treatments-xml/data/C3/F1/C1/C3F1C11A938644BBD019EE3B350174D0.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.222.3375" ID-GBIF-Dataset="8951a9f2-d604-41db-a43c-605fef3cb1af" ID-PMC="PMC3459029" ID-Pensoft-Pub="1313-2970-222-11" ID-PubMed="23129985" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-222-11" ModsDocOrigin="ZooKeys 222" ModsDocTitle="Review of the species level taxonomy of the neotropical butterfly genus Oenomaus (Lycaenidae, Theclinae, Eumaeini)" checkinTime="1451248737870" checkinUser="pensoft" docAuthor="Faynel, Christophe, Busby, Robert C. &amp; Robbins, Robert K." docDate="2012" docId="C3F1C11A938644BBD019EE3B350174D0" docLanguage="en" docName="ZooKeys 222: 11-45" docOrigin="ZooKeys 222" docSource="http://dx.doi.org/10.3897/zookeys.222.3375" docTitle="Oenomaus lea Faynel &amp; Robbins, sp. n." docType="treatment" docVersion="4" lastPageNumber="17" masterDocId="FF8FFFA2C315166F635BFFE7FFA97842" masterDocTitle="Review of the species level taxonomy of the neotropical butterfly genus Oenomaus (Lycaenidae, Theclinae, Eumaeini)" masterLastPageNumber="45" masterPageNumber="11" pageNumber="16" updateTime="1668154462762" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the species level taxonomy of the neotropical butterfly genus Oenomaus (Lycaenidae, Theclinae, Eumaeini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Faynel, Christophe</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Busby, Robert C.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Robbins, Robert K.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>222</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>11</mods:start>
<mods:end>45</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.222.3375</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.222.3375</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-222-11</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152037077" LSID="urn:lsid:zoobank.org:act:A8315354-6FFF-4C71-AEE5-92925AE8833B" httpUri="http://treatment.plazi.org/id/C3F1C11A938644BBD019EE3B350174D0" lastPageId="6" lastPageNumber="17" pageId="5" pageNumber="16">
<subSubSection pageId="5" pageNumber="16" type="nomenclature">
<paragraph pageId="5" pageNumber="16">
<taxonomicName LSID="urn:lsid:zoobank.org:act:A8315354-6FFF-4C71-AEE5-92925AE8833B" authority="Faynel &amp; Robbins" class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus lea" order="Lepidoptera" pageId="5" pageNumber="16" phylum="Arthropoda" rank="species" species="lea">Oenomaus lea Faynel &amp; Robbins</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="16">sp. n.</taxonomicNameLabel>
Figs 4, 21, 39, 47
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="17" pageId="5" pageNumber="16" type="type-locality">
<paragraph pageId="5" pageNumber="16">Type-locality.</paragraph>
<paragraph lastPageId="6" lastPageNumber="17" pageId="5" pageNumber="16">
Ecuador: Napo,
<normalizedToken originalValue="Misahuallí">Misahualli</normalizedToken>
Rd, Latas Grande, 7.7 km E Puerto Napo,
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-1.0333333">1°02.0'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.735">77°44.1'W</geoCoordinate>
, 470 m. The holotype was collected along the road from Tena to
<normalizedToken originalValue="Misahuallí">Misahualli</normalizedToken>
, which in 1991 was a patchwork of
<normalizedToken originalValue="“fincas”">&quot;fincas&quot;</normalizedToken>
and remnant second growth forest
<pageBreakToken pageId="6" pageNumber="17" start="start">.</pageBreakToken>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="17" type="type-specimen">
<paragraph pageId="6" pageNumber="17">Type-specimen.</paragraph>
<paragraph pageId="6" pageNumber="17">
Holotype ♂ (Fig. 4) labeled as &quot;ECUADOR Napo / Misahualli Rd. 470m / Latas Grande / 9 Nov.' 91 / S. S. Nicolay&quot; [rectangular, white, printed and handwritten], &quot;USNM ENT 00180040&quot; [rectangular, white, printed], &quot;GENITALIA NO. / 2011: 408♂ / C. FAYNEL&quot; [rectangular, green, printed] &quot;Holotype ♂ /
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus lea" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="lea">Oenomaus lea</taxonomicName>
/ Faynel &amp; Robbins, 2012&quot; [rectangular, red, printed]. Deposited in USNM.
