249 lines
30 KiB
XML
249 lines
30 KiB
XML
<document id="3C7835D7B4333034049FCD87B81F0A0D" ID-CLB-Dataset="20925" ID-DOI="10.11646/zootaxa.5093.4.2" ID-GBIF-Dataset="d3abaf4f-14fd-4a8b-9362-357792cc0cbf" ID-ISSN="1175-5326" ID-Zenodo-Dep="5912678" ID-ZooBank="440E7DD2-5736-49BB-A59E-9FA494402544" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643354187259" checkinUser="plazi" docAuthor="Daniel, Gimo M., Strümpher, Werner P. & Deschodt, Christian M." docDate="2022" docId="03B087B9FFB1FFBE7DD552C96E8BFA69" docLanguage="en" docName="zootaxa.5093.4.2.pdf" docOrigin="Zootaxa 5093 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Endroedyolini Davis, Deschodt & Scholtz 2019" docType="key" docVersion="5" lastPageNumber="419" masterDocId="FF89FFC1FFB4FFBB7D4253136B17FFE9" masterDocTitle="An unexpected new flightless dung beetle species (Coleoptera: Scarabaeidae: Scarabaeinae: Endroedyolini) from the Cederberg Mountains, South Africa" masterLastPageNumber="420" masterPageNumber="414" pageNumber="419" updateTime="1699304596146" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
|
<mods:mods id="E3C8309D886C209CD9AC206398B03640" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="F4CA3CE0646C40A748E50024EEF31FBB">
|
|
<mods:title id="F706F3A24F63321C5F458311EE66ACC4">An unexpected new flightless dung beetle species (Coleoptera: Scarabaeidae: Scarabaeinae: Endroedyolini) from the Cederberg Mountains, South Africa</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="4CFC314EBBC4C4B61E7AD421F10E6468" type="personal">
|
|
<mods:role id="7071FCAC9CE776053498C91E1BDED6EF">
|
|
<mods:roleTerm id="B9D4121340E21BC2937A9835740924E3">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="6C777683B332969F3BB47028A118FB72">Daniel, Gimo M.</mods:namePart>
|
|
<mods:nameIdentifier id="779A8F78AD064711693223AE9B91AC26" type="ORCID">0000-0003-3602-5034</mods:nameIdentifier>
|
|
<mods:affiliation id="9A953D93203F00B631E2ED99D47CFD52">Department of Terrestrial Invertebrates, National Museum, 36 Aliwal Street, Bloemfontein, South Africa. & Department of Biological & Environmental Sciences, Walter Sisulu University, Mthatha, South Africa. gimo. daniel @ nasmus. co. za; https: // orcid. org / 0000 - 0003 - 3602 - 5034</mods:affiliation>
|
|
<mods:nameIdentifier id="609C616E3A726441911F90E2F62FEFAE" type="email">gimo.daniel@nasmus.co.za</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name id="B6D503455581DB1FCCA7BCBC3B05FCF6" type="personal">
|
|
<mods:role id="B5DE0F30ABA0EB2D1A4BAA2C7B15428C">
|
|
<mods:roleTerm id="95E5392DCCD43C3F6FBC5DEB7DEDA271">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="AAEA2536C439744C700CD67ED1EE241F">Strümpher, Werner P.</mods:namePart>
|
|
<mods:affiliation id="B4E2610A93B5027D7CD8B3C966DDC5FB">Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa.</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="C5C216862CCD0C7C0B27928D6DAE7F8D" type="personal">
|
|
<mods:role id="7DB59E9B55C3F4AE7E6F6B968BAA6B9E">
|
|
<mods:roleTerm id="63F8464EFA515B4AF65EC38A4370F0EA">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="B125A8473F7B0F6FAE495D9492844719">Deschodt, Christian M.</mods:namePart>
|
|
<mods:nameIdentifier id="F5ECF6DB526720A7C75C7071264AE513" type="ORCID">0000-0002-3886-1813</mods:nameIdentifier>
|
|
<mods:affiliation id="A780739B523C247B9900128037BC3CF7">Department of Zoology & Entomology, University of Pretoria, Private Bag X 20, Hatfield, 0028 South Africa. cdeschodt @ zoology. up. ac. za; https: // orcid. org / 0000 - 0002 - 3886 - 1813</mods:affiliation>
