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<mods:title id="D3F97E8B3DE7D6B172EF8561F71B0018">New species of the Microserica lineata group from Laos and Thailand (Coleoptera Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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Key to the species of the 
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<emphasis id="B911EAE4004CFFA6969DFB9398CE1458" bold="true" box="[459,694,1086,1113]" italics="true" pageId="10" pageNumber="273">Microserica lineata</emphasis>
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group
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1 Phallobase short, about twice as long as wide in right side lateral view (
<figureCitation id="135E2A73004CFFA694C2FB2899801498" box="[916,1016,1156,1178]" captionStart="FIGURES 1" captionStartId="1.[151,264,1624,1648]" captionTargetBox="[208,1380,634,1590]" captionTargetId="figure-118@1.[202,1386,630,1597]" captionTargetPageId="1" captionText="FIGURES 1A–M. A–C, Microserica lineata Moser, 1915 (syntype: Myanmar: Pegu); D–F, Microserica squamulata (Moser, 1915) (lectotype: “Siam Hinlap”); G–I, Microserica crenatostriata Ahrens, 2004 (holotype); J–M Microserica geberbauer Ahrens, 2004 (holotype). A, D, G, J, aedeagus, left side lateral view; C, F, I, M, aedeagus, right side lateral view; B, E, H, K, parameres, dorsal view. Scale: 0.5 mm. Abbreviations: llpphb—left lateral process of phallobase; ms—dorsomedian emargination (sinuation) of phallobase; phb—phallobase; rpm—right paramere." figureDoi="http://doi.org/10.5281/zenodo.4413034" httpUri="https://zenodo.org/record/4413034/files/figure.png" pageId="10" pageNumber="273">Figs. 1F, I</figureCitation>
). Pronotum yellowish with one or two dark spots................................................................................................ 2
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- Phallobase long, more than three times longer than wide in right side lateral view (
<figureCitation id="135E2A73004CFFA694ADFB129E0A14D5" box="[1019,1138,1214,1236]" captionStart="FIGURES 1" captionStartId="1.[151,264,1624,1648]" captionTargetBox="[208,1380,634,1590]" captionTargetId="figure-118@1.[202,1386,630,1597]" captionTargetPageId="1" captionText="FIGURES 1A–M. A–C, Microserica lineata Moser, 1915 (syntype: Myanmar: Pegu); D–F, Microserica squamulata (Moser, 1915) (lectotype: “Siam Hinlap”); G–I, Microserica crenatostriata Ahrens, 2004 (holotype); J–M Microserica geberbauer Ahrens, 2004 (holotype). A, D, G, J, aedeagus, left side lateral view; C, F, I, M, aedeagus, right side lateral view; B, E, H, K, parameres, dorsal view. Scale: 0.5 mm. Abbreviations: llpphb—left lateral process of phallobase; ms—dorsomedian emargination (sinuation) of phallobase; phb—phallobase; rpm—right paramere." figureDoi="http://doi.org/10.5281/zenodo.4413034" httpUri="https://zenodo.org/record/4413034/files/figure.png" pageId="10" pageNumber="273">Figs. 1C, M</figureCitation>
). Pronotum uniformly brown ................................................................................................... 3
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2 Pronotum yellowish with one median dark spot. Right paramere long, its apex strongly widened apically (
<figureCitation id="135E2A73004CFFA693B1FB559F4A150E" box="[1255,1330,1273,1295]" captionStart="FIGURES 1" captionStartId="1.[151,264,1624,1648]" captionTargetBox="[208,1380,634,1590]" captionTargetId="figure-118@1.[202,1386,630,1597]" captionTargetPageId="1" captionText="FIGURES 1A–M. A–C, Microserica lineata Moser, 1915 (syntype: Myanmar: Pegu); D–F, Microserica squamulata (Moser, 1915) (lectotype: “Siam Hinlap”); G–I, Microserica crenatostriata Ahrens, 2004 (holotype); J–M Microserica geberbauer Ahrens, 2004 (holotype). A, D, G, J, aedeagus, left side lateral view; C, F, I, M, aedeagus, right side lateral view; B, E, H, K, parameres, dorsal view. Scale: 0.5 mm. Abbreviations: llpphb—left lateral process of phallobase; ms—dorsomedian emargination (sinuation) of phallobase; phb—phallobase; rpm—right paramere." figureDoi="http://doi.org/10.5281/zenodo.4413034" httpUri="https://zenodo.org/record/4413034/files/figure.png" pageId="10" pageNumber="273">Fig. 1B</figureCitation>
). Antennal club little longer than the remaining antennomeres combined................................ 
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<emphasis id="B911EAE4004CFFA693E0FABB9F63152D" bold="true" box="[1206,1307,1302,1324]" italics="true" pageId="10" pageNumber="273">M. lineata</emphasis>
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- Pronotum yellowish with two symmetric dark spots beside the dark midline. Right paramere short, its apex weakly widened apically (
<figureCitation id="135E2A73004CFFA6966DFAFD9BFF1566" box="[315,391,1361,1383]" captionStart="FIGURES 1" captionStartId="1.[151,264,1624,1648]" captionTargetBox="[208,1380,634,1590]" captionTargetId="figure-118@1.[202,1386,630,1597]" captionTargetPageId="1" captionText="FIGURES 1A–M. A–C, Microserica lineata Moser, 1915 (syntype: Myanmar: Pegu); D–F, Microserica squamulata (Moser, 1915) (lectotype: “Siam Hinlap”); G–I, Microserica crenatostriata Ahrens, 2004 (holotype); J–M Microserica geberbauer Ahrens, 2004 (holotype). A, D, G, J, aedeagus, left side lateral view; C, F, I, M, aedeagus, right side lateral view; B, E, H, K, parameres, dorsal view. Scale: 0.5 mm. Abbreviations: llpphb—left lateral process of phallobase; ms—dorsomedian emargination (sinuation) of phallobase; phb—phallobase; rpm—right paramere." figureDoi="http://doi.org/10.5281/zenodo.4413034" httpUri="https://zenodo.org/record/4413034/files/figure.png" pageId="10" pageNumber="273">Fig. 1K</figureCitation>
). Antennal club twice as long as the remaining antennomeres combined..... 
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<emphasis id="B911EAE4004CFFA693D6FAFD9F691567" bold="true" box="[1152,1297,1360,1382]" italics="true" pageId="10" pageNumber="273">M. geberbauer</emphasis>
Ahrens, 2004
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3 Left lateral process of phallobase strongly widened apically but straight (
<figureCitation id="135E2A73004CFFA694C8FAC299911585" box="[926,1001,1390,1412]" captionStart="FIGURES 1" captionStartId="1.[151,264,1624,1648]" captionTargetBox="[208,1380,634,1590]" captionTargetId="figure-118@1.[202,1386,630,1597]" captionTargetPageId="1" captionText="FIGURES 1A–M. A–C, Microserica lineata Moser, 1915 (syntype: Myanmar: Pegu); D–F, Microserica squamulata (Moser, 1915) (lectotype: “Siam Hinlap”); G–I, Microserica crenatostriata Ahrens, 2004 (holotype); J–M Microserica geberbauer Ahrens, 2004 (holotype). A, D, G, J, aedeagus, left side lateral view; C, F, I, M, aedeagus, right side lateral view; B, E, H, K, parameres, dorsal view. Scale: 0.5 mm. Abbreviations: llpphb—left lateral process of phallobase; ms—dorsomedian emargination (sinuation) of phallobase; phb—phallobase; rpm—right paramere." figureDoi="http://doi.org/10.5281/zenodo.4413034" httpUri="https://zenodo.org/record/4413034/files/figure.png" pageId="10" pageNumber="273">Fig. 1D</figureCitation>
)................................... 5
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<keyLead id="309495C3004CFFA697C1FA209FE415A0" box="[151,1436,1420,1442]" pageId="10" pageNumber="273">- Left lateral process of phallobase narrow and not widened apically...............................................4</keyLead>
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4 Left lateral process of phallobase strongly hooked ventrally at apex (
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). Body densely covered with small white setae. Body smaller, 
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. Anterior margin of metafemur with serrated line.............. 
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Ahrens, 2004
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- Left lateral process of phallobase evenly narrowed towards apex (
<figureCitation id="135E2A73004CFFA69437FA4899DD15FB" box="[865,933,1508,1530]" captionStart="FIGURES 3" captionStartId="6.[151,264,1909,1933]" captionTargetBox="[212,1376,186,1882]" captionTargetId="figure-27@6.[203,1384,181,1885]" captionTargetPageId="6" captionText="FIGURES 3A–L. A–D, Microserica banvaneue Bohacz &amp; Ahrens, new species (holotype); E–H, M. helferi Bohacz &amp; Ahrens, new species (holotype); I–L, M. tenasserimensis Bohacz &amp; Ahrens, new species (holotype). A, E, I, aedeagus, left side lateral view; C, G, K, aedeagus, right side lateral view; B, F, J, parameres, dorsal view; D, H, L, habitus (not to scale). Scale: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4413038" httpUri="https://zenodo.org/record/4413038/files/figure.png" pageId="10" pageNumber="273">Fig. 3I</figureCitation>
). Serrated line beside anterior margin of metafemur and beside dorsal margin of metatibia lacking...................... 
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<emphasis id="B911EAE4004CFFA694F5F9AD9E191617" bold="true" box="[931,1121,1537,1558]" italics="true" pageId="10" pageNumber="273">M. tenasserimensis</emphasis>
Bohacz &amp; Ahrens
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, 
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5 Ventral lamina of left side of phallobase strongly produced ventrally, phallobase results distally strongly widened (
<figureCitation id="135E2A73004CFFA6927BF9B29F021635" box="[1325,1402,1566,1588]" captionStart="FIGURES 2" captionStartId="3.[151,264,1661,1685]" captionTargetBox="[185,1403,281,1632]" captionTargetId="figure-60@3.[177,1411,273,1638]" captionTargetPageId="3" captionText="FIGURES 2A–F. A–D, Microserica ventrosa Bohacz &amp; Ahrens, new species (holotype); E–H, M. ginae Bohacz &amp; Ahrens, new species (holotype). A, E, aedeagus, left side lateral view; C, G, aedeagus, right side lateral view; B, F, parameres, dorsal view; D, H, habitus (not to scale). Scale: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4413036" httpUri="https://zenodo.org/record/4413036/files/figure.png" pageId="10" pageNumber="273">Fig. 2A</figureCitation>
).................................................................. 
<emphasis id="B911EAE4004CFFA694BDF9909FE51650" bold="true" box="[1003,1437,1595,1617]" pageId="10" pageNumber="273">
<taxonomicName id="4C654D75004CFFA694BDF9909F641651" authority="Bohacz &amp; Ahrens" authorityName="Bohacz &amp; Ahrens" authorityYear="2020" box="[1003,1308,1595,1617]" class="Insecta" family="Melolonthidae" genus="Microserica" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="ventrosa" status="sp. nov.">
<emphasis id="B911EAE4004CFFA694BDF9909E191650" bold="true" box="[1003,1121,1596,1617]" italics="true" pageId="10" pageNumber="273">M. ventrosa</emphasis>
Bohacz &amp; Ahrens
</taxonomicName>
, 
<taxonomicNameLabel id="A222579F004CFFA69270F9909FE51650" box="[1318,1437,1595,1617]" pageId="10" pageNumber="273" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BDA36F6004CFFA697C1F9F59FE4166E" blockId="10.[151,1437,1156,1911]" box="[151,1436,1625,1647]" pageId="10" pageNumber="273">
<keyLead id="309495C3004CFFA697C1F9F59FE4166E" box="[151,1436,1625,1647]" pageId="10" pageNumber="273">
- Ventral lamina of left side of phallobase not produced (
<figureCitation id="135E2A73004CFFA695A5F9F59947166E" box="[755,831,1625,1647]" captionStart="FIGURES 1" captionStartId="1.[151,264,1624,1648]" captionTargetBox="[208,1380,634,1590]" captionTargetId="figure-118@1.[202,1386,630,1597]" captionTargetPageId="1" captionText="FIGURES 1A–M. A–C, Microserica lineata Moser, 1915 (syntype: Myanmar: Pegu); D–F, Microserica squamulata (Moser, 1915) (lectotype: “Siam Hinlap”); G–I, Microserica crenatostriata Ahrens, 2004 (holotype); J–M Microserica geberbauer Ahrens, 2004 (holotype). A, D, G, J, aedeagus, left side lateral view; C, F, I, M, aedeagus, right side lateral view; B, E, H, K, parameres, dorsal view. Scale: 0.5 mm. Abbreviations: llpphb—left lateral process of phallobase; ms—dorsomedian emargination (sinuation) of phallobase; phb—phallobase; rpm—right paramere." figureDoi="http://doi.org/10.5281/zenodo.4413034" httpUri="https://zenodo.org/record/4413034/files/figure.png" pageId="10" pageNumber="273">Fig. 1D</figureCitation>
), phallobase distally not widened....................... 6
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<paragraph id="8BDA36F6004CFFA697C1F9DA9FE5168D" blockId="10.[151,1437,1156,1911]" box="[151,1437,1654,1676]" pageId="10" pageNumber="273">
<keyLead id="309495C3004CFFA697C1F9DA9FE5168D" box="[151,1437,1654,1676]" pageId="10" pageNumber="273">
6 Body densely covered with small white scales. Right paramere short and wide............… 
<taxonomicName id="4C654D75004CFFA69320F9DB9FE5168D" authority="(Moser, 1915)" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[1142,1437,1654,1676]" class="Insecta" family="Melolonthidae" genus="Microserica" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="squamulata">
<emphasis id="B911EAE4004CFFA69320F9DB9FE5168D" bold="true" box="[1142,1437,1654,1676]" pageId="10" pageNumber="273">
<emphasis id="B911EAE4004CFFA69320F9DB9F73168D" bold="true" box="[1142,1291,1654,1676]" italics="true" pageId="10" pageNumber="273">M. squamulata</emphasis>
(Moser, 1915)
</emphasis>
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</paragraph>
<paragraph id="8BDA36F6004CFFA697C1F9389FE416A8" blockId="10.[151,1437,1156,1911]" box="[151,1436,1684,1706]" pageId="10" pageNumber="273">
<keyLead id="309495C3004CFFA697C1F9389FE416A8" box="[151,1436,1684,1706]" pageId="10" pageNumber="273">- Body sparsely covered with small white scales, only with dense, white minute setae................................. 7</keyLead>
</paragraph>
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<paragraph id="8BDA36F6004CFFA697C1F91D9FE416C7" blockId="10.[151,1437,1156,1911]" box="[151,1436,1713,1735]" pageId="10" pageNumber="273">
<keyLead id="309495C3004CFFA697C1F91D9FE416C7" box="[151,1436,1713,1735]" pageId="10" pageNumber="273">7 Right paramere robust and elongate, width subequal to maximum phallobase width................................. 8</keyLead>
</paragraph>
<paragraph id="8BDA36F6004CFFA697C1F9639FE51700" blockId="10.[151,1437,1156,1911]" pageId="10" pageNumber="273">
<keyLead id="309495C3004CFFA697C1F9639FE51700" pageId="10" pageNumber="273">
- Right paramere narrow and curved, width much narrower than maximum phallobase width.............................................................................................. 
<emphasis id="B911EAE4004CFFA694A8F9409FE51700" bold="true" box="[1022,1437,1771,1793]" pageId="10" pageNumber="273">
<taxonomicName id="4C654D75004CFFA694A8F9409F641701" authority="Bohacz &amp; Ahrens" authorityName="Bohacz &amp; Ahrens" authorityYear="2020" box="[1022,1308,1771,1793]" class="Insecta" family="Melolonthidae" genus="Microserica" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="B911EAE4004CFFA694A8F9409E181700" bold="true" box="[1022,1120,1771,1793]" italics="true" pageId="10" pageNumber="273">M. helferi</emphasis>
Bohacz &amp; Ahrens
</taxonomicName>
, 
<taxonomicNameLabel id="A222579F004CFFA69270F9409FE51700" box="[1318,1437,1771,1793]" pageId="10" pageNumber="273" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8BDA36F6004CFFA697C1F8A59FE5173D" blockId="10.[151,1437,1156,1911]" pageId="10" pageNumber="273">
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8 Base of labroclypeus dull. Dorsal margin of right paramere evenly curved (lateral view)................................................................................................ 
<emphasis id="B911EAE4004CFFA6935FF88B9FE5173D" bold="true" box="[1033,1437,1830,1852]" pageId="10" pageNumber="273">
<taxonomicName id="4C654D75004CFFA6935FF88B9F64173A" authority="Bohacz &amp; Ahrens" authorityName="Bohacz &amp; Ahrens" authorityYear="2020" box="[1033,1308,1830,1852]" class="Insecta" family="Melolonthidae" genus="Microserica" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="ginae" status="sp. nov.">
<emphasis id="B911EAE4004CFFA6935FF88B9E19173A" bold="true" box="[1033,1121,1830,1852]" italics="true" pageId="10" pageNumber="273">M. ginae</emphasis>
Bohacz &amp; Ahrens
</taxonomicName>
, 
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</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BDA36F6004CFFA697C1F8E89FE51776" blockId="10.[151,1437,1156,1911]" pageId="10" pageNumber="273">
<keyLead id="309495C3004CFFA697C1F8E89FE51776" pageId="10" pageNumber="273">
- Base of labroclypeus shiny. Dorsal margin of right paramere strongly bent at middle and concave basally (lateral view).................................................................. .. 
<emphasis id="B911EAE4004CFFA69485F8CD9FE51776" bold="true" box="[979,1437,1888,1911]" pageId="10" pageNumber="273">
<taxonomicName id="4C654D75004CFFA69485F8CD9F641777" authority="Bohacz &amp; Ahrens" authorityName="Bohacz &amp; Ahrens" authorityYear="2020" box="[979,1308,1888,1911]" class="Insecta" family="Melolonthidae" genus="Microserica" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="banvaneue" status="sp. nov.">
<emphasis id="B911EAE4004CFFA69485F8CD9E191777" bold="true" box="[979,1121,1888,1910]" italics="true" pageId="10" pageNumber="273">M. banvaneue</emphasis>
Bohacz &amp; Ahrens
</taxonomicName>
, 
<taxonomicNameLabel id="A222579F004CFFA69270F8CD9FE51776" box="[1318,1437,1889,1911]" pageId="10" pageNumber="273" rank="species">new species</taxonomicNameLabel>
</emphasis>
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