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<document id="C8B4E09B6B396BD576F37269F9B0596C" ID-DOI="10.11646/zootaxa.5481.5.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="12783058" ID-ZooBank="B8DC6181-5F90-4071-AE4A-B121F5BF7C90" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1721383172061" checkinUser="plazi" docAuthor="Acosta, Luis E." docDate="2024" docId="DD5F54119F66FF8E70AFFF28FB8FF871" docLanguage="en" docName="zootaxa.5481.5.4.pdf" docOrigin="Zootaxa 5481 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.5.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acrographinotus tariquiae Acosta, 2024, sp. nov." docType="treatment" docUuid="58394B7D-EFC6-4DC9-A945-098BB5D75101" docUuidSource="ZooBank" docVersion="3" lastPageNumber="555" masterDocId="21662C699F63FF867038FFBFFF80FFAE" masterDocTitle="The two southernmost species of the Andean genus Acrographinotus Holmgren (Arachnida, Opiliones, Gonyleptidae) described from Bolivia and Argentina" masterLastPageNumber="562" masterPageNumber="547" pageNumber="552" updateTime="1721436848161" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="84D68D47AD82B57E13C7B090F435A897">The two southernmost species of the Andean genus Acrographinotus Holmgren (Arachnida, Opiliones, Gonyleptidae) described from Bolivia and Argentina</mods:title>
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<taxonomicName id="92F69E849F66FF8370AFFF28FE59FF1F" authorityName="Acosta" authorityYear="2024" box="[151,473,151,178]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="552" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F66FF8370AFFF28FE59FF1F" bold="true" box="[151,473,151,178]" italics="true" pageId="5" pageNumber="552">Acrographinotus tariquiae</emphasis>
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<figureCitation id="CDCDF9829F66FF8370AFFF60FE93FF57" box="[151,275,223,249]" captionStart="FIGURE 3" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[155,1431,313,1749]" captionTargetId="figure-90@6.[151,1435,309,1753]" captionTargetPageId="6" captionText="FIGURE 3. Acrographinotus tariquiae sp. nov. AF: HOlOtYpe ♂ (CBF). A. DORsal scutum, fRee teRgites (alsO shOwing dORsal anal operculum and VAP apophyses), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, VAP with apophyses (also showing apophysis of FT-III), right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. GJ: Armature variations on FT-III and VAP in tOpOtYpical ♂ paRatYpes. GH: Smallest α-male (CBF), IJ: β-male (CDA 000.914). G, I: ventRal view; H, J: lateral view. Scale lines: 2 mm (AE, GJ), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783064" httpUri="https://zenodo.org/record/12783064/files/figure.png" pageId="5" pageNumber="552">Figs. 3AJ</figureCitation>
,
<figureCitation id="CDCDF9829F66FF837127FF60FEE1FF57" box="[287,353,223,249]" captionStart="FIGURE 4" captionStartId="8.[151,250,1558,1582]" captionTargetBox="[277,1309,493,1531]" captionTargetId="figure-189@8.[273,1313,489,1535]" captionTargetPageId="8" captionText="FIGURE 4. Acrographinotus tariquiae sp. nov., distal end Of penis, HOlOtYpe ♂ (CBF). A: dORsal view; B: lateRal view; C: detail of stylus and ventral process. Scale lines: 0.1 mm (A, B), 0.05 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.12783067" httpUri="https://zenodo.org/record/12783067/files/figure.png" pageId="5" pageNumber="552">4AC</figureCitation>
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<taxonomicName id="92F69E849F66FF8370AFFE9BFEC2FE92" ID-CoL="7NMYV" authorityName="Holmgren" authorityYear="1917" box="[151,322,292,316]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="552" phylum="Arthropoda" rank="genus">
<emphasis id="678239159F66FF8370AFFE9BFEC2FE92" box="[151,322,292,316]" italics="true" pageId="5" pageNumber="552">Acrographinotus</emphasis>
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[sp. from
<collectingRegion id="97322BE59F66FF837193FE9BFE69FE92" box="[427,489,292,316]" country="Bolivia" name="Tarija" pageId="5" pageNumber="552">Tarija</collectingRegion>
, S
<collectingCountry id="2DE1A5979F66FF83723FFE9BFDD4FE92" box="[519,596,292,316]" name="Bolivia" pageId="5" pageNumber="552">Bolivia</collectingCountry>
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<treatmentCitation id="D457C3169F66FF837267FE9BFC87FE95" author="Acosta, L. E." box="[607,775,291,316]" page="58" pageId="5" pageNumber="552" year="2001">
<bibRefCitation id="316798F69F66FF837267FE9BFC87FE95" author="Acosta, L. E." box="[607,775,291,316]" pageId="5" pageNumber="552" pagination="58 - 66" refId="ref10602" refString="Acosta, L. E. (2001) The identity of Acrographinotus erectispina, with a review of the generic diagnosis, and the description of a new species (Opiliones, Gonyleptidae, Pachylinae). Bulletin of the British Arachnological Society, 12 (2), 58 - 66." type="journal article" year="2001">Acosta 2001: 58</bibRefCitation>
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, 60.
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<emphasis id="678239159F66FF8370AFFED5FEA5FE2D" bold="true" box="[151,293,361,388]" pageId="5" pageNumber="552">Type series.</emphasis>
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,
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<collectingRegion id="97322BE59F66FF837189FED4FD06FE2A" box="[433,646,363,388]" country="Bolivia" name="Tarija" pageId="5" pageNumber="552">Tarija Department</collectingRegion>
.
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(
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),
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<specimenCount id="43F02E8E9F66FF837342FED5FC28FE2A" box="[890,936,362,388]" count="3" pageId="5" pageNumber="552" type="male">3 ♂</specimenCount>
,
<specimenCount id="43F02E8E9F66FF83738EFED5FC62FE2A" box="[950,994,362,388]" count="3" pageId="5" pageNumber="552" type="female">3 ♀</specimenCount>
,
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(
<collectionCode id="33E77DC29F66FF83748FFED5FB71FE2A" box="[1207,1265,362,388]" country="Bolivia" httpUri="http://grbio.org/cool/ky5m-b42n" name="Coleccion Boliviana de Fauna" pageId="5" pageNumber="552">CBF</collectionCode>
),
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<specimenCount id="43F02E8E9F66FF83753FFED5FAB3FE2A" box="[1287,1331,362,388]" count="1" pageId="5" pageNumber="552" type="male">1 ♂</specimenCount>
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(
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),
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<typeStatus id="8A4D5BA59F66FF837198FE31FD80FE06" box="[416,512,398,424]" pageId="5" pageNumber="552" type="paratype">paRatYpe</typeStatus>
(
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),
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<materialsCitation id="E59EEF5A9F66FF8372E7FE31FC78FE62" collectingDate="1995-12-11" collectingDateMax="1995-12-13" collectingDateMin="1995-12-11" collectionCode="CBF, CDA" collectorName="P. Goloboff" country="Bolivia" county="Subida de La EscaleRa" elevation="2700" latitude="-21.991533" location="Fauna National Reserve" longLatPrecision="1" longitude="-64.6216" municipality="TaRiquia Flora" pageId="5" pageNumber="552" specimenCode="CDA 000.914" specimenCount="2" specimenCount-female="5" specimenCount-male="2" stateProvince="Tarija" typeStatus="paratype">
<specimenCount id="43F02E8E9F66FF8372E7FE31FC88FE06" box="[735,776,398,424]" count="2" pageId="5" pageNumber="552" type="male">2 ♂</specimenCount>
<typeStatus id="8A4D5BA59F66FF837337FE31FCFAFE06" box="[783,890,398,424]" pageId="5" pageNumber="552" type="paratype">paRatYpes</typeStatus>
(
<specimenCode id="05504D7C9F66FF8373BFFE31FBAAFE06" box="[903,1066,398,424]" pageId="5" pageNumber="552">CDA 000.914</specimenCode>
):
<collectingCounty id="BC289D8B9F66FF837403FE31FAB6FE06" box="[1083,1334,398,424]" pageId="5" pageNumber="552">Subida de La EscaleRa</collectingCounty>
,
<collectingMunicipality id="B52D7F7D9F66FF837507FE31FF52FE62" pageId="5" pageNumber="552">TaRiquía Flora</collectingMunicipality>
and
<location id="5029B3DC9F66FF83713FFE0DFD95FE62" LSID="urn:lsid:plazi:treatment:DD5F54119F66FF8E70AFFF28FB8FF871:5029B3DC9F66FF83713FFE0DFD95FE62" box="[263,533,434,460]" country="Bolivia" county="Subida de La EscaleRa" latitude="-21.991533" longLatPrecision="1" longitude="-64.6216" municipality="TaRiquia Flora" name="Fauna National Reserve" pageId="5" pageNumber="552" stateProvince="Tarija">Fauna National Reserve</location>
,
<date id="2148C3C79F66FF837218FE0DFC9FFE63" box="[544,799,434,461]" pageId="5" pageNumber="552" value="1995-12-11" valueMax="1995-12-13" valueMin="1995-12-11">
<collectingDate id="310C3A2F9F66FF837218FE0DFC9FFE63" box="[544,799,434,461]" pageId="5" pageNumber="552" value="1995-12-11" valueMax="1995-12-13" valueMin="1995-12-11">1113 December 1995</collectingDate>
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,
<collectorName id="F80380D19F66FF837311FE0DFC29FE62" box="[809,937,434,460]" pageId="5" pageNumber="552">P. Goloboff</collectorName>
<emphasis id="678239159F66FF837397FE0DFC51FE62" box="[943,977,434,460]" italics="true" pageId="5" pageNumber="552">leg</emphasis>
.—
</materialsCitation>
<materialsCitation id="E59EEF5A9F66FF8373C1FE0DFB27FE5E" collectingDate="1995-12-10" collectingDateMax="1995-12-13" collectingDateMin="1995-12-10" collectionCode="CBF" collectorName="P. Goloboff" country="Bolivia" county="Subida de La EscaleRa" elevation="2700" latitude="-21.991533" location="AbRa de La Hondura" longLatPrecision="1" longitude="-64.6216" municipality="TaRiquia Flora" pageId="5" pageNumber="552" specimenCode="MACN-AR 9597, MACN-AR 9598" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Tarija" typeStatus="paratype">
<specimenCount id="43F02E8E9F66FF8373C1FE0DFBA0FE62" box="[1017,1056,434,460]" count="1" pageId="5" pageNumber="552" type="male">1 ♂</specimenCount>
,
<specimenCount id="43F02E8E9F66FF837413FE0DFBD4FE62" box="[1067,1108,434,460]" count="1" pageId="5" pageNumber="552" type="female">1 ♀</specimenCount>
<typeStatus id="8A4D5BA59F66FF837462FE0DFB45FE62" box="[1114,1221,434,460]" pageId="5" pageNumber="552" type="paratype">paRatYpes</typeStatus>
(
<collectionCode id="33E77DC29F66FF8374ECFE0DFA8FFE62" box="[1236,1295,434,460]" country="Bolivia" httpUri="http://grbio.org/cool/ky5m-b42n" name="Coleccion Boliviana de Fauna" pageId="5" pageNumber="552">CBF</collectionCode>
):
<location id="5029B3DC9F66FF837518FE0DFF79FE5E" LSID="urn:lsid:plazi:treatment:DD5F54119F66FF8E70AFFF28FB8FF871:5029B3DC9F66FF837518FE0DFF79FE5E" country="Bolivia" county="Subida de La EscaleRa" latitude="-21.991533" longLatPrecision="1" longitude="-64.6216" municipality="TaRiquia Flora" name="AbRa de La Hondura" pageId="5" pageNumber="552" stateProvince="Tarija">AbRa de La Hondura</location>
[
<emphasis id="678239159F66FF837131FE68FEABFE5E" box="[265,299,471,496]" italics="true" pageId="5" pageNumber="552">ca.</emphasis>
<geoCoordinate id="30C283C09F66FF83710AFE69FE3FFE5F" box="[306,447,470,497]" degrees="21" direction="south" minutes="59.492" orientation="latitude" pageId="5" pageNumber="552" precision="1" value="-21.991533">21°59.492S</geoCoordinate>
<geoCoordinate id="30C283C09F66FF8371FEFE69FDE3FE5E" box="[454,611,470,496]" degrees="64" direction="west" minutes="37.296" orientation="longitude" pageId="5" pageNumber="552" precision="1" value="-64.6216">64°37.296W</geoCoordinate>
],
<elevation id="DEDB02349F66FF83724DFE69FC87FE5E" box="[629,775,470,496]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="552" unit="m" value="2700.0">
<quantity id="920E48E29F66FF83724DFE69FD4AFE5E" box="[629,714,470,496]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="552" unit="m" value="2700.0">2700 m</quantity>
a.s.l.