</paragraph>
<paragraph pageId="6" pageNumber="17">
Paratypes: Ecuador.1 ♂: La Merced on
<normalizedToken originalValue="Río">Rio</normalizedToken>
Pastaza below
<normalizedToken originalValue="Baños">Banos</normalizedToken>
, Alt. 4000 ft. [= 1220 m], W. J. Coxey, III.1930, A.N.S. Lot 217, genitalia NO. 1992: 12 ♂ R.K. Robbins (ANSP). Peru. 3 ♂: UC, Pucallpa, 200 m, X.2007, Michael
<normalizedToken originalValue="Büche">Bueche</normalizedToken>
leg. (CF); LO, Contamana,
<normalizedToken originalValue="Río">Rio</normalizedToken>
Ucayali, 300 m,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-7.3166666">7°19'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-74.8">74°48'W</geoCoordinate>
, IX.2010, leg J.
<normalizedToken originalValue="Ramírez">Ramirez</normalizedToken>
(LYD); SM, Juanjui, upper Huallaga River, IX. 1934, collector G. Klug, collection E.I. Huntington NO. 1055 (AMNH).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="17" type="description">
<paragraph pageId="6" pageNumber="17">Description, diagnosis and recognition as a distinct species.</paragraph>
<paragraph pageId="6" pageNumber="17">
Male FW length: 18.2 mm (SD = 0.8,
<taxonomicName genus="N" lsidName="N" pageId="6" pageNumber="17" rank="genus">N</taxonomicName>
= 4). Wing pattern (Fig. 4) and genitalia (Fig. 21) illustrated.
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus lea" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="lea">Oenomaus lea</taxonomicName>
and
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus atesa" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="atesa">Oenomaus atesa</taxonomicName>
(Hewitson, 1867)are the only two
<taxonomicName lsidName="" pageId="6" pageNumber="17" rank="tribe" tribe="Eumaeini">Eumaeini</taxonomicName>
sharing the striking underside wing pattern with two transverse brown bands on the VHW. However, males of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus lea" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="lea">Oenomaus lea</taxonomicName>
differ from males of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus atesa" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="atesa">Oenomaus atesa</taxonomicName>
(Figs 5, 6 in
<bibRefCitation pageId="6" pageNumber="17">Faynel 2006</bibRefCitation>
) by (1) a greater expanse of the DFW blue, especially in the area from vein R3 to vein Cu1, (2) two brown patches on the VFW instead of a single median brown band; one patch is located along the costa, and the other is triangular and situated in the basal part of cell Cu1-Cu2, and (3) a lighter dorsal blue color with a different hue of blue along the HW veins M2, M3, Cu1 and Cu2. Moreover, the black spot in VHW cell Cu1-Cu2 is usually more apparent in
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus atesa" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="atesa">Oenomaus atesa</taxonomicName>
than in
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus lea" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="lea">Oenomaus lea</taxonomicName>
. Males of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus lea" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="lea">Oenomaus lea</taxonomicName>
alsodiffer genitalically from those of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus atesa" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="atesa">Oenomaus atesa</taxonomicName>
(Fig. 26 in
<bibRefCitation pageId="6" pageNumber="17">Faynel 2006</bibRefCitation>
) by (1) a longer and wider saccus, (2) the dorsal part of the valvae in lateral aspect shorter than the ventral part, and not pointed at the posterior end, (3) a straight penis in lateral view, and (4) no tooth at the end of the penis. The eighth tergum shows no difference from that of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus atesa" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="atesa">Oenomaus atesa</taxonomicName>
. Lastly, the divergence of
<normalizedToken originalValue="“barcode”">&quot;barcode&quot;</normalizedToken>
DNA sequence data between
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus lea" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="lea">Oenomaus lea</taxonomicName>
(CF-LYC-005) and
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus atesa" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="atesa">Oenomaus atesa</taxonomicName>
(CF-LYC-003) is more than 6% (Table 1). The female of
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus lea" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="lea">Oenomaus lea</taxonomicName>
is unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="17" type="etymology">
<paragraph pageId="6" pageNumber="17">Etymology.</paragraph>
<paragraph pageId="6" pageNumber="17">
This species is named for
<normalizedToken originalValue="Léa">Lea</normalizedToken>
Faynel, daughter of Christophe Faynel. The name is a feminine noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="17" type="habitat and distribution">
<paragraph pageId="6" pageNumber="17">Habitat and distribution.</paragraph>
<paragraph pageId="6" pageNumber="17">
<taxonomicName class="Insecta" family="Lycaenidae" genus="Oenomaus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenomaus lea" order="Lepidoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="lea">Oenomaus lea</taxonomicName>
occurs in wet lowland forest up to 1200 m elevation in eastern Ecuador and eastern Peru (Fig. 47).
</paragraph>
</subSubSection>
</treatment>
</document>