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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.62.47854" ID-GBIF-Dataset="af20103b-7bf8-4a68-92c2-1a88288c65d1" ID-PMC="PMC7010847" ID-Pensoft-Pub="1314-4049-62-97" ID-Pensoft-UUID="0CDEF6D5DE4451A181F8818FFC66ACD9" ID-PubMed="32076384" ModsDocID="1314-4049-62-97" checkinTime="1580810382597" checkinUser="pensoft" docAuthor="Zhu, Haiyan, Pan, Meng, Bezerra, Jadson D. P., Tian, Chengming &amp; Fan, Xinlei" docDate="2020" docId="37DD8FA7A93E510EBD0B0C4C2E4998C2" docLanguage="en" docName="MycoKeys 62: 97-121" docOrigin="MycoKeys 62" docSource="http://dx.doi.org/10.3897/mycokeys.62.47854" docTitle="Cytospora spiraeicola H. Y. Zhu &amp; X. L. Fan 2020, sp. nov." docType="treatment" docVersion="4" id="0CDEF6D5DE4451A181F8818FFC66ACD9" lastPageNumber="97" masterDocId="0CDEF6D5DE4451A181F8818FFC66ACD9" masterDocTitle="Discovery of Cytospora species associated with canker disease of tree hosts from Mount Dongling of China" masterLastPageNumber="121" masterPageNumber="97" pageNumber="97" updateTime="1668136383112" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Discovery of Cytospora species associated with canker disease of tree hosts from Mount Dongling of China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhu, Haiyan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pan, Meng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bezerra, Jadson D. P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tian, Chengming</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fan, Xinlei</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>62</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>97</mods:start>
<mods:end>121</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.62.47854</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.62.47854</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-62-97</mods:identifier>
<mods:identifier type="Pensoft-UUID">0CDEF6D5DE4451A181F8818FFC66ACD9</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="161939146" LSID="urn:lsid:plazi:treatment:37DD8FA7A93E510EBD0B0C4C2E4998C2" httpUri="http://treatment.plazi.org/id/37DD8FA7A93E510EBD0B0C4C2E4998C2" lastPageNumber="97" pageId="0" pageNumber="97">
<subSubSection pageId="0" pageNumber="97" type="nomenclature">
<paragraph pageId="0" pageNumber="97">
<taxonomicName LSID="37DD8FA7-A93E-510E-BD0B-0C4C2E4998C2" authority="H. Y. Zhu &amp; X. L. Fan" authorityName="H. Y. Zhu &amp; X. L. Fan" authorityYear="2020" class="Sordariomycetes" family="Valsaceae" genus="Cytospora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cytospora spiraeicola" order="Diaporthales" pageId="0" pageNumber="97" phylum="Ascomycota" rank="species" species="spiraeicola" status="sp. nov.">Cytospora spiraeicola H.Y. Zhu &amp; X.L. Fan</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="97">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 7" captionStartId="F6" captionText="Figure 7. Cytospora spiraeicola from Spiraea salicifolia (CF 2019803). A, B habit of ascomata on twig C transverse section of ascoma D longitudinal section through ascoma E asci and ascospores F, G ascus H ascospores I colonies on PDA at 3 days (left) and 30 days (right). Scale bars: 1 mm (A, B); 500 μm (C, D); 10 μm (E-H)." figureDoi="10.3897/mycokeys.62.47854.figure7" httpUri="https://binary.pensoft.net/fig/375799" pageId="0" pageNumber="97">Fig. 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="97" type="etymology">
<paragraph pageId="0" pageNumber="97">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="97">
Named after the host genus on which it was collected,
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Spiraea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Spiraea" order="Rosales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="97">Spiraea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="97" type="holotype">
<paragraph pageId="0" pageNumber="97">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="97">
China, Beijing City, Mentougou District, Mount Dongling, Xiaolongmen Forestry Centre (
<geoCoordinate degrees="115" direction="east" minutes="28" orientation="longitude" precision="1" seconds="28.52" value="115.47459">115°28'28.52&quot;E</geoCoordinate>
,
<geoCoordinate degrees="39" direction="north" minutes="55" orientation="latitude" precision="1" seconds="49.42" value="39.930397">39°55'49.42&quot;N</geoCoordinate>
), from branches of
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Spiraea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Spiraea salicifolia" order="Rosales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="salicifolia">
<emphasis italics="true" pageId="0" pageNumber="97">Spiraea salicifolia</emphasis>
</taxonomicName>
, 17 Aug 2017, H.Y. Zhu &amp; X.L. Fan, holotype CF 2019803, ex-type living culture CFCC 53138.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="97" type="description">
<paragraph pageId="0" pageNumber="97">Description.</paragraph>
<paragraph pageId="0" pageNumber="97">
<emphasis italics="true" pageId="0" pageNumber="97">Necrotrophic</emphasis>
on branches of
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Spiraea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Spiraea salicifolia" order="Rosales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="salicifolia">
<emphasis italics="true" pageId="0" pageNumber="97">Spiraea salicifolia</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Rehder &amp; E. H. Wilson" authorityYear="1915" class="Magnoliopsida" family="Malvaceae" genus="Tilia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tilia nobilis" order="Malvales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="nobilis">
<emphasis italics="true" pageId="0" pageNumber="97">Tilia nobilis</emphasis>
</taxonomicName>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="97">Sexual morph</emphasis>
:
<emphasis italics="true" pageId="0" pageNumber="97">Ascostromata</emphasis>
immersed in the bark, erumpent through the surface of bark, scattered, with 3-5 perithecia arranged regularly, 660-890
<normalizedToken originalValue="µm">µm</normalizedToken>
in diam.
