
<taxonomicName id="0CB28182FF8DFF9DFCC9D5FEFB81F9C5" authority="Lin &amp; Li, 2022" authorityName="Lin &amp; Li" authorityYear="2022" box="[831,1147,1517,1541]" class="Arachnida" family="Halonoproctidae" genus="Latouchia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="yuanjingae">
<emphasis id="F9C62613FF8DFF9DFCC9D5FEFC29F9C5" box="[831,979,1518,1541]" italics="true" pageId="11" pageNumber="12">L. yuanjingae</emphasis>
Lin &amp; Li, 2022
</taxonomicName>
by the straight embolus and the length of embolus is as long as the bulb, but it can be distinguished by the serrated embolus in ventral view (
<emphasis id="F9C62613FF8DFF9DFACFD600FAACF9E7" box="[1337,1366,1552,1575]" italics="true" pageId="11" pageNumber="12">vs.</emphasis>
smooth in
<taxonomicName id="0CB28182FF8DFF9DFF52D623FECDF98A" authorityName="Lin &amp; Li" authorityYear="2022" box="[164,311,1587,1610]" class="Arachnida" family="Halonoproctidae" genus="Latouchia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="yuanjingae">
<emphasis id="F9C62613FF8DFF9DFF52D623FECDF98A" box="[164,311,1587,1610]" italics="true" pageId="11" pageNumber="12">L. yuanjingae</emphasis>
</taxonomicName>
) and lacking bristles laterally on the tibia tip (
<emphasis id="F9C62613FF8DFF9DFCDFD623FCBCF98A" box="[809,838,1587,1610]" italics="true" pageId="11" pageNumber="12">vs.</emphasis>
present in
<taxonomicName id="0CB28182FF8DFF9DFC4AD623FBB5F98A" authorityName="Lin &amp; Li" authorityYear="2022" box="[956,1103,1587,1610]" class="Arachnida" family="Halonoproctidae" genus="Latouchia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="yuanjingae">
<emphasis id="F9C62613FF8DFF9DFC4AD623FBB5F98A" box="[956,1103,1587,1610]" italics="true" pageId="11" pageNumber="12">L. yuanjingae</emphasis>
</taxonomicName>
). The female of the new species can be distinguished from those of
<taxonomicName id="0CB28182FF8DFF9DFDFDD646FCA2F9AD" authority="Zhang &amp; Wang, 2021" authorityName="Zhang &amp; Wang" authorityYear="2021" box="[523,856,1621,1645]" class="Arachnida" family="Halonoproctidae" genus="Latouchia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="yejiei">
<emphasis id="F9C62613FF8DFF9DFDFDD646FD99F9AC" box="[523,611,1621,1645]" italics="true" pageId="11" pageNumber="12">L. yejiei</emphasis>
<bibRefCitation id="AF2387F0FF8DFF9DFD9DD645FCA2F9AD" author="Zhang &amp; Wang &amp; Two new species of the genus Latouchia Pocock &amp; China &amp; Araneae &amp; Mygalomorphae &amp; Halonoproctidae &amp; Biodiversity Data Journal" box="[619,856,1621,1645]" pageId="11" pageNumber="12" pagination="1 - 11" refId="ref35386" refString="Zhang, S., Wang, C. 2021. Two new species of the genus Latouchia Pocock, 1901 from southeast China (Araneae: Mygalomorphae: Halonoproctidae). Biodiversity Data Journal, 9 (e 72456): 1 - 11. doi: 10.3897 / BDJ. 9. e 72456" type="journal article" year="2021">Zhang &amp; Wang, 2021</bibRefCitation>
</taxonomicName>
by the prominent spermathecal heads (
<emphasis id="F9C62613FF8DFF9DFAF0D646FAD9F9AD" box="[1286,1315,1622,1645]" italics="true" pageId="11" pageNumber="12">vs.</emphasis>
absent in