|
|
<mods:nameIdentifier id="C12C1AD9E55ECF12ED00AEEF1131A940" type="email">cdeschodt@zoology.up.ac.za</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:typeOfResource id="58EE3A9A029630EBDF66DB217E0195DB">text</mods:typeOfResource>
|
|
<mods:relatedItem id="3AF645F737014739741DD0DBE7DDBF47" type="host">
|
|
<mods:titleInfo id="5E6BB6E239CF0F7356CC26E9B228D8F9">
|
|
<mods:title id="4BE239CDEF24FA0EA3871F3C77AE0B96">Zootaxa</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="5C59AFA613B86B8D092B06AC2B7582CF">
|
|
<mods:date id="0DABCC26AAF6F649109AAFF0C199B1DE">2022</mods:date>
|
|
<mods:detail id="87C63777F1BF1A9ED6320869F1660D04" type="pubDate">
|
|
<mods:number id="D42523920817A66A0D1622DBD618B573">2022-01-28</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="374B36EBBC4C4006B63ED928B008FC8B" type="volume">
|
|
<mods:number id="924B8752E27644228F6453B0CC1DB261">5093</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="B090210C6B57E12EE02D59EAA3CDEAE8" type="issue">
|
|
<mods:number id="EC7A3F98FDED463BF6414D4E9C094435">4</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="A37AEEA42F4FCE82531A14B365E0BFBA" unit="page">
|
|
<mods:start id="88CC431A94A34DEE48CA0BAEE7A9EF77">414</mods:start>
|
|
<mods:end id="03F6BD59D33A066684FC3E9FC20A8F6D">420</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification id="6E9BAEF6833258ADD3EDD969F833A2DE">journal article</mods:classification>
|
|
<mods:identifier id="336CAF9E311FF7163729B414FF0ED425" type="CLB-Dataset">20925</mods:identifier>
|
|
<mods:identifier id="862045F00B963B62463F55CD7B4D252F" type="DOI">10.11646/zootaxa.5093.4.2</mods:identifier>
|
|
<mods:identifier id="AAD6EA76D1B34484B1BF9C004290F5DE" type="GBIF-Dataset">d3abaf4f-14fd-4a8b-9362-357792cc0cbf</mods:identifier>
|
|
<mods:identifier id="893F73FF67D7AC26AB34708BC81104DD" type="ISSN">1175-5326</mods:identifier>
|
|
<mods:identifier id="9678CE11104AF725E6ED9F34F98256BC" type="Zenodo-Dep">5912678</mods:identifier>
|
|
<mods:identifier id="F6729331A57D5F65E6BDE7F6B51DDE4C" type="ZooBank">440E7DD2-5736-49BB-A59E-9FA494402544</mods:identifier>
|
|
</mods:mods>
|
|
<subSection id="E2962F44FFB1FFBE7DD552C96E8BFA69" pageId="5" pageNumber="419" type="multiple">
|
|
<treatment id="03B087B9FFB1FFBE7DD552C96E8BFA69" ID-DOI="http://doi.org/10.5281/zenodo.5914082" ID-GBIF-Taxon="192700782" ID-Zenodo-Dep="5914082" LSID="urn:lsid:plazi:treatment:03B087B9FFB1FFBE7DD552C96E8BFA69" httpUri="http://treatment.plazi.org/id/03B087B9FFB1FFBE7DD552C96E8BFA69" lastPageNumber="419" pageId="5" pageNumber="419">
|
|
<subSubSection id="C3036524FFB1FFBE7DD552C96FDAFE1D" box="[151,1229,474,501]" pageId="5" pageNumber="419" type="nomenclature">
|
|
<paragraph id="8BA636AFFFB1FFBE7DD552C96FDAFE1D" blockId="5.[151,1229,474,501]" box="[151,1229,474,501]" pageId="5" pageNumber="419">
|
|
<heading id="D0EE81C3FFB1FFBE7DD552C96FDAFE1D" bold="true" box="[151,1229,474,501]" fontSize="11" level="1" pageId="5" pageNumber="419" reason="1">
|
|
<emphasis id="B96DEABDFFB1FFBE7DD552C96FDAFE1D" bold="true" box="[151,1229,474,501]" pageId="5" pageNumber="419">
|
|
Identification key to
|
|
<taxonomicName id="4C194D2CFFB1FFBE7CD952C96947FE1D" ID-CoL="628D9" authorityName="Davis, Deschodt & Scholtz" authorityYear="2019" box="[411,592,474,500]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="tribe" tribe="Endroedyolini">Endroedyolini</taxonomicName>
