265 lines
30 KiB
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265 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e85571" ID-Pensoft-Pub="1864-8312-81-409" ID-Pensoft-UUID="1F7030AA7B0B54EAB1846C6B1339DB56" ID-ZooBank="764845DB2D5D4E7E9E12DCA816F5BCD8" ModsDocID="1864-8312-81-409" checkinTime="1682026883038" checkinUser="pensoft" docAuthor="Garcia, Andres F. & Villarreal, Osvaldo" docDate="2023" docId="FB35BEA7C56150DCA491D659C3684B8E" docLanguage="en" docName="ArthropodSystPhyl 81: 409-437" docOrigin="Arthropod Systematics & amp; Phylogeny 81" docPubDate="2023-04-20" docSource="http://dx.doi.org/10.3897/asp.81.e85571" docTitle="Paravima lokura García & Villarreal 2023, sp. nov." docType="treatment" docUuid="4F6A4D10-9436-4F98-AD28-5B7E3D0B1B4A" docUuidSource="ZooBank" docVersion="1" id="1F7030AA7B0B54EAB1846C6B1339DB56" lastPageNumber="409" masterDocId="1F7030AA7B0B54EAB1846C6B1339DB56" masterDocTitle="Looks are deceiving: a cladistic analysis, three new species, and a new diagnosis of Paravima Caporiacco, 1951 (Opiliones: Agoristenidae)" masterLastPageNumber="437" masterPageNumber="409" pageNumber="409" updateTime="1682026883038" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Looks are deceiving: a cladistic analysis, three new species, and a new diagnosis of Paravima Caporiacco, 1951 (Opiliones: Agoristenidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Garcia, Andres F.</mods:namePart>
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<mods:affiliation>Departamento de Invertebrados, Museu Nacional / Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, Sao Cristovao, 20.940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">agarciarinc@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Villarreal, Osvaldo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5355-3723</mods:nameIdentifier>
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<mods:affiliation>Departamento de Invertebrados, Museu Nacional / Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, Sao Cristovao, 20.940 - 040, Rio de Janeiro, RJ, Brazil & Museo del Instituto de Zoologia Agricola, Facultad de Agronomia, Universidad Central de Venezuela, Maracay, Aragua, Venezuela</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>81</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>409</mods:start>
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<mods:end>437</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.81.e85571</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.81.e85571</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-81-409</mods:identifier>
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<mods:identifier type="ZooBank">764845DB2D5D4E7E9E12DCA816F5BCD8</mods:identifier>
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<mods:identifier type="Pensoft-UUID">1F7030AA7B0B54EAB1846C6B1339DB56</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:4F6A4D10-9436-4F98-AD28-5B7E3D0B1B4A" httpUri="http://treatment.plazi.org/id/FB35BEA7C56150DCA491D659C3684B8E" lastPageNumber="409" pageId="0" pageNumber="409">
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<subSubSection pageId="0" pageNumber="409" type="nomenclature">
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<paragraph pageId="0" pageNumber="409">
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<taxonomicName LSID="https://zoobank.org/4F6A4D10-9436-4F98-AD28-5B7E3D0B1B4A" authority="García & Villarreal, 2023" authorityName="García & Villarreal" authorityYear="2023" class="Arachnida" family="Agoristenidae" genus="Paravima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paravima lokura" order="Opiliones" pageId="0" pageNumber="409" phylum="Arthropoda" rank="species" species="lokura" status="sp. nov.">Paravima lokura</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="409">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="409" type="description">
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<paragraph pageId="0" pageNumber="409">
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. Habitus: A Panoramic view; B dorsal view; C ventral view; D lateral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409">Figs 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">, 6</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Map showing the geographic distribution of Paravima in Northern South America. The background colored shapes refer to the biogeographic provinces of Morrone et al. (2022)." figureDoi="10.3897/asp.81.e85571.figure18" httpUri="https://binary.pensoft.net/fig/840450" pageId="0" pageNumber="409">, 18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="409" type="diagnosis">
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<paragraph pageId="0" pageNumber="409">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="409">
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Ocularium tuberculate. Mesotergal area III with large conical paramedian tubercles, close to each other (Figs
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. Habitus: A Panoramic view; B dorsal view; C ventral view; D lateral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409">5D, E</figureCitation>
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) (in
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<taxonomicName lsidName="P. propespelunca" pageId="0" pageNumber="409" rank="species" species="propespelunca">
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<emphasis italics="true" pageId="0" pageNumber="409">P. propespelunca</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="P. flumencaurimarensis" pageId="0" pageNumber="409" rank="species" species="flumencaurimarensis">
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<emphasis italics="true" pageId="0" pageNumber="409">P. flumencaurimarensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="P. goodnightorum" pageId="0" pageNumber="409" rank="species" species="goodnightorum">
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<emphasis italics="true" pageId="0" pageNumber="409">P. goodnightorum</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="P. locumida" pageId="0" pageNumber="409" rank="species" species="locumida">
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<emphasis italics="true" pageId="0" pageNumber="409">P. locumida</emphasis>
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</taxonomicName>
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, and
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<taxonomicName lsidName="P. morritomacairensis" pageId="0" pageNumber="409" rank="species" species="morritomacairensis">