</elevation>
,
<date id="2148C3C79F66FF83732BFE69FC6BFE5F" box="[787,1003,470,497]" pageId="5" pageNumber="552" value="1995-12-10">
<collectingDate id="310C3A2F9F66FF83732BFE69FC6BFE5F" box="[787,1003,470,497]" pageId="5" pageNumber="552" value="1995-12-10">10 December 1995</collectingDate>
</date>
,
<collectorName id="F80380D19F66FF8373CEFE69FBF7FE5E" box="[1014,1143,470,496]" pageId="5" pageNumber="552">P. Goloboff</collectorName>
<emphasis id="678239159F66FF837446FE69FB20FE5E" box="[1150,1184,470,496]" italics="true" pageId="5" pageNumber="552">leg</emphasis>
.
</materialsCitation>
</paragraph>
<paragraph id="5549E5079F66FF8370FFFE45FD1AFD96" blockId="5.[151,1437,361,785]" pageId="5" pageNumber="552">
<materialsCitation id="E59EEF5A9F66FF8370FFFE45FD16FD96" country="Bolivia" county="Fauna National Reserve" elevation="2150" latitude="-22.01615" location="Subida de La Escalera" longLatPrecision="1" longitude="-64.57816" municipality="Tariquia Flora" pageId="5" pageNumber="552" specimenCount="1" stateProvince="Tarija Department" typeStatus="holotype">
<emphasis id="678239159F66FF8370FFFE45FEE5FDBD" bold="true" box="[199,357,505,532]" pageId="5" pageNumber="552">Type locality.</emphasis>
<location id="5029B3DC9F66FF837153FE45FDE4FDBA" LSID="urn:lsid:plazi:treatment:DD5F54119F66FF8E70AFFF28FB8FF871:5029B3DC9F66FF837153FE45FDE4FDBA" box="[363,612,506,532]" country="Bolivia" county="Fauna National Reserve" latitude="-22.01615" longLatPrecision="1" longitude="-64.57816" municipality="Tariquia Flora" name="Subida de La Escalera" pageId="5" pageNumber="552" stateProvince="Tarija Department">Subida de La Escalera</location>
(
<emphasis id="678239159F66FF83724BFE44FD15FDBA" box="[627,661,507,532]" italics="true" pageId="5" pageNumber="552">ca.</emphasis>
<geoCoordinate id="30C283C09F66FF8372A3FE45FC9AFDBA" box="[667,794,506,532]" degrees="22" direction="south" minutes="0.969" orientation="latitude" pageId="5" pageNumber="552" precision="1" value="-22.01615">22°0.969S</geoCoordinate>
<geoCoordinate id="30C283C09F66FF837319FE45FC39FDBA" box="[801,953,506,532]" degrees="64" direction="west" minutes="34.690" orientation="longitude" pageId="5" pageNumber="552" precision="1" value="-64.57816">64°34.690W</geoCoordinate>
, ~
<quantity id="920E48E29F66FF8373E9FE45FBF1FDBA" box="[977,1137,506,532]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="5" pageNumber="552" unit="m" value="2150.0" valueMax="2300.0" valueMin="2000.0">
<elevation id="DEDB02349F66FF8373E9FE45FBF1FDBA" box="[977,1137,506,532]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="5" pageNumber="552" unit="m" value="2150.0" valueMax="2300.0" valueMin="2000.0">20002300 m</elevation>
</quantity>
),
<collectingMunicipality id="B52D7F7D9F66FF8374BCFE45FAA2FDBA" box="[1156,1314,506,532]" pageId="5" pageNumber="552">Tariquía Flora</collectingMunicipality>
and
<collectingCounty id="BC289D8B9F66FF837560FE45FEDCFD96" pageId="5" pageNumber="552">Fauna National Reserve</collectingCounty>
,
<collectingRegion id="97322BE59F66FF83715EFDA1FDB5FD96" box="[358,565,542,568]" country="Bolivia" name="Tarija" pageId="5" pageNumber="552">Tarija Department</collectingRegion>
,
<collectingCountry id="2DE1A5979F66FF837278FDA1FD16FD96" box="[576,662,542,568]" name="Bolivia" pageId="5" pageNumber="552">Bolivia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1DECB68C9F66FF8370FFFDFDFDEBFD2E" pageId="5" pageNumber="552" type="etymology">
<paragraph id="5549E5079F66FF8370FFFDFDFDEBFD2E" blockId="5.[151,1437,361,785]" pageId="5" pageNumber="552">
<emphasis id="678239159F66FF8370FFFDFDFECEFDF2" bold="true" box="[199,334,578,604]" pageId="5" pageNumber="552">Etymology.</emphasis>
The name
<taxonomicName id="92F69E849F66FF8371E8FDFCFDB7FDF2" authorityName="Acosta" authorityYear="2024" box="[464,567,579,604]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="552" phylum="Arthropoda" rank="species" species="tariquiae">
<emphasis id="678239159F66FF8371E8FDFCFDB7FDF2" box="[464,567,579,604]" italics="true" pageId="5" pageNumber="552">tariquiae</emphasis>
</taxonomicName>
is formed as the feminine genitive of the Bolivian protected area where the
<typeStatus id="8A4D5BA59F66FF8370AFFDD9FF47FD2E" box="[151,199,614,640]" pageId="5" pageNumber="552">type</typeStatus>
locality is placed (Tariquía Reserve).
</paragraph>
</subSubSection>
<subSubSection id="1DECB68C9F66FF8170FFFD36FD29F913" lastPageId="7" lastPageNumber="554" pageId="5" pageNumber="552" type="description">
<paragraph id="5549E5079F66FF8370FFFD36FCF4FCBE" blockId="5.[151,1437,361,785]" pageId="5" pageNumber="552">
<emphasis id="678239159F66FF8370FFFD36FED9FD0D" bold="true" box="[199,345,649,675]" pageId="5" pageNumber="552">Description.</emphasis>
<emphasis id="678239159F66FF837167FD34FD24FD0A" box="[351,676,650,676]" italics="true" pageId="5" pageNumber="552">Measurements and meristics.</emphasis>
DS length: males
<quantity id="920E48E29F66FF837357FD35FC75FD0A" box="[879,1013,650,677]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="7.2" metricValueMin="5.9" pageId="5" pageNumber="552" unit="mm" value="6.55" valueMax="7.2" valueMin="5.9">5.97.2 mm</quantity>
(mean
<quantity id="920E48E29F66FF83747FFD35FB1EFD0A" box="[1095,1182,650,676]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="5" pageNumber="552" unit="mm" value="6.4">6.4 mm</quantity>
, n=8), females
<quantity id="920E48E29F66FF837570FD35FF43FD66" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="7.4" metricValueMin="6.3" pageId="5" pageNumber="552" unit="mm" value="6.85" valueMax="7.4" valueMin="6.3">6.37.4 mm</quantity>
(mean
<quantity id="920E48E29F66FF83712FFD11FEEFFD66" box="[279,367,686,712]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="5" pageNumber="552" unit="mm" value="6.8">6.8 mm</quantity>
, n=5). FeIV/DS ratio (males): 0.861.15 (mean 1.02, n=8); in 5/
<specimenCount id="43F02E8E9F66FF837479FD11FB16FD66" box="[1089,1174,686,713]" count="8" pageId="5" pageNumber="552" type="male">8 males</specimenCount>
this ratio is above 1, in 7/8 above 0.9 (
<figureCitation id="CDCDF9829F66FF837105FD6DFE02FD42" box="[317,386,722,748]" captionStart="FIGURE 2" captionStartId="2.[151,250,1241,1265]" captionTargetBox="[155,1431,509,1214]" captionTargetId="figure-169@2.[151,1435,505,1218]" captionTargetPageId="2" captionText="FIGURE 2. Comparative morphometrics of Acrographinotus tariquiae sp. nov. and A. calilegua sp. nov., displayed as box-plots with data points (internal lines, dashed: mean, solid: median). Left: DS length (in mm) for males and females; the single outlying female from El Duraznillo (star), tentatively assigned to A. calilegua but not used in calculations, is shown for reference. Right: FeIV/DS ratio, males. In box-plots of male A. tariquiae, small aRROws fROm the left identifY the twO examined β-males." figureDoi="http://doi.org/10.5281/zenodo.12783062" httpUri="https://zenodo.org/record/12783062/files/figure.png" pageId="5" pageNumber="552">Fig. 2</figureCitation>
). Detailed measurements of the
<typeStatus id="8A4D5BA59F66FF8372D1FD6DFCCBFD42" box="[745,843,722,748]" pageId="5" pageNumber="552" type="holotype">holotype</typeStatus>
male and a female
<typeStatus id="8A4D5BA59F66FF83741BFD6DFB08FD42" box="[1059,1160,722,748]" pageId="5" pageNumber="552" type="paratype">paratype</typeStatus>
:
<tableCitation id="1874D0BC9F66FF8374AAFD6DFB65FD42" box="[1170,1253,722,748]" captionStart="TABLE 1" captionStartId="5.[151,239,831,855]" captionTargetId="graphics-230@5.[151,1436,937,1819]" captionTargetPageId="5" captionText="TABLE 1. Measurements (mm) of male holotype and a female paratype of Acrographinotus tariquiae sp. nov. and Acrographinotus calilegua sp. nov.." pageId="5" pageNumber="552">Table 1</tableCitation>
. Tarsal formula: males 56:810:7:7 (
<typeStatus id="8A4D5BA59F66FF8371BBFD49FE67FCBE" box="[387,487,758,784]" pageId="5" pageNumber="552" type="holotype">holotype</typeStatus>
with 5/6:9:7:7), females 6:79:7:7.
</paragraph>
<caption id="0189B58F9F66FF8370AFFC80FD98FCD5" pageId="5" pageNumber="552" startId="5.[151,239,831,855]" targetBox="[163,1414,907,1813]" targetIsTable="true" targetPageId="5" targetType="table">
<paragraph id="5549E5079F66FF8370AFFC80FD98FCD5" blockId="5.[151,1437,831,892]" pageId="5" pageNumber="552">
<emphasis id="678239159F66FF8370AFFC80FE8EFCF9" bold="true" box="[151,270,831,855]" pageId="5" pageNumber="552">TABLE 1.</emphasis>
Measurements (mm) of male holotype and a female paratype of
<taxonomicName id="92F69E849F66FF8373D1FCFFFA89FCF9" authorityName="Acosta" authorityYear="2024" box="[1001,1289,832,856]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="552" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F66FF8373D1FCFFFA89FCF9" box="[1001,1289,832,856]" italics="true" pageId="5" pageNumber="552">Acrographinotus tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F66FF83752BFCFFFAECFCF6" bold="true" box="[1299,1388,832,856]" pageId="5" pageNumber="552">
<taxonomicNameLabel id="7CB1846E9F66FF83752BFCFFFAECFCF6" box="[1299,1388,832,856]" pageId="5" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="92F69E849F66FF8370AFFCDBFD98FCD5" authorityName="Acosta" authorityYear="2024" box="[151,536,868,892]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="552" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F66FF8370AFFCDBFE35FCD2" box="[151,437,868,892]" italics="true" pageId="5" pageNumber="552">Acrographinotus calilegua</emphasis>
<emphasis id="678239159F66FF837183FCDBFD91FCD2" bold="true" box="[443,529,868,892]" pageId="5" pageNumber="552">sp. nov.</emphasis>
.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="5549E5079F66FF8372D2FC33FAE6F8BB" pageId="5" pageNumber="552">
<table id="27F617A79F660079709BFC34FA06F8BB" box="[163,1414,907,1813]" gridcols="5" gridrows="22" pageId="5" pageNumber="552">
<tr id="EBC6E7459F660079709BFC34FA06FC0A" box="[163,1414,907,932]" gridrow="0" pageId="5" pageNumber="552" rowspan-0="1">
<th id="A8178E399F660079729FFC34FB88FC0A" box="[679,1032,907,932]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="5" pageNumber="552">
<taxonomicName id="92F69E849F66FF8372D2FC33FC3CFC0A" authorityName="Acosta" authorityYear="2024" box="[746,956,908,932]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="552" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F66FF8372D2FC33FC3CFC0A" bold="true" box="[746,956,908,932]" pageId="5" pageNumber="552">
<emphasis id="678239159F66FF8372D2FC33FCE5FC0A" bold="true" box="[746,869,908,932]" italics="true" pageId="5" pageNumber="552">A. tariquiae</emphasis>
sp. nov.
