treatments-xml/data/03/CC/87/03CC87D3FFAD9F3DFF59FEAACB11FDE4.xml

<document id="98182140BB4F460A16854A3697B672A9" ID-CLB-Dataset="31934" ID-DOI="10.11646/zootaxa.4324.3.5" ID-GBIF-Dataset="5216886e-cb75-4427-b6b1-95236ae79662" ID-ISSN="1175-5326" ID-Zenodo-Dep="998438" ID-ZooBank="2Fd1A391-F168-4F1F-9Fa9-17D93A00418B" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506581813147" checkinUser="plazi" docAuthor="Zhang, Lanni" docDate="2017" docId="03CC87D3FFAD9F3DFF59FEAACB11FDE4" docLanguage="en" docName="zootaxa.4324.3.5.pdf" docOrigin="Zootaxa 4324 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eotetranychus spinophilus Zhang 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="508" masterDocId="FFF5FFABFFAB9F2CFFCEFF8DCE74F966" masterDocTitle="A new Australian species of Eotetranychus (Acari: Tetranychidae) from buck spinifex Triodia mitchelli (Poaceae), intraspecific variation in Eotetranychus, and the synonymy of Platytetranychus with Eotetranychus" masterLastPageNumber="517" masterPageNumber="491" pageNumber="497" updateTime="1698469141324" updateUser="ExternalLinkService">
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<mods:title id="F8975352AC0BE1FBFEED2C49ACAC45DD">A new Australian species of Eotetranychus (Acari: Tetranychidae) from buck spinifex Triodia mitchelli (Poaceae), intraspecific variation in Eotetranychus, and the synonymy of Platytetranychus with Eotetranychus</mods:title>
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<subSubSection id="C37F654EFFAD9F2AFF59FEAACF5EF805" pageId="6" pageNumber="497" type="nomenclature">
<paragraph id="8BDA36C5FFAD9F2AFF59FEAACD0FF824" blockId="6.[151,891,295,355]" box="[151,891,295,322]" pageId="6" pageNumber="497">
<heading id="D09281A9FFAD9F2AFF59FEAACD0FF824" bold="true" box="[151,891,295,322]" fontSize="11" level="1" pageId="6" pageNumber="497" reason="1">
<emphasis id="B911EAD7FFAD9F2AFF59FEAACD0FF824" bold="true" box="[151,891,295,322]" pageId="6" pageNumber="497">
<taxonomicName id="4C654D46FFAD9F2AFF59FEAACD63F824" authority="Zhang, Beard &amp; Seeman" authorityName="Zhang" authorityYear="2017" box="[151,791,295,322]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="497" phylum="Arthropoda" rank="species" species="spinophilus" status="sp. nov.">
<emphasis id="B911EAD7FFAD9F2AFF59FEAACFAEF827" bold="true" box="[151,474,295,321]" italics="true" pageId="6" pageNumber="497">Eotetranychus spinophilus</emphasis>
Zhang, Beard &amp; Seeman
</taxonomicName>
<taxonomicNameLabel id="A22257ACFFAD9F2AFCD0FEA5CD0FF824" box="[798,891,296,322]" pageId="6" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36C5FFAD9F2AFF59FEC7CF5EF805" blockId="6.[151,891,295,355]" box="[151,298,330,355]" pageId="6" pageNumber="497">
<figureCitation id="135E2A40FFAD9F2AFF59FEC7CF5EF805" box="[151,298,330,355]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 7" captionStart-2="FIGURES 8 – 14" captionStart-3="FIGURES 15 – 19" captionStart-4="FIGURES 20 – 21" captionStart-5="FIGURES 22 – 28" captionStart-6="FIGURES 29 – 37" captionStartId-0="8.[151,250,1974,1996]" captionStartId-1="9.[151,264,1958,1981]" captionStartId-2="10.[151,264,1904,1927]" captionStartId-3="11.[151,264,1982,2005]" captionStartId-4="12.[151,264,1653,1676]" captionStartId-5="13.[151,264,1943,1966]" captionStartId-6="14.[151,264,1939,1962]" captionTargetBox-0="[183,1357,312,1918]" captionTargetBox-1="[257,1312,203,1917]" captionTargetBox-2="[162,1387,217,1873]" captionTargetBox-3="[220,1342,202,1928]" captionTargetBox-4="[173,1426,238,1624]" captionTargetBox-5="[161,1418,656,1917]" captionTargetBox-6="[253,1321,209,1908]" captionTargetId-0="figure@8.[151,1436,303,1950]" captionTargetId-1="figure@9.[230,1332,193,1940]" captionTargetId-2="figure@10.[151,1436,193,1884]" captionTargetId-3="figure@11.[205,1368,193,1940]" captionTargetId-4="figure@12.[151,1436,193,1633]" captionTargetId-5="figure@13.[151,1436,639,1922]" captionTargetId-6="figure@14.[234,1353,193,1935]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="13" captionTargetPageId-6="14" captionText-0="FIGURE 1. Eotetranychus spinophilus sp. nov., female, dorsal idiosoma." captionText-1="FIGURES 2 – 7. Eotetranychus spinophilus sp. nov. female. (2) mid-ventral region; (3) posterior ventral opisthogaster; (4 – 7) spermatheca variation." captionText-2="FIGURES 8 – 14. Eotetranychus spinophilus sp. nov. female. (8 – 10) peritreme, different specimens; (11) palp, ventral view; (12) palp, dorsal view; (13) leg I, right side, dorsal to abaxial view; (14) ambulacrum I. Scale bar for ambulacrum = 40 µm." captionText-3="FIGURES 15 – 19. Eotetranychus spinophilus sp. nov. female. (15) leg II, right side, dorsal view; (16) ambulacrum II; (17) leg III, right side, abaxial view; (18) ambulacrum III; (19) leg IV, right side, abaxial view." captionText-4="FIGURES 20 – 21. Eotetranychus spinophilus sp. nov. male. (20) dorsal idiosoma; (21) posterior ventral opisthogaster." captionText-5="FIGURES 22 – 28. Eotetranychus spinophilus sp. nov. male. (22) leg I, right side, dorsal to abaxial view; (23 – 24) empodium I, different specimens and views; (25) ambulacrum I; (26) leg II, dorsal to adaxial view; (27) empodium III; (28) ambulacrum III, different specimen. Scale bar for ambulacra = 20 µm." captionText-6="FIGURES 29 – 37. Eotetranychus spinophilus sp. nov. male. (29) leg III, left side, dorsal to adaxial view; (30) leg IV, right side, abaxial view; (31) palp, dorsal view; (32 – 34) aedeagus, different aspects of same specimen; (35 – 37) aedeagus, different aspects of another specimen. Scale bar for aedeagi = 100 µm." httpUri-0="https://zenodo.org/record/998440/files/figure.png" httpUri-1="https://zenodo.org/record/998442/files/figure.png" httpUri-2="https://zenodo.org/record/998444/files/figure.png" httpUri-3="https://zenodo.org/record/998446/files/figure.png" httpUri-4="https://zenodo.org/record/998448/files/figure.png" httpUri-5="https://zenodo.org/record/998450/files/figure.png" httpUri-6="https://zenodo.org/record/998452/files/figure.png" pageId="6" pageNumber="497">Figures 1–37</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37F654EFFAD9F2AFF59FE1FCCE9FA78" pageId="6" pageNumber="497" type="diagnosis">
<paragraph id="8BDA36C5FFAD9F2AFF59FE1FCCE9FA78" blockId="6.[151,1437,402,1951]" pageId="6" pageNumber="497">
<emphasis id="B911EAD7FFAD9F2AFF59FE1FCF65F8CD" bold="true" box="[151,273,402,427]" pageId="6" pageNumber="497">Diagnosis.</emphasis>
Both sexes: Peritreme anastomosing; dorsal setae weakly barbed (each seta with minute distal fork); dorsocentral setae (
<emphasis id="B911EAD7FFAD9F2AFEBDFE3ACFF3F8B5" box="[371,391,439,467]" italics="true" pageId="6" pageNumber="497">c1</emphasis>
, 
<emphasis id="B911EAD7FFAD9F2AFE58FE3BCFD8F8B5" box="[406,428,438,467]" italics="true" pageId="6" pageNumber="497">d1</emphasis>
, 
<emphasis id="B911EAD7FFAD9F2AFE75FE3ACFBBF8B5" box="[443,463,439,467]" italics="true" pageId="6" pageNumber="497">e1</emphasis>
, 
<emphasis id="B911EAD7FFAD9F2AFE10FE3ACF99F8B5" box="[478,493,439,467]" italics="true" pageId="6" pageNumber="497">f1</emphasis>
) generally relatively short: setae 
<emphasis id="B911EAD7FFAD9F2AFCA6FE3ACD09F8B5" box="[872,893,439,467]" italics="true" pageId="6" pageNumber="497">c1</emphasis>
much shorter than distance 
<emphasis id="B911EAD7FFAD9F2AFB73FE3ACA81F8B5" box="[1213,1269,438,467]" italics="true" pageId="6" pageNumber="497">c1–d1</emphasis>
; setae 
<emphasis id="B911EAD7FFAD9F2AFA8AFE3BCB2FF8B5" box="[1348,1371,438,467]" italics="true" pageId="6" pageNumber="497">d1</emphasis>
, 
<emphasis id="B911EAD7FFAD9F2AFAA7FE3ACB09F8B5" box="[1385,1405,439,467]" italics="true" pageId="6" pageNumber="497">e1</emphasis>
, 
<emphasis id="B911EAD7FFAD9F2AFA42FE3ACBE8F8B5" box="[1420,1436,439,467]" italics="true" pageId="6" pageNumber="497">f1</emphasis>
about as long as distance to setal bases in next row. Tibiae III–IV with longitudinal striae mesally and with proximal and distal rings of transverse striae; tarsi III–IV with entirely longitudinal striae. Female: dorsal opisthosoma with mostly transverse striae; with some weakly arched striae between 
<emphasis id="B911EAD7FFAD9F2AFB96FDABCAF1FB24" box="[1112,1157,550,578]" italics="true" pageId="6" pageNumber="497">f1–f1</emphasis>
; with longitudinal striae laterad 
<emphasis id="B911EAD7FFAD9F2AFF3CFDC0CF72FB0F" box="[242,262,589,617]" italics="true" pageId="6" pageNumber="497">c2</emphasis>
and 
<emphasis id="B911EAD7FFAD9F2AFE82FDC1CFF0FB0F" box="[332,388,588,617]" italics="true" pageId="6" pageNumber="497">d1–e1</emphasis>
. Venter with mostly transverse striae; pregenital striae transverse, weak; genital striae transverse to weakly arched. Leg setal counts: trochanters 1, 1, 1, 1; femora 10, 7, 2, 2; genua 5, 5, 3, 3; tibiae 9 (1), 5, 5, 5; tarsi 15 (3+3), 12 (2+3), 8 (1+0), 8 (1+0). Male: dorsal opisthosoma with mostly transverse striae; with band of more widely-spaced transverse striae posteriad 
<emphasis id="B911EAD7FFAD9F2AFCC1FD36CD51FBBE" box="[783,805,699,728]" italics="true" pageId="6" pageNumber="497">h1</emphasis>
; with longitudinal striae laterad 
<emphasis id="B911EAD7FFAD9F2AFB57FD31CAD9FBBE" box="[1177,1197,700,728]" italics="true" pageId="6" pageNumber="497">c2</emphasis>
and 
<emphasis id="B911EAD7FFAD9F2AFB28FD36CB6AFBBE" box="[1254,1310,699,728]" italics="true" pageId="6" pageNumber="497">d1–e1</emphasis>
. Aedeagus directed dorsally, strongly sigmoid, blunt-tipped; dorsal margin with distinct angle forming a weak yet distinct distal knob; ventral margin smoothly sigmoid.
</paragraph>
</subSubSection>
<subSubSection id="C37F654EFFAD9F2AFF09FCA4CCD6FAD7" pageId="6" pageNumber="497" type="materials_examined">
<paragraph id="8BDA36C5FFAD9F2AFF09FCA4CCD6FAD7" blockId="6.[151,1437,402,1951]" pageId="6" pageNumber="497">
<emphasis id="B911EAD7FFAD9F2AFF09FCA4CFC6FA24" bold="true" box="[199,434,809,834]" pageId="6" pageNumber="497">Material examined.</emphasis>
Holotype. ♀. Australia, ex. buck spinifex 
<taxonomicName id="4C654D46FFAD9F2AFC57FCA4CB50FA24" authority="Benth" authorityName="Benth" authorityYear="1878" box="[921,1316,809,834]" class="Liliopsida" family="Poaceae" genus="Triodia" kingdom="Animalia" order="Poales" pageId="6" pageNumber="497" phylum="Tracheophyta" rank="species" species="mitchellii">
<emphasis id="B911EAD7FFAD9F2AFC57FCA4CA2AFA24" box="[921,1118,809,834]" italics="true" pageId="6" pageNumber="497">Triodia mitchellii</emphasis>
Benth (Poaceae)
</taxonomicName>
, 
<quantity id="4C9D9B20FFAD9F2AFAFEFCA7CB0DFA24" box="[1328,1401,809,834]" metricMagnitude="4" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="497" unit="km" value="90.0">90 km</quantity>
W Moonie, on Moonie Highway, Queensland, 27°56′57″S 149°31′19″E, 
<date id="FFDB1005FFAD9F2AFC6FFCDDCA70FA0F" box="[929,1028,848,873]" pageId="6" pageNumber="497" value="2007-05-05">5.v.2007</date>
, J.J. Beard. Paratypes. 4 ♀♀, 5 ♂♂, 4 deutonymphs, data same as holotype. Holotype and 9 paratypes deposited in QM; 1 ♀, 1 ♂ paratype deposited in ANIC; 1 ♀, 1 ♂ paratype deposited in USNM.
</paragraph>
</subSubSection>
<subSubSection id="C37F654EFFAD9F3DFF09FC31CC12F8B5" lastPageId="17" lastPageNumber="508" pageId="6" pageNumber="497" type="description">
<paragraph id="8BDA36C5FFAD9F2AFF09FC31CD9EFD83" blockId="6.[151,1437,402,1951]" pageId="6" pageNumber="497">
<emphasis id="B911EAD7FFAD9F2AFF09FC31CC1EFAB2" bold="true" box="[199,618,956,981]" pageId="6" pageNumber="497">
Description. Adult female. 
<emphasis id="B911EAD7FFAD9F2AFDC8FC30CC1EFAB2" bold="true" box="[518,618,957,980]" italics="true" pageId="6" pageNumber="497">Dorsum.</emphasis>
</emphasis>
(
<figureCitation id="135E2A40FFAD9F2AFDB4FC31CCCDFAB3" box="[634,697,956,981]" captionStart="FIGURE 1" captionStartId="8.[151,250,1974,1996]" captionTargetBox="[183,1357,312,1918]" captionTargetId="figure@8.[151,1436,303,1950]" captionTargetPageId="8" captionText="FIGURE 1. Eotetranychus spinophilus sp. nov., female, dorsal idiosoma." httpUri="https://zenodo.org/record/998440/files/figure.png" pageId="6" pageNumber="497">Fig. 1</figureCitation>
) Body measurements: 
<emphasis id="B911EAD7FFAD9F2AFC73FC33CD81FABF" box="[957,1013,956,985]" italics="true" pageId="6" pageNumber="497">
<date id="FFDB1005FFAD9F2AFC73FC33CDA5FABF" box="[957,977,958,985]" pageId="6" pageNumber="497">v2</date>
–h1
</emphasis>
255–261, 
<emphasis id="B911EAD7FFAD9F2AFBA5FC33CAC3FABF" box="[1131,1207,958,985]" italics="true" pageId="6" pageNumber="497">sc2–sc2</emphasis>
145–170; idiosomal length 310–360, width 220–235. Distances between other setae: 
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<emphasis id="B911EAD7FFAD9F2AFC49FC69CDC9FD66" box="[903,957,996,1024]" italics="true" pageId="6" pageNumber="497">v2–v2</emphasis>
62
</date>
–72, 
<emphasis id="B911EAD7FFAD9F2AFBD1FC69CA1FFD66" box="[1055,1131,996,1024]" italics="true" pageId="6" pageNumber="497">sc1–sc1</emphasis>
85–92, 
<emphasis id="B911EAD7FFAD9F2AFB03FC69CB70FD66" box="[1229,1284,996,1024]" italics="true" pageId="6" pageNumber="497">c1–c1</emphasis>
53–58, 
<emphasis id="B911EAD7FFAD9F2AFAABFC69CBE8FD66" box="[1381,1436,996,1024]" italics="true" pageId="6" pageNumber="497">c3–c3</emphasis>
176–213, 
<emphasis id="B911EAD7FFAD9F2AFEC6FB87CF35FD41" box="[264,321,1034,1063]" italics="true" pageId="6" pageNumber="497">d1–d1</emphasis>
44–53, 
<emphasis id="B911EAD7FFAD9F2AFE50FB87CFBBFD45" box="[414,463,1034,1063]" italics="true" pageId="6" pageNumber="497">d1–d</emphasis>
<date id="FFDB1005FFAD9F2AFE01FB94CC5CFD44" box="[463,552,1034,1063]" pageId="6" pageNumber="497" value="1932-02-29">
<emphasis id="B911EAD7FFAD9F2AFE01FB94CFA3FD41" box="[463,471,1049,1063]" italics="true" pageId="6" pageNumber="497">2</emphasis>
29–32
</date>
, 
<emphasis id="B911EAD7FFAD9F2AFDFAFB86CC1EFD41" box="[564,618,1035,1063]" italics="true" pageId="6" pageNumber="497">
e1– 
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1
</emphasis>
53–59, 
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e1– 
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2
</emphasis>
32–34, 
<emphasis id="B911EAD7FFAD9F2AFC94FB87CD0AFD44" box="[858,894,1034,1063]" italics="true" pageId="6" pageNumber="497">f1–f</emphasis>
<date id="FFDB1005FFAD9F2AFCB1FB94CDA3FD44" box="[895,983,1034,1063]" pageId="6" pageNumber="497" value="1933-01-30">
<emphasis id="B911EAD7FFAD9F2AFCB1FB94CDF3FD41" box="[895,903,1049,1063]" italics="true" pageId="6" pageNumber="497">1</emphasis>
30–33
</date>
, 
<emphasis id="B911EAD7FFAD9F2AFC2DFB87CA60FD45" box="[995,1044,1034,1063]" italics="true" pageId="6" pageNumber="497">h1–h</emphasis>
<date id="FFDB1005FFAD9F2AFBDBFB94CA19FD44" box="[1045,1133,1034,1063]" pageId="6" pageNumber="497" value="1917-01-19">
<emphasis id="B911EAD7FFAD9F2AFBDBFB94CA69FD41" box="[1045,1053,1049,1063]" italics="true" pageId="6" pageNumber="497">1</emphasis>
17–19
</date>
; 
<emphasis id="B911EAD7FFAD9F2AFBB4FB86CAC6FD41" box="[1146,1202,1034,1063]" italics="true" pageId="6" pageNumber="497">c1–d1</emphasis>
45–51, 
<emphasis id="B911EAD7FFAD9F2AFAC0FB87CB4AFD44" box="[1294,1342,1034,1063]" italics="true" pageId="6" pageNumber="497">
d1– 
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</emphasis>
<date id="FFDB1005FFAD9F2AFAF0FB94CBE3FD44" box="[1342,1431,1034,1063]" pageId="6" pageNumber="497" value="1933-01-30">
<emphasis id="B911EAD7FFAD9F2AFAF0FB94CB32FD41" box="[1342,1350,1049,1063]" italics="true" pageId="6" pageNumber="497">1</emphasis>
30–33
</date>
; 
<emphasis id="B911EAD7FFAD9F2AFF59FBBFCEBDFD28" box="[151,201,1073,1102]" italics="true" pageId="6" pageNumber="497">e1–f1</emphasis>
40–45; 
<emphasis id="B911EAD7FFAD9F2AFEF5FBBCCF12FD2C" box="[315,358,1073,1102]" italics="true" pageId="6" pageNumber="497">f1–h</emphasis>
<date id="FFDB1005FFAD9F2AFEA9FBCDCFBEFD2F" box="[359,458,1073,1102]" pageId="6" pageNumber="497" value="1932-01-28">
<emphasis id="B911EAD7FFAD9F2AFEA9FBCDCF1BFD28" box="[359,367,1088,1102]" italics="true" pageId="6" pageNumber="497">1</emphasis>
28–32
</date>
. Prodorsal striae longitudinal; opisthosomal striae transverse medially between dorsocentral setae and arched between setae 
<emphasis id="B911EAD7FFAD9F2AFD66FBD5CCA1FD12" box="[680,725,1112,1140]" italics="true" pageId="6" pageNumber="497">f1–f1</emphasis>
, striae longitudinal to oblique laterally. Dorsal setae weakly barbed, with minute distal fork. Opisthosomal setae shorter than (
<emphasis id="B911EAD7FFAD9F2AFCBEFBF2CDF0FDFD" box="[880,900,1151,1179]" italics="true" pageId="6" pageNumber="497">c1</emphasis>
, 
<emphasis id="B911EAD7FFAD9F2AFC5FFBF3CDD3FDFD" box="[913,935,1150,1179]" italics="true" pageId="6" pageNumber="497">d1</emphasis>
), or about as long as (
<emphasis id="B911EAD7FFAD9F2AFB52FBF2CB45FDFD" box="[1180,1329,1150,1179]" italics="true" pageId="6" pageNumber="497">c2, d2, e1–2, f1–2</emphasis>
) distance to base of seta in next row. Setal lengths: 
<date id="FFDB1005FFAD9F2AFDA8FB2BCCD4FDD8" box="[614,672,1189,1218]" pageId="6" pageNumber="497" value="1942-05-02">
<emphasis id="B911EAD7FFAD9F2AFDA8FB2BCC0FFDA4" box="[614,635,1190,1218]" italics="true" pageId="6" pageNumber="497">v2</emphasis>
42
</date>
–51, 
<emphasis id="B911EAD7FFAD9F2AFD1BFB2BCC81FDA4" box="[725,757,1190,1218]" italics="true" pageId="6" pageNumber="497">sc1</emphasis>
45–56, 
<emphasis id="B911EAD7FFAD9F2AFC81FB2BCD1AFDA4" box="[847,878,1190,1218]" italics="true" pageId="6" pageNumber="497">sc2</emphasis>
45–56, 
<emphasis id="B911EAD7FFAD9F2AFC07FB2BCDA1FDDB" box="[969,981,1190,1213]" italics="true" pageId="6" pageNumber="497">c</emphasis>
<date id="FFDB1005FFAD9F2AFC1BFB39CA58FDD8" box="[981,1068,1190,1218]" pageId="6" pageNumber="497" value="1942-01-31">
<emphasis id="B911EAD7FFAD9F2AFC1BFB39CDA9FDA4" box="[981,989,1204,1218]" italics="true" pageId="6" pageNumber="497">1</emphasis>
31–42
</date>
, 
<emphasis id="B911EAD7FFAD9F2AFBF6FB2BCA38FDA4" box="[1080,1100,1190,1218]" italics="true" pageId="6" pageNumber="497">c2</emphasis>
48–58, 
<emphasis id="B911EAD7FFAD9F2AFB69FB2BCACFFDA4" box="[1191,1211,1190,1218]" italics="true" pageId="6" pageNumber="497">c3</emphasis>
53–64, 
<emphasis id="B911EAD7FFAD9F2AFAD8FB28CB58FDA4" box="[1302,1324,1189,1218]" italics="true" pageId="6" pageNumber="497">d1</emphasis>
33–42, 
<emphasis id="B911EAD7FFAD9F2AFA48FB28CBE8FDA4" box="[1414,1436,1189,1218]" italics="true" pageId="6" pageNumber="497">d2</emphasis>
43–54, 
<geoCoordinate id="EE515002FFAD9F2AFF25FB40CF55FD83" box="[235,289,1228,1257]" direction="east" orientation="longitude" pageId="6" pageNumber="497" precision="925" value="1.5833333">
<emphasis id="B911EAD7FFAD9F2AFF25FB40CE8BFD8F" box="[235,255,1229,1257]" italics="true" pageId="6" pageNumber="497">e1</emphasis>
35
</geoCoordinate>
–43, 
<geoCoordinate id="EE515002FFAD9F2AFE94FB40CFE6FD83" box="[346,402,1228,1257]" direction="east" orientation="longitude" pageId="6" pageNumber="497" precision="925" value="2.7">
<emphasis id="B911EAD7FFAD9F2AFE94FB40CF1AFD8F" box="[346,366,1229,1257]" italics="true" pageId="6" pageNumber="497">e2</emphasis>
42
</geoCoordinate>
–55, 
<emphasis id="B911EAD7FFAD9F2AFE07FB40CFADFD8F" box="[457,473,1229,1257]" italics="true" pageId="6" pageNumber="497">f1</emphasis>
36–51, 
<emphasis id="B911EAD7FFAD9F2AFDFAFB40CC37FD8F" box="[564,579,1229,1257]" italics="true" pageId="6" pageNumber="497">f2</emphasis>
44–56, 
<emphasis id="B911EAD7FFAD9F2AFD50FB41CCC0FD8F" box="[670,692,1228,1257]" italics="true" pageId="6" pageNumber="497">h1</emphasis>
46–55, 
<emphasis id="B911EAD7FFAD9F2AFCC0FB41CD50FD8F" box="[782,804,1228,1257]" italics="true" pageId="6" pageNumber="497">h2</emphasis>
33–38, 
<emphasis id="B911EAD7FFAD9F2AFCB1FB41CDE1FD8F" box="[895,917,1228,1257]" italics="true" pageId="6" pageNumber="497">h3</emphasis>
32–36.