<emphasis italics="true" pageId="0" pageNumber="97">Conceptacle</emphasis>
absent.
<emphasis italics="true" pageId="0" pageNumber="97">Ectostromatic disc</emphasis>
pale grey, usually surrounded by tightly crowded ostiolar necks, quadrangular, 240-350
<normalizedToken originalValue="µm">µm</normalizedToken>
in diam., with 5-8 ostioles arranged regularly per disc.
<emphasis italics="true" pageId="0" pageNumber="97">Ostioles</emphasis>
numerous, dark grey to black, at the same or above the level as the disc, concentrated, arranged regularly in a disc, 25-40
<normalizedToken originalValue="µm">µm</normalizedToken>
in diam.
<emphasis italics="true" pageId="0" pageNumber="97">Perithecia</emphasis>
dark grey to black, flask-shaped to spherical, arranged circularly, 210-250
<normalizedToken originalValue="µm">µm</normalizedToken>
in diam.
<emphasis italics="true" pageId="0" pageNumber="97">Paraphyses</emphasis>
lacking.
<emphasis italics="true" pageId="0" pageNumber="97">Asci</emphasis>
free, clavate to elongate, obovoid, 26-37
<normalizedToken originalValue="×">x</normalizedToken>
7.5-9 (av. = 33
<normalizedToken originalValue="±">+/-</normalizedToken>
2.5
<normalizedToken originalValue="×">x</normalizedToken>
8.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9, n = 10)
<normalizedToken originalValue="μm">μm</normalizedToken>
, 8-spored.
<emphasis italics="true" pageId="0" pageNumber="97">Ascospores</emphasis>
biseriate, elongate-allantoid, thin-walled, hyaline, slightly curved, aseptate, 8.5-12
<normalizedToken originalValue="×">x</normalizedToken>
2.5-3.5 (av. = 10
<normalizedToken originalValue="±">+/-</normalizedToken>
1
<normalizedToken originalValue="×">x</normalizedToken>
3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.3, n = 30)
<normalizedToken originalValue="μm">μm</normalizedToken>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="97">Asexual morph</emphasis>
: not observed.
</paragraph>
<caption doi="10.3897/mycokeys.62.47854.figure7" httpUri="https://binary.pensoft.net/fig/375799" pageId="0" pageNumber="97" start="Figure 7" startId="F6">
<paragraph pageId="0" pageNumber="97">
<emphasis bold="true" pageId="0" pageNumber="97">Figure 7.</emphasis>
<taxonomicName authorityName="H. Y. Zhu &amp; X. L. Fan" authorityYear="2020" class="Dothideomycetes" family="Cyclothyriellaceae" genus="Cytospora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cytospora spiraeicola" order="Pleosporales" pageId="0" pageNumber="97" phylum="Ascomycota" rank="species" species="spiraeicola">
<emphasis italics="true" pageId="0" pageNumber="97">Cytospora spiraeicola</emphasis>
</taxonomicName>
from
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Spiraea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Spiraea salicifolia" order="Rosales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="salicifolia">
<emphasis italics="true" pageId="0" pageNumber="97">Spiraea salicifolia</emphasis>
</taxonomicName>
(CF 2019803).