<taxonomicName id="0CB28182FF8DFF9DFA60D646FF38F94F" authorityName="Zhang &amp; Wang" authorityYear="2021" class="Arachnida" family="Halonoproctidae" genus="Latouchia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="yejiei">
<emphasis id="F9C62613FF8DFF9DFA60D646FF38F94F" italics="true" pageId="11" pageNumber="12">L. yejiei</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="CB0DFA01FF8DFF9DFF4BD68AFE7EF8FD" blockId="11.[136,1452,1447,2061]" pageId="11" pageNumber="12">
Description. Male (
<typeStatus id="140944A3FF8DFF9DFE7AD68AFE0AF972" box="[396,496,1690,1714]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
). Total length 9.71; carapace 4.72 long, 4.16 wide, opisthosoma 4.99 long, 3.37 wide. Eye sizes and interdistances: AME 0.16, ALE 0.29, PME 0.21, PLE 0.28, AME–AME 0.07, AME–ALE 0.07, PME–PME 0.30, PME–PLE 0.02, AME–PME 0.09, ALE–PLE 0.09. Chelicerae with five promarginal and five retromarginal teeth. Leg measurements: I 14.18 (4.34, 5.08, 2.91, 1.85), II 12.59 (3.81, 4.16, 2.82, 1.80), III 10.52 (2.88, 3.05, 2.70, 1.89), IV 15.14 (4.16, 5.04, 3.85, 2.09).
</paragraph>
<paragraph id="CB0DFA01FF8DFF9DFF4BD757FB10F865" blockId="11.[136,1452,1447,2061]" pageId="11" pageNumber="12">
Coloration (
<figureCitation id="5389E684FF8DFF9DFEC9D757FE58F89F" box="[319,418,1863,1887]" captionStart="Figure 10" captionStartId="13.[155,218,2026,2048]" captionTargetBox="[189,1399,252,1991]" captionTargetId="figure-48@13.[189,1399,252,1991]" captionTargetPageId="13" captionText="Figure 10. Latouchia wenruni Lin &amp; Li, sp. nov., holotype male (B) and paratype female (A, C). A. Vulva, dorsal view; B–C. Habitus, dorsal view. Abbreviations: SH—spermathecal head; Sta—stalk. Scale bars: A=0.3mm; B–C=2.0mm." figureDoi="http://doi.org/10.5281/zenodo.10941342" httpUri="https://zenodo.org/record/10941342/files/figure.png" pageId="11" pageNumber="12">Fig. 10B</figureCitation>
). Carapace dark brown to brown, covered with black setae. Chelicerae dark brown. Endites and labium red-brown. Sternum yellow-brown. Legs brown with black setae. Opisthosoma oval, dark brown, dorsum with numerous irregular white spots, laterally brown, ventrum yellow-brown. Spinnerets short, yellow-brown.
</paragraph>
<paragraph id="CB0DFA01FF8DFF9DFF4BD7BFFDE6F82A" blockId="11.[136,1452,1447,2061]" pageId="11" pageNumber="12">
Palp (
<figureCitation id="5389E684FF8DFF9DFF0BD7BFFEBBF807" box="[253,321,1967,1991]" captionStart="Figure 9" captionStartId="12.[155,218,2026,2048]" captionTargetBox="[189,1392,633,1980]" captionTargetId="figure-194@12.[189,1399,633,1987]" captionTargetPageId="12" captionText="Figure 9. Latouchia wenruni Lin &amp; Li, sp. nov., holotype male, left palp. A. Ventral view; B. Retrolateral view; C. Embolus tip, dorsal view.Abbreviations: AE—apophysis of embolic tip; E—embolus. Scale bars: A–B=0.5mm; C=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10941337" httpUri="https://zenodo.org/record/10941337/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
). Tibia four times longer than wide, bristles on lateral side of tip are stout. Embolus slightly curved, tip with five serrations and an apophysis.