</paragraph>
<paragraph id="8BDA36C5FFAD9F2AFF09FB7ECA56FC1E" blockId="6.[151,1437,402,1951]" pageId="6" pageNumber="497">
<emphasis id="B911EAD7FFAD9F2AFF09FB7ECF64FC6C" bold="true" box="[199,272,1267,1290]" italics="true" pageId="6" pageNumber="497">Venter</emphasis>
. (
<figureCitation id="135E2A40FFAD9F2AFEE8FB7ECFFDFC6D" box="[294,393,1267,1292]" captionStart="FIGURES 2 – 7" captionStartId="9.[151,264,1958,1981]" captionTargetBox="[257,1312,203,1917]" captionTargetId="figure@9.[230,1332,193,1940]" captionTargetPageId="9" captionText="FIGURES 2 – 7. Eotetranychus spinophilus sp. nov. female. (2) mid-ventral region; (3) posterior ventral opisthogaster; (4 – 7) spermatheca variation." httpUri="https://zenodo.org/record/998442/files/figure.png" pageId="6" pageNumber="497">Figs 2–3</figureCitation>
) Striae almost entirely transverse, without lobes; coxae with fine transverse striae; pregenital striae weak, transverse; striae on genital flap transverse to weakly arched, fine; genital region flanked by oblique to longitudinal striae; all ventral setae thin, smooth: 
<emphasis id="B911EAD7FFAD9F2AFD71FAB6CCAFFC35" box="[703,731,1339,1363]" italics="true" pageId="6" pageNumber="497">1a</emphasis>
19–29, 
<emphasis id="B911EAD7FFAD9F2AFCFBFAB6CD25FC32" box="[821,849,1339,1364]" italics="true" pageId="6" pageNumber="497">1b</emphasis>
24–30, 
<emphasis id="B911EAD7FFAD9F2AFC65FAB6CDB2FC35" box="[939,966,1339,1363]" italics="true" pageId="6" pageNumber="497">1c</emphasis>
37–46, 
<emphasis id="B911EAD7FFAD9F2AFBD1FAB6CA4FFC32" box="[1055,1083,1339,1364]" italics="true" pageId="6" pageNumber="497">2b</emphasis>
28–34, 
<emphasis id="B911EAD7FFAD9F2AFB5BFAB6CADBFC35" box="[1173,1199,1339,1363]" italics="true" pageId="6" pageNumber="497">2c</emphasis>
44–55, 
<emphasis id="B911EAD7FFAD9F2AFAC4FAB6CB52FC35" box="[1290,1318,1339,1363]" italics="true" pageId="6" pageNumber="497">3a</emphasis>
13–19, 
<emphasis id="B911EAD7FFAD9F2AFA4EFAB6CBEFFC32" box="[1408,1435,1339,1364]" italics="true" pageId="6" pageNumber="497">3b</emphasis>
29–34, 
<emphasis id="B911EAD7FFAD9F2AFF25FAD2CF72FC11" box="[235,262,1375,1399]" italics="true" pageId="6" pageNumber="497">4a</emphasis>
19–22, 
<emphasis id="B911EAD7FFAD9F2AFEACFAD2CF0AFC1E" box="[354,382,1375,1400]" italics="true" pageId="6" pageNumber="497">4b</emphasis>
28–33, 
<emphasis id="B911EAD7FFAD9F2AFE17FAEDCF81FC11" box="[473,501,1376,1399]" italics="true" pageId="6" pageNumber="497">ag</emphasis>
23–26, 
<emphasis id="B911EAD7FFAD9F2AFD9EFAEDCC2AFC11" box="[592,606,1376,1399]" italics="true" pageId="6" pageNumber="497">g</emphasis>
<date id="FFDB1005FFAD9F2AFD90FAE3CCC1FC1E" box="[606,693,1375,1404]" pageId="6" pageNumber="497" value="1924-01-26">
<emphasis id="B911EAD7FFAD9F2AFD90FAE3CC12FC1A" box="[606,614,1390,1404]" italics="true" pageId="6" pageNumber="497">1</emphasis>
24–26
</date>
, 
<emphasis id="B911EAD7FFAD9F2AFD0EFAEDCCBAFC11" box="[704,718,1376,1399]" italics="true" pageId="6" pageNumber="497">g</emphasis>
<date id="FFDB1005FFAD9F2AFD00FAE3CD51FC11" box="[718,805,1375,1404]" pageId="6" pageNumber="497" value="1927-02-31">
<emphasis id="B911EAD7FFAD9F2AFD00FAE3CCA2FC1A" box="[718,726,1390,1404]" italics="true" pageId="6" pageNumber="497">2</emphasis>
27–31
</date>
, 
<emphasis id="B911EAD7FFAD9F2AFCFFFAEDCD3EFC11" box="[817,842,1376,1399]" italics="true" pageId="6" pageNumber="497">ps</emphasis>
<date id="FFDB1005FFAD9F2AFC84FAE3CDD5FC1E" box="[842,929,1375,1404]" pageId="6" pageNumber="497" value="2016-01-21">
<emphasis id="B911EAD7FFAD9F2AFC84FAE3CD26FC1A" box="[842,850,1390,1404]" italics="true" pageId="6" pageNumber="497">1</emphasis>
16–21
</date>
, 
<emphasis id="B911EAD7FFAD9F2AFC62FAEDCDB1FC11" box="[940,965,1376,1399]" italics="true" pageId="6" pageNumber="497">ps</emphasis>
<date id="FFDB1005FFAD9F2AFC0BFAE3CA56FC1E" box="[965,1058,1375,1404]" pageId="6" pageNumber="497" value="2016-02-21">
<emphasis id="B911EAD7FFAD9F2AFC0BFAE3CDB9FC1A" box="[965,973,1390,1404]" italics="true" pageId="6" pageNumber="497">2</emphasis>
16–21.
</date>
</paragraph>
<paragraph id="8BDA36C5FFAD9F2AFF09FA08CAF4FCA4" blockId="6.[151,1437,402,1951]" pageId="6" pageNumber="497">
<taxonomicName id="4C654D46FFAD9F2AFF09FA08CF2FFCF8" authorityName="Krefft" authorityYear="1871" box="[199,347,1413,1438]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="6" pageNumber="497" phylum="Nematoda" rank="genus">
<emphasis id="B911EAD7FFAD9F2AFF09FA08CF2FFCF8" bold="true" box="[199,347,1413,1438]" italics="true" pageId="6" pageNumber="497">Gnathosoma</emphasis>
</taxonomicName>
. (
<figureCitation id="135E2A40FFAD9F2AFEBFFA08CF90FCF8" box="[369,484,1413,1438]" captionStart="FIGURES 8 – 14" captionStartId="10.[151,264,1904,1927]" captionTargetBox="[162,1387,217,1873]" captionTargetId="figure@10.[151,1436,193,1884]" captionTargetPageId="10" captionText="FIGURES 8 – 14. Eotetranychus spinophilus sp. nov. female. (8 – 10) peritreme, different specimens; (11) palp, ventral view; (12) palp, dorsal view; (13) leg I, right side, dorsal to abaxial view; (14) ambulacrum I. Scale bar for ambulacrum = 40 µm." httpUri="https://zenodo.org/record/998444/files/figure.png" pageId="6" pageNumber="497">Figs 8–12</figureCitation>
) Palp tarsus with short, broad spinneret (
<emphasis id="B911EAD7FFAD9F2AFC76FA08CDA8FCF8" box="[952,988,1413,1438]" italics="true" pageId="6" pageNumber="497">suζ</emphasis>
) (2 wide, 5 long), one solenidion (4–5) and two eupathidia (
<emphasis id="B911EAD7FFAD9F2AFEB2FA24CFE6FCA4" box="[380,402,1449,1474]" italics="true" pageId="6" pageNumber="497">Μl</emphasis>
′ 4–5, 
<emphasis id="B911EAD7FFAD9F2AFE19FA24CF8DFCA4" box="[471,505,1449,1474]" italics="true" pageId="6" pageNumber="497">Μl&quot;</emphasis>
5–6) (
<figureCitation id="135E2A40FFAD9F2AFD8CFA24CCB7FCA4" box="[578,707,1449,1474]" captionStart="FIGURES 8 – 14" captionStartId="10.[151,264,1904,1927]" captionTargetBox="[162,1387,217,1873]" captionTargetId="figure@10.[151,1436,193,1884]" captionTargetPageId="10" captionText="FIGURES 8 – 14. Eotetranychus spinophilus sp. nov. female. (8 – 10) peritreme, different specimens; (11) palp, ventral view; (12) palp, dorsal view; (13) leg I, right side, dorsal to abaxial view; (14) ambulacrum I. Scale bar for ambulacrum = 40 µm." httpUri="https://zenodo.org/record/998444/files/figure.png" pageId="6" pageNumber="497">Figs 11–12</figureCitation>
). Peritremes anastomosing (
<figureCitation id="135E2A40FFAD9F2AFC31FA24CA06FCA4" box="[1023,1138,1449,1474]" captionStart="FIGURES 8 – 14" captionStartId="10.[151,264,1904,1927]" captionTargetBox="[162,1387,217,1873]" captionTargetId="figure@10.[151,1436,193,1884]" captionTargetPageId="10" captionText="FIGURES 8 – 14. Eotetranychus spinophilus sp. nov. female. (8 – 10) peritreme, different specimens; (11) palp, ventral view; (12) palp, dorsal view; (13) leg I, right side, dorsal to abaxial view; (14) ambulacrum I. Scale bar for ambulacrum = 40 µm." httpUri="https://zenodo.org/record/998444/files/figure.png" pageId="6" pageNumber="497">Figs 8–10</figureCitation>
).
</paragraph>
<paragraph id="8BDA36C5FFAD9F2AFF09FA40CD95FF6C" blockId="6.[151,1437,402,1951]" pageId="6" pageNumber="497">
<emphasis id="B911EAD7FFAD9F2AFF09FA40CF2AFC80" bold="true" box="[199,350,1485,1510]" italics="true" pageId="6" pageNumber="497">Spermatheca</emphasis>
. (
<figureCitation id="135E2A40FFAD9F2AFEB9FA40CFAAFC80" box="[375,478,1485,1510]" captionStart="FIGURES 2 – 7" captionStartId="9.[151,264,1958,1981]" captionTargetBox="[257,1312,203,1917]" captionTargetId="figure@9.[230,1332,193,1940]" captionTargetPageId="9" captionText="FIGURES 2 – 7. Eotetranychus spinophilus sp. nov. female. (2) mid-ventral region; (3) posterior ventral opisthogaster; (4 – 7) spermatheca variation." httpUri="https://zenodo.org/record/998442/files/figure.png" pageId="6" pageNumber="497">Figs 4–7</figureCitation>
) A short narrow tube ending in small rounded (dorsal aspect 
<figureCitation id="135E2A40FFAD9F2AFB7DFA40CB6CFC80" box="[1203,1304,1485,1510]" captionStart="FIGURES 2 – 7" captionStartId="9.[151,264,1958,1981]" captionTargetBox="[257,1312,203,1917]" captionTargetId="figure@9.[230,1332,193,1940]" captionTargetPageId="9" captionText="FIGURES 2 – 7. Eotetranychus spinophilus sp. nov. female. (2) mid-ventral region; (3) posterior ventral opisthogaster; (4 – 7) spermatheca variation." httpUri="https://zenodo.org/record/998442/files/figure.png" pageId="6" pageNumber="497">Figs 4–5</figureCitation>
) to weakly reniform (lateral aspect 
<figureCitation id="135E2A40FFAD9F2AFE6DFA7CCC72FF6C" box="[419,518,1521,1546]" captionStart="FIGURES 2 – 7" captionStartId="9.[151,264,1958,1981]" captionTargetBox="[257,1312,203,1917]" captionTargetId="figure@9.[230,1332,193,1940]" captionTargetPageId="9" captionText="FIGURES 2 – 7. Eotetranychus spinophilus sp. nov. female. (2) mid-ventral region; (3) posterior ventral opisthogaster; (4 – 7) spermatheca variation." httpUri="https://zenodo.org/record/998442/files/figure.png" pageId="6" pageNumber="497">Figs 6–7</figureCitation>
) bulb with honeycomb surface patterning.
</paragraph>
<paragraph id="8BDA36C5FFAD9F2AFF09F99BCD6BFEF8" blockId="6.[151,1437,402,1951]" pageId="6" pageNumber="497">
<emphasis id="B911EAD7FFAD9F2AFF09F99BCE89FF4B" bold="true" box="[199,253,1558,1581]" italics="true" pageId="6" pageNumber="497">Legs</emphasis>
. (
<figureCitation id="135E2A40FFAD9F2AFED4F998CFD4FF48" box="[282,416,1557,1582]" captionStart-0="FIGURES 8 – 14" captionStart-1="FIGURES 15 – 19" captionStartId-0="10.[151,264,1904,1927]" captionStartId-1="11.[151,264,1982,2005]" captionTargetBox-0="[162,1387,217,1873]" captionTargetBox-1="[220,1342,202,1928]" captionTargetId-0="figure@10.[151,1436,193,1884]" captionTargetId-1="figure@11.[205,1368,193,1940]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 8 – 14. Eotetranychus spinophilus sp. nov. female. (8 – 10) peritreme, different specimens; (11) palp, ventral view; (12) palp, dorsal view; (13) leg I, right side, dorsal to abaxial view; (14) ambulacrum I. Scale bar for ambulacrum = 40 µm." captionText-1="FIGURES 15 – 19. Eotetranychus spinophilus sp. nov. female. (15) leg II, right side, dorsal view; (16) ambulacrum II; (17) leg III, right side, abaxial view; (18) ambulacrum III; (19) leg IV, right side, abaxial view." httpUri-0="https://zenodo.org/record/998444/files/figure.png" httpUri-1="https://zenodo.org/record/998446/files/figure.png" pageId="6" pageNumber="497">Figs 13–19</figureCitation>
) Trochanters I–IV, femora I–IV, genua I–IV, tibiae I–II and proximal tarsi I–II with transverse striae; tibiae III–IV with mostly longitudinal striae except with proximal and distal rings of transverse striae; distal tarsi I–II with oblique–longitudinal striae; tarsi III–IV with longitudinal striae. Tarsus I (
<figureCitation id="135E2A40FFAD9F2AFAC7F9D0CB2EFF10" box="[1289,1370,1629,1654]" captionStart="FIGURES 8 – 14" captionStartId="10.[151,264,1904,1927]" captionTargetBox="[162,1387,217,1873]" captionTargetId="figure@10.[151,1436,193,1884]" captionTargetPageId="10" captionText="FIGURES 8 – 14. Eotetranychus spinophilus sp. nov. female. (8 – 10) peritreme, different specimens; (11) palp, ventral view; (12) palp, dorsal view; (13) leg I, right side, dorsal to abaxial view; (14) ambulacrum I. Scale bar for ambulacrum = 40 µm." httpUri="https://zenodo.org/record/998444/files/figure.png" pageId="6" pageNumber="497">Fig. 13</figureCitation>
) with two distal and adjacent duplex setae (solenidia: proximal 
<emphasis id="B911EAD7FFAD9F2AFCD3F90CCD43FFFC" box="[797,823,1665,1690]" italics="true" pageId="6" pageNumber="497">ω ′</emphasis>
31–33, distal 
<emphasis id="B911EAD7FFAD9F2AFC19F90FCD83FFFF" box="[983,1015,1666,1689]" italics="true" pageId="6" pageNumber="497">ω&quot;</emphasis>
42–46; companion fastigial setae: 
<emphasis id="B911EAD7FFAD9F2AFAB1F90FCBE0FFFC" box="[1407,1428,1666,1690]" italics="true" pageId="6" pageNumber="497">ft'</emphasis>
, 
<emphasis id="B911EAD7FFAD9F2AFF59F92BCEC7FFD8" box="[151,179,1702,1726]" italics="true" pageId="6" pageNumber="497">ft&quot;</emphasis>
8–10), single proximal solenidion (
<emphasis id="B911EAD7FFAD9F2AFD81F92BCC1BFFDB" box="[591,623,1702,1725]" italics="true" pageId="6" pageNumber="497">ω&quot;</emphasis>
<date id="FFDB1005FFAD9F2AFDA0F938CCBCFFD8" box="[622,712,1702,1731]" pageId="6" pageNumber="497" value="2014-12-01">
<emphasis id="B911EAD7FFAD9F2AFDA0F938CC02FFA5" box="[622,630,1717,1731]" italics="true" pageId="6" pageNumber="497">1</emphasis>
12–14
</date>
), and three distal eupathidia (
<emphasis id="B911EAD7FFAD9F2AFBD1F928CA4AFFD8" box="[1055,1086,1701,1726]" italics="true" pageId="6" pageNumber="497">p'ζ</emphasis>
, 
<emphasis id="B911EAD7FFAD9F2AFB81F928CA00FFD8" box="[1103,1140,1701,1726]" italics="true" pageId="6" pageNumber="497">p&quot;ζ</emphasis>
18–20; 
<emphasis id="B911EAD7FFAD9F2AFB1AF928CB74FFD8" box="[1236,1280,1701,1726]" italics="true" pageId="6" pageNumber="497">pv ′ζ</emphasis>
12–13); with three tactile setae and one solenidion (
<emphasis id="B911EAD7FFAD9F2AFD8AF940CC18FF8F" box="[580,620,1741,1769]" italics="true" pageId="6" pageNumber="497">ω&quot;1</emphasis>
) proximal to or overlapping with bases of proximal duplex setae. Tibia I with single short solenidion 
<emphasis id="B911EAD7FFAD9F2AFE2DF979CF86FE6D" box="[483,498,1780,1803]" italics="true" pageId="6" pageNumber="497">φ</emphasis>
9–10. Tarsus II (
<figureCitation id="135E2A40FFAD9F2AFD71F97ECD67FE6A" box="[703,787,1779,1804]" captionStart="FIGURES 15 – 19" captionStartId="11.[151,264,1982,2005]" captionTargetBox="[220,1342,202,1928]" captionTargetId="figure@11.[205,1368,193,1940]" captionTargetPageId="11" captionText="FIGURES 15 – 19. Eotetranychus spinophilus sp. nov. female. (15) leg II, right side, dorsal view; (16) ambulacrum II; (17) leg III, right side, abaxial view; (18) ambulacrum III; (19) leg IV, right side, abaxial view." httpUri="https://zenodo.org/record/998446/files/figure.png" pageId="6" pageNumber="497">Fig. 15</figureCitation>
) with one duplex seta (solenidion 
<emphasis id="B911EAD7FFAD9F2AFB62F979CABFFE6D" box="[1196,1227,1780,1803]" italics="true" pageId="6" pageNumber="497">ω&quot;</emphasis>
26–31), proximal solenidion (
<emphasis id="B911EAD7FFAD9F2AFED1F895CF4BFE49" box="[287,319,1816,1839]" italics="true" pageId="6" pageNumber="497">ω&quot;</emphasis>
<date id="FFDB1005FFAD9F2AFEF1F8ABCFFEFE56" box="[319,394,1816,1844]" pageId="6" pageNumber="497" value="2011-09-01">
<emphasis id="B911EAD7FFAD9F2AFEF1F8ABCF33FE52" box="[319,327,1830,1844]" italics="true" pageId="6" pageNumber="497">1</emphasis>
9–11
</date>
) and three distal eupathidia (
<emphasis id="B911EAD7FFAD9F2AFD2CF89ACD75FE56" box="[738,769,1815,1840]" italics="true" pageId="6" pageNumber="497">p'ζ</emphasis>
, 
<emphasis id="B911EAD7FFAD9F2AFCDDF89ACD43FE56" box="[787,823,1815,1840]" italics="true" pageId="6" pageNumber="497">p&quot;ζ</emphasis>
18–20; 
<emphasis id="B911EAD7FFAD9F2AFC55F89ACDB3FE56" box="[923,967,1815,1840]" italics="true" pageId="6" pageNumber="497">pv ′ζ</emphasis>
12–13). Tarsus III (
<figureCitation id="135E2A40FFAD9F2AFB7BF89ACB7DFE56" box="[1205,1289,1815,1840]" captionStart="FIGURES 15 – 19" captionStartId="11.[151,264,1982,2005]" captionTargetBox="[220,1342,202,1928]" captionTargetId="figure@11.[205,1368,193,1940]" captionTargetPageId="11" captionText="FIGURES 15 – 19. Eotetranychus spinophilus sp. nov. female. (15) leg II, right side, dorsal view; (16) ambulacrum II; (17) leg III, right side, abaxial view; (18) ambulacrum III; (19) leg IV, right side, abaxial view." httpUri="https://zenodo.org/record/998446/files/figure.png" pageId="6" pageNumber="497">Fig. 17</figureCitation>
) with single proximal solenidion (
<emphasis id="B911EAD7FFAD9F2AFE49F8B3CFD5FE31" box="[391,417,1854,1879]" italics="true" pageId="6" pageNumber="497">ω ′</emphasis>
14–24). Tarsus IV (
<figureCitation id="135E2A40FFAD9F2AFDB1F8B3CCA4FE31" box="[639,720,1854,1879]" captionStart="FIGURES 15 – 19" captionStartId="11.[151,264,1982,2005]" captionTargetBox="[220,1342,202,1928]" captionTargetId="figure@11.[205,1368,193,1940]" captionTargetPageId="11" captionText="FIGURES 15 – 19. Eotetranychus spinophilus sp. nov. female. (15) leg II, right side, dorsal view; (16) ambulacrum II; (17) leg III, right side, abaxial view; (18) ambulacrum III; (19) leg IV, right side, abaxial view." httpUri="https://zenodo.org/record/998446/files/figure.png" pageId="6" pageNumber="497">Fig. 19</figureCitation>
) with single proximal solenidion (
<emphasis id="B911EAD7FFAD9F2AFB9FF8B3CA1EFE31" box="[1105,1130,1854,1879]" italics="true" pageId="6" pageNumber="497">ω ′</emphasis>
17–24). Empodia split into two sets of three proximoventral hairs, with proximal pair of hairs thicker than other two pairs (
<figureCitation id="135E2A40FFAD9F2AFB75F8EFCB65FE1D" box="[1211,1297,1890,1915]" captionStart="FIGURES 8 – 14" captionStartId="10.[151,264,1904,1927]" captionTargetBox="[162,1387,217,1873]" captionTargetId="figure@10.[151,1436,193,1884]" captionTargetPageId="10" captionText="FIGURES 8 – 14. Eotetranychus spinophilus sp. nov. female. (8 – 10) peritreme, different specimens; (11) palp, ventral view; (12) palp, dorsal view; (13) leg I, right side, dorsal to abaxial view; (14) ambulacrum I. Scale bar for ambulacrum = 40 µm." httpUri="https://zenodo.org/record/998444/files/figure.png" pageId="6" pageNumber="497">Figs 14</figureCitation>
, 
<figureCitation id="135E2A40FFAD9F2AFAD2F8EFCB11FE1D" box="[1308,1381,1890,1915]" captionStart="FIGURES 15 – 19" captionStartId="11.[151,264,1982,2005]" captionTargetBox="[220,1342,202,1928]" captionTargetId="figure@11.[205,1368,193,1940]" captionTargetPageId="11" captionText="FIGURES 15 – 19. Eotetranychus spinophilus sp. nov. female. (15) leg II, right side, dorsal view; (16) ambulacrum II; (17) leg III, right side, abaxial view; (18) ambulacrum III; (19) leg IV, right side, abaxial view." httpUri="https://zenodo.org/record/998446/files/figure.png" pageId="6" pageNumber="497">16, 18</figureCitation>
). All leg setae finely barbed. Number of phaneres on legs I–IV:
</paragraph>
<paragraph id="8BDA36C5FFAC9F2BFE9EFF41CB65F8E1" pageId="7" pageNumber="498">
<table id="F965C465FFAC60D3FF51FF41CB69F8E1" box="[159,1309,204,391]" gridcols="6" gridrows="5" pageId="7" pageNumber="498">
<tr id="35553487FFAC60D3FF51FF41CB69F984" box="[159,1309,204,226]" gridrow="0" pageId="7" pageNumber="498" rowspan-0="1">
<th id="76845DFBFFAC60D3FE9EFF41CFCAF984" box="[336,446,204,226]" gridcol="1" gridrow="0" pageId="7" pageNumber="498">Trochanter</th>
<th id="76845DFBFFAC60D3FDE3FF41CC1BF984" box="[557,623,204,226]" gridcol="2" gridrow="0" pageId="7" pageNumber="498">Femur</th>
<th id="76845DFBFFAC60D3FCC5FF41CD36F984" box="[779,834,204,226]" gridcol="3" gridrow="0" pageId="7" pageNumber="498">Genu</th>
<th id="76845DFBFFAC60D3FC26FF41CA68F984" box="[1000,1052,204,226]" gridcol="4" gridrow="0" pageId="7" pageNumber="498">Tibia</th>
<th id="76845DFBFFAC60D3FB0BFF41CB69F984" box="[1221,1309,204,226]" gridcol="5" gridrow="0" pageId="7" pageNumber="498">Tarsus</th>
</tr>
<tr id="35553487FFAC60D3FF51FF78CB69F86D" box="[159,1309,245,267]" gridrow="1" pageId="7" pageNumber="498">
<th id="76845DFBFFAC60D3FF51FF78CE93F86D" box="[159,231,245,267]" gridcol="0" gridrow="1" pageId="7" pageNumber="498">Leg I</th>
<td id="76845DFBFFAC60D3FE9EFF78CFCAF86D" box="[336,446,245,267]" gridcol="1" gridrow="1" pageId="7" pageNumber="498">1</td>
<td id="76845DFBFFAC60D3FDE3FF78CC1BF86D" box="[557,623,245,267]" gridcol="2" gridrow="1" pageId="7" pageNumber="498">10</td>
<td id="76845DFBFFAC60D3FCC5FF78CD36F86D" box="[779,834,245,267]" gridcol="3" gridrow="1" pageId="7" pageNumber="498">5</td>
<td id="76845DFBFFAC60D3FC26FF78CA68F86D" box="[1000,1052,245,267]" gridcol="4" gridrow="1" pageId="7" pageNumber="498">9 (1)</td>
<td id="76845DFBFFAC60D3FB0BFF78CB69F86D" box="[1221,1309,245,267]" gridcol="5" gridrow="1" pageId="7" pageNumber="498">15 (3+3)</td>
</tr>
<tr id="35553487FFAC60D3FF51FE93CB69F852" box="[159,1309,286,308]" gridrow="2" pageId="7" pageNumber="498">
<th id="76845DFBFFAC60D3FF51FE93CE93F852" box="[159,231,286,308]" gridcol="0" gridrow="2" pageId="7" pageNumber="498">LegII</th>
<td id="76845DFBFFAC60D3FE9EFE93CFCAF852" box="[336,446,286,308]" gridcol="1" gridrow="2" pageId="7" pageNumber="498">1</td>
<td id="76845DFBFFAC60D3FDE3FE93CC1BF852" box="[557,623,286,308]" gridcol="2" gridrow="2" pageId="7" pageNumber="498">7</td>
<td id="76845DFBFFAC60D3FCC5FE93CD36F852" box="[779,834,286,308]" gridcol="3" gridrow="2" pageId="7" pageNumber="498">5</td>
<td id="76845DFBFFAC60D3FC26FE93CA68F852" box="[1000,1052,286,308]" gridcol="4" gridrow="2" pageId="7" pageNumber="498">5</td>
<td id="76845DFBFFAC60D3FB0BFE93CB69F852" box="[1221,1309,286,308]" gridcol="5" gridrow="2" pageId="7" pageNumber="498">12(2+3)</td>
</tr>
<tr id="35553487FFAC60D3FF51FEC5CB69F838" box="[159,1309,328,350]" gridrow="3" pageId="7" pageNumber="498">
<th id="76845DFBFFAC60D3FF51FEC5CE93F838" box="[159,231,328,350]" gridcol="0" gridrow="3" pageId="7" pageNumber="498">LegIII</th>
<td id="76845DFBFFAC60D3FE9EFEC5CFCAF838" box="[336,446,328,350]" gridcol="1" gridrow="3" pageId="7" pageNumber="498">1</td>
<td id="76845DFBFFAC60D3FDE3FEC5CC1BF838" box="[557,623,328,350]" gridcol="2" gridrow="3" pageId="7" pageNumber="498">2</td>
<td id="76845DFBFFAC60D3FCC5FEC5CD36F838" box="[779,834,328,350]" gridcol="3" gridrow="3" pageId="7" pageNumber="498">3</td>
<td id="76845DFBFFAC60D3FC26FEC5CA68F838" box="[1000,1052,328,350]" gridcol="4" gridrow="3" pageId="7" pageNumber="498">5</td>
<td id="76845DFBFFAC60D3FB0BFEC5CB69F838" box="[1221,1309,328,350]" gridcol="5" gridrow="3" pageId="7" pageNumber="498">8(1+0)</td>
</tr>
<tr id="35553487FFAC60D3FF51FEFCCB69F8E1" box="[159,1309,369,391]" gridrow="4" pageId="7" pageNumber="498">
<th id="76845DFBFFAC60D3FF51FEFCCE93F8E1" box="[159,231,369,391]" gridcol="0" gridrow="4" pageId="7" pageNumber="498">LegIV</th>
<td id="76845DFBFFAC60D3FE9EFEFCCFCAF8E1" box="[336,446,369,391]" gridcol="1" gridrow="4" pageId="7" pageNumber="498">1</td>
<td id="76845DFBFFAC60D3FDE3FEFCCC1BF8E1" box="[557,623,369,391]" gridcol="2" gridrow="4" pageId="7" pageNumber="498">2</td>
<td id="76845DFBFFAC60D3FCC5FEFCCD36F8E1" box="[779,834,369,391]" gridcol="3" gridrow="4" pageId="7" pageNumber="498">3</td>
<td id="76845DFBFFAC60D3FC26FEFCCA68F8E1" box="[1000,1052,369,391]" gridcol="4" gridrow="4" pageId="7" pageNumber="498">5</td>
<td id="76845DFBFFAC60D3FB0BFEFCCB69F8E1" box="[1221,1309,369,391]" gridcol="5" gridrow="4" pageId="7" pageNumber="498">8(1+0)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDA36C5FFAC9F2BFF09FE45CFA0FB4C" blockId="7.[151,1436,455,2037]" pageId="7" pageNumber="498">
Leg setation as shown in 
<figureCitation id="135E2A40FFAC9F2BFE23FE4ACC1CF886" box="[493,616,455,480]" captionStart="FIGURES 8 – 14" captionStartId="10.[151,264,1904,1927]" captionTargetBox="[162,1387,217,1873]" captionTargetId="figure@10.[151,1436,193,1884]" captionTargetPageId="10" captionText="FIGURES 8 – 14. Eotetranychus spinophilus sp. nov. female. (8 – 10) peritreme, different specimens; (11) palp, ventral view; (12) palp, dorsal view; (13) leg I, right side, dorsal to abaxial view; (14) ambulacrum I. Scale bar for ambulacrum = 40 µm." httpUri="https://zenodo.org/record/998444/files/figure.png" pageId="7" pageNumber="498">Figures 13</figureCitation>
, 
<figureCitation id="135E2A40FFAC9F2BFDB8FE4ACC99F886" box="[630,749,455,480]" captionStart="FIGURES 15 – 19" captionStartId="11.[151,264,1982,2005]" captionTargetBox="[220,1342,202,1928]" captionTargetId="figure@11.[205,1368,193,1940]" captionTargetPageId="11" captionText="FIGURES 15 – 19. Eotetranychus spinophilus sp. nov. female. (15) leg II, right side, dorsal view; (16) ambulacrum II; (17) leg III, right side, abaxial view; (18) ambulacrum III; (19) leg IV, right side, abaxial view." httpUri="https://zenodo.org/record/998446/files/figure.png" pageId="7" pageNumber="498">15, 17, 19</figureCitation>
. Anomalies: one specimen with asymmetrical loss of tibia I seta 
<emphasis id="B911EAD7FFAC9F2BFF02FE66CE96FB6E" box="[204,226,491,520]" italics="true" pageId="7" pageNumber="498">l ′1</emphasis>
, one specimen with asymmetrical addition of tibia I solenidion 
<emphasis id="B911EAD7FFAC9F2BFC09FE66CDA8FB62" box="[967,988,491,516]" italics="true" pageId="7" pageNumber="498">φ ′</emphasis>
, one specimen with duplication (bases touching) of tarsus I seta 
<emphasis id="B911EAD7FFAC9F2BFE7DFD9FCFB9FB49" box="[435,461,530,559]" italics="true" pageId="7" pageNumber="498">v ′1</emphasis>
.
</paragraph>
<paragraph id="8BDA36C5FFAC9F2BFF09FDB5CA53FF6D" blockId="7.[151,1436,455,2037]" pageId="7" pageNumber="498">
<emphasis id="B911EAD7FFAC9F2BFF09FDB5CC01FB37" bold="true" box="[199,629,568,593]" italics="true" pageId="7" pageNumber="498">Ontogenetic changes in leg setation</emphasis>
. In general, most of the setae that are suppressed on the legs of 
<taxonomicName id="4C654D46FFAC9F2BFA4AFDB7CF6CFB13" authorityName="Zhang" authorityYear="2017" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="498" phylum="Arthropoda" rank="species" species="spinophilus">
<emphasis id="B911EAD7FFAC9F2BFA4AFDB7CBE1FB37" box="[1412,1429,570,593]" italics="true" pageId="7" pageNumber="498">E</emphasis>
. 
<emphasis id="B911EAD7FFAC9F2BFF59FDD1CF6CFB13" box="[151,280,604,629]" italics="true" pageId="7" pageNumber="498">spinophilus</emphasis>
</taxonomicName>
are adult setae. Here, femur I with ten setae in the adult matches the standard tetranychid chaetotaxy, as outlined in 
<bibRefCitation id="EFF44B34FFAC9F2BFED2FD0DCF94FBFF" author="Lindquist" box="[284,480,640,665]" pageId="7" pageNumber="498" refString="Lindquist, E. E. (1985) External anatomy. In: Helle, W. &amp; Sabelis, M. W. (Eds.), Spider Mites: Their Biology, Natural Enemies and Control. Vol. 1 A. Elsevier Science Publishers B. V., Amsterdam, pp. 3 - 28." type="journal article" year="1985">Lindquist (1985)</bibRefCitation>
; however, femur II has seven setae, which has an extra seta present, 
<emphasis id="B911EAD7FFAC9F2BFACDFD0DCB63FBFF" box="[1283,1303,640,665]" italics="true" pageId="7" pageNumber="498">l&quot;</emphasis>
, that is not usually present on 
<taxonomicName id="4C654D46FFAC9F2BFEA0FD29CC78FBDB" box="[366,524,676,701]" class="Arachnida" family="Tetranychidae" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="498" phylum="Arthropoda" rank="subFamily" subFamily="Tetranychinae">Tetranychinae</taxonomicName>
spider mites. One to three of setae 
<emphasis id="B911EAD7FFAC9F2BFC6BFD29CDC7FBDB" box="[933,947,676,701]" italics="true" pageId="7" pageNumber="498">l'</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFC0CFD29CDA2FBDB" box="[962,982,676,701]" italics="true" pageId="7" pageNumber="498">l&quot;</emphasis>
and 
<emphasis id="B911EAD7FFAC9F2BFBC1FD28CA53FBDA" box="[1039,1063,677,700]" italics="true" pageId="7" pageNumber="498">v&quot;</emphasis>
are usually added to femur II of 
<taxonomicName id="4C654D46FFAC9F2BFF59FD4ACF67FB86" box="[151,275,711,736]" class="Arachnida" family="Tetranychidae" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="498" phylum="Arthropoda" rank="subFamily" subFamily="Bryobiinae">Bryobiinae</taxonomicName>
deutonymphs, but usually none are added in 
<taxonomicName id="4C654D46FFAC9F2BFCE8FD4ACDB0FB86" box="[806,964,711,736]" class="Arachnida" family="Tetranychidae" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="498" phylum="Arthropoda" rank="subFamily" subFamily="Tetranychinae">Tetranychinae</taxonomicName>
deutonymphs (
<bibRefCitation id="EFF44B34FFAC9F2BFBB8FD4ACB44FB86" author="Lindquist" box="[1142,1328,711,736]" pageId="7" pageNumber="498" refString="Lindquist, E. E. (1985) External anatomy. In: Helle, W. &amp; Sabelis, M. W. (Eds.), Spider Mites: Their Biology, Natural Enemies and Control. Vol. 1 A. Elsevier Science Publishers B. V., Amsterdam, pp. 3 - 28." type="journal article" year="1985">Lindquist, 1985</bibRefCitation>
). Parallel trends towards suppression of additional femoral setae on legs II–IV are found in other tetranychoid families, with no setae added in 
<taxonomicName id="4C654D46FFAC9F2BFEAEFC82CC77FA4E" authorityName="Baker &amp; Pritchard" authorityYear="1953" box="[352,515,783,808]" class="Arachnida" family="Linotetranidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="498" phylum="Arthropoda" rank="family">Linotetranidae</taxonomicName>
and 
<taxonomicName id="4C654D46FFAC9F2BFDF4FC82CCA8FA4E" authorityName="Berlese" authorityYear="1913" box="[570,732,783,808]" class="Arachnida" family="Tenuipalpidae" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="498" phylum="Arthropoda" rank="family">Tenuipalpidae</taxonomicName>
. However, 
<taxonomicName id="4C654D46FFAC9F2BFC94FC82CD87FA4E" authorityName="Baker &amp; Pritchard" authorityYear="1953" box="[858,1011,783,808]" class="Arachnida" family="Tuckerellidae" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="498" phylum="Arthropoda" rank="family">Tuckerellidae</taxonomicName>
is similar to the 
<taxonomicName id="4C654D46FFAC9F2BFB61FC82CB58FA4E" box="[1199,1324,783,808]" class="Arachnida" family="Tetranychidae" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="498" phylum="Arthropoda" rank="subFamily" subFamily="Bryobiinae">Bryobiinae</taxonomicName>
in adding 
<emphasis id="B911EAD7FFAC9F2BFF59FCBECED1FA2A" box="[151,165,819,844]" italics="true" pageId="7" pageNumber="498">l'</emphasis>
and 
<emphasis id="B911EAD7FFAC9F2BFF2EFCBECE80FA2A" box="[224,244,819,844]" italics="true" pageId="7" pageNumber="498">l&quot;</emphasis>
to femur II, but 
<emphasis id="B911EAD7FFAC9F2BFE76FCB9CFA5FA2D" box="[440,465,820,843]" italics="true" pageId="7" pageNumber="498">v&quot;</emphasis>
is suppressed. According to 
<bibRefCitation id="EFF44B34FFAC9F2BFCEEFCBECD95FA2A" author="Lindquist" box="[800,993,819,844]" pageId="7" pageNumber="498" refString="Lindquist, E. E. (1985) External anatomy. In: Helle, W. &amp; Sabelis, M. W. (Eds.), Spider Mites: Their Biology, Natural Enemies and Control. Vol. 1 A. Elsevier Science Publishers B. V., Amsterdam, pp. 3 - 28." type="journal article" year="1985">Lindquist (1985)</bibRefCitation>
, the standard larval and protonymphal setae on femur II are 
<emphasis id="B911EAD7FFAC9F2BFE42FCDBCFEEFA09" box="[396,410,854,879]" italics="true" pageId="7" pageNumber="498">d</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFE64FCDACFC8FA08" box="[426,444,855,878]" italics="true" pageId="7" pageNumber="498">v'</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFE02FCDBCF86FA09" box="[460,498,854,879]" italics="true" pageId="7" pageNumber="498">bv&quot;</emphasis>
, and no setae are added in the deutonymph. Therefore the deutonymph retains the larval complement on femur II, while three setae are added in the adult, 
<emphasis id="B911EAD7FFAC9F2BFC15FCF7CD85FAF1" box="[987,1009,890,919]" italics="true" pageId="7" pageNumber="498">l'1</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFBCFFCF7CA68FAF1" box="[1025,1052,890,919]" italics="true" pageId="7" pageNumber="498">l&quot;1</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFBE2FCF6CA38FAF1" box="[1068,1100,891,919]" italics="true" pageId="7" pageNumber="498">v&quot;1</emphasis>
. Here, the deutonymph of 
<taxonomicName id="4C654D46FFAC9F2BFA4AFCF6CF6CFADF" authorityName="Zhang" authorityYear="2017" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="498" phylum="Arthropoda" rank="species" species="spinophilus">
<emphasis id="B911EAD7FFAC9F2BFA4AFCF6CBE1FAF4" box="[1412,1429,891,914]" italics="true" pageId="7" pageNumber="498">E</emphasis>
. 
<emphasis id="B911EAD7FFAC9F2BFF59FC2DCF6CFADF" box="[151,280,928,953]" italics="true" pageId="7" pageNumber="498">spinophilus</emphasis>
</taxonomicName>
indeed has the standard larval complement of setae present, although we do not have a larva/ protonymph to unequivocally confirm it; and the standard three setae, along with the extra seta 
<emphasis id="B911EAD7FFAC9F2BFB09FC49CAAFFABB" box="[1223,1243,964,989]" italics="true" pageId="7" pageNumber="498">l&quot;</emphasis>
, are added in the adult. Femora III–IV retain the larval/protonymphal complement of two setae, 
<emphasis id="B911EAD7FFAC9F2BFBFDFC65CA35FD67" box="[1075,1089,1000,1025]" italics="true" pageId="7" pageNumber="498">d</emphasis>
and 
<emphasis id="B911EAD7FFAC9F2BFB4EFC64CAEBFD66" box="[1152,1183,1001,1024]" italics="true" pageId="7" pageNumber="498">ev'</emphasis>
, in all known stages, meaning that two adult setae are not expressed on femur III–IV, i.e. 
<emphasis id="B911EAD7FFAC9F2BFC75FB81CDA5FD4F" box="[955,977,1036,1065]" italics="true" pageId="7" pageNumber="498">l'1</emphasis>
and 
<emphasis id="B911EAD7FFAC9F2BFBC0FB80CA55FD42" box="[1038,1057,1037,1060]" italics="true" pageId="7" pageNumber="498">v'</emphasis>
. Genua I–II have standard adult chaetotaxy of five setae present (
<emphasis id="B911EAD7FFAC9F2BFDC0FBBFCC68FD2D" box="[526,540,1074,1099]" italics="true" pageId="7" pageNumber="498">d</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFDE2FBBFCC4EFD2D" box="[556,570,1074,1099]" italics="true" pageId="7" pageNumber="498">l'</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFD87FBBFCC28FD2D" box="[585,604,1074,1099]" italics="true" pageId="7" pageNumber="498">l&quot;</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFDA2FBBECC0BFD2C" box="[620,639,1075,1098]" italics="true" pageId="7" pageNumber="498">v'</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFD43FBBECCD1FD2C" box="[653,677,1075,1098]" italics="true" pageId="7" pageNumber="498">v&quot;</emphasis>
), with seta 
<emphasis id="B911EAD7FFAC9F2BFCE2FBBFCD4EFD2D" box="[812,826,1074,1099]" italics="true" pageId="7" pageNumber="498">d</emphasis>
assumed to be added in the deutonymph as would be standard, and the remaining setae being larval. Genu III with three setae (
<emphasis id="B911EAD7FFAC9F2BFC1FFBDBCDABFD09" box="[977,991,1110,1135]" italics="true" pageId="7" pageNumber="498">d</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFC23FBDBCD8FFD09" box="[1005,1019,1110,1135]" italics="true" pageId="7" pageNumber="498">l'</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFBC7FBDACA6FFD08" box="[1033,1051,1111,1134]" italics="true" pageId="7" pageNumber="498">v'</emphasis>
) is missing adult seta 
<emphasis id="B911EAD7FFAC9F2BFAD8FBDACB5BFD08" box="[1302,1327,1111,1134]" italics="true" pageId="7" pageNumber="498">v&quot;</emphasis>
(n.b. seta 
<emphasis id="B911EAD7FFAC9F2BFF59FBF7CED1FDF5" box="[151,165,1146,1171]" italics="true" pageId="7" pageNumber="498">d</emphasis>
is assumed to be added in the deutonymph, as is typical). Genu IV with three setae (
<emphasis id="B911EAD7FFAC9F2BFB93FBF7CA1FFDF5" box="[1117,1131,1146,1171]" italics="true" pageId="7" pageNumber="498">d</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFBB4FBF7CAFCFDF5" box="[1146,1160,1146,1171]" italics="true" pageId="7" pageNumber="498">l'</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFB58FBF6CADCFDF4" box="[1174,1192,1147,1170]" italics="true" pageId="7" pageNumber="498">v'</emphasis>
) is missing adult seta 
<emphasis id="B911EAD7FFAC9F2BFF59FB13CEDBFDD3" box="[151,175,1182,1205]" italics="true" pageId="7" pageNumber="498">v&quot;</emphasis>
(n.b. the expression of deutonymph seta 
<emphasis id="B911EAD7FFAC9F2BFDB4FB10CCFCFDD0" box="[634,648,1181,1206]" italics="true" pageId="7" pageNumber="498">d</emphasis>
is delayed until the adult stage). Two setae are normally added to tibia I in the adult; however, only one of these setae is added in this species, 
<emphasis id="B911EAD7FFAC9F2BFC60FB4FCDBCFDB8" box="[942,968,1218,1246]" italics="true" pageId="7" pageNumber="498">v'1</emphasis>
, while the other seta, 
<emphasis id="B911EAD7FFAC9F2BFB72FB4FCAA8FDB8" box="[1212,1244,1218,1246]" italics="true" pageId="7" pageNumber="498">v&quot;1</emphasis>
, is suppressed. It is assumed that the standard deutonymphal additions of 
<emphasis id="B911EAD7FFAC9F2BFCD4FB65CD5BFC63" box="[794,815,1256,1285]" italics="true" pageId="7" pageNumber="498">l'1</emphasis>
and 
<emphasis id="B911EAD7FFAC9F2BFCA7FB65CDF0FC63" box="[873,900,1256,1285]" italics="true" pageId="7" pageNumber="498">l&quot;1</emphasis>
were made to tibia I in the deutonymph stage. Tibia II–IV all retain the larval complement of five setae (
<emphasis id="B911EAD7FFAC9F2BFCE2FA83CD4EFC41" box="[812,826,1294,1319]" italics="true" pageId="7" pageNumber="498">d</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFC87FA83CD23FC41" box="[841,855,1294,1319]" italics="true" pageId="7" pageNumber="498">l'</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFCABFA83CD0DFC41" box="[869,889,1294,1319]" italics="true" pageId="7" pageNumber="498">l&quot;</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFC46FA82CDEFFC40" box="[904,923,1295,1318]" italics="true" pageId="7" pageNumber="498">v'</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFC64FA82CDB6FC40" box="[938,962,1295,1318]" italics="true" pageId="7" pageNumber="498">v&quot;</emphasis>
), with tibia II missing the two adult setae normally added, 
<emphasis id="B911EAD7FFAC9F2BFE98FABFCF18FC29" box="[342,364,1330,1359]" italics="true" pageId="7" pageNumber="498">l'1</emphasis>
and 
<emphasis id="B911EAD7FFAC9F2BFE6AFABECFCAFC29" box="[420,446,1331,1359]" italics="true" pageId="7" pageNumber="498">v'1</emphasis>
; tibia III is missing the one standard adult addition, 
<emphasis id="B911EAD7FFAC9F2BFBDDFABECA5AFC29" box="[1043,1070,1331,1359]" italics="true" pageId="7" pageNumber="498">v'1</emphasis>
; and tibia IV is missing the two standard adult additions, 
<emphasis id="B911EAD7FFAC9F2BFE61FAD5CFBFFC13" box="[431,459,1368,1397]" italics="true" pageId="7" pageNumber="498">l&quot;1</emphasis>
and 
<emphasis id="B911EAD7FFAC9F2BFDCFFAD4CC6FFC13" box="[513,539,1369,1397]" italics="true" pageId="7" pageNumber="498">v'1</emphasis>
. Tarsus I with 15 (3+3) setae is missing three setae, adult seta 
<emphasis id="B911EAD7FFAC9F2BFB1AFAD4CA9AFC13" box="[1236,1262,1369,1397]" italics="true" pageId="7" pageNumber="498">v'2</emphasis>
, and two of the four standard deutonymph setae, 
<emphasis id="B911EAD7FFAC9F2BFDDFFA0DCC46FCFA" box="[529,562,1408,1436]" italics="true" pageId="7" pageNumber="498">v&quot;1</emphasis>
and 
<emphasis id="B911EAD7FFAC9F2BFDA5FAF2CCF2FCFA" box="[619,646,1407,1436]" italics="true" pageId="7" pageNumber="498">l&quot;1</emphasis>
. Tarsus II with 12 (2+3) setae is missing four setae, three of the four adult setae 
<emphasis id="B911EAD7FFAC9F2BFEDAFA2BCF5AFCA5" box="[276,302,1446,1475]" italics="true" pageId="7" pageNumber="498">v'2</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFEF2FA2BCF28FCA5" box="[316,348,1446,1475]" italics="true" pageId="7" pageNumber="498">v&quot;1</emphasis>
, 
<emphasis id="B911EAD7FFAC9F2BFEA7FA28CF0BFCA5" box="[361,383,1445,1475]" italics="true" pageId="7" pageNumber="498">l'1</emphasis>
(
<emphasis id="B911EAD7FFAC9F2BFE40FA2BCFC2FCA5" box="[398,438,1446,1475]" italics="true" pageId="7" pageNumber="498">ω&quot;1</emphasis>
is added in adult as is standard), and the deutonymphal seta 
<emphasis id="B911EAD7FFAC9F2BFB98FA2BCA05FCA5" box="[1110,1137,1446,1475]" italics="true" pageId="7" pageNumber="498">v'1</emphasis>
. Tarsus III with 8 (1) setae is missing two setae, the single adult seta 
<emphasis id="B911EAD7FFAC9F2BFDB2FA40CCE3FC8F" box="[636,663,1485,1513]" italics="true" pageId="7" pageNumber="498">v'1</emphasis>
and the larval seta 
<emphasis id="B911EAD7FFAC9F2BFCB5FA40CDD5FC82" box="[891,929,1485,1508]" italics="true" pageId="7" pageNumber="498">pv&quot;</emphasis>
. Tarsus IV with 8 (1) setae is missing three setae, two of the three adult setae 
<emphasis id="B911EAD7FFAC9F2BFDDCFA79CC58FF76" box="[530,556,1524,1552]" italics="true" pageId="7" pageNumber="498">v'1</emphasis>
and 
<emphasis id="B911EAD7FFAC9F2BFDADFA79CCF7FF76" box="[611,643,1524,1552]" italics="true" pageId="7" pageNumber="498">v&quot;1</emphasis>
(
<emphasis id="B911EAD7FFAC9F2BFD5DFA79CCD9FF6D" box="[659,685,1524,1547]" italics="true" pageId="7" pageNumber="498">ω'</emphasis>
is added), and the larval seta 
<emphasis id="B911EAD7FFAC9F2BFC37FA79CA6BFF6D" box="[1017,1055,1524,1547]" italics="true" pageId="7" pageNumber="498">pv&quot;</emphasis>
.
</paragraph>
<paragraph id="8BDA36C5FFAC9F2BFF09F994CC1EFF54" blockId="7.[151,1436,455,2037]" box="[199,618,1561,1586]" pageId="7" pageNumber="498">
<emphasis id="B911EAD7FFAC9F2BFF09F994CF62FF54" bold="true" box="[199,278,1561,1586]" italics="true" pageId="7" pageNumber="498">Colour</emphasis>
. Yellow to pale orange in life.
</paragraph>
<paragraph id="8BDA36C5FFAC9F2BFF09F9B0CF7EFE07" blockId="7.[151,1436,455,2037]" pageId="7" pageNumber="498">
<emphasis id="B911EAD7FFAC9F2BFF09F9B0CFB4FF33" bold="true" box="[199,448,1597,1622]" pageId="7" pageNumber="498">
Adult male. 
<emphasis id="B911EAD7FFAC9F2BFE92F9B3CFB4FF33" bold="true" box="[348,448,1598,1621]" italics="true" pageId="7" pageNumber="498">Dorsum.</emphasis>
</emphasis>
(
<figureCitation id="135E2A40FFAC9F2BFE1FF9B0CC57FF30" box="[465,547,1597,1622]" captionStart="FIGURES 20 – 21" captionStartId="12.[151,264,1653,1676]" captionTargetBox="[173,1426,238,1624]" captionTargetId="figure@12.[151,1436,193,1633]" captionTargetPageId="12" captionText="FIGURES 20 – 21. Eotetranychus spinophilus sp. nov. male. (20) dorsal idiosoma; (21) posterior ventral opisthogaster." httpUri="https://zenodo.org/record/998448/files/figure.png" pageId="7" pageNumber="498">Fig. 20</figureCitation>
) Body measurements: 
<emphasis id="B911EAD7FFAC9F2BFCE7F9B3CD15FF3C" box="[809,865,1597,1626]" italics="true" pageId="7" pageNumber="498">
<date id="FFDB1005FFAC9F2BFCE7F9B3CD49FF3C" box="[809,829,1598,1626]" pageId="7" pageNumber="498">v2</date>
–h1
</emphasis>
196–228, 
<emphasis id="B911EAD7FFAC9F2BFC14F9B3CA52FF3C" box="[986,1062,1598,1626]" italics="true" pageId="7" pageNumber="498">sc2–sc2</emphasis>
110–125; idiosomal length 240– 305, width 140–180. Distances between other setae: 
<date id="FFDB1005FFAC9F2BFD26F9E8CD37FF1B" box="[744,835,1636,1664]" pageId="7" pageNumber="498" value="1952-05-02" valueMax="1952-05-02" valueMin="1952-05-02">
<emphasis id="B911EAD7FFAC9F2BFD26F9E8CD6BFFE6" box="[744,799,1637,1664]" italics="true" pageId="7" pageNumber="498">v2–v2</emphasis>
52
</date>
–60, 
<emphasis id="B911EAD7FFAC9F2BFCB4F9E8CDB3FFE6" box="[890,967,1637,1664]" italics="true" pageId="7" pageNumber="498">sc1–sc1</emphasis>
66–77, 
<emphasis id="B911EAD7FFAC9F2BFBECF9E8CA2DFFE6" box="[1058,1113,1637,1664]" italics="true" pageId="7" pageNumber="498">c1–c1</emphasis>
44–51, 
<emphasis id="B911EAD7FFAC9F2BFB7AF9E8CA9FFFE6" box="[1204,1259,1637,1664]" italics="true" pageId="7" pageNumber="498">c3–c3</emphasis>
148–216, 
<emphasis id="B911EAD7FFAC9F2BFAADF9E9CBE8FFE6" box="[1379,1436,1636,1664]" italics="true" pageId="7" pageNumber="498">d1–d1</emphasis>
41–44, 
<emphasis id="B911EAD7FFAC9F2BFF25F907CF69FFC5" box="[235,285,1674,1703]" italics="true" pageId="7" pageNumber="498">d1–d</emphasis>
<date id="FFDB1005FFAC9F2BFED3F914CF01FFC5" box="[285,373,1674,1703]" pageId="7" pageNumber="498" value="1922-02-25">
<emphasis id="B911EAD7FFAC9F2BFED3F914CF51FFC1" box="[285,293,1689,1703]" italics="true" pageId="7" pageNumber="498">2</emphasis>
22–25
</date>
, 
<emphasis id="B911EAD7FFAC9F2BFE4FF906CFC3FFC1" box="[385,439,1675,1703]" italics="true" pageId="7" pageNumber="498">
e1– 
<geoCoordinate id="EE515002FFAC9F2BFE6AF906CFDBFFC4" box="[420,431,1675,1698]" direction="east" orientation="longitude" pageId="7" pageNumber="498" precision="925" value="1.7166667">e</geoCoordinate>
1
</emphasis>
43–44, 
<emphasis id="B911EAD7FFAC9F2BFDDDF906CC35FFC4" box="[531,577,1675,1703]" italics="true" pageId="7" pageNumber="498">
e1– 
<geoCoordinate id="EE515002FFAC9F2BFDF8F906CC35FFC4" box="[566,577,1675,1698]" direction="east" orientation="longitude" pageId="7" pageNumber="498" precision="925" value="2.35">e</geoCoordinate>
</emphasis>
<date id="FFDB1005FFAC9F2BFD8FF914CCEDFFC5" box="[577,665,1675,1703]" pageId="7" pageNumber="498" value="1921-02-23">
<emphasis id="B911EAD7FFAC9F2BFD8FF914CC3DFFC1" box="[577,585,1689,1703]" italics="true" pageId="7" pageNumber="498">2</emphasis>
21–23
</date>
, 
<emphasis id="B911EAD7FFAC9F2BFD6BF906CCBEFFC5" box="[677,714,1675,1703]" italics="true" pageId="7" pageNumber="498">f1–f</emphasis>
<date id="FFDB1005FFAC9F2BFD04F914CD56FFC5" box="[714,802,1675,1703]" pageId="7" pageNumber="498" value="1924-01-28">
<emphasis id="B911EAD7FFAC9F2BFD04F914CCA6FFC1" box="[714,722,1689,1703]" italics="true" pageId="7" pageNumber="498">1</emphasis>
24–28
</date>
, 
<emphasis id="B911EAD7FFAC9F2BFCE0F907CD2BFFC5" box="[814,863,1674,1703]" italics="true" pageId="7" pageNumber="498">h1–h</emphasis>
<date id="FFDB1005FFAC9F2BFCAEF914CDCFFFC5" box="[864,955,1675,1703]" pageId="7" pageNumber="498" value="2011-01-14">
<emphasis id="B911EAD7FFAC9F2BFCAEF914CD1CFFC1" box="[864,872,1689,1703]" italics="true" pageId="7" pageNumber="498">1</emphasis>
11–14.