|
|
, adapted and modified from
|
|
<bibRefCitation id="EF884B5EFFB1FFBE7EF852C96FD0FE1D" author="Deschodt, C. M. & Sole, C. L. & Scholtz, C. H." box="[954,1223,474,501]" pageId="5" pageNumber="419" pagination="357 - 362" refId="ref4528" refString="Deschodt, C. M., Sole, C. L. & Scholtz, C. H. (2020) Upsa centennial, a new genus and species (Coleoptera: Scarabaeidae: Scarabaeinae: Endroedyolini) from the Southern Mistbelt forest in South Africa: celebrating 100 years of entomology at the University of Pretoria. Zootaxa 4728, 357 - 362. https: // doi. org / 10.11646 / zootaxa. 4728.3.4" type="journal article" year="2020">
|
|
Deschodt
|
|
<emphasis id="B96DEABDFFB1FFBE797752C86F7EFE1C" bold="true" box="[1077,1129,474,501]" italics="true" pageId="5" pageNumber="419">et al</emphasis>
|
|
. (2020)
|
|
</bibRefCitation>
|
|
.
|
|
</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3036524FFB1FFBE7DD5510A6E8BFA69" pageId="5" pageNumber="419" type="key">
|
|
<key id="FF88619EFFB1FFBE7DD5510A6E8BFA69" pageId="5" pageNumber="419">
|
|
<keyStep id="30ED2E0AFFB1FFBE7DD5510A6E8BFD6F" pageId="5" pageNumber="419">
|
|
<paragraph id="8BA636AFFFB1FFBE7DD5510A6E8BFDA2" blockId="5.[151,1437,536,1409]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD5510A6E8BFDA2" pageId="5" pageNumber="419">
|
|
1. Metaventrite anteriorly with deep fovea on each side of midline, pronotal disc on each side near posterior angle with short elevated carina ................................................
|
|
<taxonomicName id="4C194D2CFFB1FFBE7EF051256E8BFDA2" ID-CoL="39SQY" authority="Scholtz & Howden, 1987" authorityName="Scholtz & Howden" authorityYear="1987" box="[946,1436,566,588]" class="Insecta" family="Scarabaeidae" genus="Endroedyolus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="419" phylum="Arthropoda" rank="species" species="paradoxus">
|
|
<emphasis id="B96DEABDFFB1FFBE7EF051256FB5FDA2" box="[946,1186,566,587]" italics="true" pageId="5" pageNumber="419">Endroedyolus paradoxus</emphasis>
|
|
<bibRefCitation id="EF884B5EFFB1FFBE79EA51256E8BFDA2" author="Scholtz, C. H. & Howden, H. F." box="[1192,1436,566,588]" pageId="5" pageNumber="419" pagination="75 - 119" refId="ref4756" refString="Scholtz, C. H. & Howden, H. F. (1987) A revision of the African Cathonina (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the Entomological Society of Southern Africa, 50, 75 - 119." type="journal article" year="1987">Scholtz & Howden, 1987</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BA636AFFFB1FFBE7DD551476E8BFD6F" blockId="5.[151,1437,536,1409]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD551476E8BFD6F" pageId="5" pageNumber="419">- Metaventrite anteriorly without deep fovea on each side of midline, and pronotal disc with or without short elevated carina on each side near posterior angle........................................................................... 2</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30ED2E0AFFB1FFBE7DD5519D6E8BFD37" pageId="5" pageNumber="419">
|
|
<paragraph id="8BA636AFFFB1FFBE7DD5519D6E8BFD29" blockId="5.[151,1437,536,1409]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD5519D6E8BFD29" pageId="5" pageNumber="419">
|
|
2. Pronotal disc with short elevated carina on each side near posterior angle...................................................................................................
|
|
<taxonomicName id="4C194D2CFFB1FFBE7ED951B86E8BFD29" ID-CoL="BSNR" authority="Deschodt & Scholtz, 2008" authorityName="Deschodt & Scholtz" authorityYear="2008" box="[923,1436,683,705]" class="Insecta" family="Scarabaeidae" genus="Aliuscanthoniola" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="419" phylum="Arthropoda" rank="species" species="similaris">
|
|
<emphasis id="B96DEABDFFB1FFBE7ED951B86F8EFD29" box="[923,1177,683,704]" italics="true" pageId="5" pageNumber="419">Aliuscanthoniola similaris</emphasis>
|
|
<bibRefCitation id="EF884B5EFFB1FFBE79DD51B86E8BFD29" author="Deschodt, C. M. & Scholtz, C. H." box="[1183,1436,683,705]" pageId="5" pageNumber="419" pagination="91 - 106" refId="ref4469" refString="Deschodt, C. M. & Scholtz, C. H. (2008) Systematics of South African forest-endemic dung beetles: new genera and species of small Canthonini (Scarabaeidae: Scarabaeinae). African Entomology 16, 91 - 106. https: // doi. org / 10.4001 / 1021 - 3589 - 16.1.91" type="journal article" year="2008">Deschodt & Scholtz, 2008</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BA636AFFFB1FFBE7DD551DA6E8BFD37" blockId="5.[151,1437,536,1409]" box="[151,1436,712,734]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD551DA6E8BFD37" box="[151,1436,712,734]" pageId="5" pageNumber="419">- Pronotal disc without short, elevated carina on each side near posterior angle ...................................... 3</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30ED2E0AFFB1FFBE7DD551F56E8BFCF0" pageId="5" pageNumber="419">
|
|
<paragraph id="8BA636AFFFB1FFBE7DD551F56E8BFD12" blockId="5.[151,1437,536,1409]" box="[151,1436,742,764]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD551F56E8BFD12" box="[151,1436,742,764]" pageId="5" pageNumber="419">3. Mesoventrite medially wide, or wide enough for, more than three irregular rows of punctures......................... 4</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BA636AFFFB1FFBE7DD550176E8BFCF0" blockId="5.[151,1437,536,1409]" box="[151,1436,771,793]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD550176E8BFCF0" box="[151,1436,771,793]" pageId="5" pageNumber="419">- Mesoventrite medially narrow, with at most two irregular rows of punctures (punctures sometimes absent in this area)..... 5</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30ED2E0AFFB1FFBE7DD550326E8BFCBA" pageId="5" pageNumber="419">
|
|
<paragraph id="8BA636AFFFB1FFBE7DD550326E8BFCDF" blockId="5.[151,1437,536,1409]" box="[151,1436,800,822]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD550326E8BFCDF" box="[151,1436,800,822]" pageId="5" pageNumber="419">
|
|
4. Mesometaventral suture angulate at anterior apex..................
|
|
<taxonomicName id="4C194D2CFFB1FFBE7EE550336E8BFCDF" ID-CoL="4B9HD" authority="Scholtz & Howden, 1987" authorityName="Scholtz & Howden" authorityYear="1987" box="[935,1436,800,822]" class="Insecta" family="Scarabaeidae" genus="Outenikwanus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="419" phylum="Arthropoda" rank="species" species="tomentosus">
|
|
<emphasis id="B96DEABDFFB1FFBE7EE550336FB5FCDF" box="[935,1186,800,822]" italics="true" pageId="5" pageNumber="419">Outenikwanus tomentosus</emphasis>
|
|
<bibRefCitation id="EF884B5EFFB1FFBE79EA50336E8BFCDF" author="Scholtz, C. H. & Howden, H. F." box="[1192,1436,800,822]" pageId="5" pageNumber="419" pagination="75 - 119" refId="ref4756" refString="Scholtz, C. H. & Howden, H. F. (1987) A revision of the African Cathonina (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the Entomological Society of Southern Africa, 50, 75 - 119." type="journal article" year="1987">Scholtz & Howden, 1987</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BA636AFFFB1FFBE7DD5502D6E8BFCBA" blockId="5.[151,1437,536,1409]" box="[151,1436,830,852]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD5502D6E8BFCBA" box="[151,1436,830,852]" pageId="5" pageNumber="419">
|
|
- Mesometaventral suture rounded at anterior apex......................
|
|
<taxonomicName id="4C194D2CFFB1FFBE7E84502D6E8BFCBA" ID-CoL="7DM72" authority="Deschodt, Sole & Scholtz, 2020" authorityName="Deschodt, Sole & Scholtz" authorityYear="2020" box="[966,1436,830,852]" class="Insecta" family="Scarabaeidae" genus="Upsa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="419" phylum="Arthropoda" rank="species" species="centennial">
|
|
<emphasis id="B96DEABDFFB1FFBE7E84502D6F75FCBA" box="[966,1122,830,851]" italics="true" pageId="5" pageNumber="419">Upsa centennial</emphasis>
|
|
<bibRefCitation id="EF884B5EFFB1FFBE792A502D6E8BFCBA" author="Deschodt, C. M. & Sole, C. L. & Scholtz, C. H." box="[1128,1436,830,852]" pageId="5" pageNumber="419" pagination="357 - 362" refId="ref4528" refString="Deschodt, C. M., Sole, C. L. & Scholtz, C. H. (2020) Upsa centennial, a new genus and species (Coleoptera: Scarabaeidae: Scarabaeinae: Endroedyolini) from the Southern Mistbelt forest in South Africa: celebrating 100 years of entomology at the University of Pretoria. Zootaxa 4728, 357 - 362. https: // doi. org / 10.11646 / zootaxa. 4728.3.4" type="journal article" year="2020">Deschodt, Sole & Scholtz, 2020</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30ED2E0AFFB1FFBE7DD550486E8BFC67" pageId="5" pageNumber="419">
|
|
<paragraph id="8BA636AFFFB1FFBE7DD550486E8BFC98" blockId="5.[151,1437,536,1409]" box="[151,1436,859,881]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD550486E8BFC98" box="[151,1436,859,881]" pageId="5" pageNumber="419">5. Elytron in lateral view with depression on lateral side ......................................................... 6</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BA636AFFFB1FFBE7DD5506A6E8BFC67" blockId="5.[151,1437,536,1409]" box="[151,1436,888,910]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD5506A6E8BFC67" box="[151,1436,888,910]" pageId="5" pageNumber="419">- Elytron in lateral view without depression on posterolateral side ................................................. 7</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30ED2E0AFFB1FFBE7DD550856E8AFBED" pageId="5" pageNumber="419">
|
|
<paragraph id="8BA636AFFFB1FFBE7DD550856E8BFC21" blockId="5.[151,1437,536,1409]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD550856E8BFC21" pageId="5" pageNumber="419">
|
|
6. Pronotal depression on mid-basal edge present, mesometaventral suture rounded; lateral elytral spine in males present..................................................................
|
|
<taxonomicName id="4C194D2CFFB1FFBE7EFC50A06E8BFC21" ID-CoL="6YPVR" authority="Roets & Oberlander 2010" authorityName="Roets & Oberlander" authorityYear="2010" box="[958,1436,947,969]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="419" phylum="Arthropoda" rank="species" species="oubosiensis">
|
|
<emphasis id="B96DEABDFFB1FFBE7EFC50A06F88FC21" box="[958,1183,947,968]" italics="true" pageId="5" pageNumber="419">Silvaphilus oubosiensis</emphasis>
|
|
<bibRefCitation id="EF884B5EFFB1FFBE79E750A76E8BFC21" author="Roets, F. & Oberlander, K. C." box="[1189,1436,947,969]" pageId="5" pageNumber="419" pagination="369 - 373" refId="ref4697" refString="Roets, F. & Oberlander, K. C. (2010) Silvaphilus: A new relict forest-endemic Canthonini dung beetle genus from the Western Cape Province of South Africa (Coleoptera: Scarabaeidae: Scarabaeinae). African Entomology, 18, 369 - 373. https: // doi. org / 10.4001 / 003.018.0213" type="journal article" year="2010">Roets & Oberlander 2010</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BA636AFFFB1FFBE7DD550C26E8AFBED" blockId="5.[151,1437,536,1409]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD550C26E8AFBED" pageId="5" pageNumber="419">
|
|
- Pronotal depression on mid-basal edge absent, mesometaventral suture curving very slightly anteriorly; lateral elytral spine in males absent .......................................