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<emphasis italics="true" pageId="0" pageNumber="409">P. morritomacairensis</emphasis>
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</taxonomicName>
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ocularium unarmed and mesotergal area III with separated conical/mammilliform tubercles).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="409" type="description">
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<paragraph pageId="0" pageNumber="409">Description.</paragraph>
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<paragraph pageId="0" pageNumber="409">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="409">Holotype</emphasis>
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(ICN-Ao-1490*). - Measurements: CL: 1.2, AL: 1.8, CW: 2.0, AW: 2.6, BaCh: 0.4, IOD: 0.9; Leg I (Tr: 0.3, Fe: 4.0, Pa: 0.9, Ti: 3.0, Mt: 4.3, Ta: 1.7); Leg II (Tr: 0.4, Fe: 9.5, Pa: 1.1, Ti: 8.0, Mt: 10.1, Ta: 3.5); Leg III (Tr: 0.7, Fe: 8.0, Pa: 1.6, Ti: 5.0, Mt: 9.0, Ta: 1.9); Leg IV (Tr: 0.9, Fe: 10.1, Pa: 1.3, Ti: 7.0, Mt: 10.3, Ta: 2.8). -
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<emphasis italics="true" pageId="0" pageNumber="409">Dorsum</emphasis>
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: Anterior and lateral margins of DS smooth. Ocularium with granules (Figs
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. Habitus: A Panoramic view; B dorsal view; C ventral view; D lateral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409">5B, D</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6A</figureCitation>
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). Mesotergum delimited, divided into four areas, mostly smooth. Area I divided into two halves, each one with one small tubercle; area II-IV undivided; II with a pair of medium-sized tubercles close to the medial axis of the body (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6A, B</figureCitation>
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); III with a pair of conical wide tubercles close to each other (Figs
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. Habitus: A Panoramic view; B dorsal view; C ventral view; D lateral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409">5D, E</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6B</figureCitation>
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); IV with four small tubercles. Posterior border of scutum straight, with some tubercles. Free tergites I-II and anal operculum with some tubercles (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6A</figureCitation>
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). -
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<emphasis italics="true" pageId="0" pageNumber="409">Venter</emphasis>
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: Coxae I-IV with some granules (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. Habitus: A Panoramic view; B dorsal view; C ventral view; D lateral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409">5C</figureCitation>
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). Coxa I with one medial tubercle on the anterior margin, a longitudinal row of tubercles at the medial region, a group of three tubercles on the anterodistal margin, and two tubercles close to the posterodistal margin (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6C</figureCitation>
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); coxa II longer than coxa I; coxa III longer than I and II; coxa IV backward projected (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. Habitus: A Panoramic view; B dorsal view; C ventral view; D lateral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409">5C</figureCitation>
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). Sternites with a row of small tubercles each. Stigmatic area smooth. Stigmata large, oval and transverse (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. Habitus: A Panoramic view; B dorsal view; C ventral view; D lateral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409">5C</figureCitation>
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). -
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<emphasis italics="true" pageId="0" pageNumber="409">Chelicerae</emphasis>
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: Segment I rectangular, with well-marked bulla (Figs
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. Habitus: A Panoramic view; B dorsal view; C ventral view; D lateral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409">5B</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6A</figureCitation>
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), one ectal subdistal tubercle, and two tubercles on the proximal border (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paravima locumida Gonzalez-Sponga, 1987 (MAGS 324) male. Habitus: A Dorsal view; B lateral view; C anterior view; D posterior view; E ventral view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure4" httpUri="https://binary.pensoft.net/fig/840436" pageId="0" pageNumber="409">4B</figureCitation>
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). Chelicera swollen (Figs
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. Habitus: A Panoramic view; B dorsal view; C ventral view; D lateral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409">5B, E</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6B</figureCitation>
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). Hand with some sparse setiferous tubercles of different sizes in the fronto mesal portion, and a frontal row of small setiferous tubercles extending from the mesal to the ectal face of the chelicera close to the base of the fingers (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6D</figureCitation>
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). Fixed finger with the inner surface irregularly dentated. Movable finger with one trapezoid, sub-basal tooth and with the inner surface dentate (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6D</figureCitation>
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). -
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<emphasis italics="true" pageId="0" pageNumber="409">Pedipalps</emphasis>
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: Longer than DS length, smooth. Tr ventrally with one subapical setiferous tubercle. Fe with a ventromesal row of five setiferous tubercles (the third shorter than the others), and one large ventroectal setiferous tubercle in the distal portion (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. Habitus: A Panoramic view; B dorsal view; C ventral view; D lateral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409">5C</figureCitation>
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). Pa with one large mesal setiferous tubercle. Ti ectal iII, mesal IIi. Ta ectal IIi, mesal IIi (Figs
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. Habitus: A Panoramic view; B dorsal view; C ventral view; D lateral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409">5</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6B</figureCitation>
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). -
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<emphasis italics="true" pageId="0" pageNumber="409">Legs</emphasis>
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: Leg I smooth, legs II-IV with minute granules. Leg I filiform, the rest, getting steadily thicker from leg II to IV (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. Habitus: A Panoramic view; B dorsal view; C ventral view; D lateral view; E anterior view. Scale bars: 1 mm." figureDoi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409">5A</figureCitation>
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). Fe IV three times DS length. Tarsal formula: 9(3)-9(3)/20(3)-20(3)/8-9/8-8. -
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<emphasis italics="true" pageId="0" pageNumber="409">Penis</emphasis>
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: LP short and depressed, half-moon shaped, with anterolateral sharp corners dorsally pointed (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6E-G</figureCitation>
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). Malleus carrying the branched MS-A-B (three pairs of MS-A and one pair of MS-B); MS-B at the same level of MS-A1 (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6G</figureCitation>
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). MS-D1-2 long, located in a vertical line on a keel between the LP and the base of the stylus (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6G</figureCitation>
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). MS-E2 large and branched (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6F</figureCitation>
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). Stylus sinuous, surpassing the lamina parva, narrower at distal region, tip irregular (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paravima lokura sp. nov. (ICN-Ao- 1490 *) male holotype. A Habitus, dorsal view; B lateral view; C left coxa I, ventral view; D chelicera, anterior view. Apical part of the penis: E Dorsal view; F ventral view; G lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS)." figureDoi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409">6G</figureCitation>
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). -
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<emphasis italics="true" pageId="0" pageNumber="409">Color</emphasis>
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(in ethanol): Carapace and mesotergum reticulated Deep Brown (56) on Brownish Orange (54). Area III tubercles Brownish Black (65). Pedipalps and chelicerae Strong Orange Yellow (68). Posterior border and free tergites Dark Brown (59). Legs I-IV Deep Orange Yellow (69). -
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<emphasis italics="true" pageId="0" pageNumber="409">Female</emphasis>
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: Unknown.
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</paragraph>
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<caption doi="10.3897/asp.81.e85571.figure5" httpUri="https://binary.pensoft.net/fig/840437" pageId="0" pageNumber="409" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="409">
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<emphasis bold="true" pageId="0" pageNumber="409">Figure 5.</emphasis>
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<taxonomicName authorityName="García & Villarreal" authorityYear="2023" class="Arachnida" family="Agoristenidae" genus="Paravima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paravima lokura" order="Opiliones" pageId="0" pageNumber="409" phylum="Arthropoda" rank="species" species="lokura">
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<emphasis italics="true" pageId="0" pageNumber="409">Paravima lokura</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="409">sp. nov.</emphasis>
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(ICN-Ao-1490*) male holotype. Habitus:
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<emphasis bold="true" pageId="0" pageNumber="409">A</emphasis>
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Panoramic view;
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<emphasis bold="true" pageId="0" pageNumber="409">B</emphasis>
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dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="409">C</emphasis>
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ventral view;
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<emphasis bold="true" pageId="0" pageNumber="409">D</emphasis>
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lateral view;
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<emphasis bold="true" pageId="0" pageNumber="409">E</emphasis>
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anterior view. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<caption doi="10.3897/asp.81.e85571.figure6" httpUri="https://binary.pensoft.net/fig/840438" pageId="0" pageNumber="409" start="Figure 6" startId="F6">
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|
<paragraph pageId="0" pageNumber="409">
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<emphasis bold="true" pageId="0" pageNumber="409">Figure 6.</emphasis>
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<taxonomicName authorityName="García & Villarreal" authorityYear="2023" class="Arachnida" family="Agoristenidae" genus="Paravima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paravima lokura" order="Opiliones" pageId="0" pageNumber="409" phylum="Arthropoda" rank="species" species="lokura">
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<emphasis italics="true" pageId="0" pageNumber="409">Paravima lokura</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="409">sp. nov.</emphasis>
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|
(ICN-Ao-1490*) male holotype.
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<emphasis bold="true" pageId="0" pageNumber="409">A</emphasis>
|
|
Habitus, dorsal view;
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|
<emphasis bold="true" pageId="0" pageNumber="409">B</emphasis>
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lateral view;
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|
<emphasis bold="true" pageId="0" pageNumber="409">C</emphasis>
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|
left coxa I, ventral view;
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<emphasis bold="true" pageId="0" pageNumber="409">D</emphasis>
|
|
chelicera, anterior view. Apical part of the penis:
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<emphasis bold="true" pageId="0" pageNumber="409">E</emphasis>
|
|
Dorsal view;
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|
<emphasis bold="true" pageId="0" pageNumber="409">F</emphasis>
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ventral view;
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<emphasis bold="true" pageId="0" pageNumber="409">G</emphasis>
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lateral view. Scale bars: 1 mm (penis with not scaled figures). Abbreviation: Macrosetae (MS).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="409" type="etymology">
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<paragraph pageId="0" pageNumber="409">Derivatio nominis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="409">
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In the Tunebo language, spoken by the indigenous people that inhabit the region where the species was collected,
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<taxonomicName authorityName="García & Villarreal" authorityYear="2023" class="Arachnida" family="Agoristenidae" genus="Paravima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paravima lokura" order="Opiliones" pageId="0" pageNumber="409" phylum="Arthropoda" rank="species" species="lokura">
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|
<emphasis italics="true" pageId="0" pageNumber="409">Paravima lokura</emphasis>
|
|
</taxonomicName>
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|
means spider. Noun in apposition.
|
|
</paragraph>
|
|
</subSubSection>
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<subSubSection pageId="0" pageNumber="409" type="distribution">
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|
<paragraph pageId="0" pageNumber="409">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="409">
|
|
Known just from the type locality, PNN
|
|
<normalizedToken originalValue="Tamá">Tama</normalizedToken>
|
|
, Norte de Santander department, in the
|
|
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
|
biogeographic province (
|
|
<bibRefCitation DOI="https://doi.org/10.1590/0001-3765202220211167" author="Morrone, JJ" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="409" refId="B34" refString="Morrone, JJ, Escalante, T, Rodriguez-Tapia, G, Carmona, A, Arana, M, Mercado-Gomez, JD, 2022. Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias 94: e20211167. https://doi.org/10.1590/0001-3765202220211167" title="Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias 94: e 20211167." url="https://doi.org/10.1590/0001-3765202220211167" year="2022">Morrone et al. 2022</bibRefCitation>
|
|
) (Fig.
|
|
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Map showing the geographic distribution of Paravima in Northern South America. The background colored shapes refer to the biogeographic provinces of Morrone et al. (2022)." figureDoi="10.3897/asp.81.e85571.figure18" httpUri="https://binary.pensoft.net/fig/840450" pageId="0" pageNumber="409">18</figureCitation>
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|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="409" type="materials_examined">
|
|
<paragraph pageId="0" pageNumber="409">Material examined.</paragraph>
|
|
<paragraph pageId="0" pageNumber="409">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="409">Type material</emphasis>
|
|
:
|
|
<materialsCitation collectingDate="1999-06-25" collectorName="V. R. Mayusa" country="COLOMBIA" elevation="2170" location="Parque Nacional Natural Tama" specimenCount="♂" stateProvince="Norte de Santander" typeStatus="Holotype">
|
|
<typeStatus>Holotype</typeStatus>
|
|
<specimenCount>♂</specimenCount>
|
|
,
|
|
<collectingCountry name="Colombia">COLOMBIA</collectingCountry>
|
|
,
|
|
<collectingRegion country="Colombia" name="Norte de Santander">Norte de Santander</collectingRegion>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:FB35BEA7C56150DCA491D659C3684B8E:7DDAB82A40D14EB0AC7DBBBAA75C2F7F" country="COLOMBIA" name="Parque Nacional Natural Tama" stateProvince="Norte de Santander">
|
|
Parque Nacional Natural
|
|
<normalizedToken originalValue="Tamá">Tama</normalizedToken>
|
|
</location>
|
|
| [7.26214° -72.25064°] |
|
|
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.17" unit="m" value="2170.0">
|
|
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.17" unit="m" value="2170.0">2170 m</elevation>
|
|
</quantity>
|
|
|
|
|
<collectingDate value="1999-06-25">25.vi.1999</collectingDate>
|
|
,
|
|
<collectorName>V.R. Mayusa</collectorName>
|
|
leg. (ICN-Ao-1490*).
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |