159 lines
17 KiB
XML
159 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.96.101873" ID-Pensoft-Pub="1314-2607-96-241" ID-Pensoft-UUID="A5722C06212C5BA7A2099B77C6A8DF54" ID-ZooBank="15A2B06B92F34E70AC8F6FEABF365E71" ModsDocID="1314-2607-96-241" checkinTime="1684817117864" checkinUser="pensoft" docAuthor="Wood, Thomas J." docDate="2023" docId="91D35C680A880B854DCC768FCFC2E252" docLanguage="en" docName="JourHymenoptRes 96: 241-484" docOrigin="Journal of Hymenoptera Research 96" docPubDate="2023-05-22" docSource="http://dx.doi.org/10.3897/jhr.96.101873" docTitle="Pruinosandrena farinosoides subsp. subgen. nov." docType="treatment" docVersion="1" id="A5722C06212C5BA7A2099B77C6A8DF54" lastPageNumber="241" masterDocId="A5722C06212C5BA7A2099B77C6A8DF54" masterDocTitle="The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera, Andrenidae)" masterLastPageNumber="484" masterPageNumber="241" pageNumber="241" updateTime="1684817117864" updateUser="pensoft">
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<mods:titleInfo id="6F725AA47BA529D9D5C1E8A90E2AE600">
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<mods:title id="BEDFBE9AD2AFE7506F7107774FF80480">The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera, Andrenidae)</mods:title>
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<mods:namePart id="D58845C59F692BAE5660EFC124DE63E5">Wood, Thomas J.</mods:namePart>
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<mods:affiliation id="F511FE5D82AB61FAABE0D915FA4BC4D6">University of Mons, Research Institute for Biosciences, Laboratory of Zoology, Place du Parc 20, 7000, Mons, Belgium</mods:affiliation>
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<mods:nameIdentifier id="83142285CBB673DD8A3C2DC36D5C2EE9" type="email">thomasjames.wood@umons.ac.be</mods:nameIdentifier>
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<mods:typeOfResource id="0E7839C3FB49DED3F5971C54F8767E63">text</mods:typeOfResource>
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<mods:title id="B17BBAAC70AAC15CA9C653E00031CB08">Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part id="CA8A796EA16B549D968EAAF2BECF5494">
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<mods:date id="B6CE51B0DFAA65A9557DE30A38ADE2FA">2023</mods:date>
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<mods:number id="C45CCB909813782014CF5476530A6854">2023-05-22</mods:number>
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<mods:detail id="118D5ECAEBC98374DBDB3926AFD9D10E" type="volume">
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<mods:number id="DA942031A3D0DF7FCDB28ADA523E9F41">96</mods:number>
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<mods:start id="4EC7A35B57FFCA896667B08C771298F6">241</mods:start>
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<mods:end id="C8FDF4CF363DF82571056CB49886DFE1">484</mods:end>
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<mods:url id="E37F6954E2E5ECC5F893C1FD6B881DA0">http://dx.doi.org/10.3897/jhr.96.101873</mods:url>
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<mods:classification id="8623E0083A836606B4966EB30DB5C471">journal article</mods:classification>
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<mods:identifier id="2BAEDA32EA86BEB70B55F8FF64EFCB10" type="DOI">http://dx.doi.org/10.3897/jhr.96.101873</mods:identifier>
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<mods:identifier id="B0A3C70C6EB36317D46AB1AC92E2D8D7" type="Pensoft-Pub">1314-2607-96-241</mods:identifier>
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<treatment id="91D35C680A880B854DCC768FCFC2E252" LSID="urn:lsid:zoobank.org:act:62A1E8AF-01AD-4D69-8CD5-E20843A286D5" httpUri="http://treatment.plazi.org/id/91D35C680A880B854DCC768FCFC2E252" lastPageNumber="241" pageId="0" pageNumber="241">
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<subSubSection id="FD0F9AAE7DD8D395A58D7C23C30A6CD6" pageId="0" pageNumber="241" type="nomenclature">
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<paragraph id="7C615C6D915DFA68F5BCF877C4A53A84" pageId="0" pageNumber="241">
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Subgenus
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<taxonomicName id="51229B5D3CC616584567FA22BBC804E0" LSID="https://zoobank.org/62A1E8AF-01AD-4D69-8CD5-E20843A286D5" family="Andrenidae" genus="Pruinosandrena" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pruinosandrena subsp. subgen. nov." order="Hymenoptera" pageId="0" pageNumber="241" rank="subSpecies" species="farinosoides" subSpecies="subgen. nov.">Pruinosandrena subgen. nov.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="A7279377CC5BB8F274C00B018B4F7028" pageId="0" pageNumber="241" type="type species">
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<paragraph id="FE4D5882C554C8A8BAE2C8056905EC5F" pageId="0" pageNumber="241">Type species.</paragraph>
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<paragraph id="A80AC27C67F26849DEC51A5CACBEA091" pageId="0" pageNumber="241">
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<taxonomicName id="0996E37B74DD19E2018EE04264F6DCD0" authorityName="Erichson" authorityYear="1835" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena pruinosa" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="pruinosa">
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<emphasis id="2A44E6609D2F05564D1CE4D33580A9F6" italics="true" pageId="0" pageNumber="241">Andrena pruinosa</emphasis>
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</taxonomicName>
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Erichson, 1835.
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</paragraph>
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</subSubSection>
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<subSubSection id="C579A77133BAF7E9A070F28851929D95" pageId="0" pageNumber="241" type="diagnosis">
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<paragraph id="196A67817673579789D21CDC726B2E66" pageId="0" pageNumber="241">Diagnosis.</paragraph>
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<paragraph id="D7F7837FD08A9EC96450BA0BCD7B7EEC" pageId="0" pageNumber="241">
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This group of species was formerly placed in the subgenus
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<taxonomicName id="A6FD2CDADFA84EDF2BD8E1DB6A3F2586" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Campylogaster)" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="subGenus" subGenus="Campylogaster">Andrena Campylogaster</taxonomicName>
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due to the unusual character of the strongly and densely punctate mesepisternum and dorsolateral parts of the propodeum (punctures separated by <0.5 puncture diameters). However, as discussed above,
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<taxonomicName id="C23B9D663820120CF1E26363880B056A" authorityName="Dours" authorityYear="1873" class="Insecta" family="Andrenidae" genus="Campylogaster" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Campylogaster" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="genus">
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<emphasis id="855F2585290010F72C83E61305265D10" italics="true" pageId="0" pageNumber="241">Campylogaster</emphasis>
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</taxonomicName>
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sensu Warncke is polyphyletic and can be broken into three distinct clades (
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<taxonomicName id="64D69505039C32A7096255C99C5CA100" authorityName="Dours" authorityYear="1873" class="Insecta" family="Andrenidae" genus="Campylogaster" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Campylogaster" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="genus">
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<emphasis id="A03D21F8FE03ECFD67252C9C68C52E6A" italics="true" pageId="0" pageNumber="241">Campylogaster</emphasis>
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</taxonomicName>
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s. str.,
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<taxonomicName id="3B76FA45BCAF9C4D0162B262EB19949A" family="Andrenidae" genus="Pruinosandrena" kingdom="Animalia" lsidName="Pruinosandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="3FB94D45A906BD11DEA3A477FA5614C6" italics="true" pageId="0" pageNumber="241">Pruinosandrena</emphasis>
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</taxonomicName>
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, and the
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<taxonomicName id="3ED4958BB41F85C15873FAFF375A17D2" authorityName="Eversmann" authorityYear="1852" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena incisa" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="incisa">
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<emphasis id="8D5AADBB7CD2877F961B4AAD5D9BE64B" italics="true" pageId="0" pageNumber="241">Andrena incisa</emphasis>
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</taxonomicName>
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-group). All three share the distinctly punctate mesepisternum and also an extremely long ocelloccipital distance exceeding three times the diameter of the lateral ocellus. However, separation is straightforward. The true
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<taxonomicName id="C559BED39E202BD27191A573F4C13B9D" authorityName="Dours" authorityYear="1873" class="Insecta" family="Andrenidae" genus="Campylogaster" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Campylogaster" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="genus">
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<emphasis id="55B2BC94BBA93A71080ACAD724FC8904" italics="true" pageId="0" pageNumber="241">Campylogaster</emphasis>
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</taxonomicName>
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have the marginal area of the terga clearly and distinctly impressed with the apical margin reflexed; the impression therefore forms a latitudinal depressed furrow. In
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<taxonomicName id="6B0F856378EB7B52AE15B6609CF0D05C" family="Andrenidae" genus="Pruinosandrena" kingdom="Animalia" lsidName="Pruinosandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="B2301EF2BB9CB3865B057EE985B8770D" italics="true" pageId="0" pageNumber="241">Pruinosandrena</emphasis>
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</taxonomicName>
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, the marginal areas of the terga are flat, without any kind of depression. In the
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<taxonomicName id="78D1388F6D536936F3404315D48CA8C5" authorityName="Eversmann" authorityYear="1852" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena incisa" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="incisa">
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<emphasis id="14AE259B018844FDAA1D8D63FDD521FB" italics="true" pageId="0" pageNumber="241">Andrena incisa</emphasis>
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</taxonomicName>
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-group, the tergal margins are flat and the mesepisternum is densely punctate, but the dorsolateral parts of the propodeum have only raised reticulation, without punctures. The propodeal triangle is also clearly delineated by raised lateral carinae, whereas these are absent in
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<taxonomicName id="612623574821C948C0F8DBC86F815314" family="Andrenidae" genus="Pruinosandrena" kingdom="Animalia" lsidName="Pruinosandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="90B7479BB331BC2B544CFB2007DCB0E6" italics="true" pageId="0" pageNumber="241">Pruinosandrena</emphasis>
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</taxonomicName>
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. The combination of extremely long ocelloccipital distance, dense punctures on the mesepisternum and dorsolateral parts of the propodeum, and flat tergal marginal areas is therefore unique and characterises
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<taxonomicName id="855F12B7AE249A56B5B8EFD0FD3A2618" family="Andrenidae" genus="Pruinosandrena" kingdom="Animalia" lsidName="Pruinosandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="C511E1A6F32F5CC2A4BB19A3CC35278E" italics="true" pageId="0" pageNumber="241">Pruinosandrena</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="9CDC4FD5152AFD456841A6E0645F93A7" pageId="0" pageNumber="241" type="description">
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<paragraph id="835913942F308F5BEFED20C670290B36" pageId="0" pageNumber="241">Description.</paragraph>
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<paragraph id="3AC6D226979D50943D9931110D9D053F" pageId="0" pageNumber="241">
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Medium-sized bees (10-14 mm). Integument variable, from dark with at most tergal margins lightened hyaline-yellow to entirety of metasoma and legs red-marked; male clypeus yellow-marked in one species. Head moderately broad, 1.2 times broader than long. Gena broad, exceeding width of compound eye; ocelloccipital distance extremely long, at least 3 times diameter of lateral ocellus. Facial fovea variable, from narrow to occupying entirety of distance between lateral ocellus and inner margin of compound eye. Female scutum, scutellum, and metanotum with pubescence variable, in some species with extremely short squamous hairs, hairs longer and non-squamous in other species. Pronotum laterally with humeral angle. Mesepisternum and dorsolateral parts of propodeum densely and clearly punctate, punctures confluent to separated by <0.5 puncture diameters. Propodeal triangle without lateral carinae, internal surface with dense network of irregularly raised rugosity, thus contrasting punctate dorsolateral surface. Forewing with nervulus interstitial. Hind tibial spurs simple, not broadened basally or medially. Terga typically densely and finely punctate, punctures separated by 1 puncture diameter, at least on T2-5. Male genital capsule simple, compact, with gonocoxae apically truncate to produced into weak rounded teeth. Gonostyli apically spatulate, penis valves more or less narrow, parallel-sided, occupying less than
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<normalizedToken id="233033D8DF88701FB539D64677C041B5" originalValue="½">1/2</normalizedToken>
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space between gonostyli.
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</paragraph>
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</subSubSection>
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<subSubSection id="41ADA2702010B6B4100713ABD1AAC1D2" pageId="0" pageNumber="241" type="etymology">
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<paragraph id="B7909DC6A938D139126DEE74E82ED679" pageId="0" pageNumber="241">Etymology.</paragraph>
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<paragraph id="61279A3655414FD0F94767ECD2B7C6F5" pageId="0" pageNumber="241">
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The name is taken from the type species for the subgenus,
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<taxonomicName id="C9D53BD999DA6D3A14268EAD473BE92A" class="Insecta" family="Plantaginaceae" kingdom="Animalia" lsidName="A. pruinosa" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="pruinosa">
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<emphasis id="D546D8A13203FA6C6201456E6DBCE39C" italics="true" pageId="0" pageNumber="241">A. pruinosa</emphasis>
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</taxonomicName>
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. The Latin word
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<taxonomicName id="CA424C720752526AD0379AEF872C2C22" authorityName="Erichson" authorityYear="1835" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena pruinosa" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="pruinosa">
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<emphasis id="3908918E67C50C0BEA962E32EE557FDA" italics="true" pageId="0" pageNumber="241">Andrena pruinosa</emphasis>
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</taxonomicName>
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is the feminine singular of
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<emphasis id="1E916F286F3FA5837C41CE2B3C32CEE8" italics="true" pageId="0" pageNumber="241">pruinosus</emphasis>
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which means
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<normalizedToken id="E75E0F611F4DE86CF4D7EE98209B4D92" originalValue="‘frosty’">'frosty'</normalizedToken>
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, in reference to the squamous hairs of the mesosoma. The gender is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="EE361A39EA87D61661952FE77E495D61" pageId="0" pageNumber="241" type="included species">
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<paragraph id="96E97A0FD3E94ABBA22179C86C33C1DD" pageId="0" pageNumber="241">Included species.</paragraph>
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<paragraph id="F3F988DA48C6DD5A4CC9099FC1FE71E6" pageId="0" pageNumber="241">
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<taxonomicName id="2EF1217E24796B04A7E38154260F6D6D" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena caroli" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="caroli">
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<emphasis id="D85735AD879B0B05E5DD7D7649277DD4" italics="true" pageId="0" pageNumber="241">Andrena caroli</emphasis>
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</taxonomicName>
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(Morocco to Israel);
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<taxonomicName id="4A805850E53651681700CF4914F522C4" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena nilotica" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="nilotica">
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<emphasis id="49552381CEB4C216ADED4A4A5E1A68A4" italics="true" pageId="0" pageNumber="241">Andrena nilotica</emphasis>
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</taxonomicName>
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Warncke, 1967 (Spain);
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<taxonomicName id="0B946EE15502265D69F9033657A8E6F3" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena parata" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="parata">
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<emphasis id="8C987660AFF4D3CF615066C355FD8631" italics="true" pageId="0" pageNumber="241">Andrena parata</emphasis>
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</taxonomicName>
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(Spain);
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<taxonomicName id="E2508CF87673B861B6F899F6135328B2" authorityName="Erichson" authorityYear="1835" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena pruinosa" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="pruinosa">
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<emphasis id="CDD3CD826C3DB41422EA43A881D7C902" italics="true" pageId="0" pageNumber="241">Andrena pruinosa</emphasis>
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</taxonomicName>
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(Spain);
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<taxonomicName id="4E1A0C52C195A0CC643B5AD370DBD300" authorityName="Wood" authorityYear="2020" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena sparsipunctata" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="sparsipunctata">
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<emphasis id="D2F3E8C73AC54DCCFC60D83AB5C37521" italics="true" pageId="0" pageNumber="241">Andrena sparsipunctata</emphasis>
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</taxonomicName>
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Wood, 2020 (Morocco);
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<taxonomicName id="9871B016F810224E7E015B945127593A" authorityName="Dours" authorityYear="1872" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena succinea" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="succinea">
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<emphasis id="B6AF44E482FBB4206FC496F8FEF36AFC" italics="true" pageId="0" pageNumber="241">Andrena succinea</emphasis>
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</taxonomicName>
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(Morocco to Iran and Saudi Arabia). The centre of diversity is therefore Spain + Morocco, with all six species occurring here.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |