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<document id="5C06409985AA607F47C40A292727E96C" ID-DOI="10.5281/zenodo.189670" ID-GBIF-Dataset="cac989cb-05f4-4993-a1ae-9cfb71e6d2e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="189670" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461103756872" checkinUser="plazi" docAuthor="Valais, Silvina De" docDate="2009" docId="03B01A20BB01FFAEFF2E6F4DFF1687D8" docLanguage="en" docName="zt02203p021.pdf" docOrigin="Zootaxa 2203" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Amphibiopus patagonicus Casamiquela 1961" docType="treatment" docVersion="9" lastPageNumber="12" masterDocId="FF896258BB09FFA5FFB96C02FFDA8013" masterDocTitle="Ichnotaxonomic revision of Ameghinichnus, a mammalian ichnogenus from the Middle Jurassic La Matilde Formation, Santa Cruz province, Argentina" masterLastPageNumber="21" masterPageNumber="1" pageNumber="9" updateTime="1698592280095" updateUser="plazi">
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<mods:title id="C4A78C4F2BE97C07EBC790645C6877C8">Ichnotaxonomic revision of Ameghinichnus, a mammalian ichnogenus from the Middle Jurassic La Matilde Formation, Santa Cruz province, Argentina</mods:title>
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<mods:date id="020FF38CBC5529A03886E7DDC9471746">2009</mods:date>
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<mods:identifier id="CB11F65D5DB31882D08961C3C75477C5" type="DOI">10.5281/zenodo.189670</mods:identifier>
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<emphasis id="B96D7724BB01FFADFF2E6F4DFE8D837A" bold="true" box="[151,343,847,873]" italics="true" pageId="8" pageNumber="9">Ameghinichnus</emphasis>
<taxonomicName id="4C19D0B5BB01FFADFEE76F4DFD3E837A" authority="Casamiquela, 1961" authorityName="Casamiquela" authorityYear="1961" box="[350,740,847,873]" class="Amphibia" genus="Amphibiopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="patagonicus">
<emphasis id="B96D7724BB01FFADFEE76F4DFE2A837A" bold="true" box="[350,496,847,873]" italics="true" pageId="8" pageNumber="9">patagonicus</emphasis>
<bibRefCitation id="EF88D6C7BB01FFADFE416F4DFD3E837A" author="Casamiquela" box="[504,740,847,873]" pageId="8" pageNumber="19" refString="Casamiquela, R. M. (1961) Sobre la presencia de un mamifero en el primer elenco (icnologico) de vertebrados del Jurasico de la Patagonia. Physis, 22, 225 - 233." type="journal article" year="1961">Casamiquela, 1961</bibRefCitation>
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<figureCitation id="1322B7B3BB01FFADFF2E6F70FEDC839F" box="[151,262,882,908]" captionStart="FIGURE 3" captionStartId="9.[151,255,1863,1887]" captionTargetBox="[187,1413,202,1830]" captionTargetId="figure@9.[171,1429,194,1842]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of the type series of Ameghinichnus patagonicus. A, MLP 60 - X- 31 - 1; B, MLP 60 - X- 31 - 14 B, holotype (right trackway: 1) and MLP 60 - X- 31 - 14 A, paratype (left trackway: 2), the black arrows indicate the moving direction of the trackmaker; C, PVL 2303; D, PVL 2302, the white arrow indicates the bilobate outline; E, detail of the two first manus-pes sets of MLP 60 - X- 31 - 14 B; F, MLP 60 - X- 31 - 15; G, MLP 60 - X- 31 - 3. A-C, D, F and G, paratypes. Scale bars: 2 cm in D and E, 5 cm in the rest. m, manus imprint; p, pes imprint." httpUri="https://zenodo.org/record/189673/files/figure.png" pageId="8" pageNumber="9">Figures 3</figureCitation>
,
<figureCitation id="1322B7B3BB01FFADFEAA6F70FEF8839F" box="[275,290,882,908]" captionStart="FIGURE 4" captionStartId="10.[151,256,1837,1861]" captionTargetBox="[261,1332,181,1802]" captionTargetId="figure@10.[245,1348,168,1812]" captionTargetPageId="10" captionText="FIGURE 4. Photographs of specimens of Ameghinichnus patagonicus. A, MPM-PIc- 3768 (first manus-pes set to the left) and 3769 (first manus-pes set to the right); B, MPEF-IC 1016 at left, indicated with a 1, MPEF-IC 1015 at rigth, indicated with a 2; C, MPEF-IC 1023; D, MACN 18619; E, MPEF-IC 1041; F, MACN-SC 4354; G, MACN 18622, the white arrows indicate the bilobate outline; H, detail of the third manus-pes sets of MPM-PIc- 3952; I, MPM-PIc- 3952. Scale bars: 2 cm in C, E, F, and H, 5 cm in A, D, G, and I, and 10 cm in B. The black arrow in C, E, and F indicates the tail mark. m, manus imprint; p, pes imprint." httpUri="https://zenodo.org/record/189674/files/figure.png" pageId="8" pageNumber="9">4</figureCitation>
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<emphasis id="B96D7724BB01FFADFF2E6FC2FEE183C4" box="[151,315,960,983]" italics="true" pageId="8" pageNumber="9">Ameghinichnus</emphasis>
<taxonomicName id="4C19D0B5BB01FFADFEFD6FC2FD1C83C4" authority="Casamiquela, 1960: 11" authorityName="Casamiquela" authorityPageNumber="11" authorityYear="1960" box="[324,710,960,983]" class="Amphibia" genus="Amphibiopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="patagonicus">
<emphasis id="B96D7724BB01FFADFEFD6FC2FE1C83C4" box="[324,454,960,983]" italics="true" pageId="8" pageNumber="9">patagonicus</emphasis>
<treatmentCitation id="0AB88D27BB01FFADFE776FC2FD1C83C4" author="Casamiquela" box="[462,710,960,983]" page="11" pageId="8" pageNumber="9" year="1960">
<bibRefCitation id="EF88D6C7BB01FFADFE776FC2FD7A83C4" author="Casamiquela" box="[462,672,960,983]" pageId="8" pageNumber="19" refString="Casamiquela, R. M. (1960) El hallazgo del primer elenco (icnologico) jurasico de vertebrados terrestres de Latinoamerica (noticia). Revista de la Asociacion Geologica Argentina, 15, 5 - 14." type="journal article" year="1960">Casamiquela, 1960</bibRefCitation>
: 11
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(
<emphasis id="B96D7724BB01FFADFD6F6FC2FCB583C4" box="[726,879,960,983]" italics="true" pageId="8" pageNumber="9">nomen nudum</emphasis>
);
<treatmentCitation id="0AB88D27BB01FFADFC3E6FC2FB4A83C4" author="Casamiquela" box="[903,1168,960,983]" page="226" pageId="8" pageNumber="9" year="1961">
<bibRefCitation id="EF88D6C7BB01FFADFC3E6FC2FB8383C4" author="Casamiquela" box="[903,1113,960,983]" pageId="8" pageNumber="19" refString="Casamiquela, R. M. (1961) Sobre la presencia de un mamifero en el primer elenco (icnologico) de vertebrados del Jurasico de la Patagonia. Physis, 22, 225 - 233." type="journal article" year="1961">Casamiquela, 1961</bibRefCitation>
: 226
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;
<treatmentCitation id="0AB88D27BB01FFADFB256FC2FA4283C4" author="Casamiquela" box="[1180,1432,960,983]" page="90" pageId="8" pageNumber="9" year="1964">
<bibRefCitation id="EF88D6C7BB01FFADFB256FC2FAB483C4" author="Casamiquela" box="[1180,1390,960,983]" pageId="8" pageNumber="19" refString="Casamiquela, R. M. (1964) Estudios icnologicos. Problemas y metodos de la icnologia con aplicacion al estudio de pisadas mesozoicas (Reptilia, Mammalia) de la Patagonia. Colegio Industrial Pio IX, Buenos Aires, Argentina, 229 pp." type="book" year="1964">Casamiquela, 1964</bibRefCitation>
: 90
</treatmentCitation>
;
<treatmentCitation id="0AB88D27BB01FFADFF7E6FE2FE6A83E4" author="Bonaparte" box="[199,432,992,1015]" page="451" pageId="8" pageNumber="9" year="1978">
<bibRefCitation id="EF88D6C7BB01FFADFF7E6FE2FEA183E4" author="Bonaparte" box="[199,379,992,1015]" pageId="8" pageNumber="19" refString="Bonaparte, J. F. (1978) El Mesozoico de America del Sur y sus tetrapodos. Fundacion Miguel Lillo, Opera Lilloana, 26, Tucuman, 596 pp." type="book" year="1978">Bonaparte, 1978</bibRefCitation>
: 451
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;
<treatmentCitation id="0AB88D27BB01FFADFE026FE2FD4C83E4" author="Bonaparte" box="[443,662,992,1015]" page="81" pageId="8" pageNumber="9" year="1983">
<bibRefCitation id="EF88D6C7BB01FFADFE026FE2FDB483E4" author="Bonaparte" box="[443,622,992,1015]" pageId="8" pageNumber="19" refString="Bonaparte, J. F. (1983) Jurassic tetrapods from South America and dispersal routes. In: Jacobs, L. L. (Ed.), Aspects of vertebrate history: Essay in honor of Edwin Harris Colbert. Museum of Northern Arizona Press, pp. 73 - 98." type="book chapter" year="1983">Bonaparte, 1983</bibRefCitation>
: 81
</treatmentCitation>
; Leonardi &amp; de
<treatmentCitation id="0AB88D27BB01FFADFCFA6FE2FB8E83E4" author="Oliveira" box="[835,1108,992,1015]" page="220" pageId="8" pageNumber="9" year="1990">
<bibRefCitation id="EF88D6C7BB01FFADFCFA6FE2FBC583E4" author="Oliveira" box="[835,1055,992,1015]" pageId="8" pageNumber="20" refString="Leonardi, G. &amp; de Oliveira Lima, F. H. (1990) A revision of the Triassic and Jurassic tetrapod footprints of Argentina and a new approach on the age and meaning of the Botucatu Formation footprints (Brazil). Revista Brasileira de Geociencias, 20, 216 - 229." type="journal article" year="1990">Oliveira Lima, 1990</bibRefCitation>
: 220
</treatmentCitation>
, 222, 225;
<treatmentCitation id="0AB88D27BB01FFADFB706FE2FA4D83E4" author="Leonardi" box="[1225,1431,992,1015]" page="27" pageId="8" pageNumber="9" year="1994">
<bibRefCitation id="EF88D6C7BB01FFADFB706FE2FAAA83E4" author="Leonardi" box="[1225,1392,992,1015]" pageId="8" pageNumber="20" refString="Leonardi, G. (1994) Annotated atlas of South America tetrapod footprints (Devonian to Holocene). Companhia de Pesquisa de Recursos Minerais, Rio de Janeiro, Brasil, 247 pp." type="book" year="1994">Leonardi, 1994</bibRefCitation>
: 27
</treatmentCitation>
;
<treatmentCitation id="0AB88D27BB01FFADFF7E6802FDFA8404" author="Rainforth" box="[199,544,1024,1047]" page="266" pageId="8" pageNumber="9" year="1996">
<bibRefCitation id="EF88D6C7BB01FFADFF7E6802FE318404" author="Rainforth" box="[199,491,1024,1047]" pageId="8" pageNumber="21" refString="Rainforth, E. C. &amp; Lockley, M. G. (1996) Tracks of diminutive dinosaurs and hopping mammals from the Jurassic of North and South America. In: Morales, M. (Ed.), The continental Jurassic. Museum of Northern Arizona Bulletin 60, pp. 265 - 269." type="journal article" year="1996">Rainforth &amp; Lockley, 1996</bibRefCitation>
: 266
</treatmentCitation>
;
<treatmentCitation id="0AB88D27BB01FFADFD926802FD248404" author="Sarjeant" box="[555,766,1024,1047]" page="160" pageId="8" pageNumber="9" year="2000">
<bibRefCitation id="EF88D6C7BB01FFADFD926802FD138404" author="Sarjeant" box="[555,713,1024,1047]" pageId="8" pageNumber="21" refString="Sarjeant, W. A. S. (2000) The mesozoic mammal footprint record reconsidered: with an account of new discoveries in the Cretaceous of Northwestern Alberta, Canada. Paleontology Society of Korea Special Publication, 4, 153 - 168." type="journal article" year="2000">Sarjeant, 2000</bibRefCitation>
: 160
</treatmentCitation>
;
<treatmentCitation id="0AB88D27BB01FFADFCB06802FBD58404" author="Casamiquela" box="[777,1039,1024,1047]" page="434" pageId="8" pageNumber="9" year="2002">
<bibRefCitation id="EF88D6C7BB01FFADFCB06802FC038404" author="Casamiquela" box="[777,985,1024,1047]" pageId="8" pageNumber="19" refString="Casamiquela, R. M. (2002) Icnitas de vertebrados (Mamiferos, Dinosaurios) del Mesojurasico. In: Haller, M. J. (Ed.), Geologia y Recursos Naturales de Santa Cruz, Relatorio del 15 Congreso Geologico Argentino, II, pp. 433 - 438." type="journal article" year="2002">Casamiquela, 2002</bibRefCitation>
: 434
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;
<treatmentCitation id="0AB88D27BB01FFADFBA36802FAC58404" author="Melchor" box="[1050,1311,1024,1047]" page="51" pageId="8" pageNumber="9" year="2004">
<bibRefCitation id="EF88D6C7BB01FFADFBA36802FB2C8404" author="Melchor" box="[1050,1270,1024,1047]" pageId="8" pageNumber="21" refString="Melchor, R. N., de Valais, S. &amp; Genise, J. F. (2004) Middle Jurassic mammalian and dinosaur footprints and petrified forests from the volcaniclastic La Matilde Formation. In: Bellosi, E. S. &amp; Melchor, R. N (Eds.), Fieldtrip guidebook. I International Congress on Ichnology, Museo Egidio Feruglio, Trelew, pp. 47 - 63." type="book chapter" year="2004">
Melchor
<emphasis id="B96D7724BB01FFADFBC26802FB708404" box="[1147,1194,1024,1047]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
., 2004
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: 51
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.
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</paragraph>
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<typeStatus id="54A21594BB01FFADFF2E683DFEC8844A" box="[151,274,1087,1113]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED0833F3BB01FFADFEA66842FE878449" box="[287,349,1088,1114]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="8" pageNumber="9">MLP</collectionCode>
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A, trackways with six manus-pes sets with opposite arrangement about the midline (associated with the
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B) (trackway
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<figureCitation id="1322B7B3BB01FFADFC4E6864FB9C8493" box="[1015,1094,1126,1152]" captionStart="FIGURE 3" captionStartId="9.[151,255,1863,1887]" captionTargetBox="[187,1413,202,1830]" captionTargetId="figure@9.[171,1429,194,1842]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of the type series of Ameghinichnus patagonicus. A, MLP 60 - X- 31 - 1; B, MLP 60 - X- 31 - 14 B, holotype (right trackway: 1) and MLP 60 - X- 31 - 14 A, paratype (left trackway: 2), the black arrows indicate the moving direction of the trackmaker; C, PVL 2303; D, PVL 2302, the white arrow indicates the bilobate outline; E, detail of the two first manus-pes sets of MLP 60 - X- 31 - 14 B; F, MLP 60 - X- 31 - 15; G, MLP 60 - X- 31 - 3. A-C, D, F and G, paratypes. Scale bars: 2 cm in D and E, 5 cm in the rest. m, manus imprint; p, pes imprint." httpUri="https://zenodo.org/record/189673/files/figure.png" pageId="8" pageNumber="9">Figs. 3</figureCitation>
b, 3e).
</paragraph>
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<typeStatus id="54A21594BB01FFADFF7C688FFE9F84B4" box="[197,325,1165,1191]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
.
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,
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,
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,
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,
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,
<date id="FFA78DF6BB01FFADFBE4688FFB3384B4" box="[1117,1257,1165,1191]" pageId="8" pageNumber="9" value="1960-10-31" valueMax="1960-10-14" valueMin="1960-10-31">60-X-31-14</date>
B,
<date id="FFA78DF6BB01FFADFAB3688FFA4D84B4" box="[1290,1431,1165,1191]" pageId="8" pageNumber="9" value="1960-10-31" valueMax="1960-10-15" valueMin="1960-10-31">60-X-31-15</date>
,
<collectionCode id="ED0833F3BB01FFADFF2E68B6FF1484DD" box="[151,206,1204,1230]" httpUri="http://grbio.org/cool/pbkd-hmeb" name="Paleontologia de Vertebrados Lillo" pageId="8" pageNumber="9">PVL</collectionCode>
2300, 2301, 2302, 2303, 2304 (
<figureCitation id="1322B7B3BB01FFADFDFC68B6FD4C84DD" box="[581,662,1204,1230]" captionStart="FIGURE 3" captionStartId="9.[151,255,1863,1887]" captionTargetBox="[187,1413,202,1830]" captionTargetId="figure@9.[171,1429,194,1842]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of the type series of Ameghinichnus patagonicus. A, MLP 60 - X- 31 - 1; B, MLP 60 - X- 31 - 14 B, holotype (right trackway: 1) and MLP 60 - X- 31 - 14 A, paratype (left trackway: 2), the black arrows indicate the moving direction of the trackmaker; C, PVL 2303; D, PVL 2302, the white arrow indicates the bilobate outline; E, detail of the two first manus-pes sets of MLP 60 - X- 31 - 14 B; F, MLP 60 - X- 31 - 15; G, MLP 60 - X- 31 - 3. A-C, D, F and G, paratypes. Scale bars: 2 cm in D and E, 5 cm in the rest. m, manus imprint; p, pes imprint." httpUri="https://zenodo.org/record/189673/files/figure.png" pageId="8" pageNumber="9">Figs. 3</figureCitation>
a–d,f,g).
</paragraph>
<paragraph id="8BA6AB36BB01FFADFF7C68D8FE5B8639" blockId="8.[151,1437,1087,1926]" pageId="8" pageNumber="9">
<emphasis id="B96D7724BB01FFADFF7C68D8FE1984E7" bold="true" box="[197,451,1242,1268]" pageId="8" pageNumber="9">Additional material.</emphasis>
<collectionCode id="ED0833F3BB01FFADFE7068D8FDFB84E7" box="[457,545,1242,1268]" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="8" pageNumber="9">MACN</collectionCode>
18528, 18616, 18617, 18618, 18619, 18620, 18621, 18622, 18623, 18525b, 3823, MACN-SC 3824, 3825, 3826, 3827, 3828, 3829, 3830, 3831, 3833, 3834, 3835, 3836, 3837, 3838, 4354,
<collectionCode id="ED0833F3BB01FFADFF66692AFECC8551" box="[223,278,1320,1346]" httpUri="http://grbio.org/cool/pbkd-hmeb" name="Paleontologia de Vertebrados Lillo" pageId="8" pageNumber="9">PVL</collectionCode>
2742, 2743, 3684, 3685, 3686, 3688, 3689, 3691, 3692, 3693, 3694, 3697, 3698, 3699, 4354,
<collectionCode id="ED0833F3BB01FFADFAE6692AFA418551" box="[1375,1435,1320,1346]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="8" pageNumber="9">MLP</collectionCode>
<date id="FFA78DF6BB01FFADFF2E694CFEC6857B" box="[151,284,1358,1384]" pageId="8" pageNumber="9" value="1965-11-12" valueMax="1965-11-01" valueMin="1965-11-12">65-XI-12-1</date>
(more than thirty slabs with the same collection number),
<date id="FFA78DF6BB01FFADFC71694CFB8B857B" box="[968,1105,1358,1384]" pageId="8" pageNumber="9" value="1965-11-12" valueMax="1965-11-04" valueMin="1965-11-12">65-XI-12-4</date>
,
<date id="FFA78DF6BB01FFADFBE5694CFB38857B" box="[1116,1250,1358,1384]" pageId="8" pageNumber="9" value="1965-11-13" valueMax="1965-11-01" valueMin="1965-11-13">65-XI-13-1</date>
(two slabs with the same collection number), one slab with no collection number, MPEF-IC 1015, 1016, 1023, 1030, 1031, 1033, 1034, 1035, 1036, 1037, 1038, 1040, 1041, MPM-PIc-173, 175, 176, 177, 178, 179, 180, 181, 182, 183, 184, 184, 185, 186, 187, 188, 189, 194, 195, 196, 197, 198, 199, 201, 202, 203, 204, 205, 206, 207, 208, 209, 210, 3763, 3764, 3765, 3766, 3767, 3768, 3769, 3770, 3771, 3772, 3773, 3774, 3775, 3776, 3952, 3953, 3954, 3955, 3956 (
<figureCitation id="1322B7B3BB01FFADFE926A12FEAE8639" box="[299,372,1552,1578]" captionStart="FIGURE 4" captionStartId="10.[151,256,1837,1861]" captionTargetBox="[261,1332,181,1802]" captionTargetId="figure@10.[245,1348,168,1812]" captionTargetPageId="10" captionText="FIGURE 4. Photographs of specimens of Ameghinichnus patagonicus. A, MPM-PIc- 3768 (first manus-pes set to the left) and 3769 (first manus-pes set to the right); B, MPEF-IC 1016 at left, indicated with a 1, MPEF-IC 1015 at rigth, indicated with a 2; C, MPEF-IC 1023; D, MACN 18619; E, MPEF-IC 1041; F, MACN-SC 4354; G, MACN 18622, the white arrows indicate the bilobate outline; H, detail of the third manus-pes sets of MPM-PIc- 3952; I, MPM-PIc- 3952. Scale bars: 2 cm in C, E, F, and H, 5 cm in A, D, G, and I, and 10 cm in B. The black arrow in C, E, and F indicates the tail mark. m, manus imprint; p, pes imprint." httpUri="https://zenodo.org/record/189674/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BA6AB36BB01FFADFF7C6A34FE098795" blockId="8.[151,1437,1087,1926]" pageId="8" pageNumber="9">
<emphasis id="B96D7724BB01FFADFF7C6A34FE658643" bold="true" box="[197,447,1590,1616]" pageId="8" pageNumber="9">Emended diagnosis.</emphasis>
<emphasis id="B96D7724BB01FFADFE7E6A35FDA68643" box="[455,636,1591,1616]" italics="true" pageId="8" pageNumber="9">Ameghinichnus</emphasis>
preserved as manus and pes tracks with a length:width ratio of 0.7. Manus digit impressions are subequal in length, with a relative digit length: I&lt;II=V&lt;IV&lt;III, angles between III-IV&lt;I-II≈II-III≈IV-V, I-V averages 151º. Manus impressions display an inward rotation of about -16º in relation to the midline, and are located aligned-to-posterior to pes impressions. Pes digit impressions are subequal in length, with relative length: I&lt;IV≈III=V&lt;II, angles between III-IV&lt;II-III&lt;I-II&lt;IV-V, I-V averages 152º. Pes display an outward rotation of about 31º in relation to the midline. Claw marks are absent and there are no marked phalangeal pads. When manus-pes sets display alternate arrangement about the midline, manus and pes pace angulation is about 143º and 114º, respectively, and manus and pes stride length is about
<quantity id="4CE106D3BB01FFADFAF06B47FA46874C" box="[1353,1436,1861,1887]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" pageId="8" pageNumber="9" unit="mm" value="83.0">83 mm</quantity>
and 81.2 mm, respectively.
</paragraph>
</subSubSection>
<caption id="DF66FBBEBB00FFACFF2E6B45FC3E87FA" httpUri="https://zenodo.org/record/189673/files/figure.png" pageId="9" pageNumber="10" targetBox="[187,1413,202,1830]" targetPageId="9">
<paragraph id="8BA6AB36BB00FFACFF2E6B45FC3E87FA" blockId="9.[151,1437,1863,2025]" pageId="9" pageNumber="10">
<emphasis id="B96D7724BB00FFACFF2E6B45FEC0874C" bold="true" box="[151,282,1863,1887]" pageId="9" pageNumber="10">FIGURE 3.</emphasis>
Photographs of the type series of
<emphasis id="B96D7724BB00FFACFD326B4AFC63874C" box="[651,953,1864,1887]" italics="true" pageId="9" pageNumber="10">
Ameghinichnus
<taxonomicName id="4C19D0B5BB00FFACFC8E6B4AFC63874C" box="[823,953,1864,1887]" class="Amphibia" genus="Amphibiopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="patagonicus">patagonicus</taxonomicName>
</emphasis>
.
<emphasis id="B96D7724BB00FFACFC716B45FC01874C" bold="true" box="[968,987,1863,1887]" pageId="9" pageNumber="10">A</emphasis>
, MLP 60-X-31-1;
<emphasis id="B96D7724BB00FFACFB1E6B45FB1A874C" bold="true" box="[1191,1216,1863,1887]" pageId="9" pageNumber="10">B,</emphasis>
MLP 60-X-31-14B, holotype (right trackway:
<emphasis id="B96D7724BB00FFACFE086B68FE648791" bold="true" box="[433,446,1898,1922]" pageId="9" pageNumber="10">1</emphasis>
) and MLP 60-X-31-14A, paratype (left trackway:
<emphasis id="B96D7724BB00FFACFC5F6B68FC298791" bold="true" box="[998,1011,1898,1922]" pageId="9" pageNumber="10">2</emphasis>
), the black arrows indicate the moving direction of the trackmaker;
<emphasis id="B96D7724BB00FFACFE706B8EFE0687B7" bold="true" box="[457,476,1932,1956]" pageId="9" pageNumber="10">C</emphasis>
, PVL 2303;
<emphasis id="B96D7724BB00FFACFDD16B8EFDA187B7" bold="true" box="[616,635,1932,1956]" pageId="9" pageNumber="10">D</emphasis>
, PVL 2302, the white arrow indicates the bilobate outline;
<emphasis id="B96D7724BB00FFACFB426B8EFAD787B7" bold="true" box="[1275,1293,1932,1956]" pageId="9" pageNumber="10">E</emphasis>
, detail of the two first manus-pes sets of MLP 60-X-31-14B;
<emphasis id="B96D7724BB00FFACFD186BADFD7587D4" bold="true" box="[673,687,1967,1991]" pageId="9" pageNumber="10">F</emphasis>
, MLP 60-X-31-15;
<emphasis id="B96D7724BB00FFACFC336BADFC4687D4" bold="true" box="[906,924,1967,1991]" pageId="9" pageNumber="10">G</emphasis>
, MLP 60-X-31-3.
<emphasis id="B96D7724BB00FFACFBD16BADFB4D87D4" bold="true" box="[1128,1175,1967,1991]" pageId="9" pageNumber="10">A-C</emphasis>
,
<emphasis id="B96D7724BB00FFACFB1F6BADFB6387D4" bold="true" box="[1190,1209,1967,1991]" pageId="9" pageNumber="10">D</emphasis>
,
<emphasis id="B96D7724BB00FFACFB716BADFB0387D4" bold="true" box="[1224,1241,1967,1991]" pageId="9" pageNumber="10">F</emphasis>
and
<emphasis id="B96D7724BB00FFACFAB76BADFAFA87D4" bold="true" box="[1294,1312,1967,1991]" pageId="9" pageNumber="10">G</emphasis>
, paratypes. Scale bars: 2 cm in D and E, 5 cm in the rest. m, manus imprint; p, pes imprint.
</paragraph>
</caption>
<caption id="DF66FBBEBB03FFAFFF2E6B2FFDB087E1" httpUri="https://zenodo.org/record/189674/files/figure.png" pageId="10" pageNumber="11" targetBox="[261,1332,181,1802]" targetPageId="10">
<paragraph id="8BA6AB36BB03FFAFFF2E6B2FFDB087E1" blockId="10.[151,1437,1837,2034]" pageId="10" pageNumber="11">
<emphasis id="B96D7724BB03FFAFFF2E6B2FFEC68756" bold="true" box="[151,284,1837,1861]" pageId="10" pageNumber="11">FIGURE 4.</emphasis>
Photographs of specimens of
<emphasis id="B96D7724BB03FFAFFDD16B2CFC468756" box="[616,924,1838,1861]" italics="true" pageId="10" pageNumber="11">
Ameghinichnus
<taxonomicName id="4C19D0B5BB03FFAFFCAE6B2CFC468756" box="[791,924,1838,1861]" class="Amphibia" genus="Amphibiopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="patagonicus">patagonicus</taxonomicName>
</emphasis>
.
<emphasis id="B96D7724BB03FFAFFC126B2FFC648756" bold="true" box="[939,958,1837,1861]" pageId="10" pageNumber="11">A</emphasis>
, MPM-PIc-3768 (first manus-pes set to the left) and 3769 (first manus-pes set to the right);
<emphasis id="B96D7724BB03FFAFFD1E6B52FD63877B" bold="true" box="[679,697,1872,1896]" pageId="10" pageNumber="11">B</emphasis>
, MPEF-IC 1016 at left, indicated with a
<emphasis id="B96D7724BB03FFAFFBC26B52FB52877B" bold="true" box="[1147,1160,1872,1896]" pageId="10" pageNumber="11">1</emphasis>
, MPEF-IC 1015 at rigth, indicated with a
<emphasis id="B96D7724BB03FFAFFEF06B70FE8C8799" bold="true" box="[329,342,1906,1930]" pageId="10" pageNumber="11">2</emphasis>
;
<emphasis id="B96D7724BB03FFAFFEDD6B70FEAD8799" bold="true" box="[356,375,1906,1930]" pageId="10" pageNumber="11">C</emphasis>
, MPEF-IC 1023;
<emphasis id="B96D7724BB03FFAFFD806B70FD968799" bold="true" box="[569,588,1906,1930]" pageId="10" pageNumber="11">D</emphasis>
, MACN 18619;
<emphasis id="B96D7724BB03FFAFFCBB6B70FCCE8799" bold="true" box="[770,788,1906,1930]" pageId="10" pageNumber="11">E</emphasis>
, MPEF-IC 1041;
<emphasis id="B96D7724BB03FFAFFC6C6B70FC398799" bold="true" box="[981,995,1906,1930]" pageId="10" pageNumber="11">F</emphasis>
, MACN-SC 4354;
<emphasis id="B96D7724BB03FFAFFB0C6B70FB1D8799" bold="true" box="[1205,1223,1906,1930]" pageId="10" pageNumber="11">G</emphasis>
, MACN 18622, the white arrows indicate the bilobate outline;
<emphasis id="B96D7724BB03FFAFFDD06B97FDA487BE" bold="true" box="[617,638,1941,1965]" pageId="10" pageNumber="11">H</emphasis>
, detail of the third manus-pes sets of MPM-PIc-3952;
<emphasis id="B96D7724BB03FFAFFB6B6B97FB0687BE" bold="true" box="[1234,1244,1941,1965]" pageId="10" pageNumber="11">I</emphasis>
, MPM-PIc-3952. Scale bars: 2 cm in C, E, F, and H, 5 cm in A, D, G, and I, and 10 cm in B. The black arrow in
<emphasis id="B96D7724BB03FFAFFB296BBAFB7987C3" bold="true" box="[1168,1187,1976,2000]" pageId="10" pageNumber="11">C</emphasis>
,
<emphasis id="B96D7724BB03FFAFFB086BBAFB1987C3" bold="true" box="[1201,1219,1976,2000]" pageId="10" pageNumber="11">E</emphasis>
, and
<emphasis id="B96D7724BB03FFAFFB476BBAFAD487C3" bold="true" box="[1278,1294,1976,2000]" pageId="10" pageNumber="11">F</emphasis>
indicates the tail mark. m, manus imprint; p, pes imprint.
</paragraph>
</caption>
<subSubSection id="C303F8BDBB02FFAEFF7C6C95FCF08227" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8BA6AB36BB02FFAEFF7C6C95FBCC815F" blockId="11.[151,1437,151,1995]" pageId="11" pageNumber="12">
<emphasis id="B96D7724BB02FFAEFF7C6C95FDC680A2" bold="true" box="[197,540,151,177]" pageId="11" pageNumber="12">Remarks and comparisons.</emphasis>
The ichnospecies
<emphasis id="B96D7724BB02FFAEFD4E6C9AFC7680A2" box="[759,940,152,177]" italics="true" pageId="11" pageNumber="12">Ameghinichnus</emphasis>
<taxonomicName id="4C19D0B5BB02FFAEFC0D6C9AFAD080A1" authority="(Ellenberger, 1970)" authorityName="Ellenberger" authorityYear="1970" baseAuthorityName="Ellenberger" baseAuthorityYear="1970" box="[948,1290,152,178]" class="Amphibia" genus="Amphibiopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="mirabilis">
<emphasis id="B96D7724BB02FFAEFC0D6C9AFBC480A2" box="[948,1054,152,177]" italics="true" pageId="11" pageNumber="12">mirabilis</emphasis>
(
<bibRefCitation id="EF88D6C7BB02FFAEFB976C9AFAD880A1" author="Ellenberger" box="[1070,1282,152,178]" lastPageId="11" lastPageNumber="20" pageId="11" pageNumber="19" refString="Ellenberger, P. (1970) Les niveaux paleontologiques de premiere apparition des mammiferes primordiaux en Afrique du Sud et leur ichnologie. Establissement de zones stratigraphiques detaillees dans le Stormberg du Lesotho (Afrique du Sud) (Trias superieur a Jurassique). Abstracts 2 Gondwana Symposium, South Africa, 1970, pp. 343 - 370." type="journal article" year="1970">Ellenberger, 1970</bibRefCitation>
)
</taxonomicName>
nov. comb., from the Elliot Formation in the Karoo basin, is composed of large-sized, symmetric manus and pes impressions subequal in size, with digit imprints splayed and subequal in length.
<emphasis id="B96D7724BB02FFAEFBE96CE7FA4680ED" box="[1104,1436,229,254]" italics="true" pageId="11" pageNumber="12">
Ameghinichnus
<taxonomicName id="4C19D0B5BB02FFAEFAB46CE7FA4680ED" box="[1293,1436,229,254]" class="Amphibia" genus="Amphibiopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="patagonicus">patagonicus</taxonomicName>
</emphasis>
displays a track morphology comparable to
<taxonomicName id="4C19D0B5BB02FFAEFD246D0EFCF38136" box="[669,809,268,294]" class="Amphibia" genus="Amphibiopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="mirabilis">
<emphasis id="B96D7724BB02FFAEFD246D0EFD758136" box="[669,687,268,293]" italics="true" pageId="11" pageNumber="12">A</emphasis>
.
<emphasis id="B96D7724BB02FFAEFD066D0EFCF38136" box="[703,809,268,293]" italics="true" pageId="11" pageNumber="12">mirabilis</emphasis>
</taxonomicName>
, but as it lacks the trackway parameters and is larger in size. It is not possible to assure the synonymy between both ichnospecies.
</paragraph>
<paragraph id="8BA6AB36BB02FFAEFF7C6D5BFD4381D3" blockId="11.[151,1437,151,1995]" pageId="11" pageNumber="12">
<emphasis id="B96D7724BB02FFAEFF7C6D5BFDFB8161" box="[197,545,345,370]" italics="true" pageId="11" pageNumber="12">
Ameghinichnus
<taxonomicName id="4C19D0B5BB02FFAEFE326D5BFDFB8161" box="[395,545,345,370]" class="Amphibia" genus="Amphibiopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="patagonicus">patagonicus</taxonomicName>
</emphasis>
can be distinguished from the specimens from the Newark basin,
<collectingCountry id="F30EEBA6BB02FFAEFAE16D5BFA428160" box="[1368,1432,345,371]" name="United States of America" pageId="11" pageNumber="12">USA</collectingCountry>
, mentioned as
<emphasis id="B96D7724BB02FFAEFE866D82FE2D818A" box="[319,503,384,409]" italics="true" pageId="11" pageNumber="12">Ameghinichnus</emphasis>
n. isp. by
<bibRefCitation id="EF88D6C7BB02FFAEFDCE6D82FBFE8189" author="Olsen" box="[631,1060,384,410]" pageId="11" pageNumber="21" refString="Olsen, P. E. &amp; Rainforth, E. C. (2001) The &quot; Age of Dinosaurs &quot; in the Newark Basin, with special reference to the Lower Hudson Valley. New York State Geological Association Guidebook, pp. 59 - 176." type="book chapter" year="2001">Olsen and Rainforth (2001: fig. 59)</bibRefCitation>
by their lack of the sharp claw marks, as well as being almost half the size.
</paragraph>
<paragraph id="8BA6AB36BB02FFAEFF7C6DCFFCF08227" blockId="11.[151,1437,151,1995]" pageId="11" pageNumber="12">
The specimen presented by
<bibRefCitation id="EF88D6C7BB02FFAEFDB56DCFFCCB81F4" author="Gierlinski" box="[524,785,461,487]" pageId="11" pageNumber="20" refString="Gierlinski, G., Pienkowski, G. &amp; Niedzwiedzki, G. (2004) Tetrapod track assemblage in the Hettangian of Soltykow, Poland, and its paleoenvironmental background. Ichnos, 11, 195 - 213." type="journal article" year="2004">
Gierliński
<emphasis id="B96D7724BB02FFAEFD316DCFFD6181F5" box="[648,699,461,486]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. (2004
</bibRefCitation>
,
<bibRefCitation id="EF88D6C7BB02FFAEFCA46DCFFC8381F4" author="Gierlinski" box="[797,857,461,487]" pageId="11" pageNumber="20" refString="Gierlinski, G., Niedwiedzki, G. &amp; Pienkowski, G. (2005) Early Hettangian vertebrate ichnoassemblage from Poland. Abstracts of Tracking dinosaur origins. The Triassic / Jurassic terretrial transition, Conference, Utah, 3 - 4." type="book chapter" year="2005">2005</bibRefCitation>
) from the Lower Jurassic of Sołtyków,
<collectingCountry id="F30EEBA6BB02FFAEFA916DCFFAA681F4" box="[1320,1404,461,487]" name="Poland" pageId="11" pageNumber="12">Poland</collectingCountry>
, is interpreted here as a left track. It lacks sufficient details to make a comparison with
<emphasis id="B96D7724BB02FFAEFB636DF6FEFC8220" italics="true" pageId="11" pageNumber="12">
Ameghinichnus
<taxonomicName id="4C19D0B5BB02FFAEFF2E6E18FEFC8220" box="[151,294,538,563]" class="Amphibia" genus="Amphibiopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="patagonicus">patagonicus</taxonomicName>
</emphasis>
to define an ichnotaxonomical relationship.
</paragraph>
</subSubSection>
<subSubSection id="C303F8BDBB02FFAEFF7C6E42FF1687D8" pageId="11" pageNumber="12" type="description">
<paragraph id="8BA6AB36BB02FFAEFF7C6E42FE3F82BB" blockId="11.[151,1437,151,1995]" pageId="11" pageNumber="12">
<emphasis id="B96D7724BB02FFAEFF7C6E42FEBE8249" bold="true" box="[197,356,576,602]" pageId="11" pageNumber="12">Description.</emphasis>
The specimens of
<emphasis id="B96D7724BB02FFAEFDF66E43FC738249" box="[591,937,577,602]" italics="true" pageId="11" pageNumber="12">
Ameghinichnus
<taxonomicName id="4C19D0B5BB02FFAEFCAD6E43FC738249" box="[788,937,577,602]" class="Amphibia" genus="Amphibiopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="patagonicus">patagonicus</taxonomicName>
</emphasis>
are represented by numerous tracks and trackways produced by a quadrupedal trackmaker.
<tableCitation id="C69B9E8DBB02FFAEFD496E6AFC9D8291" box="[752,839,616,642]" captionStart="TABLE 1" captionStartId="3.[151,244,809,833]" captionTargetBox="[151,1436,984,1780]" captionTargetPageId="3" captionText="TABLE 1. Summary of measurements of track and trackway parameters of Ameghinichnus patagonicus and A. manantialensis. Linear measurements in millimeters. For abbreviations, see Figure 2." httpUri="http://table.plazi.org/id/DF66FBBEBB0AFFA6FF2E6F2BFBFF8375" pageId="11" pageNumber="12" tableUuid="DF66FBBEBB0AFFA6FF2E6F2BFBFF8375">Table 1</tableCitation>
summarizes the measurements from 28 trackways accounting for 405 imprints.
</paragraph>
<paragraph id="8BA6AB36BB02FFAEFF7C6EB7FA4F830F" blockId="11.[151,1437,151,1995]" pageId="11" pageNumber="12">The trackways are composed of manus-pes sets, both pentadactyl tracks, which are nearly symmetrical in relation to the major imprint axis. Manus and pes digit impressions are slender, with a maximum width up to 1.5 mm and may be parallel-sided. They lack phalangeal pads and claw marks, though have a distal swelling.</paragraph>
<paragraph id="8BA6AB36BB02FFAEFF7C6F2BFADF83CD" blockId="11.[151,1437,151,1995]" pageId="11" pageNumber="12">The average width and length of the hand impressions are 11.5 mm and 8.2 mm, respectively. Digit impressions are subequal in size and shape, with an average length, in decreasing order, of: I: 2.8 mm, II and V: 3.2 mm, IV: 3.4 mm, and III: 3.5 mm. The angles formed by them are relatively equidistant, having an average of: I-II: 42º, II-III: 42º, III-IV: 37º, and IV-V: 42º; the I-V displays an angle with a mean of 151º. Manus imprints show an outward (negative) rotation relative to the midline, with a mean of -16º.</paragraph>
<paragraph id="8BA6AB36BB02FFAEFF7C6FE8FDDC848C" blockId="11.[151,1437,151,1995]" pageId="11" pageNumber="12">
The average width of the footprints is
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, while the average length is
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. Digit impressions are subequal as in the case of the manus, with an average length of: I: 3.0 mm, IV: 3.6 mm, III and V: 3.7 mm, and II: 4.0 mm. The average angles formed by them are as follows: I-II: 42º, II-III: 30º, III-IV: 26º, and IV-V: 58º; the I–V displays an angle with a mean of 152º. Pes imprints show an inward (positive) rotation relative to the midline with an average of 31º.
</paragraph>
<paragraph id="8BA6AB36BB02FFAEFF7C68AEFD108529" blockId="11.[151,1437,151,1995]" pageId="11" pageNumber="12">
Both manus and pes sole display metacarpal and metatarsal imprints, a posterocentered depression, and a posterior bilobate outline which is only visible in the best- preserved tracks. When this posterior bilobated outline is present, the length of the track (named here as total length, see figure 2) is about 9.2 mm for hand prints and 10.9 mm for footprints (
<figureCitation id="1322B7B3BB02FFAEFD946922FDA78529" box="[557,637,1312,1338]" captionStart="FIGURE 3" captionStartId="9.[151,255,1863,1887]" captionTargetBox="[187,1413,202,1830]" captionTargetId="figure@9.[171,1429,194,1842]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of the type series of Ameghinichnus patagonicus. A, MLP 60 - X- 31 - 1; B, MLP 60 - X- 31 - 14 B, holotype (right trackway: 1) and MLP 60 - X- 31 - 14 A, paratype (left trackway: 2), the black arrows indicate the moving direction of the trackmaker; C, PVL 2303; D, PVL 2302, the white arrow indicates the bilobate outline; E, detail of the two first manus-pes sets of MLP 60 - X- 31 - 14 B; F, MLP 60 - X- 31 - 15; G, MLP 60 - X- 31 - 3. A-C, D, F and G, paratypes. Scale bars: 2 cm in D and E, 5 cm in the rest. m, manus imprint; p, pes imprint." httpUri="https://zenodo.org/record/189673/files/figure.png" pageId="11" pageNumber="12">Figs. 3</figureCitation>
d,
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).
</paragraph>
<paragraph id="8BA6AB36BB02FFAEFF7C6944FECC8631" blockId="11.[151,1437,151,1995]" pageId="11" pageNumber="12">
The manus-pes sets can display two arrangements: either ipsilateral with the manus and pes imprints on the same side of the body of the trackmaker in alternate arrangement (sensu
<bibRefCitation id="EF88D6C7BB02FFAEFBA7696FFB628594" author="Trewin" box="[1054,1208,1389,1415]" pageId="11" pageNumber="21" refString="Trewin, N. H. (1994) A draft system for the identification and description of arthropod trackways. Palaeontology, 37, 811 - 823." type="journal article" year="1994">Trewin 1994</bibRefCitation>
; e.g.