|
|
<taxonomicName id="4C194D2CFFB1FFBE7E6850FD6E36FBED" ID-CoL="8V23B" authority="Daniel, Strumpher & Deschodt" authorityName="Daniel, Strumpher & Deschodt" authorityYear="2022" box="[810,1313,1006,1028]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="419" phylum="Arthropoda" rank="species" species="joselmae" status="sp. nov.">
|
|
<emphasis id="B96DEABDFFB1FFBE7E6850FD68F8FBEA" box="[810,1007,1006,1027]" italics="true" pageId="5" pageNumber="419">Silvaphilus joselmae</emphasis>
|
|
Daniel, Strümpher & Deschodt
|
|
</taxonomicName>
|
|
<emphasis id="B96DEABDFFB1FFBE786450FD6E8AFBED" bold="true" box="[1318,1437,1006,1028]" pageId="5" pageNumber="419">
|
|
<taxonomicNameLabel id="A25E57C6FFB1FFBE786450FD6E8AFBED" box="[1318,1437,1006,1028]" pageId="5" pageNumber="419" rank="species">new species</taxonomicNameLabel>
|
|
</emphasis>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30ED2E0AFFB1FFBE7DD5571F6E8BFBD7" pageId="5" pageNumber="419">
|
|
<paragraph id="8BA636AFFFB1FFBE7DD5571F6E8BFBC9" blockId="5.[151,1437,536,1409]" box="[151,1436,1035,1057]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD5571F6E8BFBC9" box="[151,1436,1035,1057]" pageId="5" pageNumber="419">7. Mesometaventral suture rounded at anterior apex............................................................ 8</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BA636AFFFB1FFBE7DD5573A6E8BFBD7" blockId="5.[151,1437,536,1409]" box="[151,1436,1064,1086]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD5573A6E8BFBD7" box="[151,1436,1064,1086]" pageId="5" pageNumber="419">- Mesometaventral suture angulate at anterior apex........................................................... 10</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30ED2E0AFFB1FFBE7DD557556E8BFB91" pageId="5" pageNumber="419">
|
|
<paragraph id="8BA636AFFFB1FFBE7DD557556E8BFBB2" blockId="5.[151,1437,536,1409]" box="[151,1436,1094,1116]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD557556E8BFBB2" box="[151,1436,1094,1116]" pageId="5" pageNumber="419">
|
|
8. Protibial teeth 2 and 3 small, almost half the size of protibial tooth 1...........
|
|
<taxonomicName id="4C194D2CFFB1FFBE794657556E8BFBB2" ID-CoL="75VW2" authority="Scholtz & Howden, 1987" authorityName="Scholtz & Howden" authorityYear="1987" box="[1028,1436,1094,1116]" class="Insecta" family="Scarabaeidae" genus="Peckolus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="419" phylum="Arthropoda" rank="species" species="parvus">
|
|
<emphasis id="B96DEABDFFB1FFBE794657556FB5FBB2" box="[1028,1186,1094,1115]" italics="true" pageId="5" pageNumber="419">Peckolus parvus</emphasis>
|
|
<bibRefCitation id="EF884B5EFFB1FFBE79EA57556E8BFBB2" author="Scholtz, C. H. & Howden, H. F." box="[1192,1436,1094,1116]" pageId="5" pageNumber="419" pagination="75 - 119" refId="ref4756" refString="Scholtz, C. H. & Howden, H. F. (1987) A revision of the African Cathonina (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the Entomological Society of Southern Africa, 50, 75 - 119." type="journal article" year="1987">Scholtz & Howden, 1987</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BA636AFFFB1FFBE7DD557776E8BFB91" blockId="5.[151,1437,536,1409]" box="[151,1436,1123,1145]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD557776E8BFB91" box="[151,1436,1123,1145]" pageId="5" pageNumber="419">- Protibial teeth 2 and 3 minute, much less than half the size of protibial tooth 1..................................... 9</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30ED2E0AFFB1FFBE7DD557926E8BFB5A" pageId="5" pageNumber="419">
|
|
<paragraph id="8BA636AFFFB1FFBE7DD557926E8BFB7F" blockId="5.[151,1437,536,1409]" box="[151,1436,1152,1174]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD557926E8BFB7F" box="[151,1436,1152,1174]" pageId="5" pageNumber="419">