</date>
Body tapering caudally, striation and setal form similar to that of female; dorsal opisthosoma with mostly transverse striae; with band of more widely–spaced transverse striae posteriad 
<emphasis id="B911EAD7FFAC9F2BFE08F959CFA8FF97" box="[454,476,1748,1777]" italics="true" pageId="7" pageNumber="498">h1</emphasis>
; with longitudinal striae laterad 
<emphasis id="B911EAD7FFAC9F2BFC9DF95BCD13FF97" box="[851,871,1750,1777]" italics="true" pageId="7" pageNumber="498">c2</emphasis>
and 
<emphasis id="B911EAD7FFAC9F2BFC6FF959CDADFF97" box="[929,985,1748,1777]" italics="true" pageId="7" pageNumber="498">d1–e1</emphasis>
; dorsal setae weakly barbed, each with minute distal fork. Setal lengths: 
<date id="FFDB1005FFAC9F2BFDC2F971CC30FE72" box="[524,580,1787,1816]" pageId="7" pageNumber="498" value="1939-05-02">
<emphasis id="B911EAD7FFAC9F2BFDC2F971CC55FE7E" box="[524,545,1788,1816]" italics="true" pageId="7" pageNumber="498">v2</emphasis>
39
</date>
–46, 
<emphasis id="B911EAD7FFAC9F2BFDB2F971CCEFFE7E" box="[636,667,1788,1816]" italics="true" pageId="7" pageNumber="498">sc1</emphasis>
37–51, 
<emphasis id="B911EAD7FFAC9F2BFD38F971CD62FE7E" box="[758,790,1788,1816]" italics="true" pageId="7" pageNumber="498">sc2</emphasis>
48–57, 
<emphasis id="B911EAD7FFAC9F2BFCBFF971CD09FE75" box="[881,893,1788,1811]" italics="true" pageId="7" pageNumber="498">c</emphasis>
<date id="FFDB1005FFAC9F2BFCB0F887CDA1FE75" box="[894,981,1787,1816]" pageId="7" pageNumber="498" value="1937-01-28">
<emphasis id="B911EAD7FFAC9F2BFCB0F887CDF2FE7E" box="[894,902,1802,1816]" italics="true" pageId="7" pageNumber="498">1</emphasis>
28–37
</date>
, 
<emphasis id="B911EAD7FFAC9F2BFC2FF971CD81FE7E" box="[993,1013,1788,1816]" italics="true" pageId="7" pageNumber="498">c2</emphasis>
44–54, 
<emphasis id="B911EAD7FFAC9F2BFB9EF971CA11FE7E" box="[1104,1125,1788,1816]" italics="true" pageId="7" pageNumber="498">c3</emphasis>
56–63, 
<emphasis id="B911EAD7FFAC9F2BFB0FF976CAA2FE7E" box="[1217,1238,1787,1816]" italics="true" pageId="7" pageNumber="498">d1</emphasis>
32–38, 
<emphasis id="B911EAD7FFAC9F2BFAFCF976CB3CFE7E" box="[1330,1352,1787,1816]" italics="true" pageId="7" pageNumber="498">d2</emphasis>
46–53, 
<geoCoordinate id="EE515002FFAC9F2BFF59F8AECEB9FE5C" box="[151,205,1826,1854]" direction="east" orientation="longitude" pageId="7" pageNumber="498" precision="925" value="1.55">
<emphasis id="B911EAD7FFAC9F2BFF59F8AECEDFFE58" box="[151,171,1827,1854]" italics="true" pageId="7" pageNumber="498">e1</emphasis>
33
</geoCoordinate>
–40, 
<geoCoordinate id="EE515002FFAC9F2BFECBF8AECF48FE5C" box="[261,316,1825,1854]" direction="east" orientation="longitude" pageId="7" pageNumber="498" precision="925" value="2.65">
<emphasis id="B911EAD7FFAC9F2BFECBF8AECF6EFE58" box="[261,282,1827,1854]" italics="true" pageId="7" pageNumber="498">e2</emphasis>
39
</geoCoordinate>
–52, 
<emphasis id="B911EAD7FFAC9F2BFEBAF8AFCFF7FE58" box="[372,387,1826,1854]" italics="true" pageId="7" pageNumber="498">f1</emphasis>
36–40, 
<emphasis id="B911EAD7FFAC9F2BFE13F8AFCF99FE58" box="[477,493,1826,1854]" italics="true" pageId="7" pageNumber="498">f2</emphasis>
46–49, 
<emphasis id="B911EAD7FFAC9F2BFD89F8ACCC29FE58" box="[583,605,1825,1854]" italics="true" pageId="7" pageNumber="498">h1</emphasis>
33–39, 
<emphasis id="B911EAD7FFAC9F2BFD79F8ACCCB1FE5C" box="[695,709,1825,1850]" italics="true" pageId="7" pageNumber="498">h</emphasis>
<date id="FFDB1005FFAC9F2BFD0AF8BDCD68FE5C" box="[708,796,1826,1854]" pageId="7" pageNumber="498" value="1921-02-23">
<emphasis id="B911EAD7FFAC9F2BFD0AF8BDCCB8FE58" box="[708,716,1840,1854]" italics="true" pageId="7" pageNumber="498">2</emphasis>
21–23
</date>
, 
<emphasis id="B911EAD7FFAC9F2BFCE8F8ACCD40FE5C" box="[806,820,1825,1850]" italics="true" pageId="7" pageNumber="498">h</emphasis>
<date id="FFDB1005FFAC9F2BFCFAF8BDCDE4FE5C" box="[820,912,1826,1854]" pageId="7" pageNumber="498" value="1921-03-23">
<emphasis id="B911EAD7FFAC9F2BFCFAF8BDCD48FE58" box="[820,828,1840,1854]" italics="true" pageId="7" pageNumber="498">3</emphasis>
21–23.
</date>
Setae 
<emphasis id="B911EAD7FFAC9F2BFC14F8ACCD84FE58" box="[986,1008,1825,1854]" italics="true" pageId="7" pageNumber="498">h3</emphasis>
often inserted ventrally, depending on mounting.
</paragraph>
<paragraph id="8BDA36C5FFAC9F2BFF09F8E1CF7EFE93" blockId="7.[151,1436,455,2037]" pageId="7" pageNumber="498">
<emphasis id="B911EAD7FFAC9F2BFF09F8E1CF64FEE5" bold="true" box="[199,272,1900,1923]" italics="true" pageId="7" pageNumber="498">Venter</emphasis>
. (
<figureCitation id="135E2A40FFAC9F2BFEE4F8E1CF0BFEE2" box="[298,383,1900,1925]" captionStart="FIGURES 20 – 21" captionStartId="12.[151,264,1653,1676]" captionTargetBox="[173,1426,238,1624]" captionTargetId="figure@12.[151,1436,193,1633]" captionTargetPageId="12" captionText="FIGURES 20 – 21. Eotetranychus spinophilus sp. nov. male. (20) dorsal idiosoma; (21) posterior ventral opisthogaster." httpUri="https://zenodo.org/record/998448/files/figure.png" pageId="7" pageNumber="498">Fig. 21</figureCitation>
) Body finely striate, striae almost entirely transverse with no lobes; all ventral setae thin, smooth: 
<emphasis id="B911EAD7FFAC9F2BFF37F81DCF61FECE" box="[249,277,1936,1960]" italics="true" pageId="7" pageNumber="498">1a</emphasis>
23–26, 
<emphasis id="B911EAD7FFAC9F2BFEBFF802CFF9FECE" box="[369,397,1935,1960]" italics="true" pageId="7" pageNumber="498">1b</emphasis>
23–26, 
<emphasis id="B911EAD7FFAC9F2BFE27F81DCC70FECE" box="[489,516,1936,1960]" italics="true" pageId="7" pageNumber="498">1c</emphasis>
36–41, 
<emphasis id="B911EAD7FFAC9F2BFDAEF802CC08FECE" box="[608,636,1935,1960]" italics="true" pageId="7" pageNumber="498">2b</emphasis>
30–31, 
<emphasis id="B911EAD7FFAC9F2BFD16F81DCC80FECE" box="[728,756,1936,1960]" italics="true" pageId="7" pageNumber="498">3a</emphasis>
14–15, 
<emphasis id="B911EAD7FFAC9F2BFC9EF802CD18FECE" box="[848,876,1935,1960]" italics="true" pageId="7" pageNumber="498">3b</emphasis>
26–30, 
<emphasis id="B911EAD7FFAC9F2BFC07F81DCD91FECE" box="[969,997,1936,1960]" italics="true" pageId="7" pageNumber="498">4a</emphasis>
19–20, 
<emphasis id="B911EAD7FFAC9F2BFB8FF802CA29FECE" box="[1089,1117,1935,1960]" italics="true" pageId="7" pageNumber="498">4b</emphasis>
25–28, 
<emphasis id="B911EAD7FFAC9F2BFB77F81CCAA1FECE" box="[1209,1237,1937,1960]" italics="true" pageId="7" pageNumber="498">ag</emphasis>
21–24, 
<emphasis id="B911EAD7FFAC9F2BFAFCF81CCB34FECE" box="[1330,1344,1937,1960]" italics="true" pageId="7" pageNumber="498">g</emphasis>
<date id="FFDB1005FFAC9F2BFAF1F813CBECFECE" box="[1343,1432,1936,1964]" pageId="7" pageNumber="498" value="2014-01-17">
<emphasis id="B911EAD7FFAC9F2BFAF1F813CB33FECA" box="[1343,1351,1950,1964]" italics="true" pageId="7" pageNumber="498">1</emphasis>
14–17
</date>
, 
<emphasis id="B911EAD7FFAC9F2BFF59F83ACED1FEA8" box="[151,165,1975,1998]" italics="true" pageId="7" pageNumber="498">g</emphasis>
<date id="FFDB1005FFAC9F2BFF6BF848CE8BFEA9" box="[165,255,1974,2003]" pageId="7" pageNumber="498" value="2015-12-02">
<emphasis id="B911EAD7FFAC9F2BFF6BF848CED9FEB5" box="[165,173,1989,2003]" italics="true" pageId="7" pageNumber="498">2</emphasis>
12–15
</date>
, 
<emphasis id="B911EAD7FFAC9F2BFEC1F83ACF5CFEA8" box="[271,296,1975,1998]" italics="true" pageId="7" pageNumber="498">ps</emphasis>
<date id="FFDB1005FFAC9F2BFEE6F848CFF7FEA9" box="[296,387,1974,2003]" pageId="7" pageNumber="498" value="2015-12-01">
<emphasis id="B911EAD7FFAC9F2BFEE6F848CF44FEB5" box="[296,304,1989,2003]" italics="true" pageId="7" pageNumber="498">1</emphasis>
12–15
</date>
, 
<emphasis id="B911EAD7FFAC9F2BFE5CF83ACFDFFEA8" box="[402,427,1975,1998]" italics="true" pageId="7" pageNumber="498">ps</emphasis>
<date id="FFDB1005FFAC9F2BFE65F848CC7FFEA9" box="[427,523,1974,2003]" pageId="7" pageNumber="498" value="2012-02-15">
<emphasis id="B911EAD7FFAC9F2BFE65F848CFC7FEB5" box="[427,435,1989,2003]" italics="true" pageId="7" pageNumber="498">2</emphasis>
12–15.
</date>
Setae 
<emphasis id="B911EAD7FFAC9F2BFD93F83ACC0AFEB5" box="[605,638,1975,2003]" italics="true" pageId="7" pageNumber="498">ps1</emphasis>
often inserted dorsally to dorsolaterally (
<figureCitation id="135E2A40FFAC9F2BFBADF83BCA9DFEA9" box="[1123,1257,1974,1999]" captionStart="FIGURES 20 – 21" captionStartId="12.[151,264,1653,1676]" captionTargetBox="[173,1426,238,1624]" captionTargetId="figure@12.[151,1436,193,1633]" captionTargetPageId="12" captionText="FIGURES 20 – 21. Eotetranychus spinophilus sp. nov. male. (20) dorsal idiosoma; (21) posterior ventral opisthogaster." httpUri="https://zenodo.org/record/998448/files/figure.png" pageId="7" pageNumber="498">Figs 20–21</figureCitation>
), depending on mounting.
</paragraph>
<paragraph id="8BDA36C5FFA39F24FF09FF1ACDA0F99E" blockId="8.[151,1436,151,248]" pageId="8" pageNumber="499">
<taxonomicName id="4C654D46FFA39F24FF09FF1ACF2FF9D6" authorityName="Krefft" authorityYear="1871" box="[199,347,151,176]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="8" pageNumber="499" phylum="Nematoda" rank="genus">
<emphasis id="B911EAD7FFA39F24FF09FF1ACF2FF9D6" bold="true" box="[199,347,151,176]" italics="true" pageId="8" pageNumber="499">Gnathosoma</emphasis>
</taxonomicName>
. (
<figureCitation id="135E2A40FFA39F24FEBFFF1ACFB7F9D6" box="[369,451,151,176]" captionStart="FIGURES 29 – 37" captionStartId="14.[151,264,1939,1962]" captionTargetBox="[253,1321,209,1908]" captionTargetId="figure@14.[234,1353,193,1935]" captionTargetPageId="14" captionText="FIGURES 29 – 37. Eotetranychus spinophilus sp. nov. male. (29) leg III, left side, dorsal to adaxial view; (30) leg IV, right side, abaxial view; (31) palp, dorsal view; (32 – 34) aedeagus, different aspects of same specimen; (35 – 37) aedeagus, different aspects of another specimen. Scale bar for aedeagi = 100 µm." httpUri="https://zenodo.org/record/998452/files/figure.png" pageId="8" pageNumber="499">Fig. 31</figureCitation>
) Short spinneret (
<emphasis id="B911EAD7FFA39F24FD41FF1ACCC7F9D6" box="[655,691,151,176]" italics="true" pageId="8" pageNumber="499">suζ</emphasis>
) on palp tarsus slightly smaller than that of female (1.5–1.8 wide, 4 long); single solenidion (4–5) and two eupathidia (
<emphasis id="B911EAD7FFA39F24FD28FF31CC88F9B3" box="[742,764,188,213]" italics="true" pageId="8" pageNumber="499">Μl</emphasis>
′ 4, 
<emphasis id="B911EAD7FFA39F24FCEBFF31CD33F9B3" box="[805,839,188,213]" italics="true" pageId="8" pageNumber="499">Μl&quot;</emphasis>
5). Dorsal femoral seta 
<emphasis id="B911EAD7FFA39F24FB98FF31CAE7F9B3" box="[1110,1171,188,213]" italics="true" pageId="8" pageNumber="499">dPFe</emphasis>
modified as short thick peg-like seta, as occurs in males of other genera. Peritreme anastomosing.
</paragraph>
<caption id="DF1A664DFFA39F24FF59F83BCDF0FEAA" box="[151,900,1974,1996]" httpUri="https://zenodo.org/record/998440/files/figure.png" pageId="8" pageNumber="499" targetBox="[183,1357,312,1918]" targetPageId="8">
<paragraph id="8BDA36C5FFA39F24FF59F83BCDF0FEAA" blockId="8.[151,900,1974,1996]" box="[151,900,1974,1996]" pageId="8" pageNumber="499">
<emphasis id="B911EAD7FFA39F24FF59F83BCF67FEAA" bold="true" box="[151,275,1974,1996]" pageId="8" pageNumber="499">FIGURE 1.</emphasis>
<taxonomicName id="4C654D46FFA39F24FED7F83BCC52FEAA" authority="Zhang, 2017" authorityName="Zhang" authorityYear="2017" box="[281,550,1974,1996]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="499" phylum="Arthropoda" rank="species" species="spinophilus" status="sp. nov.">
<emphasis id="B911EAD7FFA39F24FED7F83BCC52FEAA" box="[281,550,1974,1996]" italics="true" pageId="8" pageNumber="499">Eotetranychus spinophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257ACFFA39F24FDE2F83ACC09FEAA" box="[556,637,1975,1996]" pageId="8" pageNumber="499" rank="species">
<emphasis id="B911EAD7FFA39F24FDE2F83ACC09FEAA" bold="true" box="[556,637,1975,1996]" pageId="8" pageNumber="499">sp. nov.</emphasis>
</taxonomicNameLabel>
, female, dorsal idiosoma.
</paragraph>
</caption>
<caption id="DF1A664DFFA29F25FF59F82BCF09FEBD" httpUri="https://zenodo.org/record/998442/files/figure.png" pageId="9" pageNumber="500" targetBox="[257,1312,203,1917]" targetPageId="9">
<paragraph id="8BDA36C5FFA29F25FF59F82BCF09FEBD" blockId="9.[151,1435,1958,2011]" pageId="9" pageNumber="500">
<emphasis id="B911EAD7FFA29F25FF59F82BCF48FEDA" bold="true" box="[151,316,1958,1981]" pageId="9" pageNumber="500">FIGURES 2–7.</emphasis>
<taxonomicName id="4C654D46FFA29F25FE8DF82BCC26FEDA" authority="Zhang, 2017" authorityName="Zhang" authorityYear="2017" box="[323,594,1958,1980]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="500" phylum="Arthropoda" rank="species" species="spinophilus" status="sp. nov.">
<emphasis id="B911EAD7FFA29F25FE8DF82BCC26FEDA" box="[323,594,1958,1980]" italics="true" pageId="9" pageNumber="500">Eotetranychus spinophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257ACFFA29F25FD94F82ACCD8FEDA" box="[602,684,1959,1980]" pageId="9" pageNumber="500" rank="species">
<emphasis id="B911EAD7FFA29F25FD94F82ACCD8FEDA" bold="true" box="[602,684,1959,1980]" pageId="9" pageNumber="500">sp. nov.</emphasis>
</taxonomicNameLabel>
female. (2) mid-ventral region; (3) posterior ventral opisthogaster; (4–7) spermatheca variation.
</paragraph>
</caption>
<caption id="DF1A664DFFA19F26FF59F8FDCB0CFEC0" httpUri="https://zenodo.org/record/998444/files/figure.png" pageId="10" pageNumber="501" targetBox="[162,1387,217,1873]" targetPageId="10">
<paragraph id="8BDA36C5FFA19F26FF59F8FDCB0CFEC0" blockId="10.[151,1436,1904,1958]" pageId="10" pageNumber="501">
<emphasis id="B911EAD7FFA19F26FF59F8FDCF3DFEE0" bold="true" box="[151,329,1904,1927]" pageId="10" pageNumber="501">FIGURES 8–14.</emphasis>
<taxonomicName id="4C654D46FFA19F26FE9FF8FCCC14FEE1" authority="Zhang, 2017" authorityName="Zhang" authorityYear="2017" box="[337,608,1905,1927]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="501" phylum="Arthropoda" rank="species" species="spinophilus" status="sp. nov.">
<emphasis id="B911EAD7FFA19F26FE9FF8FCCC14FEE1" box="[337,608,1905,1927]" italics="true" pageId="10" pageNumber="501">Eotetranychus spinophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257ACFFA19F26FDA6F8FCCCCEFEE0" box="[616,698,1905,1926]" pageId="10" pageNumber="501" rank="species">
<emphasis id="B911EAD7FFA19F26FDA6F8FCCCCEFEE0" bold="true" box="[616,698,1905,1926]" pageId="10" pageNumber="501">sp. nov.</emphasis>
</taxonomicNameLabel>
female. (8–10) peritreme, different specimens; (11) palp, ventral view; (12) palp, dorsal view; (13) leg I, right side, dorsal to abaxial view; (14) ambulacrum I. Scale bar for ambulacrum = 40 µm.
</paragraph>
</caption>
<caption id="DF1A664DFFA09F27FF59F833CD8AFE95" httpUri="https://zenodo.org/record/998446/files/figure.png" pageId="11" pageNumber="502" targetBox="[220,1342,202,1928]" targetPageId="11">
<paragraph id="8BDA36C5FFA09F27FF59F833CD8AFE95" blockId="11.[151,1436,1982,2035]" pageId="11" pageNumber="502">
<emphasis id="B911EAD7FFA09F27FF59F833CF20FEB2" bold="true" box="[151,340,1982,2005]" pageId="11" pageNumber="502">FIGURES 15–19.</emphasis>
<taxonomicName id="4C654D46FFA09F27FE94F833CC13FEB2" authority="Zhang, 2017" authorityName="Zhang" authorityYear="2017" box="[346,615,1982,2004]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="502" phylum="Arthropoda" rank="species" species="spinophilus" status="sp. nov.">
<emphasis id="B911EAD7FFA09F27FE94F833CC13FEB2" box="[346,615,1982,2004]" italics="true" pageId="11" pageNumber="502">Eotetranychus spinophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257ACFFA09F27FDA3F832CCCAFEB2" box="[621,702,1983,2004]" pageId="11" pageNumber="502" rank="species">
<emphasis id="B911EAD7FFA09F27FDA3F832CCCAFEB2" bold="true" box="[621,702,1983,2004]" pageId="11" pageNumber="502">sp. nov.</emphasis>
</taxonomicNameLabel>
female. (15) leg II, right side, dorsal view; (16) ambulacrum II; (17) leg III, right side, abaxial view; (18) ambulacrum III; (19) leg IV, right side, abaxial view.
</paragraph>
</caption>
<caption id="DF1A664DFFA79F20FF59F9F8CB38FFEA" box="[151,1356,1653,1676]" httpUri="https://zenodo.org/record/998448/files/figure.png" pageId="12" pageNumber="503" targetBox="[173,1426,238,1624]" targetPageId="12">
<paragraph id="8BDA36C5FFA79F20FF59F9F8CB38FFEA" blockId="12.[151,1356,1653,1676]" box="[151,1356,1653,1676]" pageId="12" pageNumber="503">
<emphasis id="B911EAD7FFA79F20FF59F9F8CF27FFED" bold="true" box="[151,339,1653,1676]" pageId="12" pageNumber="503">FIGURES 20–21.</emphasis>
<taxonomicName id="4C654D46FFA79F20FE94F9FBCC13FFEA" authority="Zhang, 2017" authorityName="Zhang" authorityYear="2017" box="[346,615,1654,1676]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="spinophilus" status="sp. nov.">
<emphasis id="B911EAD7FFA79F20FE94F9FBCC13FFEA" box="[346,615,1654,1676]" italics="true" pageId="12" pageNumber="503">Eotetranychus spinophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257ACFFA79F20FDA3F9FACCCAFFEA" box="[621,702,1655,1676]" pageId="12" pageNumber="503" rank="species">
<emphasis id="B911EAD7FFA79F20FDA3F9FACCCAFFEA" bold="true" box="[621,702,1655,1676]" pageId="12" pageNumber="503">sp. nov.</emphasis>
</taxonomicNameLabel>
male. (20) dorsal idiosoma; (21) posterior ventral opisthogaster.
</paragraph>
</caption>
<paragraph id="8BDA36C5FFA79F20FF09F935CC71FF93" blockId="12.[151,1437,1720,2038]" pageId="12" pageNumber="503">
<emphasis id="B911EAD7FFA79F20FF09F935CF43FFB7" bold="true" box="[199,311,1720,1745]" italics="true" pageId="12" pageNumber="503">Aedeagus</emphasis>
. (
<figureCitation id="135E2A40FFA79F20FE83F934CFBAFFB7" box="[333,462,1721,1746]" captionStart="FIGURES 29 – 37" captionStartId="14.[151,264,1939,1962]" captionTargetBox="[253,1321,209,1908]" captionTargetId="figure@14.[234,1353,193,1935]" captionTargetPageId="14" captionText="FIGURES 29 – 37. Eotetranychus spinophilus sp. nov. male. (29) leg III, left side, dorsal to adaxial view; (30) leg IV, right side, abaxial view; (31) palp, dorsal view; (32 – 34) aedeagus, different aspects of same specimen; (35 – 37) aedeagus, different aspects of another specimen. Scale bar for aedeagi = 100 µm." httpUri="https://zenodo.org/record/998452/files/figure.png" pageId="12" pageNumber="503">Figs 32–37</figureCitation>
) Directed dorsally, strongly sigmoid, thickened with blunt tip. Measurements (see 
<figureCitation id="135E2A40FFA79F20FABEF934CEC3FF93" captionStart="FIGURES 29 – 37" captionStartId="14.[151,264,1939,1962]" captionTargetBox="[253,1321,209,1908]" captionTargetId="figure@14.[234,1353,193,1935]" captionTargetPageId="14" captionText="FIGURES 29 – 37. Eotetranychus spinophilus sp. nov. male. (29) leg III, left side, dorsal to adaxial view; (30) leg IV, right side, abaxial view; (31) palp, dorsal view; (32 – 34) aedeagus, different aspects of same specimen; (35 – 37) aedeagus, different aspects of another specimen. Scale bar for aedeagi = 100 µm." httpUri="https://zenodo.org/record/998452/files/figure.png" pageId="12" pageNumber="503">Fig. 32</figureCitation>
): a: 5–7, b: 10–11, c: 5, d: 30.
</paragraph>
<paragraph id="8BDA36C5FFA79F21FF09F88CCB44F8C9" blockId="12.[151,1437,1720,2038]" lastBlockId="13.[151,1436,151,588]" lastPageId="13" lastPageNumber="504" pageId="12" pageNumber="503">
<emphasis id="B911EAD7FFA79F20FF09F88CCE89FE7E" bold="true" box="[199,253,1793,1816]" italics="true" pageId="12" pageNumber="503">Legs</emphasis>
. (
<figureCitation id="135E2A40FFA79F20FEDBF88CCFE3FE7F" box="[277,407,1793,1818]" captionStart-0="FIGURES 22 – 28" captionStart-1="FIGURES 29 – 37" captionStartId-0="13.[151,264,1943,1966]" captionStartId-1="14.[151,264,1939,1962]" captionTargetBox-0="[161,1418,656,1917]" captionTargetBox-1="[253,1321,209,1908]" captionTargetId-0="figure@13.[151,1436,639,1922]" captionTargetId-1="figure@14.[234,1353,193,1935]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 22 – 28. Eotetranychus spinophilus sp. nov. male. (22) leg I, right side, dorsal to abaxial view; (23 – 24) empodium I, different specimens and views; (25) ambulacrum I; (26) leg II, dorsal to adaxial view; (27) empodium III; (28) ambulacrum III, different specimen. Scale bar for ambulacra = 20 µm." captionText-1="FIGURES 29 – 37. Eotetranychus spinophilus sp. nov. male. (29) leg III, left side, dorsal to adaxial view; (30) leg IV, right side, abaxial view; (31) palp, dorsal view; (32 – 34) aedeagus, different aspects of same specimen; (35 – 37) aedeagus, different aspects of another specimen. Scale bar for aedeagi = 100 µm." httpUri-0="https://zenodo.org/record/998450/files/figure.png" httpUri-1="https://zenodo.org/record/998452/files/figure.png" pageId="12" pageNumber="503">Figs 22–30</figureCitation>
) Leg striation same as female. Empodia I split into two sets of three proximoventral hairs with middle pair of hairs significantly thickened, uncinate, appearing as a pair of “claws” when viewed dorsally (
<figureCitation id="135E2A40FFA79F20FF51F8C4CF6BFE04" box="[159,287,1865,1890]" captionStart="FIGURES 22 – 28" captionStartId="13.[151,264,1943,1966]" captionTargetBox="[161,1418,656,1917]" captionTargetId="figure@13.[151,1436,639,1922]" captionTargetPageId="13" captionText="FIGURES 22 – 28. Eotetranychus spinophilus sp. nov. male. (22) leg I, right side, dorsal to abaxial view; (23 – 24) empodium I, different specimens and views; (25) ambulacrum I; (26) leg II, dorsal to adaxial view; (27) empodium III; (28) ambulacrum III, different specimen. Scale bar for ambulacra = 20 µm." httpUri="https://zenodo.org/record/998450/files/figure.png" pageId="12" pageNumber="503">Figs 23–25</figureCitation>
) (depending on angle of mounting, empodium I can appear to be uncinate with a dorsal spur, 
<figureCitation id="135E2A40FFA79F20FAF5F8C4CEC3FEE3" captionStart="FIGURES 22 – 28" captionStartId="13.[151,264,1943,1966]" captionTargetBox="[161,1418,656,1917]" captionTargetId="figure@13.[151,1436,639,1922]" captionTargetPageId="13" captionText="FIGURES 22 – 28. Eotetranychus spinophilus sp. nov. male. (22) leg I, right side, dorsal to abaxial view; (23 – 24) empodium I, different specimens and views; (25) ambulacrum I; (26) leg II, dorsal to adaxial view; (27) empodium III; (28) ambulacrum III, different specimen. Scale bar for ambulacra = 20 µm." httpUri="https://zenodo.org/record/998450/files/figure.png" pageId="12" pageNumber="503">Figs 23– 24</figureCitation>
). Empodia II–IV, similar to female empodia, split into two sets of three proximoventral hairs, with proximal pair of hairs thicker than other two pairs (
<figureCitation id="135E2A40FFA79F20FD84F81CCCA7FECF" box="[586,723,1937,1962]" captionStart="FIGURES 22 – 28" captionStartId="13.[151,264,1943,1966]" captionTargetBox="[161,1418,656,1917]" captionTargetId="figure@13.[151,1436,639,1922]" captionTargetPageId="13" captionText="FIGURES 22 – 28. Eotetranychus spinophilus sp. nov. male. (22) leg I, right side, dorsal to abaxial view; (23 – 24) empodium I, different specimens and views; (25) ambulacrum I; (26) leg II, dorsal to adaxial view; (27) empodium III; (28) ambulacrum III, different specimen. Scale bar for ambulacra = 20 µm." httpUri="https://zenodo.org/record/998450/files/figure.png" pageId="12" pageNumber="503">Figs 27, 28</figureCitation>
). Tarsus I (
<figureCitation id="135E2A40FFA79F20FC99F81CCDDFFECF" box="[855,939,1937,1962]" captionStart="FIGURES 22 – 28" captionStartId="13.[151,264,1943,1966]" captionTargetBox="[161,1418,656,1917]" captionTargetId="figure@13.[151,1436,639,1922]" captionTargetPageId="13" captionText="FIGURES 22 – 28. Eotetranychus spinophilus sp. nov. male. (22) leg I, right side, dorsal to abaxial view; (23 – 24) empodium I, different specimens and views; (25) ambulacrum I; (26) leg II, dorsal to adaxial view; (27) empodium III; (28) ambulacrum III, different specimen. Scale bar for ambulacra = 20 µm." httpUri="https://zenodo.org/record/998450/files/figure.png" pageId="12" pageNumber="503">Fig. 22</figureCitation>
) with two distal and adjacent duplex setae (solenidia: proximal 
<emphasis id="B911EAD7FFA79F20FE4BF838CFEBFEA8" box="[389,415,1973,1998]" italics="true" pageId="12" pageNumber="503">ω ′</emphasis>
30–33, distal 
<emphasis id="B911EAD7FFA79F20FD88F83BCC12FEAB" box="[582,614,1974,1997]" italics="true" pageId="12" pageNumber="503">ω&quot;</emphasis>
42–44; companion fastigial setae: 
<emphasis id="B911EAD7FFA79F20FC35F838CA65FEAB" box="[1019,1041,1973,1997]" italics="true" pageId="12" pageNumber="503">ft'</emphasis>
, 
<emphasis id="B911EAD7FFA79F20FBEFF838CA48FEAB" box="[1057,1084,1973,1997]" italics="true" pageId="12" pageNumber="503">ft&quot;</emphasis>
6–9), a single, spindle-shaped proximal solenidion (
<emphasis id="B911EAD7FFA79F20FE46F857CFDCFE97" box="[392,424,2010,2033]" italics="true" pageId="12" pageNumber="503">ω&quot;</emphasis>
<date id="FFDB1005FFA79F20FE66F865CF8AFE97" box="[424,510,2009,2038]" pageId="12" pageNumber="503" value="2010-01-11">
<emphasis id="B911EAD7FFA79F20FE66F865CFC4FE90" box="[424,432,2024,2038]" italics="true" pageId="12" pageNumber="503">1</emphasis>
10–11
</date>
), and three distal eupathidia (
<emphasis id="B911EAD7FFA79F20FC84F854CD1DFE94" box="[842,873,2009,2034]" italics="true" pageId="12" pageNumber="503">p'ζ</emphasis>
, 
<emphasis id="B911EAD7FFA79F20FCB6F854CDE8FE94" box="[888,924,2009,2034]" italics="true" pageId="12" pageNumber="503">p&quot;ζ</emphasis>
17–18; 
<emphasis id="B911EAD7FFA79F20FC37F854CA51FE94" box="[1017,1061,2009,2034]" italics="true" pageId="12" pageNumber="503">pv'ζ</emphasis>
7–9). Tibia I with three solenidia (
<emphasis id="B911EAD7FFA69F21FF6EFF15CEBFF9C9" box="[160,203,152,175]" italics="true" pageId="13" pageNumber="504">φ'Ƌ</emphasis>
6, 
<emphasis id="B911EAD7FFA69F21FF38FF15CF52F9C9" box="[246,294,152,175]" italics="true" pageId="13" pageNumber="504">φ&quot;Ƌ</emphasis>
7 (both spindle-shaped), 
<emphasis id="B911EAD7FFA69F21FD9BFF14CC10F9D6" box="[597,612,153,176]" italics="true" pageId="13" pageNumber="504">φ</emphasis>
8–9). Tarsus II (
<figureCitation id="135E2A40FFA69F21FCFFFF1ACDF3F9D6" box="[817,903,151,176]" captionStart="FIGURES 22 – 28" captionStartId="13.[151,264,1943,1966]" captionTargetBox="[161,1418,656,1917]" captionTargetId="figure@13.[151,1436,639,1922]" captionTargetPageId="13" captionText="FIGURES 22 – 28. Eotetranychus spinophilus sp. nov. male. (22) leg I, right side, dorsal to abaxial view; (23 – 24) empodium I, different specimens and views; (25) ambulacrum I; (26) leg II, dorsal to adaxial view; (27) empodium III; (28) ambulacrum III, different specimen. Scale bar for ambulacra = 20 µm." httpUri="https://zenodo.org/record/998450/files/figure.png" pageId="13" pageNumber="504">Fig. 26</figureCitation>
) with one duplex seta (solenidion 
<emphasis id="B911EAD7FFA69F21FAEDFF15CB37F9C9" box="[1315,1347,152,175]" italics="true" pageId="13" pageNumber="504">ω&quot;</emphasis>
29–30; companion seta 
<emphasis id="B911EAD7FFA69F21FE9BFF31CF04F9B2" box="[341,368,188,212]" italics="true" pageId="13" pageNumber="504">ft&quot;</emphasis>
5–7), a pair of short spindle-shaped proximal solenidia (
<emphasis id="B911EAD7FFA69F21FBDDFF31CA3EF9B2" box="[1043,1098,188,217]" italics="true" pageId="13" pageNumber="504">ω ′1Ƌ</emphasis>
, 
<emphasis id="B911EAD7FFA69F21FB92FF30CA08F9B2" box="[1116,1148,189,212]" italics="true" pageId="13" pageNumber="504">ω&quot;</emphasis>
<date id="FFDB1005FFA69F21FBB2FF46CAA0F9B2" box="[1148,1236,188,217]" pageId="13" pageNumber="504" value="2011-01-12">
<emphasis id="B911EAD7FFA69F21FBB2FF46CAF0F9BF" box="[1148,1156,203,217]" italics="true" pageId="13" pageNumber="504">1</emphasis>
11–12
</date>
) and three distal eupathidia (
<emphasis id="B911EAD7FFA69F21FED0FF6FCF49F99D" box="[286,317,226,251]" italics="true" pageId="13" pageNumber="504">p'ζ</emphasis>
, 
<emphasis id="B911EAD7FFA69F21FE83FF6FCF06F99D" box="[333,370,226,251]" italics="true" pageId="13" pageNumber="504">p&quot;ζ</emphasis>
16–17, 
<emphasis id="B911EAD7FFA69F21FE1CFF6FCF89F99D" box="[466,509,226,251]" italics="true" pageId="13" pageNumber="504">pv ′ζ</emphasis>
6–7). Tarsus III (
<figureCitation id="135E2A40FFA69F21FD07FF6FCD69F99D" box="[713,797,226,251]" captionStart="FIGURES 29 – 37" captionStartId="14.[151,264,1939,1962]" captionTargetBox="[253,1321,209,1908]" captionTargetId="figure@14.[234,1353,193,1935]" captionTargetPageId="14" captionText="FIGURES 29 – 37. Eotetranychus spinophilus sp. nov. male. (29) leg III, left side, dorsal to adaxial view; (30) leg IV, right side, abaxial view; (31) palp, dorsal view; (32 – 34) aedeagus, different aspects of same specimen; (35 – 37) aedeagus, different aspects of another specimen. Scale bar for aedeagi = 100 µm." httpUri="https://zenodo.org/record/998452/files/figure.png" pageId="13" pageNumber="504">Fig. 29</figureCitation>
) with single proximal solenidion (
<emphasis id="B911EAD7FFA69F21FB64FF6FCAB0F99D" box="[1194,1220,226,251]" italics="true" pageId="13" pageNumber="504">ω ′</emphasis>
18–21). Tarsus IV (
<figureCitation id="135E2A40FFA69F21FF51FE8BCE87F879" box="[159,243,262,287]" captionStart="FIGURES 29 – 37" captionStartId="14.[151,264,1939,1962]" captionTargetBox="[253,1321,209,1908]" captionTargetId="figure@14.[234,1353,193,1935]" captionTargetPageId="14" captionText="FIGURES 29 – 37. Eotetranychus spinophilus sp. nov. male. (29) leg III, left side, dorsal to adaxial view; (30) leg IV, right side, abaxial view; (31) palp, dorsal view; (32 – 34) aedeagus, different aspects of same specimen; (35 – 37) aedeagus, different aspects of another specimen. Scale bar for aedeagi = 100 µm." httpUri="https://zenodo.org/record/998452/files/figure.png" pageId="13" pageNumber="504">Fig. 30</figureCitation>
) with single proximal solenidion (
<emphasis id="B911EAD7FFA69F21FD4FFE8BCCEFF879" box="[641,667,262,287]" italics="true" pageId="13" pageNumber="504">ω ′</emphasis>
16–19). All leg setae finely barbed. Number of setae on legs I–IV respectively: trochanters 1, 1, 1, 1; femora 10, 7, 2, 2; genua 5, 5, 3, 3; tibiae 11 (3), 5, 5, 5; tarsi 15 (3+3), 13 (3+3), 8 (1+0), 8 (1+0). Leg setation as presented on 
<figureCitation id="135E2A40FFA69F21FD57FEC3CD4FF801" box="[665,827,334,359]" captionStart="FIGURES 22 – 28" captionStartId="13.[151,264,1943,1966]" captionTargetBox="[161,1418,656,1917]" captionTargetId="figure@13.[151,1436,639,1922]" captionTargetPageId="13" captionText="FIGURES 22 – 28. Eotetranychus spinophilus sp. nov. male. (22) leg I, right side, dorsal to abaxial view; (23 – 24) empodium I, different specimens and views; (25) ambulacrum I; (26) leg II, dorsal to adaxial view; (27) empodium III; (28) ambulacrum III, different specimen. Scale bar for ambulacra = 20 µm." httpUri="https://zenodo.org/record/998450/files/figure.png" pageId="13" pageNumber="504">Figures 22, 26</figureCitation>
, 
<figureCitation id="135E2A40FFA69F21FC89FEC2CDE4F801" box="[839,912,335,359]" captionStart="FIGURES 29 – 37" captionStartId="14.[151,264,1939,1962]" captionTargetBox="[253,1321,209,1908]" captionTargetId="figure@14.[234,1353,193,1935]" captionTargetPageId="14" captionText="FIGURES 29 – 37. Eotetranychus spinophilus sp. nov. male. (29) leg III, left side, dorsal to adaxial view; (30) leg IV, right side, abaxial view; (31) palp, dorsal view; (32 – 34) aedeagus, different aspects of same specimen; (35 – 37) aedeagus, different aspects of another specimen. Scale bar for aedeagi = 100 µm." httpUri="https://zenodo.org/record/998452/files/figure.png" pageId="13" pageNumber="504">29–30</figureCitation>
, with same chaetotaxy as female except for the presence of additional solenidia on certain segments. One specimen with three anomalies: lacking seta 
<emphasis id="B911EAD7FFA69F21FAD9FEFFCB5BF8ED" box="[1303,1327,370,395]" italics="true" pageId="13" pageNumber="504">v″</emphasis>
on femur I and seta 
<emphasis id="B911EAD7FFA69F21FEC7FE1BCF63F8C9" box="[265,279,406,431]" italics="true" pageId="13" pageNumber="504">d</emphasis>
on tibia II on left-hand side, and with symmetrical addition of two extra solenidia on tarsus I.
</paragraph>
<paragraph id="8BDA36C5FFA69F21FF09FE37CA2EFB2E" blockId="13.[151,1436,151,588]" pageId="13" pageNumber="504">
<emphasis id="B911EAD7FFA69F21FF09FE37CF17F8B5" bold="true" box="[199,355,442,467]" pageId="13" pageNumber="504">Deutonymph</emphasis>
. 
<emphasis id="B911EAD7FFA69F21FEBDFE36CF9BF8B4" box="[371,495,443,466]" italics="true" pageId="13" pageNumber="504">
<emphasis id="B911EAD7FFA69F21FEBDFE36CFA3F8B4" bold="true" box="[371,471,443,466]" italics="true" pageId="13" pageNumber="504">Dorsum.</emphasis>
S
</emphasis>
etal form and body striation similar to adult. Body measurements: 
<emphasis id="B911EAD7FFA69F21FB3FFE31CB5EF8B1" box="[1265,1322,442,471]" italics="true" pageId="13" pageNumber="504">
<date id="FFDB1005FFA69F21FB3FFE31CB72F8B1" box="[1265,1286,444,471]" pageId="13" pageNumber="504">v2</date>
–h1
</emphasis>
210–230, 
<emphasis id="B911EAD7FFA69F21FF59FE6FCE97F898" box="[151,227,482,510]" italics="true" pageId="13" pageNumber="504">sc2–sc2</emphasis>
110–135. Distances between setae: 
<emphasis id="B911EAD7FFA69F21FDB5FE6FCCC5F898" box="[635,689,482,510]" italics="true" pageId="13" pageNumber="504">c1–c1</emphasis>
46–48, 
<emphasis id="B911EAD7FFA69F21FCC3FE6FCD30F898" box="[781,836,482,510]" italics="true" pageId="13" pageNumber="504">c3–c3</emphasis>
175–200, 
<emphasis id="B911EAD7FFA69F21FC72FE6CCD81F898" box="[956,1013,481,510]" italics="true" pageId="13" pageNumber="504">d1–d1</emphasis>
37–41, 
<emphasis id="B911EAD7FFA69F21FB9CFE6CCAF7F89C" box="[1106,1155,481,510]" italics="true" pageId="13" pageNumber="504">d1–d</emphasis>
<date id="FFDB1005FFA69F21FB4DFE7DCAAFF89C" box="[1155,1243,482,510]" pageId="13" pageNumber="504" value="1928-02-31">
<emphasis id="B911EAD7FFA69F21FB4DFE7DCAFFF898" box="[1155,1163,496,510]" italics="true" pageId="13" pageNumber="504">2</emphasis>
28–31
</date>
, 
<emphasis id="B911EAD7FFA69F21FB29FE6FCB6AF898" box="[1255,1310,482,510]" italics="true" pageId="13" pageNumber="504">e1–e1</emphasis>
45–48, 
<emphasis id="B911EAD7FFA69F21FAB7FE6FCED7FB46" italics="true" pageId="13" pageNumber="504">e1– e</emphasis>
<date id="FFDB1005FFA69F21FF6DFD9ACE88FB47" box="[163,252,521,549]" pageId="13" pageNumber="504" value="1927-02-29">
<emphasis id="B911EAD7FFA69F21FF6DFD9ACEDFFB43" box="[163,171,535,549]" italics="true" pageId="13" pageNumber="504">2</emphasis>
27–29
</date>
, 
<emphasis id="B911EAD7FFA69F21FEC6FD84CF59FB47" box="[264,301,521,549]" italics="true" pageId="13" pageNumber="504">f1–f</emphasis>
<date id="FFDB1005FFA69F21FEE3FD9ACFF2FB47" box="[301,390,521,549]" pageId="13" pageNumber="504" value="1922-01-24">
<emphasis id="B911EAD7FFA69F21FEE3FD9ACF41FB43" box="[301,309,535,549]" italics="true" pageId="13" pageNumber="504">1</emphasis>
22–24
</date>
, 
<emphasis id="B911EAD7FFA69F21FE5CFD85CFB7FB47" box="[402,451,520,549]" italics="true" pageId="13" pageNumber="504">h1–h</emphasis>
<date id="FFDB1005FFA69F21FE0AFD9ACC55FB47" box="[452,545,521,549]" pageId="13" pageNumber="504" value="2013-01-14">
<emphasis id="B911EAD7FFA69F21FE0AFD9ACFB8FB43" box="[452,460,535,549]" italics="true" pageId="13" pageNumber="504">1</emphasis>
13–14.
</date>
Setal lengths: 
<date id="FFDB1005FFA69F21FD07FD84CD75FB47" box="[713,769,521,549]" pageId="13" pageNumber="504" value="1932-05-02">
<emphasis id="B911EAD7FFA69F21FD07FD84CCA9FB43" box="[713,733,521,549]" italics="true" pageId="13" pageNumber="504">v2</emphasis>
32
</date>
–40, 
<emphasis id="B911EAD7FFA69F21FCF4FD84CD2EFB43" box="[826,858,521,549]" italics="true" pageId="13" pageNumber="504">sc1</emphasis>
38–43, 
<emphasis id="B911EAD7FFA69F21FC78FD84CDA2FB43" box="[950,982,521,549]" italics="true" pageId="13" pageNumber="504">sc2</emphasis>
40–49, 
<emphasis id="B911EAD7FFA69F21FBFDFD84CA4BFB46" box="[1075,1087,521,544]" italics="true" pageId="13" pageNumber="504">c</emphasis>
<date id="FFDB1005FFA69F21FBF1FD9ACAECFB47" box="[1087,1176,521,549]" pageId="13" pageNumber="504" value="1924-01-29">
<emphasis id="B911EAD7FFA69F21FBF1FD9ACA33FB43" box="[1087,1095,535,549]" italics="true" pageId="13" pageNumber="504">1</emphasis>
24–29
</date>
, 
<emphasis id="B911EAD7FFA69F21FB6AFD84CACCFB43" box="[1188,1208,521,549]" italics="true" pageId="13" pageNumber="504">c2</emphasis>
42–43, 
<emphasis id="B911EAD7FFA69F21FADBFD84CB5DFB43" box="[1301,1321,521,549]" italics="true" pageId="13" pageNumber="504">c3</emphasis>
50–57, 
<emphasis id="B911EAD7FFA69F21FA48FD85CBE0FB47" box="[1414,1428,520,545]" italics="true" pageId="13" pageNumber="504">d</emphasis>
<date id="FFDB1005FFA69F21FA5AFD9ACEABFB2E" pageId="13" pageNumber="504" value="1935-01-28">
<emphasis id="B911EAD7FFA69F21FA5AFD9ACBE8FB43" box="[1428,1436,535,549]" italics="true" pageId="13" pageNumber="504">1</emphasis>
28–35
</date>
, 
<emphasis id="B911EAD7FFA69F21FF25FDA2CF75FB2A" box="[235,257,559,588]" italics="true" pageId="13" pageNumber="504">d2</emphasis>
39–47, 
<emphasis id="B911EAD7FFA69F21FE92FDBDCF1CFB21" box="[348,360,560,583]" italics="true" pageId="13" pageNumber="504">e</emphasis>
<date id="FFDB1005FFA69F21FEA6FDB3CFCBFB2E" box="[360,447,559,588]" pageId="13" pageNumber="504" value="1935-01-28">
<emphasis id="B911EAD7FFA69F21FEA6FDB3CF04FB2A" box="[360,368,574,588]" italics="true" pageId="13" pageNumber="504">1</emphasis>
28–35
</date>
, 
<emphasis id="B911EAD7FFA69F21FE05FDBDCFABFB2A" box="[459,479,560,588]" italics="true" pageId="13" pageNumber="504">e2</emphasis>
36–45, 
<emphasis id="B911EAD7FFA69F21FDF4FDA2CC36FB21" box="[570,578,559,583]" italics="true" pageId="13" pageNumber="504">f</emphasis>
<date id="FFDB1005FFA69F21FD8FFDB3CCEDFB2E" box="[577,665,559,588]" pageId="13" pageNumber="504" value="1936-01-31">
<emphasis id="B911EAD7FFA69F21FD8FFDB3CC3DFB2A" box="[577,585,574,588]" italics="true" pageId="13" pageNumber="504">1</emphasis>
31–36
</date>
, 
<emphasis id="B911EAD7FFA69F21FD6AFDA2CCC0FB2A" box="[676,692,559,588]" italics="true" pageId="13" pageNumber="504">f2</emphasis>
39–44, 
<emphasis id="B911EAD7FFA69F21FCC1FDA2CD50FB2A" box="[783,804,559,588]" italics="true" pageId="13" pageNumber="504">h1</emphasis>
42–45, 
<emphasis id="B911EAD7FFA69F21FCB1FDA2CDF9FB2E" box="[895,909,559,584]" italics="true" pageId="13" pageNumber="504">h</emphasis>
<date id="FFDB1005FFA69F21FC43FDB3CD90FB21" box="[909,996,559,588]" pageId="13" pageNumber="504" value="1928-02-31">
<emphasis id="B911EAD7FFA69F21FC43FDB3CDE1FB2A" box="[909,917,574,588]" italics="true" pageId="13" pageNumber="504">2</emphasis>
28–31
</date>
, 
<emphasis id="B911EAD7FFA69F21FC3EFDA2CD8AFB2E" box="[1008,1022,559,584]" italics="true" pageId="13" pageNumber="504">h</emphasis>
<date id="FFDB1005FFA69F21FC30FDB3CA2EFB2E" box="[1022,1114,559,588]" pageId="13" pageNumber="504" value="1926-03-30">
<emphasis id="B911EAD7FFA69F21FC30FDB3CA72FB2A" box="[1022,1030,574,588]" italics="true" pageId="13" pageNumber="504">3</emphasis>
26–30.
</date>
</paragraph>
<caption id="DF1A664DFFA69F21FF59F81ACCC0FE8D" httpUri="https://zenodo.org/record/998450/files/figure.png" pageId="13" pageNumber="504" targetBox="[161,1418,656,1917]" targetPageId="13">
<paragraph id="8BDA36C5FFA69F21FF59F81ACCC0FE8D" blockId="13.[151,1436,1943,2027]" pageId="13" pageNumber="504">
<emphasis id="B911EAD7FFA69F21FF59F81ACF20FECB" bold="true" box="[151,340,1943,1966]" pageId="13" pageNumber="504">FIGURES 22–28.</emphasis>
<taxonomicName id="4C654D46FFA69F21FE97F815CC13FEC8" authority="Zhang, 2017" authorityName="Zhang" authorityYear="2017" box="[345,615,1944,1966]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="504" phylum="Arthropoda" rank="species" species="spinophilus" status="sp. nov.">
<emphasis id="B911EAD7FFA69F21FE97F815CC13FEC8" box="[345,615,1944,1966]" italics="true" pageId="13" pageNumber="504">Eotetranychus spinophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257ACFFA69F21FDA3F815CCC9FECB" box="[621,701,1944,1965]" pageId="13" pageNumber="504" rank="species">
<emphasis id="B911EAD7FFA69F21FDA3F815CCC9FECB" bold="true" box="[621,701,1944,1965]" pageId="13" pageNumber="504">sp. nov.</emphasis>
</taxonomicNameLabel>
male. (22) leg I, right side, dorsal to abaxial view; (23–24) empodium I, different specimens and views; (25) ambulacrum I; (26) leg II, dorsal to adaxial view; (27) empodium III; (28) ambulacrum III, different specimen. Scale bar for ambulacra = 20 µm.
</paragraph>
</caption>
<caption id="DF1A664DFFA59F22FF59F81ECCC1FE81" httpUri="https://zenodo.org/record/998452/files/figure.png" pageId="14" pageNumber="505" targetBox="[253,1321,209,1908]" targetPageId="14">
<paragraph id="8BDA36C5FFA59F22FF59F81ECCC1FE81" blockId="14.[151,1436,1939,2023]" pageId="14" pageNumber="505">
<emphasis id="B911EAD7FFA59F22FF59F81ECF27FECF" bold="true" box="[151,339,1939,1962]" pageId="14" pageNumber="505">FIGURES 29–37.</emphasis>
<taxonomicName id="4C654D46FFA59F22FE96F819CC11FECC" authority="Zhang, 2017" authorityName="Zhang" authorityYear="2017" box="[344,613,1940,1962]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="505" phylum="Arthropoda" rank="species" species="spinophilus" status="sp. nov.">
<emphasis id="B911EAD7FFA59F22FE96F819CC11FECC" box="[344,613,1940,1962]" italics="true" pageId="14" pageNumber="505">Eotetranychus spinophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257ACFFA59F22FDA5F819CCCFFECF" box="[619,699,1940,1961]" pageId="14" pageNumber="505" rank="species">
<emphasis id="B911EAD7FFA59F22FDA5F819CCCFFECF" bold="true" box="[619,699,1940,1961]" pageId="14" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
male. (29) leg III, left side, dorsal to adaxial view; (30) leg IV, right side, abaxial view; (31) palp, dorsal view; (32–34) aedeagus, different aspects of same specimen; (35–37) aedeagus, different aspects of another specimen. Scale bar for aedeagi = 100 µm.
</paragraph>
</caption>
<caption id="DF1A664DFFA49F23FF59F8DDCAFDFEA5" httpUri="https://zenodo.org/record/998454/files/figure.png" pageId="15" pageNumber="506" targetBox="[195,1383,281,1840]" targetPageId="15">
<paragraph id="8BDA36C5FFA49F23FF59F8DDCAFDFEA5" blockId="15.[151,1436,1872,1987]" pageId="15" pageNumber="506">
<emphasis id="B911EAD7FFA49F23FF59F8DDCF20FE00" bold="true" box="[151,340,1872,1895]" pageId="15" pageNumber="506">FIGURES 38–39.</emphasis>
<taxonomicName id="4C654D46FFA49F23FE94F8DCCCB6FE01" authority="(Hermann)" authorityName="Hermann" baseAuthorityName="Hermann" baseAuthorityYear="1804" box="[346,706,1873,1895]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="506" phylum="Arthropoda" rank="species" species="tiliarium">
<emphasis id="B911EAD7FFA49F23FE94F8DCCC38FE01" box="[346,588,1873,1895]" italics="true" pageId="15" pageNumber="506">Eotetranychus tiliarium</emphasis>
(Hermann)
</taxonomicName>
female. (38) leg I, left side, dorsal to adaxial view. (39) leg II, left side, dorsal to adaxial view. Leg setae in red (and marked with an asterisk *) were sometimes absent—as an anomaly—in a survey of nine species of 
<taxonomicName id="4C654D46FFA49F23FEFBF803CFB3FEC2" baseAuthorityName="Pritchard &amp; Baker" baseAuthorityYear="1955" box="[309,455,1934,1956]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAD7FFA49F23FEFBF803CFB3FEC2" box="[309,455,1934,1956]" italics="true" pageId="15" pageNumber="506">Eotetranychus</emphasis>
</taxonomicName>
. Ontogeny (Ad = adult; DN = deutonymph; PN = protonymph; unlabelled = larva or, on leg IV, protonymph) determined from Lindquist (1985) and examination of other species of 
<taxonomicName id="4C654D46FFA49F23FC3FF820CAF7FEA5" baseAuthorityName="Pritchard &amp; Baker" baseAuthorityYear="1955" box="[1009,1155,1965,1987]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAD7FFA49F23FC3FF820CAF7FEA5" box="[1009,1155,1965,1987]" italics="true" pageId="15" pageNumber="506">Eotetranychus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF1A664DFFBB9F3CFF59F9A4CF3EFFDC" httpUri="https://zenodo.org/record/998456/files/figure.png" pageId="16" pageNumber="507" targetBox="[159,1428,246,1477]" targetPageId="16">
<paragraph id="8BDA36C5FFBB9F3CFF59F9A4CF3EFFDC" blockId="16.[151,1436,1577,1722]" pageId="16" pageNumber="507">
<emphasis id="B911EAD7FFBB9F3CFF59F9A4CF20FF59" bold="true" box="[151,340,1577,1600]" pageId="16" pageNumber="507">FIGURES 40–41.</emphasis>
<taxonomicName id="4C654D46FFBB9F3CFE95F9A7CCB6FF26" authority="(Hermann)" authorityName="Hermann" baseAuthorityName="Hermann" baseAuthorityYear="1804" box="[347,706,1578,1600]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="507" phylum="Arthropoda" rank="species" species="tiliarium">
<emphasis id="B911EAD7FFBB9F3CFE95F9A7CC39FF26" box="[347,589,1578,1600]" italics="true" pageId="16" pageNumber="507">Eotetranychus tiliarium</emphasis>
(Hermann)
</taxonomicName>
female. (40) leg III, right side, dorsal to abaxial view. (41) leg IV, right side, dorsal to abaxial view. Leg setae in red (and marked with an asterisk *) were sometimes absent—as an anomaly from a species’ typical leg setation—in a survey of nine species of 
<taxonomicName id="4C654D46FFBB9F3CFCCDF9EACDE1FF1B" baseAuthorityName="Pritchard &amp; Baker" baseAuthorityYear="1955" box="[771,917,1639,1661]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="507" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAD7FFBB9F3CFCCDF9EACDE1FF1B" box="[771,917,1639,1661]" italics="true" pageId="16" pageNumber="507">Eotetranychus</emphasis>
</taxonomicName>
. Ontogeny (Ad = adult; DN = deutonymph; PN = protonymph; unlabelled = larva or, on leg IV, protonymph) determined from Lindquist (1985) and examination of other species of 
<taxonomicName id="4C654D46FFBB9F3CFF7CF929CF30FFDC" baseAuthorityName="Pritchard &amp; Baker" baseAuthorityYear="1955" box="[178,324,1700,1722]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="507" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAD7FFBB9F3CFF7CF929CF30FFDC" box="[178,324,1700,1722]" italics="true" pageId="16" pageNumber="507">Eotetranychus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BDA36C5FFBB9F3CFF09F96FCC38FE23" blockId="16.[151,1437,1762,2008]" pageId="16" pageNumber="507">
<emphasis id="B911EAD7FFBB9F3CFF09F96FCF64FF9F" bold="true" box="[199,272,1762,1785]" italics="true" pageId="16" pageNumber="507">Venter</emphasis>
. Striation similar to adult, except longitudinal–oblique striae from aggenital setae (
<emphasis id="B911EAD7FFBB9F3CFB05F96ECA93FF9C" box="[1227,1255,1763,1786]" italics="true" pageId="16" pageNumber="507">ag</emphasis>
) to setae 
<emphasis id="B911EAD7FFBB9F3CFA97F96FCB1BFF99" box="[1369,1391,1762,1791]" italics="true" pageId="16" pageNumber="507">h2</emphasis>
, 
<emphasis id="B911EAD7FFBB9F3CFAB1F96FCBE1FF99" box="[1407,1429,1762,1791]" italics="true" pageId="16" pageNumber="507">h3</emphasis>
. Setal lengths: 
<emphasis id="B911EAD7FFBB9F3CFEF6F884CF27FE47" box="[312,339,1801,1825]" italics="true" pageId="16" pageNumber="507">1a</emphasis>
22–24, 
<emphasis id="B911EAD7FFBB9F3CFE7DF885CFBBFE47" box="[435,463,1800,1825]" italics="true" pageId="16" pageNumber="507">1b</emphasis>
22–25, 
<emphasis id="B911EAD7FFBB9F3CFDE2F884CC33FE47" box="[556,583,1801,1825]" italics="true" pageId="16" pageNumber="507">1c</emphasis>
39–40, 
<emphasis id="B911EAD7FFBB9F3CFD6BF885CCB5FE47" box="[677,705,1800,1825]" italics="true" pageId="16" pageNumber="507">2b</emphasis>
23–30, 
<emphasis id="B911EAD7FFBB9F3CFCD1F884CD4EFE47" box="[799,826,1801,1825]" italics="true" pageId="16" pageNumber="507">2c</emphasis>
41–45, 
<emphasis id="B911EAD7FFBB9F3CFC56F884CDC7FE47" box="[920,947,1801,1825]" italics="true" pageId="16" pageNumber="507">3a</emphasis>
13–17, 
<emphasis id="B911EAD7FFBB9F3CFBDCF885CA5AFE47" box="[1042,1070,1800,1825]" italics="true" pageId="16" pageNumber="507">3b</emphasis>
27–38, 
<emphasis id="B911EAD7FFBB9F3CFB42F884CADCFE47" box="[1164,1192,1801,1825]" italics="true" pageId="16" pageNumber="507">4a</emphasis>
17–19, 
<emphasis id="B911EAD7FFBB9F3CFAC8F885CB56FE47" box="[1286,1314,1800,1825]" italics="true" pageId="16" pageNumber="507">4b</emphasis>
25–27, 
<emphasis id="B911EAD7FFBB9F3CFA4EF887CBE8FE47" box="[1408,1436,1802,1825]" italics="true" pageId="16" pageNumber="507">ag</emphasis>
20–24, 
<emphasis id="B911EAD7FFBB9F3CFF25F8A3CE8DFE23" box="[235,249,1838,1861]" italics="true" pageId="16" pageNumber="507">g</emphasis>
<date id="FFDB1005FFBB9F3CFF37F8B1CF24FE23" box="[249,336,1837,1866]" pageId="16" pageNumber="507" value="1917-01-19">
<emphasis id="B911EAD7FFBB9F3CFF37F8B1CF75FE2C" box="[249,257,1852,1866]" italics="true" pageId="16" pageNumber="507">1</emphasis>
17–19
</date>
, 
<emphasis id="B911EAD7FFBB9F3CFE92F8A3CF01FE23" box="[348,373,1838,1861]" italics="true" pageId="16" pageNumber="507">ps</emphasis>
<date id="FFDB1005FFBB9F3CFEBAF8B1CFBFFE23" box="[372,459,1837,1866]" pageId="16" pageNumber="507" value="1917-01-18">
<emphasis id="B911EAD7FFBB9F3CFEBAF8B1CF08FE2C" box="[372,380,1852,1866]" italics="true" pageId="16" pageNumber="507">1</emphasis>
17–18
</date>
, 
<emphasis id="B911EAD7FFBB9F3CFE19F8A3CF84FE23" box="[471,496,1838,1861]" italics="true" pageId="16" pageNumber="507">ps</emphasis>
<date id="FFDB1005FFBB9F3CFE3EF8B1CC38FE23" box="[496,588,1837,1866]" pageId="16" pageNumber="507" value="1917-02-20">
<emphasis id="B911EAD7FFBB9F3CFE3EF8B1CF8CFE2C" box="[496,504,1852,1866]" italics="true" pageId="16" pageNumber="507">2</emphasis>
17–20.
</date>
</paragraph>
<paragraph id="8BDA36C5FFBB9F3CFF09F8DECC47FE0A" blockId="16.[151,1437,1762,2008]" box="[199,563,1875,1900]" pageId="16" pageNumber="507">
<taxonomicName id="4C654D46FFBB9F3CFF09F8DECF2FFE0A" authorityName="Krefft" authorityYear="1871" box="[199,347,1875,1900]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="16" pageNumber="507" phylum="Nematoda" rank="genus">
<emphasis id="B911EAD7FFBB9F3CFF09F8DECF2FFE0A" bold="true" box="[199,347,1875,1900]" italics="true" pageId="16" pageNumber="507">Gnathosoma</emphasis>
</taxonomicName>
. Similar to female.
</paragraph>
<paragraph id="8BDA36C5FFBB9F3CFF09F8F5CAA0FED5" blockId="16.[151,1437,1762,2008]" pageId="16" pageNumber="507">
<emphasis id="B911EAD7FFBB9F3CFF09F8F5CE89FEE9" bold="true" box="[199,253,1912,1935]" italics="true" pageId="16" pageNumber="507">Legs</emphasis>
. Leg striation and form of setae similar to female. Number of setae on legs I–IV respectively: trochanters 1, 1, 1, 0; femora 6, 3, 2, 2; genua 5, 5, 3, 2; tibiae 8 (1), 5, 5, 5; tarsi 15 (3+3), 11 (1+3), 8 (1), 7.
</paragraph>
<paragraph id="8BDA36C5FFBB9F3DFF09F832CF6FF8C9" blockId="16.[151,1437,1762,2008]" lastBlockId="17.[151,1437,151,1154]" lastPageId="17" lastPageNumber="508" pageId="16" pageNumber="507">
<emphasis id="B911EAD7FFBB9F3CFF09F832CC1CFEBE" bold="true" box="[199,616,1983,2008]" italics="true" pageId="16" pageNumber="507">Ontogenetic changes in leg setation</emphasis>
. Without having a larva or protonymph of 
<taxonomicName id="4C654D46FFBB9F3CFB9BF84CCA83FEBE" authorityName="Zhang" authorityYear="2017" box="[1109,1271,1983,2008]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="507" phylum="Arthropoda" rank="species" species="spinophilus">
<emphasis id="B911EAD7FFBB9F3CFB9BF84CCA12FEBE" box="[1109,1126,1985,2008]" italics="true" pageId="16" pageNumber="507">E</emphasis>
. 
<emphasis id="B911EAD7FFBB9F3CFBB8F832CA83FEBE" box="[1142,1271,1983,2008]" italics="true" pageId="16" pageNumber="507">spinophilus</emphasis>
</taxonomicName>
to examine, it was difficult to determine whether the setae we have labelled as 
<emphasis id="B911EAD7FFBA9F3DFCBCFF15CDEDF9C9" box="[882,921,152,175]" italics="true" pageId="17" pageNumber="508">pv&quot;</emphasis>
on both tarsus I and II are indeed 
<emphasis id="B911EAD7FFBA9F3DFAEEFF15CB33F9C9" box="[1312,1351,152,175]" italics="true" pageId="17" pageNumber="508">pv&quot;</emphasis>
or if in fact they are 
<emphasis id="B911EAD7FFBA9F3DFEF9FF30CF2CF9BF" box="[311,344,189,217]" italics="true" pageId="17" pageNumber="508">v&quot;1</emphasis>
and 
<emphasis id="B911EAD7FFBA9F3DFE56FF30CFC6F9BF" box="[408,434,189,217]" italics="true" pageId="17" pageNumber="508">v'1</emphasis>
respectively, which are normally added in the deutonymph. However, by examining different developmental stages of other species, we are confident the setae are 
<emphasis id="B911EAD7FFBA9F3DFBC1FF6ECA40F99C" box="[1039,1076,227,250]" italics="true" pageId="17" pageNumber="508">pv&quot;</emphasis>
. Tarsus III should have 9 (1+0) setae (
<quantity id="4C9D9B20FFBA9F3DFF2EFE8ACF68F879" box="[224,284,262,287]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="17" pageNumber="508" unit="in" value="10.0">10 in</quantity>
adult) and tarsus IV should have 8 setae (
<quantity id="4C9D9B20FFBA9F3DFCCCFE8ACD48F879" box="[770,828,262,287]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="17" pageNumber="508" unit="in" value="11.0">11 in</quantity>
adult) in the deutonymph; however 
<taxonomicName id="4C654D46FFBA9F3DFB10FE85CBF4F879" authorityName="Zhang" authorityYear="2017" box="[1246,1408,262,287]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="species" species="spinophilus">
<emphasis id="B911EAD7FFBA9F3DFB10FE85CA9BF879" box="[1246,1263,264,287]" italics="true" pageId="17" pageNumber="508">E</emphasis>
. 
<emphasis id="B911EAD7FFBA9F3DFB31FE8BCBF4F879" box="[1279,1408,262,287]" italics="true" pageId="17" pageNumber="508">spinophilus</emphasis>
</taxonomicName>
is missing the same seta on each of these segments, 
<emphasis id="B911EAD7FFBA9F3DFD16FEA6CC8AF824" box="[728,766,299,322]" italics="true" pageId="17" pageNumber="508">pv&quot;</emphasis>
, and consequently seta 
<emphasis id="B911EAD7FFBA9F3DFBDCFEA6CA4CF824" box="[1042,1080,299,322]" italics="true" pageId="17" pageNumber="508">pv&quot;</emphasis>
is missing on tarsi III–IV. The absence of setae 
<emphasis id="B911EAD7FFBA9F3DFE98FEC2CF0FF800" box="[342,379,335,358]" italics="true" pageId="17" pageNumber="508">pv&quot;</emphasis>
is unusual, as larval setae are normally present. This seta is also absent in 
<taxonomicName id="4C654D46FFBA9F3DFB02FEC3CB1CF801" authorityName="Baker &amp; Pritchard" authorityYear="1953" box="[1228,1384,334,359]" class="Arachnida" family="Tuckerellidae" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="family">Tuckerellidae</taxonomicName>
, and both primiventrals are absent in 
<taxonomicName id="4C654D46FFBA9F3DFDCCFEFFCCD4F8ED" authorityName="Berlese" authorityYear="1913" box="[514,672,370,395]" class="Arachnida" family="Tenuipalpidae" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="family">Tenuipalpidae</taxonomicName>
and 
<taxonomicName id="4C654D46FFBA9F3DFD16FEFFCD0BF8ED" authorityName="Baker &amp; Pritchard" authorityYear="1953" box="[728,895,370,395]" class="Arachnida" family="Linotetranidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="family">Linotetranidae</taxonomicName>
. For additional comments, see section in female description.
</paragraph>
<paragraph id="8BDA36C5FFBA9F3DFF09FE37CC12F8B5" blockId="17.[151,1437,151,1154]" box="[199,614,442,467]" pageId="17" pageNumber="508">
<emphasis id="B911EAD7FFBA9F3DFF09FE37CF9CF8B5" bold="true" box="[199,488,442,467]" pageId="17" pageNumber="508">Protonymph and Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37F654EFFBA9F3DFF09FE53CCBBFB90" pageId="17" pageNumber="508" type="diagnosis">
<paragraph id="8BDA36C5FFBA9F3DFF09FE53CCBBFB90" blockId="17.[151,1437,151,1154]" pageId="17" pageNumber="508">
<emphasis id="B911EAD7FFBA9F3DFF09FE53CFBAF891" bold="true" box="[199,462,478,503]" pageId="17" pageNumber="508">Differential diagnosis.</emphasis>
Few species of 
<taxonomicName id="4C654D46FFBA9F3DFD4DFE53CD51F891" baseAuthorityName="Pritchard &amp; Baker" baseAuthorityYear="1955" box="[643,805,478,503]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAD7FFBA9F3DFD4DFE53CD51F891" box="[643,805,478,503]" italics="true" pageId="17" pageNumber="508">Eotetranychus</emphasis>
</taxonomicName>
are like 
<taxonomicName id="4C654D46FFBA9F3DFC48FE6DCA53F891" authorityName="Zhang" authorityYear="2017" box="[902,1063,478,503]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="species" species="spinophilus">
<emphasis id="B911EAD7FFBA9F3DFC48FE6DCDE3F891" box="[902,919,480,503]" italics="true" pageId="17" pageNumber="508">E</emphasis>
. 
<emphasis id="B911EAD7FFBA9F3DFC68FE53CA53F891" box="[934,1063,478,503]" italics="true" pageId="17" pageNumber="508">spinophilus</emphasis>
</taxonomicName>
. The species 
<taxonomicName id="4C654D46FFBA9F3DFB0EFE6DCF6AFB7D" authority="Baker &amp; Tuttle, 1994" authorityName="Baker &amp; Tuttle" authorityYear="1994" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="species" species="thewkei">
<emphasis id="B911EAD7FFBA9F3DFB0EFE6DCAA5F891" box="[1216,1233,480,503]" italics="true" pageId="17" pageNumber="508">E</emphasis>
. 
<emphasis id="B911EAD7FFBA9F3DFB11FE53CB40F891" box="[1247,1332,478,503]" italics="true" pageId="17" pageNumber="508">thewkei</emphasis>
<bibRefCitation id="EFF44B34FFBA9F3DFAF2FE53CF6AFB7D" author="Baker" pageId="17" pageNumber="508" refString="Baker, E. W. &amp; Tuttle, D. M. (1994) A Guide to the Spider Mites (Tetranychidae) of the United States. Indira Publishing House, West Bloomfield, 347 pp." type="book" year="1994">Baker &amp; Tuttle, 1994</bibRefCitation>
</taxonomicName>
shares the short dorsal setae and a reduced leg setal chaetome (e.g. genu III with three setae, tibia II–III with five setae) with 
<taxonomicName id="4C654D46FFBA9F3DFE5BFDA5CC4EFB59" authorityName="Zhang" authorityYear="2017" box="[405,570,550,575]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="species" species="spinophilus">
<emphasis id="B911EAD7FFBA9F3DFE5BFDA5CFD2FB59" box="[405,422,552,575]" italics="true" pageId="17" pageNumber="508">E</emphasis>
. 
<emphasis id="B911EAD7FFBA9F3DFE77FDABCC4EFB59" box="[441,570,550,575]" italics="true" pageId="17" pageNumber="508">spinophilus</emphasis>
</taxonomicName>
, but has a peritreme terminating in a small bulb, transverse striae on the prodorsum, and much shorter dorsocentral setae 
<emphasis id="B911EAD7FFBA9F3DFD71FDC1CCA7FB01" box="[703,723,588,615]" italics="true" pageId="17" pageNumber="508">c1</emphasis>
, 
<emphasis id="B911EAD7FFBA9F3DFD2CFDC7CC8CFB01" box="[738,760,586,615]" italics="true" pageId="17" pageNumber="508">d1</emphasis>
and 
<emphasis id="B911EAD7FFBA9F3DFCFEFDC1CD31FB01" box="[816,837,588,615]" italics="true" pageId="17" pageNumber="508">e1</emphasis>
. As with the new species, the host of 
<taxonomicName id="4C654D46FFBA9F3DFB39FDC1CB18FB05" authorityName="Baker &amp; Tuttle" authorityYear="1994" box="[1271,1388,586,611]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="species" species="thewkei">
<emphasis id="B911EAD7FFBA9F3DFB39FDC1CB7CFB05" box="[1271,1288,588,611]" italics="true" pageId="17" pageNumber="508">E</emphasis>
. 
<emphasis id="B911EAD7FFBA9F3DFAD9FDC7CB18FB05" box="[1303,1388,586,611]" italics="true" pageId="17" pageNumber="508">thewkei</emphasis>
</taxonomicName>
is a grass, 
<taxonomicName id="4C654D46FFBA9F3DFF11FDFCCC3DFBEC" box="[223,585,625,650]" class="Liliopsida" family="Poaceae" genus="Aristida" kingdom="Plantae" order="Poales" pageId="17" pageNumber="508" phylum="Tracheophyta" rank="species" species="adscensionis">
<emphasis id="B911EAD7FFBA9F3DFF11FDFCCFA4FBEC" box="[223,464,625,650]" italics="true" pageId="17" pageNumber="508">Aristida adscensionis</emphasis>
(Poaceae)
</taxonomicName>
. The only other species of 
<taxonomicName id="4C654D46FFBA9F3DFCB7FDFCCA6FFBEC" baseAuthorityName="Pritchard &amp; Baker" baseAuthorityYear="1955" box="[889,1051,625,650]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAD7FFBA9F3DFCB7FDFCCA6FFBEC" box="[889,1051,625,650]" italics="true" pageId="17" pageNumber="508">Eotetranychus</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C654D46FFBA9F3DFBAEFDFFCAC8FBEF" authorityName="" authorityYear="1895" box="[1120,1212,626,649]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="17" pageNumber="508" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
are 
<taxonomicName id="4C654D46FFBA9F3DFB23FDFFCF45FBC8" authority="(Reck, 1956)" authorityName="Reck" authorityYear="1956" baseAuthorityName="Reck" baseAuthorityYear="1956" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="species" species="pomeranzevi">
<emphasis id="B911EAD7FFBA9F3DFB23FDFFCA8AFBEF" box="[1261,1278,626,649]" italics="true" pageId="17" pageNumber="508">E</emphasis>
. 
<emphasis id="B911EAD7FFBA9F3DFAC3FDFFCBE8FBEF" box="[1293,1436,626,649]" italics="true" pageId="17" pageNumber="508">pomeranzevi</emphasis>
(
<bibRefCitation id="EFF44B34FFBA9F3DFF50FD18CF5DFBC8" author="Reck" box="[158,297,661,686]" pageId="17" pageNumber="508" refString="Reck, G. F. (1956) Novye vidy tetranihovyh klescej iz Vostocnoj Gruzii. Trudy Instituta Zoologyi Akademyi Nauk Gruzinskoi S. S. R, 15, 5 - 28." type="journal article" year="1956">Reck, 1956</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C654D46FFBA9F3DFE8AFD1BCCFFFBC8" authority="Wang, 1980" authorityName="Wang" authorityYear="1980" box="[324,651,661,686]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="species" species="qinlingensis">
<emphasis id="B911EAD7FFBA9F3DFE8AFD1BCF21FBCB" box="[324,341,662,685]" italics="true" pageId="17" pageNumber="508">E</emphasis>
. 
<emphasis id="B911EAD7FFBA9F3DFEA7FD18CF84FBC8" box="[361,496,661,686]" italics="true" pageId="17" pageNumber="508">qinlingensis</emphasis>
<bibRefCitation id="EFF44B34FFBA9F3DFE33FD1BCCFFFBC8" author="Wang" box="[509,651,662,686]" pageId="17" pageNumber="508" refString="Wang, H. F. (1980) Notes on the genus Eotetranychus in China with descriptions of four new species (Acarina: Tetranychidae). Acta Entomologica Sinica, 23, 216 - 223." type="journal article" year="1980">Wang, 1980</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C654D46FFBA9F3DFD03FD1BCA62FBC8" authority="Gutierrez, 1967" authorityName="Gutierrez" authorityYear="1967" box="[717,1046,661,686]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="species" species="roederei">
<emphasis id="B911EAD7FFBA9F3DFD03FD1BCCAAFBCB" box="[717,734,662,685]" italics="true" pageId="17" pageNumber="508">E</emphasis>
. 
<emphasis id="B911EAD7FFBA9F3DFD3DFD18CD3BFBC8" box="[755,847,661,686]" italics="true" pageId="17" pageNumber="508">roederei</emphasis>
<bibRefCitation id="EFF44B34FFBA9F3DFC93FD18CA62FBC8" author="Gutierrez" box="[861,1046,661,686]" pageId="17" pageNumber="508" refString="Gutierrez, J. (1967) Huit nouvelles especes du genre Eotetranychus Oudemans (Acariens: Tetranychidae) de Madagascar. Acarologia, 9, 370 - 394." type="journal article" year="1967">Gutierrez, 1967</bibRefCitation>
</taxonomicName>
, none of which share the above mentioned features of 
<taxonomicName id="4C654D46FFBA9F3DFE53FD37CC35FBB4" authorityName="Zhang" authorityYear="2017" box="[413,577,697,722]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="species" species="spinophilus">
<emphasis id="B911EAD7FFBA9F3DFE53FD37CFDAFBB7" box="[413,430,698,721]" italics="true" pageId="17" pageNumber="508">E</emphasis>
. 
<emphasis id="B911EAD7FFBA9F3DFE0EFD34CC35FBB4" box="[448,577,697,722]" italics="true" pageId="17" pageNumber="508">spinophilus</emphasis>
</taxonomicName>
, and all of which are more typical 
<taxonomicName id="4C654D46FFBA9F3DFC26FD34CAFEFBB4" baseAuthorityName="Pritchard &amp; Baker" baseAuthorityYear="1955" box="[1000,1162,697,722]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAD7FFBA9F3DFC26FD34CAFEFBB4" box="[1000,1162,697,722]" italics="true" pageId="17" pageNumber="508">Eotetranychus</emphasis>
</taxonomicName>
with long dorsal setae, complete setal leg chaetome and linear peritremes.
</paragraph>
</subSubSection>
<subSubSection id="C37F654EFFBA9F3DFF09FC8CCC39FD5C" pageId="17" pageNumber="508" type="discussion">
<paragraph id="8BDA36C5FFBA9F3DFF09FC8CCC39FD5C" blockId="17.[151,1437,151,1154]" pageId="17" pageNumber="508">
<emphasis id="B911EAD7FFBA9F3DFF09FC8CCF4FFA7C" bold="true" box="[199,315,769,794]" pageId="17" pageNumber="508">Remarks.</emphasis>
The tribe of grasses, 
<taxonomicName id="4C654D46FFBA9F3DFDFBFC8CCCD6FA7C" box="[565,674,769,794]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="17" pageNumber="508" phylum="Tracheophyta" rank="tribe" tribe="Triodieae">Triodieae</taxonomicName>
, known as spinifex grasses, is one of the most characteristic of all Australian arid zone plant groups. Spinifex forms a dominant vegetation community known as ‘hummock grassland’ on over 22% of the continent of 
<collectingCountry id="F3727655FFBA9F3DFD6BFCC4CD7AFA04" box="[677,782,841,866]" name="Australia" pageId="17" pageNumber="508">Australia</collectingCountry>
, with vast tracts of central and north-western 
<collectingCountry id="F3727655FFBA9F3DFAFBFCC4CBE8FA04" box="[1333,1436,841,866]" name="Australia" pageId="17" pageNumber="508">Australia</collectingCountry>
dominated by spinifex grasses. Hummock grassland is the most widely distributed biome on the Australian continent (Dickman 
<emphasis id="B911EAD7FFBA9F3DFE4EFC1FCFC5FACC" box="[384,433,913,938]" italics="true" pageId="17" pageNumber="508">et al</emphasis>
., 
<emphasis id="B911EAD7FFBA9F3DFE07FC1FCFA3FACC" box="[457,471,914,938]" italics="true" pageId="17" pageNumber="508">2</emphasis>
014), and this group of grasses is of key importance to the ecology of arid and semiarid 
<collectingCountry id="F3727655FFBA9F3DFF01FC38CF41FAA8" box="[207,309,949,974]" name="Australia" pageId="17" pageNumber="508">Australia</collectingCountry>
(
<bibRefCitation id="EFF44B34FFBA9F3DFE84FC38CCA5FAA8" author="Griffin" box="[330,721,949,974]" pageId="17" pageNumber="508" refString="Griffin, G. F. (1992) Will it burn-should it burn?: management of the Spinifex grasslands of inland Australia. In: Chapman, G. P. (Ed.), Desertified Grasslands: Their Biology and Management. Academic Press, London, pp. 63 - 76." type="book chapter" year="1992">
Griffin 1992; Beard 
<emphasis id="B911EAD7FFBA9F3DFD8DFC3BCC0EFAA8" box="[579,634,949,974]" italics="true" pageId="17" pageNumber="508">et al</emphasis>
., 
<emphasis id="B911EAD7FFBA9F3DFD5BFC3BCCD7FAA8" box="[661,675,950,974]" italics="true" pageId="17" pageNumber="508">2</emphasis>
013
</bibRefCitation>
). The new spider mite species 
<taxonomicName id="4C654D46FFBA9F3DFB82FC3BCA86FAA8" authorityName="Zhang" authorityYear="2017" box="[1100,1266,949,974]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="species" species="spinophilus">
<emphasis id="B911EAD7FFBA9F3DFB82FC3BCA29FAAB" box="[1100,1117,950,973]" italics="true" pageId="17" pageNumber="508">E</emphasis>
. 
<emphasis id="B911EAD7FFBA9F3DFBBFFC38CA86FAA8" box="[1137,1266,949,974]" italics="true" pageId="17" pageNumber="508">spinophilus</emphasis>
</taxonomicName>
was found in association with four other species of mites (
<taxonomicName id="4C654D46FFBA9F3DFD05FC54CD2DFA94" authorityName="Nalepa" authorityYear="1898" box="[715,857,985,1010]" class="Arachnida" family="Eriophyidae" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="family">Eriophyidae</taxonomicName>
, 
<taxonomicName id="4C654D46FFBA9F3DFCA2FC54CD8AFA94" authorityName="Berlese" authorityYear="1916" box="[876,1022,985,1010]" class="Arachnida" family="Phytoseiidae" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="family">Phytoseiidae</taxonomicName>
, 
<taxonomicName id="4C654D46FFBA9F3DFBDCFC54CA0FFA94" authorityName="Kramer" authorityYear="1877" box="[1042,1147,985,1010]" class="Arachnida" family="Tydeidae" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="family">Tydeidae</taxonomicName>
, and a new species of 
<taxonomicName id="4C654D46FFBA9F3DFF59FC70CF00FD70" authorityName="Sayed" authorityYear="1938" box="[151,372,1021,1046]" class="Arachnida" family="Tenuipalpidae" genus="Dolichotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAD7FFBA9F3DFF59FC70CF00FD70" box="[151,372,1021,1046]" italics="true" pageId="17" pageNumber="508">Dolichotetranychus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C654D46FFBA9F3DFE47FC70CC5FFD70" authorityName="Berlese" authorityYear="1913" box="[393,555,1021,1046]" class="Arachnida" family="Tenuipalpidae" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="family">Tenuipalpidae</taxonomicName>
)) and a species of thrips (
<taxonomicName id="4C654D46FFBA9F3DFCBAFC70CD93FD70" box="[884,999,1021,1046]" class="Insecta" family="Thripidae" kingdom="Animalia" order="Thysanoptera" pageId="17" pageNumber="508" phylum="Arthropoda" rank="family">Thripidae</taxonomicName>
), suggesting that Australian spinifex grasses host a rich fauna of arthropods.
</paragraph>
</subSubSection>
<subSubSection id="C37F654EFFBA9F3DFF09FBC8CB11FDE4" pageId="17" pageNumber="508" type="etymology">
<paragraph id="8BDA36C5FFBA9F3DFF09FBC8CB11FDE4" blockId="17.[151,1437,151,1154]" pageId="17" pageNumber="508">
<emphasis id="B911EAD7FFBA9F3DFF09FBC8CF3AFD38" bold="true" box="[199,334,1093,1118]" pageId="17" pageNumber="508">Etymology.</emphasis>
The specific name 
<taxonomicName id="4C654D46FFBA9F3DFDE4FBC8CCDFFD38" authorityName="Zhang" authorityYear="2017" box="[554,683,1093,1118]" class="Arachnida" family="Tetranychidae" genus="Eotetranychus" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="508" phylum="Arthropoda" rank="species" species="spinophilus">
<emphasis id="B911EAD7FFBA9F3DFDE4FBC8CCDFFD38" box="[554,683,1093,1118]" italics="true" pageId="17" pageNumber="508">spinophilus</emphasis>
</taxonomicName>
(spine-lover) refers to the exceptionally sharply pointed tips of the leaves of the host plant genus 
<taxonomicName id="4C654D46FFBA9F3DFE26FBE4CC4DFDE4" authorityName="Hubner" authorityYear="1820" box="[488,569,1129,1154]" class="Liliopsida" family="Poaceae" genus="Triodia" kingdom="Animalia" order="Poales" pageId="17" pageNumber="508" phylum="Tracheophyta" rank="genus">
<emphasis id="B911EAD7FFBA9F3DFE26FBE4CC4DFDE4" box="[488,569,1129,1154]" italics="true" pageId="17" pageNumber="508">Triodia</emphasis>
</taxonomicName>
that make collecting from these plants a memorably painful experience.
</paragraph>
</subSubSection>
</treatment>
</document>