<figureCitation id="1322B7B3BB02FFAEFB4E696FFA928594" box="[1271,1352,1389,1415]" captionStart="FIGURE 3" captionStartId="9.[151,255,1863,1887]" captionTargetBox="[187,1413,202,1830]" captionTargetId="figure@9.[171,1429,194,1842]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of the type series of Ameghinichnus patagonicus. A, MLP 60 - X- 31 - 1; B, MLP 60 - X- 31 - 14 B, holotype (right trackway: 1) and MLP 60 - X- 31 - 14 A, paratype (left trackway: 2), the black arrows indicate the moving direction of the trackmaker; C, PVL 2303; D, PVL 2302, the white arrow indicates the bilobate outline; E, detail of the two first manus-pes sets of MLP 60 - X- 31 - 14 B; F, MLP 60 - X- 31 - 15; G, MLP 60 - X- 31 - 3. A-C, D, F and G, paratypes. Scale bars: 2 cm in D and E, 5 cm in the rest. m, manus imprint; p, pes imprint." httpUri="https://zenodo.org/record/189673/files/figure.png" pageId="11" pageNumber="12">Figs. 3</figureCitation>
a,g), or both sets accounting the four limbs of the trackmaker in a transverse group as opposite arrangement (sensu
<bibRefCitation id="EF88D6C7BB02FFAEFF2E69B9FEEA85C6" author="Trewin" box="[151,304,1467,1493]" pageId="11" pageNumber="21" refString="Trewin, N. H. (1994) A draft system for the identification and description of arthropod trackways. Palaeontology, 37, 811 - 823." type="journal article" year="1994">Trewin 1994</bibRefCitation>
), with one manus and pes set on one side, opposite the corresponding pair on the other side (e.g.
<figureCitation id="1322B7B3BB02FFAEFF2E69E3FF0685E8" box="[151,220,1505,1531]" captionStart="FIGURE 3" captionStartId="9.[151,255,1863,1887]" captionTargetBox="[187,1413,202,1830]" captionTargetId="figure@9.[171,1429,194,1842]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of the type series of Ameghinichnus patagonicus. A, MLP 60 - X- 31 - 1; B, MLP 60 - X- 31 - 14 B, holotype (right trackway: 1) and MLP 60 - X- 31 - 14 A, paratype (left trackway: 2), the black arrows indicate the moving direction of the trackmaker; C, PVL 2303; D, PVL 2302, the white arrow indicates the bilobate outline; E, detail of the two first manus-pes sets of MLP 60 - X- 31 - 14 B; F, MLP 60 - X- 31 - 15; G, MLP 60 - X- 31 - 3. A-C, D, F and G, paratypes. Scale bars: 2 cm in D and E, 5 cm in the rest. m, manus imprint; p, pes imprint." httpUri="https://zenodo.org/record/189673/files/figure.png" pageId="11" pageNumber="12">Fig. 3</figureCitation>
e, trackway
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4b). In both cases, the footprints are located externally and overstep or slightly overlap the manus.
</paragraph>
<paragraph id="8BA6AB36BB02FFAEFF7C6A2CFE4A86F0" blockId="11.[151,1437,151,1995]" pageId="11" pageNumber="12">
When the sets have an alternate arrangement, the trackways display an average pace angulation of 143º for the manus, and of 114º for the pes. The average stride length of the manus and pes imprints is
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and 81.2 mm, respectively. The inner trackway width between manus imprints averages 2.7 mm, and the width between footprints averages of 17.5 mm, whereas the average outer trackways width is 23.6 mm for manus and 38.8 mm for pes.
</paragraph>
<paragraph id="8BA6AB36BB02FFAEFF7C6AF2FCBA8723" blockId="11.[151,1437,151,1995]" pageId="11" pageNumber="12">When the sets have a transverse or opposite arrangement, the distance between two consecutive groups, named herein as a hop or jump, displays a mean of 91.6 mm.</paragraph>
<paragraph id="8BA6AB36BB02FFAEFF7C6B3FFF1687D8" blockId="11.[151,1437,151,1995]" pageId="11" pageNumber="12">
The trackways may show tail marks, in a continuous and curved or sinuous pattern when the sets have an alternative arrangement (e.g.
<figureCitation id="1322B7B3BB02FFAEFE4F6B66FDE4876D" box="[502,574,1892,1918]" captionStart="FIGURE 4" captionStartId="10.[151,256,1837,1861]" captionTargetBox="[261,1332,181,1802]" captionTargetId="figure@10.[245,1348,168,1812]" captionTargetPageId="10" captionText="FIGURE 4. Photographs of specimens of Ameghinichnus patagonicus. A, MPM-PIc- 3768 (first manus-pes set to the left) and 3769 (first manus-pes set to the right); B, MPEF-IC 1016 at left, indicated with a 1, MPEF-IC 1015 at rigth, indicated with a 2; C, MPEF-IC 1023; D, MACN 18619; E, MPEF-IC 1041; F, MACN-SC 4354; G, MACN 18622, the white arrows indicate the bilobate outline; H, detail of the third manus-pes sets of MPM-PIc- 3952; I, MPM-PIc- 3952. Scale bars: 2 cm in C, E, F, and H, 5 cm in A, D, G, and I, and 10 cm in B. The black arrow in C, E, and F indicates the tail mark. m, manus imprint; p, pes imprint." httpUri="https://zenodo.org/record/189674/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
c, trackway
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4b), or discrete (average length of 7.7 mm) and straight when they are opposite (e.g.
<figureCitation id="1322B7B3BB02FFAEFE506B89FDF587B6" box="[489,559,1931,1957]" captionStart="FIGURE 4" captionStartId="10.[151,256,1837,1861]" captionTargetBox="[261,1332,181,1802]" captionTargetId="figure@10.[245,1348,168,1812]" captionTargetPageId="10" captionText="FIGURE 4. Photographs of specimens of Ameghinichnus patagonicus. A, MPM-PIc- 3768 (first manus-pes set to the left) and 3769 (first manus-pes set to the right); B, MPEF-IC 1016 at left, indicated with a 1, MPEF-IC 1015 at rigth, indicated with a 2; C, MPEF-IC 1023; D, MACN 18619; E, MPEF-IC 1041; F, MACN-SC 4354; G, MACN 18622, the white arrows indicate the bilobate outline; H, detail of the third manus-pes sets of MPM-PIc- 3952; I, MPM-PIc- 3952. Scale bars: 2 cm in C, E, F, and H, 5 cm in A, D, G, and I, and 10 cm in B. The black arrow in C, E, and F indicates the tail mark. m, manus imprint; p, pes imprint." httpUri="https://zenodo.org/record/189674/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
e, trackway
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4b). Both kinds of tail marks have an average width of 4.1 mm.
</paragraph>
</subSubSection>
</treatment>
</document>
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