</paragraph>
<paragraph id="CB0DFA01FF8DFF9AFF4BD7E5FE7EFEAC" blockId="11.[136,1452,1447,2061]" lastBlockId="12.[136,1452,236,573]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">Female (IZCAS-Ar43840). Total length 18.23; carapace 8.54 long, 7.23 wide, opisthosoma 9.69 long, 6.61 wide. Eye sizes and interdistances: AME 0.26, ALE 0.43, PME 0.21, PLE 0.34, AME–AME 0.16, AME–ALE 0.22, PME–PME 0.58, PME–PLE 0.14, AME–PME 0.16, ALE–PLE 0.33. Chelicerae with six promarginal and eight retromarginal teeth. Leg measurements: I 18.64 (6.27, 7.39, 3.14, 1.84), II 15.28 (5.32, 5.57, 2.67, 1.72), III 14.22 (4.62, 5.23, 2.22, 2.15), IV 20.51 (6.29, 7.54, 4.19, 2.49).</paragraph>
<paragraph id="CB0DFA01FF8AFF9AFF4BD167FCCBFE4F" blockId="12.[136,1452,236,573]" box="[189,817,375,399]" pageId="12" pageNumber="13">
Coloration (
<figureCitation id="5389E684FF8AFF9AFEC8D167FE5AFE4F" box="[318,416,375,399]" captionStart="Figure 10" captionStartId="13.[155,218,2026,2048]" captionTargetBox="[189,1399,252,1991]" captionTargetId="figure-48@13.[189,1399,252,1991]" captionTargetPageId="13" captionText="Figure 10. Latouchia wenruni Lin &amp; Li, sp. nov., holotype male (B) and paratype female (A, C). A. Vulva, dorsal view; B–C. Habitus, dorsal view. Abbreviations: SH—spermathecal head; Sta—stalk. Scale bars: A=0.3mm; B–C=2.0mm." figureDoi="http://doi.org/10.5281/zenodo.10941342" httpUri="https://zenodo.org/record/10941342/files/figure.png" pageId="12" pageNumber="13">Fig. 10C</figureCitation>
). Similar to that of male except paler.
</paragraph>
<paragraph id="CB0DFA01FF8AFF9AFF4BD18AFA56FE72" blockId="12.[136,1452,236,573]" box="[189,1452,410,434]" pageId="12" pageNumber="13">
Endogyne (
<figureCitation id="5389E684FF8AFF9AFEC1D18AFE63FE72" box="[311,409,410,434]" captionStart="Figure 10" captionStartId="13.[155,218,2026,2048]" captionTargetBox="[189,1399,252,1991]" captionTargetId="figure-48@13.[189,1399,252,1991]" captionTargetPageId="13" captionText="Figure 10. Latouchia wenruni Lin &amp; Li, sp. nov., holotype male (B) and paratype female (A, C). A. Vulva, dorsal view; B–C. Habitus, dorsal view. Abbreviations: SH—spermathecal head; Sta—stalk. Scale bars: A=0.3mm; B–C=2.0mm." figureDoi="http://doi.org/10.5281/zenodo.10941342" httpUri="https://zenodo.org/record/10941342/files/figure.png" pageId="12" pageNumber="13">Fig. 10A</figureCitation>
). Spermathecae sock shaped, with numerous glandular pores; stalks almost as wide as spermatheca.
</paragraph>
<paragraph id="CB0DFA01FF8AFF9AFF4BD1ACFCADFDDA" blockId="12.[136,1452,236,573]" pageId="12" pageNumber="13">
<materialsCitation id="7BDAF05CFF8AFF9AFF4BD1ACFCA9FDDA" collectingDate="2022-09-13" collectorName="Run Wen &amp; Chengbin Wen &amp; Neng Guo" country="China" elevation="603" latitude="19.2815" location="Material" longLatPrecision="7" longitude="109.0987" pageId="12" pageNumber="13" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="holotype">
<location id="CE6DACDAFF8AFF9AFF4BD1ACFEEDFE14" LSID="urn:lsid:plazi:treatment:431B4B17FF8DFF9AFF7ED560FD39FDFC:CE6DACDAFF8AFF9AFF4BD1ACFEEDFE14" box="[189,279,444,468]" country="China" latitude="19.2815" longLatPrecision="7" longitude="109.0987" name="Material" pageId="12" pageNumber="13">Material</location>
examined.
<typeStatus id="140944A3FF8AFF9AFE56D1ACFDF9FE14" box="[416,515,444,468]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
<specimenCount id="DDB43188FF8AFF9AFDF9D1ADFDD9FE14" box="[527,547,445,468]" count="1" pageId="12" pageNumber="13" type="male">♂</specimenCount>
(IZCAS-Ar43837),
<collectingCountry id="B3A5BA91FF8AFF9AFCFCD1ACFCB4FE14" box="[778,846,444,468]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
:
<taxonomicName id="0CB28182FF8AFF9AFCA8D1ACFEFEFE37" authority=", Changjiang Li Autonomous County, Bangxi Baomeiling" authorityName="Changjiang Li Autonomous County, Bangxi Baomeiling" class="Arachnida" family="Liocranidae" genus="Sphingius" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="hainan">Hainan, Changjiang Li Autonomous County, Bangxi Baomeiling</taxonomicName>
(
<geoCoordinate id="AE869CC6FF8AFF9AFEE1D1CFFE6AFE37" box="[279,400,479,503]" degrees="19.2815" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="5" value="19.2815">19.2815°N</geoCoordinate>
,
<geoCoordinate id="AE869CC6FF8AFF9AFE68D1CFFDDAFE37" box="[414,544,479,503]" degrees="109.0987" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="5" value="109.0987">109.0987°E</geoCoordinate>
, elev.
<quantity id="0C4A57E4FF8AFF9AFD9DD1CFFD4AFE37" box="[619,688,479,503]" metricMagnitude="2" metricUnit="m" metricValue="6.029999999999999" pageId="12" pageNumber="13" unit="m" value="603.0">
<elevation id="409F1D32FF8AFF9AFD9DD1CFFD4AFE37" box="[619,688,479,503]" metricMagnitude="2" metricUnit="m" metricValue="6.029999999999999" pageId="12" pageNumber="13" unit="m" value="603.0">603 m</elevation>
</quantity>
),
<date id="BF0CDCC1FF8AFF9AFD31D1CFFC58FE37" box="[711,930,479,503]" pageId="12" pageNumber="13" value="2022-09-13">
<collectingDate id="AF482529FF8AFF9AFD31D1CFFC58FE37" box="[711,930,479,503]" pageId="12" pageNumber="13" value="2022-09-13">13 September 2022</collectingDate>
</date>
,
<collectorName id="66479FD7FF8AFF9AFC47D1F0FBE0FE37" box="[945,1050,480,503]" pageId="12" pageNumber="13">Run Wen</collectorName>
,
<collectorName id="66479FD7FF8AFF9AFBDFD1CFFB31FE37" box="[1065,1227,479,503]" pageId="12" pageNumber="13">Chengbin Wen</collectorName>
and
<collectorName id="66479FD7FF8AFF9AFAF1D1F0FA80FE37" box="[1287,1402,479,504]" pageId="12" pageNumber="13">Neng Guo</collectorName>
leg.
<typeStatus id="140944A3FF8AFF9AFF7ED212FF09FDDA" box="[136,243,514,538]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="DDB43188FF8AFF9AFF0BD212FEE5FDDA" box="[253,287,514,538]" count="3" pageId="12" pageNumber="13" type="female">3♀</specimenCount>
(IZCAS-Ar43838–Ar43840), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="CB0DFA01FF8AFF9AFF4BD234FD39FDFC" blockId="12.[136,1452,236,573]" box="[189,707,548,573]" pageId="12" pageNumber="13">
Distribution. Known only from the
<typeStatus id="140944A3FF8AFF9AFDCFD235FD9DFDFD" box="[569,615,549,573]" pageId="12" pageNumber="13">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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