|
|
9. The recurved setae on the body almost half the length of the straight setae......
|
|
<taxonomicName id="4C194D2CFFB1FFBE7EBD57936E8BFB7F" ID-CoL="75VVQ" authority="Howden & Scholtz, 1988" authorityName="Howden & Scholtz" authorityYear="1988" box="[1023,1436,1152,1174]" class="Insecta" family="Scarabaeidae" genus="Peckolus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="419" phylum="Arthropoda" rank="species" species="alpinus">
|
|
<emphasis id="B96DEABDFFB1FFBE7EBD57936FB4FB7C" box="[1023,1187,1152,1173]" italics="true" pageId="5" pageNumber="419">Peckolus alpinus</emphasis>
|
|
<bibRefCitation id="EF884B5EFFB1FFBE79EA57936E8BFB7F" author="Howden, H. & Scholtz, C. H." box="[1192,1436,1152,1174]" pageId="5" pageNumber="419" pagination="133 - 136" refId="ref4607" refString="Howden, H. & Scholtz, C. H. (1988) A second species of the African genus Peckolus Scholtz & Howden (Scarabaeidae: Scarabaeini). Journal of the Entomological Society of Southern Africa 51, 133 - 136." type="journal article" year="1988">Howden & Scholtz, 1988</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BA636AFFFB1FFBE7DD5578D6E8BFB5A" blockId="5.[151,1437,536,1409]" box="[151,1436,1182,1204]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD5578D6E8BFB5A" box="[151,1436,1182,1204]" pageId="5" pageNumber="419">
|
|
- The recurved setae on the body barely discernible.....................
|
|
<taxonomicName id="4C194D2CFFB1FFBE7E84578D6E8BFB5A" ID-CoL="75W7Q" authority="Deschodt & Scholtz, 2008" authorityName="Deschodt & Scholtz" authorityYear="2008" box="[966,1436,1182,1204]" class="Insecta" family="Scarabaeidae" genus="Peckolus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="419" phylum="Arthropoda" rank="species" species="poenskopius">
|
|
<emphasis id="B96DEABDFFB1FFBE7E84578D6F8EFB5A" box="[966,1177,1182,1203]" italics="true" pageId="5" pageNumber="419">Peckolus poenskopius</emphasis>
|
|
<bibRefCitation id="EF884B5EFFB1FFBE79DD578D6E8BFB5A" author="Deschodt, C. M. & Scholtz, C. H." box="[1183,1436,1182,1204]" pageId="5" pageNumber="419" pagination="91 - 106" refId="ref4469" refString="Deschodt, C. M. & Scholtz, C. H. (2008) Systematics of South African forest-endemic dung beetles: new genera and species of small Canthonini (Scarabaeidae: Scarabaeinae). African Entomology 16, 91 - 106. https: // doi. org / 10.4001 / 1021 - 3589 - 16.1.91" type="journal article" year="2008">Deschodt & Scholtz, 2008</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30ED2E0AFFB1FFBE7DD557A86E8BFAE2" pageId="5" pageNumber="419">
|
|
<paragraph id="8BA636AFFFB1FFBE7DD557A86E8BFB07" blockId="5.[151,1437,536,1409]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD557A86E8BFB07" pageId="5" pageNumber="419">
|
|
10. Lateral part of mesometaventral suture (between mesocoxae) converging anteriorly ...........................................................................................
|
|
<taxonomicName id="4C194D2CFFB1FFBE7EDA57CB6E8BFB07" ID-CoL="6TSHV" authority="Deschodt & Scholtz, 2008" authorityName="Deschodt & Scholtz" authorityYear="2008" box="[920,1436,1240,1262]" class="Insecta" family="Scarabaeidae" genus="Parvuhowdenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="419" phylum="Arthropoda" rank="species" species="harrisoni">
|
|
<emphasis id="B96DEABDFFB1FFBE7EDA57CB6F8EFB04" box="[920,1177,1240,1261]" italics="true" pageId="5" pageNumber="419">Parvuhowdenius harrisoni</emphasis>
|
|
<bibRefCitation id="EF884B5EFFB1FFBE79DD57CB6E8BFB07" author="Deschodt, C. M. & Scholtz, C. H." box="[1183,1436,1240,1262]" pageId="5" pageNumber="419" pagination="91 - 106" refId="ref4469" refString="Deschodt, C. M. & Scholtz, C. H. (2008) Systematics of South African forest-endemic dung beetles: new genera and species of small Canthonini (Scarabaeidae: Scarabaeinae). African Entomology 16, 91 - 106. https: // doi. org / 10.4001 / 1021 - 3589 - 16.1.91" type="journal article" year="2008">Deschodt & Scholtz, 2008</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BA636AFFFB1FFBE7DD557E56E8BFAE2" blockId="5.[151,1437,536,1409]" box="[151,1436,1270,1292]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD557E56E8BFAE2" box="[151,1436,1270,1292]" pageId="5" pageNumber="419">- Lateral part of mesometaventral suture (between mesocoxae) parallel........................................... 11</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30ED2E0AFFB1FFBE7DD556006E8BFA69" pageId="5" pageNumber="419">
|
|
<paragraph id="8BA636AFFFB1FFBE7DD556006E8BFAAF" blockId="5.[151,1437,536,1409]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD556006E8BFAAF" pageId="5" pageNumber="419">
|
|
11. Punctures on mesoventrite large and clear, separated by less than one puncture diameter..........................................................................................
|
|
<taxonomicName id="4C194D2CFFB1FFBE7EFE56236E8BFAAF" ID-CoL="45ZH3" authority="Deschodt & Scholtz, 2008" authorityName="Deschodt & Scholtz" authorityYear="2008" box="[956,1436,1328,1350]" class="Insecta" family="Scarabaeidae" genus="Nebulasilvius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="419" phylum="Arthropoda" rank="species" species="insularis">
|
|
<emphasis id="B96DEABDFFB1FFBE7EFE56236F8EFAAC" box="[956,1177,1328,1349]" italics="true" pageId="5" pageNumber="419">Nebulasilvius insularis</emphasis>
|
|
<bibRefCitation id="EF884B5EFFB1FFBE79DD56236E8BFAAF" author="Deschodt, C. M. & Scholtz, C. H." box="[1183,1436,1328,1350]" pageId="5" pageNumber="419" pagination="91 - 106" refId="ref4469" refString="Deschodt, C. M. & Scholtz, C. H. (2008) Systematics of South African forest-endemic dung beetles: new genera and species of small Canthonini (Scarabaeidae: Scarabaeinae). African Entomology 16, 91 - 106. https: // doi. org / 10.4001 / 1021 - 3589 - 16.1.91" type="journal article" year="2008">Deschodt & Scholtz, 2008</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BA636AFFFB1FFBE7DD5565D6E8BFA69" blockId="5.[151,1437,536,1409]" pageId="5" pageNumber="419">
|
|
<keyLead id="30E8959AFFB1FFBE7DD5565D6E8BFA69" pageId="5" pageNumber="419">
|
|
- Punctures on mesoventrite small and separated by at least one puncture diameter ..................................................................................................
|
|
<taxonomicName id="4C194D2CFFB1FFBE7E9156786E8BFA69" ID-CoL="45ZH4" authority="Deschodt & Scholtz, 2008" authorityName="Deschodt & Scholtz" authorityYear="2008" box="[979,1436,1387,1409]" class="Insecta" family="Scarabaeidae" genus="Nebulasilvius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="419" phylum="Arthropoda" rank="species" species="johani">
|
|
<emphasis id="B96DEABDFFB1FFBE7E9156786F8EFA69" box="[979,1177,1387,1408]" italics="true" pageId="5" pageNumber="419">Nebulasilvius johani</emphasis>
|
|
<bibRefCitation id="EF884B5EFFB1FFBE79DD56786E8BFA69" author="Deschodt, C. M. & Scholtz, C. H." box="[1183,1436,1387,1409]" pageId="5" pageNumber="419" pagination="91 - 106" refId="ref4469" refString="Deschodt, C. M. & Scholtz, C. H. (2008) Systematics of South African forest-endemic dung beetles: new genera and species of small Canthonini (Scarabaeidae: Scarabaeinae). African Entomology 16, 91 - 106. https: // doi. org / 10.4001 / 1021 - 3589 - 16.1.91" type="journal article" year="2008">Deschodt & Scholtz, 2008</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
</key>
|
|
</subSubSection>
|
|
</treatment>
|
|
</subSection>
|
|
</document> |