<emphasis bold="true" pageId="0" pageNumber="97">A, B</emphasis>
habit of ascomata on twig
<emphasis bold="true" pageId="0" pageNumber="97">C</emphasis>
transverse section of ascoma
<emphasis bold="true" pageId="0" pageNumber="97">D</emphasis>
longitudinal section through ascoma
<emphasis bold="true" pageId="0" pageNumber="97">E</emphasis>
asci and ascospores
<emphasis bold="true" pageId="0" pageNumber="97">F, G</emphasis>
ascus
<emphasis bold="true" pageId="0" pageNumber="97">H</emphasis>
ascospores
<emphasis bold="true" pageId="0" pageNumber="97">I</emphasis>
colonies on PDA at 3 days (left) and 30 days (right). Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="97">A, B</emphasis>
); 500
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="97">C, D</emphasis>
); 10
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="97">E-H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="97" type="description">
<paragraph pageId="0" pageNumber="97">Culture characteristics.</paragraph>
<paragraph pageId="0" pageNumber="97">
<emphasis italics="true" pageId="0" pageNumber="97">Cultures</emphasis>
are white, growing up to 4 cm in diam. with irregular margin after 3 days, covering the 9 cm Petri dish after 6 days, becoming vinaceous buff to hazel after 7-10 days. In reverse, the cultures are the same as the upper colour after 3 days, becoming isabelline to umber after 7-10 days.
<emphasis italics="true" pageId="0" pageNumber="97">Colonies</emphasis>
are felty with a heterogeneous texture, lacking aerial mycelium.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="97" type="habitat">
<paragraph pageId="0" pageNumber="97">Habitat and distribution.</paragraph>
<paragraph pageId="0" pageNumber="97">
Known from
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Spiraea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Spiraea salicifolia" order="Rosales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="salicifolia">
<emphasis italics="true" pageId="0" pageNumber="97">Spiraea salicifolia</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Rehder &amp; E. H. Wilson" authorityYear="1915" class="Magnoliopsida" family="Malvaceae" genus="Tilia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tilia nobilis" order="Malvales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="nobilis">
<emphasis italics="true" pageId="0" pageNumber="97">Tilia nobilis</emphasis>
</taxonomicName>
in Mount Dongling, China.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="97" type="materials_examined">
<paragraph pageId="0" pageNumber="97">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="97">
China, Beijing City, Mentougou District, Mount Dongling, Xiaolongmen Forestry Centre (
<geoCoordinate degrees="115" direction="east" minutes="29" orientation="longitude" precision="1" seconds="20.49" value="115.48902">115°29'20.49&quot;E</geoCoordinate>
,
<geoCoordinate degrees="39" direction="north" minutes="57" orientation="latitude" precision="1" seconds="47.43" value="39.963177">39°57'47.43&quot;N</geoCoordinate>
), from branches of
<taxonomicName authorityName="Rehder &amp; E. H. Wilson" authorityYear="1915" class="Magnoliopsida" family="Malvaceae" genus="Tilia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tilia nobilis" order="Malvales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="nobilis">
<emphasis italics="true" pageId="0" pageNumber="97">Tilia nobilis</emphasis>
</taxonomicName>
, 17 Aug 2017, H.Y. Zhu &amp; X.L. Fan, CF 2019804, living culture CFCC 53139.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="97" type="notes">
<paragraph pageId="0" pageNumber="97">Notes.</paragraph>
<paragraph pageId="0" pageNumber="97">
<taxonomicName authorityName="H. Y. Zhu &amp; X. L. Fan" authorityYear="2020" class="Dothideomycetes" family="Cyclothyriellaceae" genus="Cytospora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cytospora spiraeicola" order="Pleosporales" pageId="0" pageNumber="97" phylum="Ascomycota" rank="species" species="spiraeicola">
<emphasis italics="true" pageId="0" pageNumber="97">Cytospora spiraeicola</emphasis>
</taxonomicName>
is associated with canker disease of
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Spiraea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Spiraea salicifolia" order="Rosales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="salicifolia">
<emphasis italics="true" pageId="0" pageNumber="97">Spiraea salicifolia</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Rehder &amp; E. H. Wilson" authorityYear="1915" class="Magnoliopsida" family="Malvaceae" genus="Tilia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tilia nobilis" order="Malvales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="species" species="nobilis">
<emphasis italics="true" pageId="0" pageNumber="97">Tilia nobilis</emphasis>
</taxonomicName>
in China, with characteristics similar to
<taxonomicName class="Sordariomycetes" family="Valsaceae" genus="Cytospora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cytospora elaeagnicola" order="Diaporthales" pageId="0" pageNumber="97" phylum="Ascomycota" rank="species" species="elaeagnicola">
<emphasis italics="true" pageId="0" pageNumber="97">Cytospora elaeagnicola</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="C. spiraeae" pageId="0" pageNumber="97" rank="species" species="spiraeae">
<emphasis italics="true" pageId="0" pageNumber="97">C. spiraeae</emphasis>
</taxonomicName>
in phylogram (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F7" captionText="Figure 2. Phylogram of Cytospora, based on combined ITS, LSU, act, rpb 2, tef 1 - α and tub 2 genes. The MP and ML bootstrap support values above 50 % are shown at the first and second positions, respectively. Thickened branches represent posterior probabilities above 0.95 from the BI. Ex-type strains are in bold. Strains from the current study are in blue." figureDoi="10.3897/mycokeys.62.47854.figure2" httpUri="https://binary.pensoft.net/fig/375794" pageId="0" pageNumber="97">2</figureCitation>
). Morphologically, it differs from
<taxonomicName lsidName="C. spiraeae" pageId="0" pageNumber="97" rank="species" species="spiraeae">
<emphasis italics="true" pageId="0" pageNumber="97">C. spiraeae</emphasis>
</taxonomicName>
by the smaller perithecia (210-250 vs. 270-400
<normalizedToken originalValue="µm">µm</normalizedToken>
in diam.) and longer ascospores (8.5-12
<normalizedToken originalValue="×">x</normalizedToken>
2.5-3.5 vs. 7-8
<normalizedToken originalValue="×">x</normalizedToken>
2-2.5
<normalizedToken originalValue="µm">µm</normalizedToken>
) (
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.338.1.4" author="Zhu, HY" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="97" pagination="49 - 62" refId="B50" refString="Zhu, HY, Tian, CM, Fan, XL, 2018a. Multigene phylogeny and morphology reveal Cytospora spiraeae sp. nov. (Diaporthales, Ascomycota) in China. Phytotaxa 338: 49 - 62, DOI: https://doi.org/10.11646/phytotaxa.338.1.4" title="Multigene phylogeny and morphology reveal Cytospora spiraeae sp. nov. (Diaporthales, Ascomycota) in China." url="https://doi.org/10.11646/phytotaxa.338.1.4" volume="338" year="2018 a">Zhu et al. 2018a</bibRefCitation>
). Phylogenetically,
<taxonomicName lsidName="C. spiraeicola" pageId="0" pageNumber="97" rank="species" species="spiraeicola">
<emphasis italics="true" pageId="0" pageNumber="97">C. spiraeicola</emphasis>
</taxonomicName>
(CFCC 53138) differs from
<taxonomicName lsidName="C. elaeagnicola" pageId="0" pageNumber="97" rank="species" species="elaeagnicola">
<emphasis italics="true" pageId="0" pageNumber="97">C. elaeagnicola</emphasis>
</taxonomicName>
(CFCC 52882) by ITS (8/665),
<emphasis italics="true" pageId="0" pageNumber="97">rpb2</emphasis>
(44/730),
<emphasis italics="true" pageId="0" pageNumber="97">tef1-α</emphasis>
(75/771) and
<emphasis italics="true" pageId="0" pageNumber="97">tub2</emphasis>
(42/624) and
<taxonomicName lsidName="C. spiraeae" pageId="0" pageNumber="97" rank="species" species="spiraeae">
<emphasis italics="true" pageId="0" pageNumber="97">C. spiraeae</emphasis>
</taxonomicName>
(CFCC 50049) by ITS (4/665),
<emphasis italics="true" pageId="0" pageNumber="97">rpb2</emphasis>
(38/730),
<emphasis italics="true" pageId="0" pageNumber="97">tef1-α</emphasis>
(63/771) and
<emphasis italics="true" pageId="0" pageNumber="97">tub2</emphasis>
(44/624) (
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.338.1.4" author="Zhu, HY" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="97" pagination="49 - 62" refId="B50" refString="Zhu, HY, Tian, CM, Fan, XL, 2018a. Multigene phylogeny and morphology reveal Cytospora spiraeae sp. nov. (Diaporthales, Ascomycota) in China. Phytotaxa 338: 49 - 62, DOI: https://doi.org/10.11646/phytotaxa.338.1.4" title="Multigene phylogeny and morphology reveal Cytospora spiraeae sp. nov. (Diaporthales, Ascomycota) in China." url="https://doi.org/10.11646/phytotaxa.338.1.4" volume="338" year="2018 a">Zhu et al. 2018a</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.1080/12298093.2019.1633902" author="Zhang, LX" journalOrPublisher="Mycobiology" pageId="0" pageNumber="97" pagination="319 - 328" refId="B52" refString="Zhang, LX, Alvarez, LV, Bonthond, G, Tian, CM, Fan, XL, 2019. Cytospora elaeagnicola sp. nov. associated with narrow-leaved oleaster canker disease in China. Mycobiology 47: 319 - 328, DOI: https://doi.org/10.1080/12298093.2019.1633902" title="Cytospora elaeagnicola sp. nov. associated with narrow-leaved oleaster canker disease in China." url="https://doi.org/10.1080/12298093.2019.1633902" volume="47" year="2019">Zhang et al. 2019</bibRefCitation>
). Therefore, we describe it as a novel species.
</paragraph>
</subSubSection>
</treatment>
</document>