treatments-xml/data/03/B0/87/03B087E55B56FFA0FF79F099FE4CF84D.xml
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<document id="EA1162FE9BB32F63A5BDD6FA3C7247A6" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B56FFA0FF79F099FE4CF84D" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena huonica Perkins 2011, new species" docType="treatment" docVersion="6" lastPageNumber="52" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="52" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="71825E2C307A35DB1FBE3F3E03EF4811">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="C2A711ADF301F2FF027005B2AC95753A">Perkins, Philip D.</mods:namePart>
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<treatment id="03B087E55B56FFA0FF79F099FE4CF84D" ID-DOI="http://doi.org/10.5281/zenodo.5302722" ID-Zenodo-Dep="5302722" LSID="urn:lsid:plazi:treatment:03B087E55B56FFA0FF79F099FE4CF84D" httpUri="http://treatment.plazi.org/id/03B087E55B56FFA0FF79F099FE4CF84D" lastPageNumber="52" pageId="51" pageNumber="52">
<subSubSection id="C30365785B56FFA0FF79F099FDAEFD5B" box="[151,536,691,718]" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="8BA636F35B56FFA0FF79F099FDAEFD5B" blockId="51.[151,536,691,750]" box="[151,536,691,718]" pageId="51" pageNumber="52">
<heading id="D0EE819F5B56FFA0FF79F099FDAEFD5B" bold="true" box="[151,536,691,718]" fontSize="11" level="1" pageId="51" pageNumber="52" reason="1">
<emphasis id="B96DEAE15B56FFA0FF79F099FDAEFD5B" bold="true" box="[151,536,691,718]" pageId="51" pageNumber="52">
<taxonomicName id="4C194D705B56FFA0FF79F099FECFFD58" ID-CoL="3N3B3" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,377,692,718]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="huonica" status="sp. nov.">
<emphasis id="B96DEAE15B56FFA0FF79F099FECFFD58" bold="true" box="[151,377,692,718]" italics="true" pageId="51" pageNumber="52">Hydraena huonica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B56FFA0FE67F09EFDAEFD5B" box="[393,536,691,717]" pageId="51" pageNumber="52" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30365785B56FFA0FF79F0FBFED0FD78" box="[151,358,726,750]" pageId="51" pageNumber="52" type="description">
<paragraph id="8BA636F35B56FFA0FF79F0FBFED0FD78" blockId="51.[151,536,691,750]" box="[151,358,726,750]" pageId="51" pageNumber="52">
(
<figureCitation id="13222A765B56FFA0FF71F0FBFF4BFD78" box="[159,253,726,750]" captionStart="FIGURE 71" captionStartId="198.[151,250,1753,1776]" captionTargetBox="[207,1379,193,1732]" captionTargetId="figure-29@198.[207,1379,193,1732]" captionTargetPageId="198" captionText="FIGURE 71. Hydraena huonica, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283535" httpUri="https://zenodo.org/record/5283535/files/figure.png" pageId="51" pageNumber="52">Figs. 71</figureCitation>
,
<figureCitation id="13222A765B56FFA0FEE6F0FBFE90FD78" box="[264,294,726,750]" captionStart="FIGURES 7273" captionStartId="199.[151,264,1618,1641]" captionTargetBox="[249,1341,205,1584]" captionTargetId="figure-29@199.[226,1360,193,1597]" captionTargetPageId="199" captionText="FIGURES 7273. Aedeagi of holotypes. 72. H. tricosipes. 73. H. huonica." figureDoi="http://doi.org/10.5281/zenodo.5283539" httpUri="https://zenodo.org/record/5283539/files/figure.png" pageId="51" pageNumber="52">73</figureCitation>
,
<figureCitation id="13222A765B56FFA0FEDCF0FBFEE8FD78" box="[306,350,726,750]" captionStart="FIGURES 467470" captionStartId="392.[151,264,1650,1673]" captionTargetBox="[165,1421,238,1599]" captionTargetId="figure-23@392.[165,1421,193,1629]" captionTargetPageId="392" captionText="FIGURES 467470. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5295735" httpUri="https://zenodo.org/record/5295735/files/figure.png" pageId="51" pageNumber="52">467</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B56FFA0FF79F132FD9FFCE8" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="8BA636F35B56FFA0FF79F132FD9FFCE8" blockId="51.[151,1437,798,2011]" pageId="51" pageNumber="52">
Type Material.
<materialsCitation id="3B713CAE5B56FFA0FEB7F132FAA6FCCC" collectingDate="1975-07-01" collectingDateMax="1975-07-30" collectingDateMin="1975-07-01" collectionCode="NMNZ" collectorName="R. W. Hornabrook" county="Kewensa" elevation="300" latitude="-5.8166666" location="Yupna" longLatPrecision="1305" longitude="146.76666" municipality="Finisterre Range" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<typeStatus id="54A288515B56FFA0FEB7F132FE70FCA1" box="[345,454,799,823]" pageId="51" pageNumber="52" type="holotype">
<emphasis id="B96DEAE15B56FFA0FEB7F132FE70FCA1" bold="true" box="[345,454,799,823]" pageId="51" pageNumber="52">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B56FFA0FDD2F132FCA8FCA1" bold="true" box="[572,798,798,823]" pageId="51" pageNumber="52">
<collectingRegion id="49DDF8115B56FFA0FDD2F132FCAEFCA1" box="[572,792,798,823]" country="Papua New Guinea" name="Madang" pageId="51" pageNumber="52">Madang Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62C74E7F5B56FFA0FCD8F132FC16FCA0" box="[822,928,799,822]" pageId="51" pageNumber="52">Kewensa</collectingCounty>
,
<collectingMunicipality id="6BC2AC895B56FFA0FC5FF133FBC4FCA1" box="[945,1138,798,823]" pageId="51" pageNumber="52">Finisterre Range</collectingMunicipality>
,
<location id="8EC660285B56FFA0FB6DF132FB7BFCA0" LSID="urn:lsid:plazi:treatment:03B087E55B56FFA0FF79F099FE4CF84D:8EC660285B56FFA0FB6DF132FB7BFCA0" box="[1155,1229,799,822]" county="Kewensa" latitude="-5.8166666" longLatPrecision="1305" longitude="146.76666" municipality="Finisterre Range" name="Yupna" pageId="51" pageNumber="52" stateProvince="Madang">Yupna</location>
,
<location id="8EC660285B56FFA0FB30F132FA21FCA0" LSID="urn:lsid:plazi:treatment:03B087E55B56FFA0FF79F099FE4CF84D:8EC660285B56FFA0FB30F132FA21FCA0" box="[1246,1431,798,822]" county="Kewensa" latitude="-5.8166666" longLatPrecision="1305" longitude="146.76666" municipality="Finisterre Range" name="Huon Peninsula" pageId="51" pageNumber="52" stateProvince="Madang">Huon Peninsula</location>
, stream margin, [GE est.]
<quantity id="4CE19B165B56FFA0FE5CF16FFD85FCCC" box="[434,563,834,858]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="51" pageNumber="52" unit="m" value="300.0" valueMax="500.0" valueMin="100.0">
<elevation id="0034D1C05B56FFA0FE5CF16FFD85FCCC" box="[434,563,834,858]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="51" pageNumber="52" unit="m" value="300.0" valueMax="500.0" valueMin="100.0">100500 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B56FFA0FDD1F16FFD2EFCCC" box="[575,664,834,858]" degrees="5" direction="south" minutes="49" orientation="latitude" pageId="51" pageNumber="52" precision="925" value="-5.8166666">5° 49' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B56FFA0FD4BF16FFCAAFCCC" box="[677,796,834,858]" degrees="146" direction="east" minutes="46" orientation="longitude" pageId="51" pageNumber="52" precision="925" value="146.76666">146° 46' E</geoCoordinate>
,
<date id="FFA710335B56FFA0FCC7F16FFC70FCCC" box="[809,966,834,858]" pageId="51" pageNumber="52" value="1975-07-01" valueMax="1975-07-30" valueMin="1975-07-01">
<collectingDate id="EFE3E9DB5B56FFA0FCC7F16FFC70FCCC" box="[809,966,834,858]" pageId="51" pageNumber="52" value="1975-07-01" valueMax="1975-07-30" valueMin="1975-07-01">130 vii 1975</collectingDate>
</date>
,
<collectorName id="26EC53255B56FFA0FC3CF16EFB16FCCC" box="[978,1184,834,858]" pageId="51" pageNumber="52">R. W. Hornabrook</collectorName>
(
<collectionCode id="ED08AE365B56FFA0FB41F16FFABEFCCC" box="[1199,1288,834,858]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="51" pageNumber="52" type="Museum">NMNZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5B56FFA0FAF5F16EFD93FCE8" collectingDate="1975-07-01" collectingDateMax="1975-07-30" collectingDateMin="1975-07-01" collectionCode="NMNZ" collectorName="R. W. Hornabrook" county="Kewensa" elevation="300" latitude="-5.8166666" location="Yupna" longLatPrecision="1305" longitude="146.76666" municipality="Finisterre Range" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<emphasis id="B96DEAE15B56FFA0FAF5F16EFA2AFCCD" bold="true" box="[1307,1436,835,859]" pageId="51" pageNumber="52">
<typeStatus id="54A288515B56FFA0FAF5F16EFA21FCCD" box="[1307,1431,835,859]" pageId="51" pageNumber="52" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same data as holotype (12
<collectionCode id="ED08AE365B56FFA0FE2BF14BFDABFCE8" box="[453,541,870,894]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="51" pageNumber="52" type="Museum">NMNZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B56FFA0FF29F1A7FBDEFBA4" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="8BA636F35B56FFA0FF29F1A7FBDEFBA4" blockId="51.[151,1437,798,2011]" pageId="51" pageNumber="52">
<emphasis id="B96DEAE15B56FFA0FF29F1A7FE64FC35" bold="true" box="[199,466,906,931]" pageId="51" pageNumber="52">Differential Diagnosis.</emphasis>
Similar to
<taxonomicName id="4C194D705B56FFA0FDA1F1A6FD5BFC35" authorityName="Perkins" authorityYear="2011" box="[591,749,907,931]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="tarsotricha">
<emphasis id="B96DEAE15B56FFA0FDA1F1A6FDD5FC34" box="[591,611,907,930]" italics="true" pageId="51" pageNumber="52">H</emphasis>
.
<emphasis id="B96DEAE15B56FFA0FD9FF1A6FD5BFC35" box="[625,749,907,931]" italics="true" pageId="51" pageNumber="52">tarsotricha</emphasis>
</taxonomicName>
in habitus and male sexual dimorphisms (no tubercle on first abdominal ventrite, fifth ventrite with transverse ridge, meso- and metatarsi with long hair-like setae) (
<figureCitation id="13222A765B56FFA0FAD9F183FA21FC50" box="[1335,1431,942,966]" captionStart="FIGURE 67" captionStartId="195.[151,250,1709,1732]" captionTargetBox="[209,1386,214,1676]" captionTargetId="figure-26@195.[186,1401,193,1688]" captionTargetPageId="195" captionText="FIGURE 67. Hydraena tarsotricha, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283528" httpUri="https://zenodo.org/record/5283528/files/figure.png" pageId="51" pageNumber="52">Figs. 67</figureCitation>
,
<figureCitation id="13222A765B56FFA0FF79F1FFFF00FC7C" box="[151,182,978,1002]" captionStart="FIGURE 71" captionStartId="198.[151,250,1753,1776]" captionTargetBox="[207,1379,193,1732]" captionTargetId="figure-29@198.[207,1379,193,1732]" captionTargetPageId="198" captionText="FIGURE 71. Hydraena huonica, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283535" httpUri="https://zenodo.org/record/5283535/files/figure.png" pageId="51" pageNumber="52">71</figureCitation>
); differing therefrom by the narrower mesoventral intercoxal process P2 (ratios ca 2/4/13/1 vs. 3/4/13/1). The male genitalia are also quite similar, differing most notably in the shapes of the parameres, and in details of the distal piece; reliable determinations will require examination of the aedeagi (
<figureCitation id="13222A765B56FFA0FC3CF637FB86FBA4" box="[978,1072,1050,1074]" captionStart="FIGURES 6869" captionStartId="196.[151,264,1650,1673]" captionText="FIGURES 6869. Aedeagi of holotypes. 68. H. tetana. 69. H. tarsotricha." pageId="51" pageNumber="52">Figs. 69</figureCitation>
,
<figureCitation id="13222A765B56FFA0FBD2F637FBEAFBA4" box="[1084,1116,1050,1074]" captionStart="FIGURES 7273" captionStartId="199.[151,264,1618,1641]" captionTargetBox="[249,1341,205,1584]" captionTargetId="figure-29@199.[226,1360,193,1597]" captionTargetPageId="199" captionText="FIGURES 7273. Aedeagi of holotypes. 72. H. tricosipes. 73. H. huonica." figureDoi="http://doi.org/10.5281/zenodo.5283539" httpUri="https://zenodo.org/record/5283539/files/figure.png" pageId="51" pageNumber="52">73</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B56FFA0FF29F613FDC7F8F8" pageId="51" pageNumber="52" type="description">
<paragraph id="8BA636F35B56FFA0FF29F613FAC5FBEC" blockId="51.[151,1437,798,2011]" pageId="51" pageNumber="52">
<emphasis id="B96DEAE15B56FFA0FF29F613FEEEFBC1" bold="true" box="[199,344,1086,1111]" inLineHeading="true" pageId="51" pageNumber="52" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B56FFA0FE42F613FDB8FBC0" box="[428,526,1086,1110]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.73/0.70; head 0.28/0.40; pronotum 0.41/0.49, PA 0.43, PB 0.43; elytra 1.07/0.70. Dorsum piceous; legs and maxillary palpi dark brown.
</paragraph>
<paragraph id="8BA636F35B56FFA0FF29F6AAFEE8FAC4" blockId="51.[151,1437,798,2011]" pageId="51" pageNumber="52">Frons punctures ca. 11.5xef, slightly larger near eyes than medially; interstices shining, 15xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum very sparsely very finely punctulate, shining; postmentum densely micropunctulate, dull. Genae raised, shining, without posterior ridge. Pronotum cordiform, median 2/3 of anterior margin emarginate; punctures on disc ca. 1xpd those of frons, interstices shining, 2 4xpd, punctures slightly larger and denser at anterior and posterior; PF1 absent; PF2 very shallow, obsolete; PF3 deep; PF4 absent.</paragraph>
<paragraph id="8BA636F35B56FFA0FF29F773FF47F990" blockId="51.[151,1437,798,2011]" pageId="51" pageNumber="52">Elytra with summit of posterior declivity slightly past midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 12xpd, as are interstices between punctures of a row, a few punctures subserial. Apices in dorsal aspect separately rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BA636F35B56FFA0FF29F43FFDC7F8F8" blockId="51.[151,1437,798,2011]" pageId="51" pageNumber="52">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 2/4/13/1. P1 laminate; median carina sinuate in profile. P2 raised, l/w ca. 3/1, sides converging toward blunt apex. Plaques very large, very narrowly separated, slightly more narrowly separated posteriorly than anteriorly, weakly raised, slightly slanting toward midline. Metaventrite without midlongitudinal ridge. AIS width at straight posterior margin ca. 2x P2. All legs long and very slender. Profemur (male) with small, sharply pointed tubercle next to trochanter; protibia straight or very slightly arcuate, distal 1/3 widened on anterior surface, cluster of short spines on medial surface at apex, moderately long spine on medial surface at base of widened area. Mesotibia very slightly arcuate. Metatibia straight, medial margin in distal 1/2 very slightly emarginate. Meso- and metatarsi with long hair-like setae along lower margin of basal tarsomeres. Fifth abdominal ventrite posteriorly transversely raised. Abdominal apex symmetrical; last tergite (male) deeply notched. Aedeagus as illustrated (
<figureCitation id="13222A765B56FFA0FDFFF57BFDD3F8F8" box="[529,613,1878,1902]" captionStart="FIGURES 7273" captionStartId="199.[151,264,1618,1641]" captionTargetBox="[249,1341,205,1584]" captionTargetId="figure-29@199.[226,1360,193,1597]" captionTargetPageId="199" captionText="FIGURES 7273. Aedeagi of holotypes. 72. H. tricosipes. 73. H. huonica." figureDoi="http://doi.org/10.5281/zenodo.5283539" httpUri="https://zenodo.org/record/5283539/files/figure.png" pageId="51" pageNumber="52">Fig. 73</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B56FFA0FF29F556FC81F804" box="[199,823,1914,1939]" pageId="51" pageNumber="52" type="etymology">
<paragraph id="8BA636F35B56FFA0FF29F556FC81F804" blockId="51.[151,1437,798,2011]" box="[199,823,1914,1939]" pageId="51" pageNumber="52">
<emphasis id="B96DEAE15B56FFA0FF29F556FEFBF805" bold="true" box="[199,333,1915,1939]" inLineHeading="true" pageId="51" pageNumber="52" reason="1">Etymology.</emphasis>
Named in reference to the Huon Peninsula.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B56FFA0FF29F5B3FE4CF84D" pageId="51" pageNumber="52" type="distribution">
<paragraph id="8BA636F35B56FFA0FF29F5B3FE4CF84D" blockId="51.[151,1437,798,2011]" pageId="51" pageNumber="52">
<emphasis id="B96DEAE15B56FFA0FF29F5B3FED6F821" bold="true" box="[199,352,1950,1975]" inLineHeading="true" pageId="51" pageNumber="52" reason="1">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54A288515B56FFA0FD31F5B2FCB9F821" box="[735,783,1951,1975]" pageId="51" pageNumber="52">type</typeStatus>
locality on the Huon Peninsula (Area 5); elevation estimated at
<quantity id="4CE19B165B56FFA0FF13F5EFFECAF84C" box="[253,380,1986,2010]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="51" pageNumber="52" unit="m" value="300.0" valueMax="500.0" valueMin="100.0">100500 m</quantity>
(
<figureCitation id="13222A765B56FFA0FE65F5EFFE5DF84C" box="[395,491,1986,2010]" captionStart="FIGURES 467470" captionStartId="392.[151,264,1650,1673]" captionTargetBox="[165,1421,238,1599]" captionTargetId="figure-23@392.[165,1421,193,1629]" captionTargetPageId="392" captionText="FIGURES 467470. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5295735" httpUri="https://zenodo.org/record/5295735/files/figure.png" pageId="51" pageNumber="52">Fig. 467</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>