</emphasis>
</taxonomicName>
</th>
<th id="A8178E399F660079740EFC34FA06FC0A" box="[1078,1414,907,932]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="5" pageNumber="552">
<emphasis id="678239159F66FF83744AFC33FAC6FC0A" bold="true" box="[1138,1350,907,932]" pageId="5" pageNumber="552">
<taxonomicName id="92F69E849F66FF83744AFC33FB70FC0D" authorityName="Acosta" authorityYear="2024" box="[1138,1264,907,932]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="552" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F66FF83744AFC33FB70FC0D" bold="true" box="[1138,1264,907,932]" italics="true" pageId="5" pageNumber="552">A. calilegua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CB1846E9F66FF8374CEFC33FAC6FC0A" box="[1270,1350,908,932]" pageId="5" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="EBC6E7459F660079709BFC0BFA06FC5E" box="[163,1414,948,1008]" gridrow="1" pageId="5" pageNumber="552" rowspan-0="1">
<td id="A8178E399F660079729FFC0BFCADFC5E" box="[679,813,948,1008]" gridcol="1" gridrow="1" pageId="5" pageNumber="552">HOLOTYPE ♂ CBF</td>
<td id="A8178E399F6600797367FC0BFB88FC5E" box="[863,1032,948,1008]" gridcol="2" gridrow="1" pageId="5" pageNumber="552">PARATYPE ♀ MACN-Ar 9598</td>
<td id="A8178E399F660079740EFC0BFB43FC5E" box="[1078,1219,948,1008]" gridcol="3" gridrow="1" pageId="5" pageNumber="552">HOLOTYPE ♂ CDA 000.912</td>
<td id="A8178E399F66007974CBFC0BFA06FC5E" box="[1267,1414,948,1008]" gridcol="4" gridrow="1" pageId="5" pageNumber="552">PARATYPE ♀ CDA 000. 913</td>
</tr>
<tr id="EBC6E7459F660079709BFBBFFA06FBB6" box="[163,1414,1024,1048]" gridrow="2" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFBBFFDFEFBB6" box="[163,638,1024,1048]" gridcol="0" gridrow="2" pageId="5" pageNumber="552">Total body length, apophysis included</th>
<td id="A8178E399F660079729FFBBFFCADFBB6" box="[679,813,1024,1048]" gridcol="1" gridrow="2" pageId="5" pageNumber="552">10.8</td>
<td id="A8178E399F6600797367FBBFFB88FBB6" box="[863,1032,1024,1048]" gridcol="2" gridrow="2" pageId="5" pageNumber="552">9.8</td>
<td id="A8178E399F660079740EFBBFFB43FBB6" box="[1078,1219,1024,1048]" gridcol="3" gridrow="2" pageId="5" pageNumber="552">8.2</td>
<td id="A8178E399F66007974CBFBBFFA06FBB6" box="[1267,1414,1024,1048]" gridcol="4" gridrow="2" pageId="5" pageNumber="552">9.1</td>
</tr>
<tr id="EBC6E7459F660079709BFB97FA06FBEF" box="[163,1414,1064,1089]" gridrow="3" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFB97FDFEFBEF" box="[163,638,1064,1089]" gridcol="0" gridrow="3" pageId="5" pageNumber="552">DS length / maximal width</th>
<td id="A8178E399F660079729FFB97FCADFBEF" box="[679,813,1064,1089]" gridcol="1" gridrow="3" pageId="5" pageNumber="552">7.0 / 7.7</td>
<td id="A8178E399F6600797367FB97FB88FBEF" box="[863,1032,1064,1089]" gridcol="2" gridrow="3" pageId="5" pageNumber="552">7.0 / 5.9</td>
<td id="A8178E399F660079740EFB97FB43FBEF" box="[1078,1219,1064,1089]" gridcol="3" gridrow="3" pageId="5" pageNumber="552">5.7 / 6.3</td>
<td id="A8178E399F66007974CBFB97FA06FBEF" box="[1267,1414,1064,1089]" gridcol="4" gridrow="3" pageId="5" pageNumber="552">6.7 / 6.5</td>
</tr>
<tr id="EBC6E7459F660079709BFBEEFA06FBC7" box="[163,1414,1105,1129]" gridrow="4" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFBEEFDFEFBC7" box="[163,638,1105,1129]" gridcol="0" gridrow="4" pageId="5" pageNumber="552">Prosoma length / width</th>
<td id="A8178E399F660079729FFBEEFCADFBC7" box="[679,813,1105,1129]" gridcol="1" gridrow="4" pageId="5" pageNumber="552">2.7 / 3.5</td>
<td id="A8178E399F6600797367FBEEFB88FBC7" box="[863,1032,1105,1129]" gridcol="2" gridrow="4" pageId="5" pageNumber="552">2.4 / 3.1</td>
<td id="A8178E399F660079740EFBEEFB43FBC7" box="[1078,1219,1105,1129]" gridcol="3" gridrow="4" pageId="5" pageNumber="552">2.5 / 3.0</td>
<td id="A8178E399F66007974CBFBEEFA06FBC7" box="[1267,1414,1105,1129]" gridcol="4" gridrow="4" pageId="5" pageNumber="552">2.5 / 3.2</td>
</tr>
<tr id="EBC6E7459F660079709BFBC6FA06FB3F" box="[163,1414,1145,1169]" gridrow="5" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFBC6FDFEFB3F" box="[163,638,1145,1169]" gridcol="0" gridrow="5" pageId="5" pageNumber="552">Leg I, total length</th>
<td id="A8178E399F660079729FFBC6FCADFB3F" box="[679,813,1145,1169]" gridcol="1" gridrow="5" pageId="5" pageNumber="552">14.5</td>
<td id="A8178E399F6600797367FBC6FB88FB3F" box="[863,1032,1145,1169]" gridcol="2" gridrow="5" pageId="5" pageNumber="552">12.1</td>
<td id="A8178E399F660079740EFBC6FB43FB3F" box="[1078,1219,1145,1169]" gridcol="3" gridrow="5" pageId="5" pageNumber="552">12.3</td>
<td id="A8178E399F66007974CBFBC6FA06FB3F" box="[1267,1414,1145,1169]" gridcol="4" gridrow="5" pageId="5" pageNumber="552">12.8</td>
</tr>
<tr id="EBC6E7459F660079709BFB1EFA06FB17" box="[163,1414,1185,1209]" gridrow="6" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFB1EFDFEFB17" box="[163,638,1185,1209]" gridcol="0" gridrow="6" pageId="5" pageNumber="552">trochanter / femur / patella I length</th>
<td id="A8178E399F660079729FFB1EFCADFB17" box="[679,813,1185,1209]" gridcol="1" gridrow="6" pageId="5" pageNumber="552">0.8 / 3.7 / 1.5</td>
<td id="A8178E399F6600797367FB1EFB88FB17" box="[863,1032,1185,1209]" gridcol="2" gridrow="6" pageId="5" pageNumber="552">0.7 / 3.1 / 1.2</td>
<td id="A8178E399F660079740EFB1EFB43FB17" box="[1078,1219,1185,1209]" gridcol="3" gridrow="6" pageId="5" pageNumber="552">0.8 / 3.1 / 1.2</td>
<td id="A8178E399F66007974CBFB1EFA06FB17" box="[1267,1414,1185,1209]" gridcol="4" gridrow="6" pageId="5" pageNumber="552">0.8 / 3.2 / 1.3</td>
</tr>
<tr id="EBC6E7459F660079709BFB76FA06FB4C" box="[163,1414,1225,1250]" gridrow="7" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFB76FDFEFB4C" box="[163,638,1225,1250]" gridcol="0" gridrow="7" pageId="5" pageNumber="552">tibia / metatarsus / tarsus I length</th>
<td id="A8178E399F660079729FFB76FCADFB4C" box="[679,813,1225,1250]" gridcol="1" gridrow="7" pageId="5" pageNumber="552">2.7 / 3.6 / 2.2</td>
<td id="A8178E399F6600797367FB76FB88FB4C" box="[863,1032,1225,1250]" gridcol="2" gridrow="7" pageId="5" pageNumber="552">2.2 / 2.9 / 2.0</td>
<td id="A8178E399F660079740EFB76FB43FB4C" box="[1078,1219,1225,1250]" gridcol="3" gridrow="7" pageId="5" pageNumber="552">2.2 / 3.0 / 2.0</td>
<td id="A8178E399F66007974CBFB76FA06FB4C" box="[1267,1414,1225,1250]" gridcol="4" gridrow="7" pageId="5" pageNumber="552">2.3 / 3.1 / 2.1</td>
</tr>
<tr id="EBC6E7459F660079709BFB4DFA06FAA4" box="[163,1414,1266,1290]" gridrow="8" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFB4DFDFEFAA4" box="[163,638,1266,1290]" gridcol="0" gridrow="8" pageId="5" pageNumber="552">Leg II, total length</th>
<td id="A8178E399F660079729FFB4DFCADFAA4" box="[679,813,1266,1290]" gridcol="1" gridrow="8" pageId="5" pageNumber="552">22.3</td>
<td id="A8178E399F6600797367FB4DFB88FAA4" box="[863,1032,1266,1290]" gridcol="2" gridrow="8" pageId="5" pageNumber="552">18.8</td>
<td id="A8178E399F660079740EFB4DFB43FAA4" box="[1078,1219,1266,1290]" gridcol="3" gridrow="8" pageId="5" pageNumber="552">18.3</td>
<td id="A8178E399F66007974CBFB4DFA06FAA4" box="[1267,1414,1266,1290]" gridcol="4" gridrow="8" pageId="5" pageNumber="552">19.4</td>
</tr>
<tr id="EBC6E7459F660079709BFAA5FA06FA9C" box="[163,1414,1306,1330]" gridrow="9" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFAA5FDFEFA9C" box="[163,638,1306,1330]" gridcol="0" gridrow="9" pageId="5" pageNumber="552">trochanter / femur / patella II length</th>
<td id="A8178E399F660079729FFAA5FCADFA9C" box="[679,813,1306,1330]" gridcol="1" gridrow="9" pageId="5" pageNumber="552">1.0 / 5.6 / 1.8</td>
<td id="A8178E399F6600797367FAA5FB88FA9C" box="[863,1032,1306,1330]" gridcol="2" gridrow="9" pageId="5" pageNumber="552">0.9 / 4.6 / 1.6</td>
<td id="A8178E399F660079740EFAA5FB43FA9C" box="[1078,1219,1306,1330]" gridcol="3" gridrow="9" pageId="5" pageNumber="552">0.8 / 4.6 / 1.6</td>
<td id="A8178E399F66007974CBFAA5FA06FA9C" box="[1267,1414,1306,1330]" gridcol="4" gridrow="9" pageId="5" pageNumber="552">0.9 / 4.8 / 1.7</td>
</tr>
<tr id="EBC6E7459F660079709BFAFDFA06FAF4" box="[163,1414,1346,1370]" gridrow="10" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFAFDFDFEFAF4" box="[163,638,1346,1370]" gridcol="0" gridrow="10" pageId="5" pageNumber="552">tibia / metatarsus / tarsus II length</th>
<td id="A8178E399F660079729FFAFDFCADFAF4" box="[679,813,1346,1370]" gridcol="1" gridrow="10" pageId="5" pageNumber="552">4.2 / 5.3 / 4.4</td>
<td id="A8178E399F6600797367FAFDFB88FAF4" box="[863,1032,1346,1370]" gridcol="2" gridrow="10" pageId="5" pageNumber="552">3.5 / 4.1 / 4.1</td>
<td id="A8178E399F660079740EFAFDFB43FAF4" box="[1078,1219,1346,1370]" gridcol="3" gridrow="10" pageId="5" pageNumber="552">3.6 / 4.2 / 3.5</td>
<td id="A8178E399F66007974CBFAFDFA06FAF4" box="[1267,1414,1346,1370]" gridcol="4" gridrow="10" pageId="5" pageNumber="552">3.8 / 4.5 / 3.7</td>
</tr>
<tr id="EBC6E7459F660079709BFAD5FA06FA2D" box="[163,1414,1386,1411]" gridrow="11" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFAD5FDFEFA2D" box="[163,638,1386,1411]" gridcol="0" gridrow="11" pageId="5" pageNumber="552">Leg III, total length</th>
<td id="A8178E399F660079729FFAD5FCADFA2D" box="[679,813,1386,1411]" gridcol="1" gridrow="11" pageId="5" pageNumber="552">18.9</td>
<td id="A8178E399F6600797367FAD5FB88FA2D" box="[863,1032,1386,1411]" gridcol="2" gridrow="11" pageId="5" pageNumber="552">15.7</td>
<td id="A8178E399F660079740EFAD5FB43FA2D" box="[1078,1219,1386,1411]" gridcol="3" gridrow="11" pageId="5" pageNumber="552">15.8</td>
<td id="A8178E399F66007974CBFAD5FA06FA2D" box="[1267,1414,1386,1411]" gridcol="4" gridrow="11" pageId="5" pageNumber="552">16.7</td>
</tr>
<tr id="EBC6E7459F660079709BFA2CFA06FA05" box="[163,1414,1427,1451]" gridrow="12" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFA2CFDFEFA05" box="[163,638,1427,1451]" gridcol="0" gridrow="12" pageId="5" pageNumber="552">trochanter / femur / patella III length</th>
<td id="A8178E399F660079729FFA2CFCADFA05" box="[679,813,1427,1451]" gridcol="1" gridrow="12" pageId="5" pageNumber="552">1.1 / 5.2 / 1.7</td>
<td id="A8178E399F6600797367FA2CFB88FA05" box="[863,1032,1427,1451]" gridcol="2" gridrow="12" pageId="5" pageNumber="552">0.9 / 4.2 / 1.5</td>
<td id="A8178E399F660079740EFA2CFB43FA05" box="[1078,1219,1427,1451]" gridcol="3" gridrow="12" pageId="5" pageNumber="552">0.9 / 4.2 / 1.5</td>
<td id="A8178E399F66007974CBFA2CFA06FA05" box="[1267,1414,1427,1451]" gridcol="4" gridrow="12" pageId="5" pageNumber="552">1.0 / 4.3 / 1.6</td>
</tr>
<tr id="EBC6E7459F660079709BFA04FA06FA7D" box="[163,1414,1467,1491]" gridrow="13" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFA04FDFEFA7D" box="[163,638,1467,1491]" gridcol="0" gridrow="13" pageId="5" pageNumber="552">tibia / metatarsus / tarsus III length</th>
<td id="A8178E399F660079729FFA04FCADFA7D" box="[679,813,1467,1491]" gridcol="1" gridrow="13" pageId="5" pageNumber="552">3.5 / 4.9 / 2.5</td>
<td id="A8178E399F6600797367FA04FB88FA7D" box="[863,1032,1467,1491]" gridcol="2" gridrow="13" pageId="5" pageNumber="552">3.0 / 3.9 / 2.2</td>
<td id="A8178E399F660079740EFA04FB43FA7D" box="[1078,1219,1467,1491]" gridcol="3" gridrow="13" pageId="5" pageNumber="552">2.9 / 4.1 / 2.2</td>
<td id="A8178E399F66007974CBFA04FA06FA7D" box="[1267,1414,1467,1491]" gridcol="4" gridrow="13" pageId="5" pageNumber="552">3.2 / 4.2 / 2.4</td>
</tr>
<tr id="EBC6E7459F660079709BFA5CFA06FA52" box="[163,1414,1507,1532]" gridrow="14" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BFA5CFDFEFA52" box="[163,638,1507,1532]" gridcol="0" gridrow="14" pageId="5" pageNumber="552">Leg IV, total length</th>
<td id="A8178E399F660079729FFA5CFCADFA52" box="[679,813,1507,1532]" gridcol="1" gridrow="14" pageId="5" pageNumber="552">29.2</td>
<td id="A8178E399F6600797367FA5CFB88FA52" box="[863,1032,1507,1532]" gridcol="2" gridrow="14" pageId="5" pageNumber="552">20.9</td>
<td id="A8178E399F660079740EFA5CFB43FA52" box="[1078,1219,1507,1532]" gridcol="3" gridrow="14" pageId="5" pageNumber="552">21.8</td>
<td id="A8178E399F66007974CBFA5CFA06FA52" box="[1267,1414,1507,1532]" gridcol="4" gridrow="14" pageId="5" pageNumber="552">21.7</td>
</tr>
<tr id="EBC6E7459F660079709BF9B4FA06F98A" box="[163,1414,1547,1572]" gridrow="15" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BF9B4FDFEF98A" box="[163,638,1547,1572]" gridcol="0" gridrow="15" pageId="5" pageNumber="552">trochanter / femur / patella IV length</th>
<td id="A8178E399F660079729FF9B4FCADF98A" box="[679,813,1547,1572]" gridcol="1" gridrow="15" pageId="5" pageNumber="552">2.7 / 7.7 / 2.6</td>
<td id="A8178E399F6600797367F9B4FB88F98A" box="[863,1032,1547,1572]" gridcol="2" gridrow="15" pageId="5" pageNumber="552">1.3 / 5.2 / 1.9</td>
<td id="A8178E399F660079740EF9B4FB43F98A" box="[1078,1219,1547,1572]" gridcol="3" gridrow="15" pageId="5" pageNumber="552">1.8 / 5.1 / 1.9</td>
<td id="A8178E399F66007974CBF9B4FA06F98A" box="[1267,1414,1547,1572]" gridcol="4" gridrow="15" pageId="5" pageNumber="552">1.4 / 5.2 / 2.0</td>
</tr>
<tr id="EBC6E7459F660079709BF98CFA06F9E2" box="[163,1414,1587,1612]" gridrow="16" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BF98CFDFEF9E2" box="[163,638,1587,1612]" gridcol="0" gridrow="16" pageId="5" pageNumber="552">tibia / metatarsus / tarsus IV length</th>
<td id="A8178E399F660079729FF98CFCADF9E2" box="[679,813,1587,1612]" gridcol="1" gridrow="16" pageId="5" pageNumber="552">5.6 / 7.9 / 2.7</td>
<td id="A8178E399F6600797367F98CFB88F9E2" box="[863,1032,1587,1612]" gridcol="2" gridrow="16" pageId="5" pageNumber="552">4.3 / 5.8 / 2.4</td>
<td id="A8178E399F660079740EF98CFB43F9E2" box="[1078,1219,1587,1612]" gridcol="3" gridrow="16" pageId="5" pageNumber="552">4.6 / 6.0 / 2.4</td>
<td id="A8178E399F66007974CBF98CFA06F9E2" box="[1267,1414,1587,1612]" gridcol="4" gridrow="16" pageId="5" pageNumber="552">4.4 / 6.2 / 2.5</td>
</tr>
<tr id="EBC6E7459F660079709BF9E3FA06F9DA" box="[163,1414,1628,1652]" gridrow="17" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BF9E3FDFEF9DA" box="[163,638,1628,1652]" gridcol="0" gridrow="17" pageId="5" pageNumber="552">Pedipalp, total length</th>
<td id="A8178E399F660079729FF9E3FCADF9DA" box="[679,813,1628,1652]" gridcol="1" gridrow="17" pageId="5" pageNumber="552">8.8</td>
<td id="A8178E399F6600797367F9E3FB88F9DA" box="[863,1032,1628,1652]" gridcol="2" gridrow="17" pageId="5" pageNumber="552">8.0</td>
<td id="A8178E399F660079740EF9E3FB43F9DA" box="[1078,1219,1628,1652]" gridcol="3" gridrow="17" pageId="5" pageNumber="552">7.6</td>
<td id="A8178E399F66007974CBF9E3FA06F9DA" box="[1267,1414,1628,1652]" gridcol="4" gridrow="17" pageId="5" pageNumber="552">8.4</td>
</tr>
<tr id="EBC6E7459F660079709BF93BFA06F932" box="[163,1414,1668,1692]" gridrow="18" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BF93BFDFEF932" box="[163,638,1668,1692]" gridcol="0" gridrow="18" pageId="5" pageNumber="552">Pp trochanter / femur / patella length</th>
<td id="A8178E399F660079729FF93BFCADF932" box="[679,813,1668,1692]" gridcol="1" gridrow="18" pageId="5" pageNumber="552">0.9 / 2.3 / 1.2</td>
<td id="A8178E399F6600797367F93BFB88F932" box="[863,1032,1668,1692]" gridcol="2" gridrow="18" pageId="5" pageNumber="552">0.8 / 2.0 / 1.0</td>
<td id="A8178E399F660079740EF93BFB43F932" box="[1078,1219,1668,1692]" gridcol="3" gridrow="18" pageId="5" pageNumber="552">0.7 / 2.0 / 1.0</td>
<td id="A8178E399F66007974CBF93BFA06F932" box="[1267,1414,1668,1692]" gridcol="4" gridrow="18" pageId="5" pageNumber="552">0.8 / 2.2 / 1.2</td>
</tr>
<tr id="EBC6E7459F660079709BF913FA06F96B" box="[163,1414,1708,1733]" gridrow="19" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BF913FDFEF96B" box="[163,638,1708,1733]" gridcol="0" gridrow="19" pageId="5" pageNumber="552">Pp tibia / tarsus / claw length</th>
<td id="A8178E399F660079729FF913FCADF96B" box="[679,813,1708,1733]" gridcol="1" gridrow="19" pageId="5" pageNumber="552">1.6 / 1.4 / 1.4</td>
<td id="A8178E399F6600797367F913FB88F96B" box="[863,1032,1708,1733]" gridcol="2" gridrow="19" pageId="5" pageNumber="552">1.6 / 1.3 / 1.3</td>
<td id="A8178E399F660079740EF913FB43F96B" box="[1078,1219,1708,1733]" gridcol="3" gridrow="19" pageId="5" pageNumber="552">1.5 / 1.2 / 1.2</td>
<td id="A8178E399F66007974CBF913FA06F96B" box="[1267,1414,1708,1733]" gridcol="4" gridrow="19" pageId="5" pageNumber="552">1.7 / 1.3 / 1.2</td>
</tr>
<tr id="EBC6E7459F660079709BF96BFA06F943" box="[163,1414,1748,1773]" gridrow="20" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BF96BFDFEF943" box="[163,638,1748,1773]" gridcol="0" gridrow="20" pageId="5" pageNumber="552">Chelicera, hand / bulla length</th>
<td id="A8178E399F660079729FF96BFCADF943" box="[679,813,1748,1773]" gridcol="1" gridrow="20" pageId="5" pageNumber="552">2.5 / 0.9</td>
<td id="A8178E399F6600797367F96BFB88F943" box="[863,1032,1748,1773]" gridcol="2" gridrow="20" pageId="5" pageNumber="552">2.2 / 0.9</td>
<td id="A8178E399F660079740EF96BFB43F943" box="[1078,1219,1748,1773]" gridcol="3" gridrow="20" pageId="5" pageNumber="552">2.1 / 0.7</td>
<td id="A8178E399F66007974CBF96BFA06F943" box="[1267,1414,1748,1773]" gridcol="4" gridrow="20" pageId="5" pageNumber="552">2.2 / 0.8</td>
</tr>
<tr id="EBC6E7459F660079709BF942FA06F8BB" box="[163,1414,1789,1813]" gridrow="21" pageId="5" pageNumber="552">
<th id="A8178E399F660079709BF942FDFEF8BB" box="[163,638,1789,1813]" gridcol="0" gridrow="21" pageId="5" pageNumber="552">Ocular mound, width / height</th>
<td id="A8178E399F660079729FF942FCADF8BB" box="[679,813,1789,1813]" gridcol="1" gridrow="21" pageId="5" pageNumber="552">1.0 / 0.3</td>
<td id="A8178E399F6600797367F942FB88F8BB" box="[863,1032,1789,1813]" gridcol="2" gridrow="21" pageId="5" pageNumber="552">1.1 / 0.6</td>
<td id="A8178E399F660079740EF942FB43F8BB" box="[1078,1219,1789,1813]" gridcol="3" gridrow="21" pageId="5" pageNumber="552">1.0 / 0.3</td>
<td id="A8178E399F66007974CBF942FA06F8BB" box="[1267,1414,1789,1813]" gridcol="4" gridrow="21" pageId="5" pageNumber="552">1.1 / 0.4</td>
</tr>
</table>
</paragraph>
<paragraph id="5549E5079F66FF8370FFF8D7FC27F808" blockId="5.[151,1437,1896,2031]" pageId="5" pageNumber="552">
<emphasis id="678239159F66FF8370FFF8D7FE74F82D" box="[199,500,1896,1923]" italics="true" pageId="5" pageNumber="552">Coloration in ethanol 70%.</emphasis>
General color uniform dark hazel, with chelicera, pedipalps, legs IIII and metatarsus IV yellowish; weak pigment reticles insinuate on DS and appendages.
</paragraph>
<paragraph id="5549E5079F66FF8070FFF80EFC78FEB3" blockId="5.[151,1437,1896,2031]" lastBlockId="6.[151,1437,151,285]" lastPageId="6" lastPageNumber="553" pageId="5" pageNumber="552">
<emphasis id="678239159F66FF8370FFF80EFEC6F864" bold="true" box="[199,326,1968,1995]" pageId="5" pageNumber="552">Males (α):</emphasis>
<emphasis id="678239159F66FF83716AF80EFE34F864" box="[338,436,1969,1994]" italics="true" pageId="5" pageNumber="552">Dorsum.</emphasis>
DS
<typeStatus id="8A4D5BA59F66FF8371D7F80EFD9FF865" box="[495,543,1969,1995]" pageId="5" pageNumber="552">tYpe</typeStatus>
λ (lambda, cOda shORt, caRapace with subpaRallel sides;
<figureCitation id="CDCDF9829F66FF837485F80FFA9AF865" box="[1213,1306,1968,1995]" captionStart="FIGURE 3" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[155,1431,313,1749]" captionTargetId="figure-90@6.[151,1435,309,1753]" captionTargetPageId="6" captionText="FIGURE 3. Acrographinotus tariquiae sp. nov. AF: HOlOtYpe ♂ (CBF). A. DORsal scutum, fRee teRgites (alsO shOwing dORsal anal operculum and VAP apophyses), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, VAP with apophyses (also showing apophysis of FT-III), right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. GJ: Armature variations on FT-III and VAP in tOpOtYpical ♂ paRatYpes. GH: Smallest α-male (CBF), IJ: β-male (CDA 000.914). G, I: ventRal view; H, J: lateral view. Scale lines: 2 mm (AE, GJ), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783064" httpUri="https://zenodo.org/record/12783064/files/figure.png" pageId="5" pageNumber="552">Fig. 3A</figureCitation>
), unaRmed, granulation feeble. Anterior border of carapace smooth, frontal hump negligible. Ocular mound low, with a small median apophysis (
<figureCitation id="CDCDF9829F65FF80714BFF28FE4AFF1F" box="[371,458,151,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[155,1431,313,1749]" captionTargetId="figure-90@6.[151,1435,309,1753]" captionTargetPageId="6" captionText="FIGURE 3. Acrographinotus tariquiae sp. nov. AF: HOlOtYpe ♂ (CBF). A. DORsal scutum, fRee teRgites (alsO shOwing dORsal anal operculum and VAP apophyses), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, VAP with apophyses (also showing apophysis of FT-III), right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. GJ: Armature variations on FT-III and VAP in tOpOtYpical ♂ paRatYpes. GH: Smallest α-male (CBF), IJ: β-male (CDA 000.914). G, I: ventRal view; H, J: lateral view. Scale lines: 2 mm (AE, GJ), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783064" httpUri="https://zenodo.org/record/12783064/files/figure.png" pageId="6" pageNumber="553">Fig. 3F</figureCitation>
); in some specimens the ocular mound is completely unarmed or has a median notch. DS area I divided, areas IIV entire; practically smooth except for sparse small granules insinuating a row on areas IIV and lateral areas, better defined on area V and free tergites III. FT-III with a strong median apophysis, slightly sigmoid in lateral view, and a row of small inconspicuous granules each side.
</paragraph>
<caption id="0189B58F9F65FF8070AFF94FFD3CF84F" ID-DOI="http://doi.org/10.5281/zenodo.12783064" ID-Zenodo-Dep="12783064" httpUri="https://zenodo.org/record/12783064/files/figure.png" pageId="6" pageNumber="553" startId="6.[151,250,1776,1800]" targetBox="[155,1431,313,1749]" targetPageId="6" targetType="figure">
<paragraph id="5549E5079F65FF8070AFF94FFD3CF84F" blockId="6.[151,1437,1776,2017]" pageId="6" pageNumber="553">
<emphasis id="678239159F65FF8070AFF94FFE93F8A6" bold="true" box="[151,275,1776,1800]" pageId="6" pageNumber="553">FIGURE 3.</emphasis>
<taxonomicName id="92F69E849F65FF807122F94EFDA8F8A6" authorityName="Acosta" authorityYear="2024" box="[282,552,1777,1801]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="553" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F65FF807122F94EFDA8F8A6" box="[282,552,1777,1801]" italics="true" pageId="6" pageNumber="553">Acrographinotus tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F65FF807217F94EFD00F8A7" bold="true" box="[559,640,1777,1801]" pageId="6" pageNumber="553">
<taxonomicNameLabel id="7CB1846E9F65FF807217F94EFD00F8A7" box="[559,640,1777,1801]" pageId="6" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AF: HOlOtYpe ♂ (CBF). A. DORsal scutum, fRee teRgites (alsO shOwing dORsal anal operculum and VAP apophyses), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, VAP with apophyses (also showing apophysis of FT-III), right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. GJ: Armature variations on FT-III and VAP in tOpOtYpical ♂ paRatYpes. GH: Smallest α-male (CBF), IJ: β-male (CDA 000.914). G, I: ventRal view; H, J: lateral view. Scale lines: 2 mm (AE, GJ), 1 mm (F).
</paragraph>
</caption>
<paragraph id="5549E5079F64FF8170FFFF28FB99FF57" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="554">
<emphasis id="678239159F64FF8170FFFF28FE93FF1E" box="[199,275,151,176]" italics="true" pageId="7" pageNumber="554">Venter.</emphasis>
Posterior margin of stigmatic segment gently concave; free sternites nearly smooth, with row of minute granules. VAP armed by two strong apophyses that flank a shelf-like projection in between; the anus is thereby armored by three apophyses, one dorsal (FT-III) and two ventral (
<figureCitation id="CDCDF9829F64FF8173BEFF60FB8CFF57" box="[902,1036,223,249]" captionStart="FIGURE 3" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[155,1431,313,1749]" captionTargetId="figure-90@6.[151,1435,309,1753]" captionTargetPageId="6" captionText="FIGURE 3. Acrographinotus tariquiae sp. nov. AF: HOlOtYpe ♂ (CBF). A. DORsal scutum, fRee teRgites (alsO shOwing dORsal anal operculum and VAP apophyses), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, VAP with apophyses (also showing apophysis of FT-III), right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. GJ: Armature variations on FT-III and VAP in tOpOtYpical ♂ paRatYpes. GH: Smallest α-male (CBF), IJ: β-male (CDA 000.914). G, I: ventRal view; H, J: lateral view. Scale lines: 2 mm (AE, GJ), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783064" httpUri="https://zenodo.org/record/12783064/files/figure.png" pageId="7" pageNumber="554">Figs. 3BD</figureCitation>
).
</paragraph>
<paragraph id="5549E5079F64FF8170FFFEBCFCBCFEB3" blockId="7.[151,1437,151,2013]" box="[199,828,259,285]" pageId="7" pageNumber="554">
<emphasis id="678239159F64FF8170FFFEBCFECAFEB3" box="[199,330,259,285]" italics="true" pageId="7" pageNumber="554">Chelicerae.</emphasis>
Tegument tenuously rugose, bulla unarmed.
</paragraph>
<paragraph id="5549E5079F64FF8170FFFE98FB4DFECB" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="554">
<emphasis id="678239159F64FF8170FFFE98FEBFFEEF" box="[199,319,295,321]" italics="true" pageId="7" pageNumber="554">Pedipalps.</emphasis>
Weak, dorsal tegument finely rugous; blunt ventral setigerous tubercles on trochanter and femur; no medial subapical spine on femur. Pedipalp spination: LAT: Ti i_[Ii•], Ta IiIi MED: Ti IiIi, Ta IIi.
</paragraph>
<paragraph id="5549E5079F64FF8170FFFED0FC33FE27" blockId="7.[151,1437,151,2013]" box="[199,947,367,393]" pageId="7" pageNumber="554">
<emphasis id="678239159F64FF8170FFFED0FEBEFE26" box="[199,318,367,393]" italics="true" pageId="7" pageNumber="554">Legs IIII.</emphasis>
Unarmed, all segments with tegument finely granulous.
</paragraph>
<paragraph id="5549E5079F64FF8170FFFE2CFB79FE7F" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="554">
<emphasis id="678239159F64FF8170FFFE2CFE97FE02" box="[199,279,403,428]" italics="true" pageId="7" pageNumber="554">Leg IV.</emphasis>
Coxa IV smooth, dilated and with a short, blunt prolateral apophysis, diagonal, and a small retrolateral one, better discernible from ventral. Coxa-trochanter joint slightly oblique in ventral view (
<figureCitation id="CDCDF9829F64FF8174ACFE08FB6CFE7F" box="[1172,1260,439,465]" captionStart="FIGURE 3" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[155,1431,313,1749]" captionTargetId="figure-90@6.[151,1435,309,1753]" captionTargetPageId="6" captionText="FIGURE 3. Acrographinotus tariquiae sp. nov. AF: HOlOtYpe ♂ (CBF). A. DORsal scutum, fRee teRgites (alsO shOwing dORsal anal operculum and VAP apophyses), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, VAP with apophyses (also showing apophysis of FT-III), right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. GJ: Armature variations on FT-III and VAP in tOpOtYpical ♂ paRatYpes. GH: Smallest α-male (CBF), IJ: β-male (CDA 000.914). G, I: ventRal view; H, J: lateral view. Scale lines: 2 mm (AE, GJ), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783064" httpUri="https://zenodo.org/record/12783064/files/figure.png" pageId="7" pageNumber="554">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="5549E5079F64FF8170FFFE64FCD0FD93" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="554">Trochanter IV wide, subtrapezoidal and asymmetric (retrolateral side larger than prolateral one), obliquely articulated to the sides; one blunt, tuberous prodorsal mound near the apical border, opposite to the coxal apophysis; distal margin with a pair of small acute retrodorsal apophyses.</paragraph>
<paragraph id="5549E5079F64FF8170FFFDF8FE62FCBB" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="554">
Femur IV sub-straight, diagonally diverging by around 25° (
<figureCitation id="CDCDF9829F64FF817357FDF8FC76FDCF" box="[879,1014,583,609]" captionStart="FIGURE 3" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[155,1431,313,1749]" captionTargetId="figure-90@6.[151,1435,309,1753]" captionTargetPageId="6" captionText="FIGURE 3. Acrographinotus tariquiae sp. nov. AF: HOlOtYpe ♂ (CBF). A. DORsal scutum, fRee teRgites (alsO shOwing dORsal anal operculum and VAP apophyses), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, VAP with apophyses (also showing apophysis of FT-III), right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. GJ: Armature variations on FT-III and VAP in tOpOtYpical ♂ paRatYpes. GH: Smallest α-male (CBF), IJ: β-male (CDA 000.914). G, I: ventRal view; H, J: lateral view. Scale lines: 2 mm (AE, GJ), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783064" httpUri="https://zenodo.org/record/12783064/files/figure.png" pageId="7" pageNumber="554">Figs. 3AB</figureCitation>
), with subdistal narrowing and apical end slightly widened; longitudinal rows of granules to small apophyses, the most conspicuous being the retrodorsal row, formed by truncate tubercles, which, beyond the subdistal narrowing, ends in a large conic subapical apophysis and a small apical one (
<figureCitation id="CDCDF9829F64FF817189FD0CFDBBFD63" box="[433,571,691,717]" captionStart="FIGURE 3" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[155,1431,313,1749]" captionTargetId="figure-90@6.[151,1435,309,1753]" captionTargetPageId="6" captionText="FIGURE 3. Acrographinotus tariquiae sp. nov. AF: HOlOtYpe ♂ (CBF). A. DORsal scutum, fRee teRgites (alsO shOwing dORsal anal operculum and VAP apophyses), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, VAP with apophyses (also showing apophysis of FT-III), right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. GJ: Armature variations on FT-III and VAP in tOpOtYpical ♂ paRatYpes. GH: Smallest α-male (CBF), IJ: β-male (CDA 000.914). G, I: ventRal view; H, J: lateral view. Scale lines: 2 mm (AE, GJ), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783064" httpUri="https://zenodo.org/record/12783064/files/figure.png" pageId="7" pageNumber="554">Figs. 3A, E</figureCitation>
); dorsal row of spaced 89 tubercles, those in the middle larger; pro- and retrolateral rows of sparse rounded granules; pro- and retroventral incomplete rows of small acute granules ending in small unciform apophyses.
</paragraph>
<paragraph id="5549E5079F64FF8170FFFCA0FCA8FC97" blockId="7.[151,1437,151,2013]" box="[199,808,799,825]" pageId="7" pageNumber="554">Patella IV granulous, with 23 acute ventral tubercles.</paragraph>
<paragraph id="5549E5079F64FF8170FFFCFCFEF6FC2F" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="554">Tibia IV with rows of small acute granules, they are progressively larger on the retroventral row, ending distally as small apophyses.</paragraph>
<paragraph id="5549E5079F64FF8170FFFC34FBC7FBBF" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="554">
<emphasis id="678239159F64FF8170FFFC34FE85FC0A" box="[199,261,907,932]" italics="true" pageId="7" pageNumber="554">Penis</emphasis>
(
<figureCitation id="CDCDF9829F64FF81712CFC34FE19FC0B" box="[276,409,907,933]" captionStart="FIGURE 4" captionStartId="8.[151,250,1558,1582]" captionTargetBox="[277,1309,493,1531]" captionTargetId="figure-189@8.[273,1313,489,1535]" captionTargetPageId="8" captionText="FIGURE 4. Acrographinotus tariquiae sp. nov., distal end Of penis, HOlOtYpe ♂ (CBF). A: dORsal view; B: lateRal view; C: detail of stylus and ventral process. Scale lines: 0.1 mm (A, B), 0.05 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.12783067" httpUri="https://zenodo.org/record/12783067/files/figure.png" pageId="7" pageNumber="554">Figs. 4AC</figureCitation>
). Distal end slightly higher than wide; front margin of VP almost straight, lateral sides gently concave. Basal macrosetae (A1A3 + B) arranged in a transversal row, at about the same level of the truncus-glans joint. Stylus tip straight and slightly dilated; VPS with the generic ibis head with crest shape, beak-like process curved and subtle veins perceptible through the translucid membranous expansion.
</paragraph>
<paragraph id="5549E5079F64FF8170FFFBA4FB1FFB47" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="554">
<emphasis id="678239159F64FF8170FFFBA4FEABFB9B" bold="true" box="[199,299,1051,1077]" pageId="7" pageNumber="554">β males:</emphasis>
TwO specimens (Out Of eight) shOw decided feminized featuRes and aRe cOnsideRed β males. In them, VAP armature is rudimentary: apophyses are replaced by button-like tubercles and there is no shelf (
<figureCitation id="CDCDF9829F64FF817523FB80FA10FBF7" box="[1307,1424,1087,1113]" captionStart="FIGURE 3" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[155,1431,313,1749]" captionTargetId="figure-90@6.[151,1435,309,1753]" captionTargetPageId="6" captionText="FIGURE 3. Acrographinotus tariquiae sp. nov. AF: HOlOtYpe ♂ (CBF). A. DORsal scutum, fRee teRgites (alsO shOwing dORsal anal operculum and VAP apophyses), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, VAP with apophyses (also showing apophysis of FT-III), right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. GJ: Armature variations on FT-III and VAP in tOpOtYpical ♂ paRatYpes. GH: Smallest α-male (CBF), IJ: β-male (CDA 000.914). G, I: ventRal view; H, J: lateral view. Scale lines: 2 mm (AE, GJ), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783064" httpUri="https://zenodo.org/record/12783064/files/figure.png" pageId="7" pageNumber="554">Figs. 3IJ</figureCitation>
). In addition, slenderness, granulation and curvature of FeIV look much like in females, and there is no subapical narrowing. Noteworthily, these males are not the smallest ones in the sample (both are above the mean DS length;
<figureCitation id="CDCDF9829F64FF8170AFFB14FF5AFB6B" box="[151,218,1195,1221]" captionStart="FIGURE 2" captionStartId="2.[151,250,1241,1265]" captionTargetBox="[155,1431,509,1214]" captionTargetId="figure-169@2.[151,1435,505,1218]" captionTargetPageId="2" captionText="FIGURE 2. Comparative morphometrics of Acrographinotus tariquiae sp. nov. and A. calilegua sp. nov., displayed as box-plots with data points (internal lines, dashed: mean, solid: median). Left: DS length (in mm) for males and females; the single outlying female from El Duraznillo (star), tentatively assigned to A. calilegua but not used in calculations, is shown for reference. Right: FeIV/DS ratio, males. In box-plots of male A. tariquiae, small aRROws fROm the left identifY the twO examined β-males." figureDoi="http://doi.org/10.5281/zenodo.12783062" httpUri="https://zenodo.org/record/12783062/files/figure.png" pageId="7" pageNumber="554">Fig. 2</figureCitation>
). The best exteRnal featuRe tO disceRn them fROm females is the DS shape, which Remains λ-
<typeStatus id="8A4D5BA59F64FF8174F5FB14FB7DFB6B" box="[1229,1277,1195,1221]" pageId="7" pageNumber="554">tYpe</typeStatus>
as in α-males. FOR additiOnal details On β males, see the cOmpaRisOn sectiOn Of
<taxonomicName id="92F69E849F64FF81735EFB70FC6DFB47" authorityName="Acosta" authorityYear="2024" box="[870,1005,1231,1257]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F64FF81735EFB70FC6DFB47" box="[870,1005,1231,1257]" italics="true" pageId="7" pageNumber="554">A. calilegua</emphasis>
</taxonomicName>
<emphasis id="678239159F64FF8173CCFB70FBCEFB47" bold="true" box="[1012,1102,1231,1257]" pageId="7" pageNumber="554">
<taxonomicNameLabel id="7CB1846E9F64FF8173CCFB70FBCEFB47" box="[1012,1102,1231,1257]" pageId="7" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
below.
</paragraph>
<paragraph id="5549E5079F64FF8170FFFB4CFBD8FA4B" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="554">
<emphasis id="678239159F64FF8170FFFB4CFEB2FAA3" bold="true" box="[199,306,1267,1293]" pageId="7" pageNumber="554">Females:</emphasis>
DS
<typeStatus id="8A4D5BA59F64FF817150FB4CFE18FAA3" box="[360,408,1267,1293]" pageId="7" pageNumber="554">tYpe</typeStatus>
α-K (alpha-keYhOle, cOda mOdeRatelY lOng with diveRgent sides), unaRmed, slightlY laRgeR than males (
<figureCitation id="CDCDF9829F64FF817119FAA8FEE5FA9F" box="[289,357,1303,1329]" captionStart="FIGURE 2" captionStartId="2.[151,250,1241,1265]" captionTargetBox="[155,1431,509,1214]" captionTargetId="figure-169@2.[151,1435,505,1218]" captionTargetPageId="2" captionText="FIGURE 2. Comparative morphometrics of Acrographinotus tariquiae sp. nov. and A. calilegua sp. nov., displayed as box-plots with data points (internal lines, dashed: mean, solid: median). Left: DS length (in mm) for males and females; the single outlying female from El Duraznillo (star), tentatively assigned to A. calilegua but not used in calculations, is shown for reference. Right: FeIV/DS ratio, males. In box-plots of male A. tariquiae, small aRROws fROm the left identifY the twO examined β-males." figureDoi="http://doi.org/10.5281/zenodo.12783062" httpUri="https://zenodo.org/record/12783062/files/figure.png" pageId="7" pageNumber="554">Fig. 2</figureCitation>
) and more slender than
<taxonomicName id="92F69E849F64FF81724FFAA8FD7FFA9F" authorityName="Acosta" authorityYear="2024" box="[631,767,1303,1329]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F64FF81724FFAA8FD7FFA9F" box="[631,767,1303,1329]" italics="true" pageId="7" pageNumber="554">A. calilegua</emphasis>
</taxonomicName>
<emphasis id="678239159F64FF81733FFAA8FCE1FA9F" bold="true" box="[775,865,1303,1329]" pageId="7" pageNumber="554">
<taxonomicNameLabel id="7CB1846E9F64FF81733FFAA8FCE1FA9F" box="[775,865,1303,1329]" pageId="7" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Granulation slightly more conspicuous and ocular mound higher than males. On free tergites granules are more acute, especially on FT-III, which also bears a small median apophysis, either horizontal or slightly oblique (inclination likely depends on the abdominal swelling). Margin of VAP bordered by small granules, no vestige of male armature. Coxa IV with small acute prolateral apophysis. Trochanter IV: one pair of acute retrolateral granules. FeIV slender, slightly curved at its base, then sub-straight, with rows of granules; retroapical apophysis very small, pointing backwards.
</paragraph>
<paragraph id="5549E5079F64FF8170FFFA51FD29F913" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="554">
<emphasis id="678239159F64FF8170FFFA51FECDF9A6" bold="true" box="[199,333,1518,1544]" pageId="7" pageNumber="554">Variability.</emphasis>
Except fOR β-tYpe specimens, VAP Of males alwaYs has a shelf, with laRge-acute apOphYses in 3/
<specimenCount id="43F02E8E9F64FF8175B7FA50FF5BF983" count="8" pageId="7" pageNumber="554" type="male">8 males</specimenCount>
, laRge-blunt in 2/8; in a single male (the smallest One, which OtheRwise dOes nOt have a decided β habitus) a quadrangular shelf-like projection with acute angles remains (
<figureCitation id="CDCDF9829F64FF81736BF988FC5CF9FF" box="[851,988,1591,1617]" captionStart="FIGURE 3" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[155,1431,313,1749]" captionTargetId="figure-90@6.[151,1435,309,1753]" captionTargetPageId="6" captionText="FIGURE 3. Acrographinotus tariquiae sp. nov. AF: HOlOtYpe ♂ (CBF). A. DORsal scutum, fRee teRgites (alsO shOwing dORsal anal operculum and VAP apophyses), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, VAP with apophyses (also showing apophysis of FT-III), right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. GJ: Armature variations on FT-III and VAP in tOpOtYpical ♂ paRatYpes. GH: Smallest α-male (CBF), IJ: β-male (CDA 000.914). G, I: ventRal view; H, J: lateral view. Scale lines: 2 mm (AE, GJ), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783064" httpUri="https://zenodo.org/record/12783064/files/figure.png" pageId="7" pageNumber="554">Figs. 3GH</figureCitation>
). The apophysis on FT-III is sigmoid in lateral view in 5/
<specimenCount id="43F02E8E9F64FF817160F9E5FE31F9DB" box="[344,433,1626,1653]" count="8" pageId="7" pageNumber="554" type="male">8 males</specimenCount>
, straight or curved in the rest. The diagonal articulation of leg IV is more exaggerated in larger males. In both sexes the low apophysis on the ocular mound may vanish, leaving a plain mound, even with a median notch giving an odd paired appearance.
</paragraph>
</subSubSection>
<subSubSection id="1DECB68C9F64FF8E70FFF979FBE5FE7F" lastPageId="8" lastPageNumber="555" pageId="7" pageNumber="554" type="diagnosis">
<paragraph id="5549E5079F64FF8E70FFF979FBE5FE7F" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1437,151,465]" lastPageId="8" lastPageNumber="555" pageId="7" pageNumber="554">
<emphasis id="678239159F64FF8170FFF979FD9DF94E" bold="true" box="[199,541,1734,1761]" pageId="7" pageNumber="554">Comparisons and diagnosis.</emphasis>
In general habitus and the armature of leg IV of male,
<taxonomicName id="92F69E849F64FF817494F978FAB6F94E" authorityName="Acosta" authorityYear="2024" box="[1196,1334,1735,1761]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F64FF817494F978FAB6F94E" box="[1196,1334,1735,1761]" italics="true" pageId="7" pageNumber="554">A. tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F64FF817507F978FA1CF94F" bold="true" box="[1343,1436,1735,1761]" pageId="7" pageNumber="554">
<taxonomicNameLabel id="7CB1846E9F64FF817507F978FA1CF94F" box="[1343,1436,1735,1761]" pageId="7" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles the
<typeStatus id="8A4D5BA59F64FF817107F954FEEFF8AB" box="[319,367,1771,1797]" pageId="7" pageNumber="554">type</typeStatus>
species
<taxonomicName id="92F69E849F64FF8171EEF954FDF4F8AA" authorityName="Roewer" authorityYear="1929" box="[470,628,1771,1797]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="erectispina">
<emphasis id="678239159F64FF8171EEF954FDF4F8AA" box="[470,628,1771,1797]" italics="true" pageId="7" pageNumber="554">A. erectispina</emphasis>
</taxonomicName>
, the latter being larger (DS length
<quantity id="920E48E29F64FF81742EF954FB21F8AB" box="[1046,1185,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="8.149999999999999" metricValueMax="9.2" metricValueMin="7.1" pageId="7" pageNumber="554" unit="mm" value="8.149999999999999" valueMax="9.2" valueMin="7.1">7.19.2 mm</quantity>
<emphasis id="678239159F64FF817494F954FB43F8AA" box="[1196,1219,1771,1796]" italics="true" pageId="7" pageNumber="554">vs</emphasis>
<quantity id="920E48E29F64FF8174F6F955FAD9F8AB" box="[1230,1369,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="7.2" metricValueMin="5.9" pageId="7" pageNumber="554" unit="mm" value="6.55" valueMax="7.2" valueMin="5.9">5.97.2 mm</quantity>
in
<taxonomicName id="92F69E849F64FF8175BCF954FF7EF886" authorityName="Acosta" authorityYear="2024" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F64FF8175BCF954FF7EF886" italics="true" pageId="7" pageNumber="554">A. tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F64FF81713EF8B0FEE0F887" bold="true" box="[262,352,1807,1833]" pageId="7" pageNumber="554">
<taxonomicNameLabel id="7CB1846E9F64FF81713EF8B0FEE0F887" box="[262,352,1807,1833]" pageId="7" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
); in some features (but not in the overall appearance), the new species can also be compared to
<taxonomicName id="92F69E849F64FF8170AFF88CFE97F8E2" authorityName="Acosta" authorityYear="2001" box="[151,279,1843,1869]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="niawpaq">
<emphasis id="678239159F64FF8170AFF88CFE97F8E2" box="[151,279,1843,1869]" italics="true" pageId="7" pageNumber="554">A. niawpaq</emphasis>
</taxonomicName>
. Of the three,
<taxonomicName id="92F69E849F64FF817186F88CFDC6F8E2" authorityName="Acosta" authorityYear="2024" box="[446,582,1843,1869]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F64FF817186F88CFDC6F8E2" box="[446,582,1843,1869]" italics="true" pageId="7" pageNumber="554">A. tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F64FF817276F88CFD28F8E3" bold="true" box="[590,680,1843,1869]" pageId="7" pageNumber="554">
<taxonomicNameLabel id="7CB1846E9F64FF817276F88CFD28F8E3" box="[590,680,1843,1869]" pageId="7" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has the most gracile look, with weaker development of apophyses and granulation. The presence of large grains—almost small tubercles—on the DS of
<taxonomicName id="92F69E849F64FF81744DF8E8FA91F8DE" authorityName="Roewer" authorityYear="1929" box="[1141,1297,1879,1905]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="erectispina">
<emphasis id="678239159F64FF81744DF8E8FA91F8DE" box="[1141,1297,1879,1905]" italics="true" pageId="7" pageNumber="554">A. erectispina</emphasis>
</taxonomicName>
is an easily observable difference (DS of
<taxonomicName id="92F69E849F64FF8171DAF8C4FDE8F83A" authorityName="Acosta" authorityYear="2024" box="[482,616,1915,1941]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F64FF8171DAF8C4FDE8F83A" box="[482,616,1915,1941]" italics="true" pageId="7" pageNumber="554">A. tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F64FF817257F8C4FD49F83B" bold="true" box="[623,713,1915,1941]" pageId="7" pageNumber="554">
<taxonomicNameLabel id="7CB1846E9F64FF817257F8C4FD49F83B" box="[623,713,1915,1941]" pageId="7" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
nearly smooth, with tiny granules in
<taxonomicName id="92F69E849F64FF817454F8C4FB6BF83A" authorityName="Acosta" authorityYear="2001" box="[1132,1259,1915,1941]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="niawpaq">
<emphasis id="678239159F64FF817454F8C4FB6BF83A" box="[1132,1259,1915,1941]" italics="true" pageId="7" pageNumber="554">A. niawpaq</emphasis>
</taxonomicName>
). VAP is armed in all three, but while the apophyses remain independent in
<taxonomicName id="92F69E849F64FF817373F820FC68F816" authorityName="Roewer" authorityYear="1929" box="[843,1000,1951,1977]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="erectispina">
<emphasis id="678239159F64FF817373F820FC68F816" box="[843,1000,1951,1977]" italics="true" pageId="7" pageNumber="554">A. erectispina</emphasis>
</taxonomicName>
, they delimit a kind of ventral shelf in
<taxonomicName id="92F69E849F64FF81708FF87CFEC0F872" authorityName="Acosta" authorityYear="2024" box="[183,320,1987,2013]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="tariquiae">
<emphasis id="678239159F64FF81708FF87CFEC0F872" box="[183,320,1987,2013]" italics="true" pageId="7" pageNumber="554">A. tariquiae</emphasis>
</taxonomicName>
and
<taxonomicName id="92F69E849F64FF817145F87CFE7FF872" authorityName="Acosta" authorityYear="2001" box="[381,511,1987,2013]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="554" phylum="Arthropoda" rank="species" species="niawpaq">
<emphasis id="678239159F64FF817145F87CFE7FF872" box="[381,511,1987,2013]" italics="true" pageId="7" pageNumber="554">A. niawpaq</emphasis>
</taxonomicName>
(
<figureCitation id="CDCDF9829F64FF81722AF87CFDECF873" box="[530,620,1987,2013]" captionStart="FIGURE 3" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[155,1431,313,1749]" captionTargetId="figure-90@6.[151,1435,309,1753]" captionTargetPageId="6" captionText="FIGURE 3. Acrographinotus tariquiae sp. nov. AF: HOlOtYpe ♂ (CBF). A. DORsal scutum, fRee teRgites (alsO shOwing dORsal anal operculum and VAP apophyses), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, VAP with apophyses (also showing apophysis of FT-III), right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. GJ: Armature variations on FT-III and VAP in tOpOtYpical ♂ paRatYpes. GH: Smallest α-male (CBF), IJ: β-male (CDA 000.914). G, I: ventRal view; H, J: lateral view. Scale lines: 2 mm (AE, GJ), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783064" httpUri="https://zenodo.org/record/12783064/files/figure.png" pageId="7" pageNumber="554">Fig. 3B</figureCitation>
, see also
<bibRefCitation id="316798F69F64FF8172E7F87CFCF3F873" author="Acosta, L. E." box="[735,883,1987,2013]" pageId="7" pageNumber="554" pagination="58 - 66" refId="ref10602" refString="Acosta, L. E. (2001) The identity of Acrographinotus erectispina, with a review of the generic diagnosis, and the description of a new species (Opiliones, Gonyleptidae, Pachylinae). Bulletin of the British Arachnological Society, 12 (2), 58 - 66." type="journal article" year="2001">Acosta 2001</bibRefCitation>
: figs. 2, 25). Further, the apophysis of FT-III is subhorizontal in Roewers species, sigmoid in
<taxonomicName id="92F69E849F6BFF8E7295FF28FCB5FF1E" authorityName="Acosta" authorityYear="2024" box="[685,821,151,177]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="tariquiae">
<emphasis id="678239159F6BFF8E7295FF28FCB5FF1E" box="[685,821,151,177]" italics="true" pageId="8" pageNumber="555">A. tariquiae</emphasis>
</taxonomicName>
and
<taxonomicName id="92F69E849F6BFF8E7357FF28FC70FF1E" authorityName="Acosta" authorityYear="2001" box="[879,1008,151,177]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="niawpaq">
<emphasis id="678239159F6BFF8E7357FF28FC70FF1E" box="[879,1008,151,177]" italics="true" pageId="8" pageNumber="555">A. niawpaq</emphasis>
</taxonomicName>
(much more developed in the latter).
<taxonomicName id="92F69E849F6BFF8E70AFFF04FE58FF7A" authorityName="Roewer" authorityYear="1929" box="[151,472,187,213]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="erectispina">
<emphasis id="678239159F6BFF8E70AFFF04FE58FF7A" box="[151,472,187,213]" italics="true" pageId="8" pageNumber="555">Acrographinotus erectispina</emphasis>
</taxonomicName>
bears one single large retroapical apophysis on trochanter IV, whilst in
<taxonomicName id="92F69E849F6BFF8E752CFF04FA1DFF7A" authorityName="Acosta" authorityYear="2024" box="[1300,1437,187,213]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F6BFF8E752CFF04FA1DFF7A" box="[1300,1437,187,213]" italics="true" pageId="8" pageNumber="555">A. tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F6BFF8E70AFFF60FF71FF57" bold="true" box="[151,241,223,249]" pageId="8" pageNumber="555">
<taxonomicNameLabel id="7CB1846E9F6BFF8E70AFFF60FF71FF57" box="[151,241,223,249]" pageId="8" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="92F69E849F6BFF8E711EFF60FE25FF56" authorityName="Acosta" authorityYear="2001" box="[294,421,223,249]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="niawpaq">
<emphasis id="678239159F6BFF8E711EFF60FE25FF56" box="[294,421,223,249]" italics="true" pageId="8" pageNumber="555">A. niawpaq</emphasis>
</taxonomicName>
there are two short apophyses; the bilobated prodorsal apophysis is peculiar to
<taxonomicName id="92F69E849F6BFF8E7525FF60FA1CFF56" authorityName="Acosta" authorityYear="2001" box="[1309,1436,223,249]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="niawpaq">
<emphasis id="678239159F6BFF8E7525FF60FA1CFF56" box="[1309,1436,223,249]" italics="true" pageId="8" pageNumber="555">A. niawpaq</emphasis>
</taxonomicName>
(it is simple in the others). The retrodorsal row of truncate tubercles on FeIV is similar in
<taxonomicName id="92F69E849F6BFF8E7494FEBCFAC8FEB2" authorityName="Roewer" authorityYear="1929" box="[1196,1352,259,285]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="erectispina">
<emphasis id="678239159F6BFF8E7494FEBCFAC8FEB2" box="[1196,1352,259,285]" italics="true" pageId="8" pageNumber="555">A. erectispina</emphasis>
</taxonomicName>
and
<taxonomicName id="92F69E849F6BFF8E75BCFEBCFF7EFEEE" authorityName="Acosta" authorityYear="2024" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="tariquiae">
<emphasis id="678239159F6BFF8E75BCFEBCFF7EFEEE" italics="true" pageId="8" pageNumber="555">A. tariquiae</emphasis>
</taxonomicName>
, whereas in
<taxonomicName id="92F69E849F6BFF8E71B3FE98FD8AFEEE" authorityName="Acosta" authorityYear="2001" box="[395,522,295,321]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="niawpaq">
<emphasis id="678239159F6BFF8E71B3FE98FD8AFEEE" box="[395,522,295,321]" italics="true" pageId="8" pageNumber="555">A. niawpaq</emphasis>
</taxonomicName>
the dorsal apophyses are larger, and the femur is thicker. The subdistal narrowing of FeIV is evident in
<taxonomicName id="92F69E849F6BFF8E71BBFEF4FD89FECA" authorityName="Acosta" authorityYear="2024" box="[387,521,331,357]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="tariquiae">
<emphasis id="678239159F6BFF8E71BBFEF4FD89FECA" box="[387,521,331,357]" italics="true" pageId="8" pageNumber="555">A. tariquiae</emphasis>
</taxonomicName>
and
<emphasis id="678239159F6BFF8E7206FEF4FD44FECA" box="[574,708,331,357]" italics="true" pageId="8" pageNumber="555">
<taxonomicName id="92F69E849F6BFF8E7206FEF4FD3FFECA" authorityName="Acosta" authorityYear="2001" box="[574,703,331,357]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="niawpaq">A. niawpaq</taxonomicName>
,
</emphasis>
not as much in
<taxonomicName id="92F69E849F6BFF8E734CFEF4FB8DFECA" authorityName="Roewer" authorityYear="1929" box="[884,1037,331,357]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="erectispina">
<emphasis id="678239159F6BFF8E734CFEF4FB8DFECA" box="[884,1037,331,357]" italics="true" pageId="8" pageNumber="555">A. erectispina</emphasis>
</taxonomicName>
(
<emphasis id="678239159F6BFF8E7425FEF4FBBAFECB" box="[1053,1082,331,357]" italics="true" pageId="8" pageNumber="555">cf.</emphasis>
<bibRefCitation id="316798F69F6BFF8E7406FEF4FB4EFECB" author="Acosta, L. E." box="[1086,1230,331,357]" pageId="8" pageNumber="555" pagination="58 - 66" refId="ref10602" refString="Acosta, L. E. (2001) The identity of Acrographinotus erectispina, with a review of the generic diagnosis, and the description of a new species (Opiliones, Gonyleptidae, Pachylinae). Bulletin of the British Arachnological Society, 12 (2), 58 - 66." type="journal article" year="2001">Acosta 2001</bibRefCitation>
). As evident in the identification key, VAP enables an easy separation of
<taxonomicName id="92F69E849F6BFF8E72CFFED0FCFFFE26" authorityName="Acosta" authorityYear="2024" box="[759,895,367,393]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F6BFF8E72CFFED0FCFFFE26" box="[759,895,367,393]" italics="true" pageId="8" pageNumber="555">A. tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F6BFF8E73BEFED0FC61FE27" bold="true" box="[902,993,367,393]" pageId="8" pageNumber="555">
<taxonomicNameLabel id="7CB1846E9F6BFF8E73BEFED0FC61FE27" box="[902,993,367,393]" pageId="8" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from other members of the genus: it is unarmed in the spiny-looking species (
<emphasis id="678239159F6BFF8E7265FE2CFC57FE02" box="[605,983,403,429]" italics="true" pageId="8" pageNumber="555">
<taxonomicName id="92F69E849F6BFF8E7265FE2CFD77FE02" authorityName="Roewer" authorityYear="1929" box="[605,759,403,429]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="curvispina">A. curvispina</taxonomicName>
.
<taxonomicName id="92F69E849F6BFF8E7339FE2CFCF2FE02" authorityName="Acosta" authorityYear="2002" box="[769,882,403,429]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="mitmaj">A. mitmaj</taxonomicName>
,
<taxonomicName id="92F69E849F6BFF8E7344FE2CFC57FE02" baseAuthorityName="Roewer" baseAuthorityYear="1957" box="[892,983,403,429]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="ortizi">A. ortizi</taxonomicName>
</emphasis>
) and bears large divergent apophyses in
<taxonomicName id="92F69E849F6BFF8E70AFFE08FEBFFE7E" baseAuthorityName="Soares &amp; Bauab" baseAuthorityYear="1972" box="[151,319,439,465]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="ceratopygus">
<emphasis id="678239159F6BFF8E70AFFE08FEBFFE7E" box="[151,319,439,465]" italics="true" pageId="8" pageNumber="555">A. ceratopygus</emphasis>
</taxonomicName>
. Differences with
<taxonomicName id="92F69E849F6BFF8E7237FE08FD16FE7F" authorityName="Acosta" authorityYear="2024" box="[527,662,439,465]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F6BFF8E7237FE08FD16FE7F" box="[527,662,439,465]" italics="true" pageId="8" pageNumber="555">A. calilegua</emphasis>
</taxonomicName>
<emphasis id="678239159F6BFF8E72A5FE08FD77FE7F" bold="true" box="[669,759,439,465]" pageId="8" pageNumber="555">
<taxonomicNameLabel id="7CB1846E9F6BFF8E72A5FE08FD77FE7F" box="[669,759,439,465]" pageId="8" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are addressed to in detail below.
</paragraph>
</subSubSection>
<caption id="0189B58F9F6BFF8E70AFF9A9FC27F9FC" ID-DOI="http://doi.org/10.5281/zenodo.12783067" ID-Zenodo-Dep="12783067" httpUri="https://zenodo.org/record/12783067/files/figure.png" pageId="8" pageNumber="555" startId="8.[151,250,1558,1582]" targetBox="[277,1309,493,1531]" targetPageId="8" targetType="figure">
<paragraph id="5549E5079F6BFF8E70AFF9A9FC27F9FC" blockId="8.[151,1436,1558,1618]" pageId="8" pageNumber="555">
<emphasis id="678239159F6BFF8E70AFF9A9FE95F980" bold="true" box="[151,277,1558,1582]" pageId="8" pageNumber="555">FIGURE 4.</emphasis>
<taxonomicName id="92F69E849F6BFF8E7125F9A9FDADF980" authorityName="Acosta" authorityYear="2024" box="[285,557,1558,1582]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F6BFF8E7125F9A9FDADF980" box="[285,557,1558,1582]" italics="true" pageId="8" pageNumber="555">Acrographinotus tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F6BFF8E720DF9A9FD08F980" bold="true" box="[565,648,1558,1582]" pageId="8" pageNumber="555">
<taxonomicNameLabel id="7CB1846E9F6BFF8E720DF9A9FD08F980" box="[565,648,1558,1582]" pageId="8" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, distal end Of penis, HOlOtYpe ♂ (CBF). A: dORsal view; B: lateRal view; C: detail of stylus and ventral process. Scale lines: 0.1 mm (A, B), 0.05 mm (C).
</paragraph>
</caption>
<subSubSection id="1DECB68C9F6BFF8E70FFF93FFB8FF871" pageId="8" pageNumber="555" type="distribution">
<paragraph id="5549E5079F6BFF8E70FFF93FFB8FF871" blockId="8.[151,1437,1664,2015]" pageId="8" pageNumber="555">
<emphasis id="678239159F6BFF8E70FFF93FFE72F934" bold="true" box="[199,498,1664,1691]" pageId="8" pageNumber="555">Distribution and habitat.</emphasis>
At the moment, the species is known from two sites in
<collectingRegion id="97322BE59F6BFF8E7459F93FFAB0F935" box="[1121,1328,1664,1691]" country="Bolivia" name="Tarija" pageId="8" pageNumber="555">Tarija Department</collectingRegion>
, southern
<collectingCountry id="2DE1A5979F6BFF8E70AFF91BFF69F910" box="[151,233,1700,1726]" name="Bolivia" pageId="8" pageNumber="555">Bolivia</collectingCountry>
(
<figureCitation id="CDCDF9829F6BFF8E70C1F91BFED3F911" box="[249,339,1700,1727]" captionStart="FIGURE 7" captionStartId="12.[151,250,1626,1650]" captionTargetBox="[151,1435,634,1602]" captionTargetId="figure-132@12.[151,1435,634,1602]" captionTargetPageId="12" captionText="FIGURE 7. A: Records of Acrographinotus tariquiae sp. nov. in Tarija Department, Bolivia (yellow dots) and A. calilegua sp. nov. in Jujuy Province, Argentina (red dots); blue contours: protected areas (1: Tariquía Flora and Fauna National Reserve; 2: Calilegua National Park). B: Type locality of A. calilegua sp. nov.: cloud forests at Monolito (ca. 1700 m a.s.l.), Calilegua National Park, Jujuy Province, Argentina." figureDoi="http://doi.org/10.5281/zenodo.12783073" httpUri="https://zenodo.org/record/12783073/files/figure.png" pageId="8" pageNumber="555">Fig. 7A</figureCitation>
). The
<typeStatus id="8A4D5BA59F6BFF8E71A1F91AFE49F911" box="[409,457,1701,1727]" pageId="8" pageNumber="555">type</typeStatus>
locality (Subida de La Escalera, ~
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a.s.l.) is placed on the east-faced slope of the N-S oriented mountains that delimit the Tariquía Flora and Fauna National Reserve. This area is described as having an abrupt relief, with herbaceous vegetation on higher sectors, and shrub- or woodlands of
<taxonomicName id="92F69E849F6BFF8E7515F953FF57F884" authority=", Alnus" authorityName="Alnus" class="Diplopoda" family="Platyrhacidae" genus="Polylepis" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="555" phylum="Arthropoda" rank="genus">
<emphasis id="678239159F6BFF8E7515F953FF57F884" italics="true" pageId="8" pageNumber="555">Polylepis, Alnus</emphasis>
</taxonomicName>
or
<taxonomicName id="92F69E849F6BFF8E70C7F8AFFE06F884" box="[255,390,1808,1834]" class="Magnoliopsida" family="Sapotaceae" genus="Podocarpus" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="555" phylum="Tracheophyta" rank="genus">
<emphasis id="678239159F6BFF8E70C7F8AFFE06F884" box="[255,390,1808,1834]" italics="true" pageId="8" pageNumber="555">Podocarpus</emphasis>
</taxonomicName>
at lower elevation and in ravines, nearly rainforest-like formations further below (
<bibRefCitation id="316798F69F6BFF8E7516F8AFFF78F8E1" author="Ayarde, H. R. &amp; Boero, C. &amp; Moris, M. &amp; Slanis, A. &amp; Gonzalez, J. A." pageId="8" pageNumber="555" pagination="29 - 62" refId="ref11025" refString="Ayarde, H. R., Boero, C., Moris, M., Slanis, A. &amp; Gonzalez, J. A. (1999) Flora y vegetacion de Tariquia. In: Gonzalez, J. A., Scrocchi, G. J. &amp; Lavilla E. O. (Eds.), Relevamiento de la biodiversidad de la Reserva Nacional de Flora y Fauna Tariquia (Tarija, Bolivia). Serie Conservacion de la Naturaleza. N ° 14. Fundacion Miguel Lillo, San Miguel de Tucuman, pp. 29 - 62." type="book chapter" year="1999">
Ayarde
<emphasis id="678239159F6BFF8E75B0F8AEFF34F8E0" italics="true" pageId="8" pageNumber="555">et al.</emphasis>
1999
</bibRefCitation>
). The specimens were caught on the steep walls of a stream ravine, in a humid sector with impoverished montane forest that includes arborescent ferns (P. Goloboff
<emphasis id="678239159F6BFF8E730EF8E6FCFAF8DC" box="[822,890,1880,1906]" italics="true" pageId="8" pageNumber="555">in litt.</emphasis>
). The second site (Abra de La Hondura,
<quantity id="920E48E29F6BFF8E757FF8E6FA1CF8DD" box="[1351,1436,1881,1907]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="555" unit="m" value="2700.0">2700 m</quantity>
a.s.l.), on the west slope of the same range, is more xeric than the former. It has a similar scabrous landscape and is dominated by grasslands, especially on elevated sites; small
<taxonomicName id="92F69E849F6BFF8E735DF81FFC4DF814" box="[869,973,1952,1978]" class="Diplopoda" family="Platyrhacidae" genus="Polylepis" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="555" phylum="Arthropoda" rank="genus">
<emphasis id="678239159F6BFF8E735DF81FFC4DF814" box="[869,973,1952,1978]" italics="true" pageId="8" pageNumber="555">Polylepis</emphasis>
</taxonomicName>
woodlands develop in ravines and more xerophilous vegetation in lower slopes (
<bibRefCitation id="316798F69F6BFF8E726EF87BFCA9F871" author="Ayarde, H. R. &amp; Boero, C. &amp; Moris, M. &amp; Slanis, A. &amp; Gonzalez, J. A." box="[598,809,1988,2015]" pageId="8" pageNumber="555" pagination="29 - 62" refId="ref11025" refString="Ayarde, H. R., Boero, C., Moris, M., Slanis, A. &amp; Gonzalez, J. A. (1999) Flora y vegetacion de Tariquia. In: Gonzalez, J. A., Scrocchi, G. J. &amp; Lavilla E. O. (Eds.), Relevamiento de la biodiversidad de la Reserva Nacional de Flora y Fauna Tariquia (Tarija, Bolivia). Serie Conservacion de la Naturaleza. N ° 14. Fundacion Miguel Lillo, San Miguel de Tucuman, pp. 29 - 62." type="book chapter" year="1999">
Ayarde
<emphasis id="678239159F6BFF8E7297F87AFD67F870" box="[687,743,1988,2014]" italics="true" pageId="8" pageNumber="555">et al.</emphasis>
1999
</bibRefCitation>
; P. Goloboff
<emphasis id="678239159F6BFF8E7384F87AFC7FF870" box="[956,1023,1988,2014]" italics="true" pageId="8" pageNumber="555">in litt.</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>